Advancing and retreating glaciers during Ice Ages crushed rock into dust. Meltwater pulses that occurred when climate cycles shifted to warm stages washed this dust into rivers. Then, when climate cycles shifted to arid stages and rivers shrank in size, exposing the sediment, wind blew this dust into huge hills alongside the rivers. This wind-blown dust is known as loess. Hills made from loess are located up and down the Mississippi River Valley, but today they are covered by vegetation that originally took root when climate became more moist and warmer. Land snail shells are often found in the loess. Scientists can use the snail species composition to estimate past average summer temperatures. (Land snails are mostly active during the warmer months.) Every species of snail occurs in ranges within certain temperature parameters. Scientists can also radio-carbon date snail shells. By dating the subfossil snail shells and identifying the species present, scientists can estimate past average summer temperatures.

Map showing where hills made of Ice Age loess sediment are found.

Road cuts reveal loess sediment.

This species of land snail requires warm summers and is evidence of warmer climates.

This is another species of land snail that indicates warmer summers.

The presence of this species indicates cooler summers, like those that occurred during Ice Ages. They still occur in Canada.

Another species of snail that requires cooler summers.
Scientists determined average annual summer temperatures in the upper Midwest near the glacier edge fell drastically about 27,000 years ago–the beginning of the Last Glacial Maximum. Before summer temperatures fell, species of snails favoring warm climate including Anguispira kochi, Hawaii minisculs, and Vallonia perspectiva are found in sediment dating between 30,000 years BP-27,000 years BP. After this for the next 12,000 years snail species that require cool summers are found in the sediment. These species include Columella alticola, Vertigo modesta, and Vallonia gracilicosta. In all scientists found 68 species of terrestrial snails from 4 genera at the study sites. The sites were from 14 midwestern states. Each site had from 5-30 species.
During the Last Glacial Maximum summer temperatures near the glacier margin averaged 6-18 degrees F cooler than those of today. Summer averages were 59-61 degrees F. Near the Gulf Coast summer average temperatures were 77 degrees F compared to 92 degrees F today. The results of this study are consistent with climate data from ice cores, ocean sediment, and pollen composition.
References:
Grimley, D.; and J. Conroy
“Last Glacial Maximum Summer Temperature Gradient from Terrestrial Gastropods in Peoria Silt (loess), Midwestern USA”
Quaternary Science Reviews June 2026
Grimley, D.; et al
“Last Glacial Maximum Ecology and Climate from Terrestrial Gastropod Assemblages in Peoria Loess, Western Kentucky”
Quaternary Science Reviews 35 (5) May 2025







































