Zootaxa 4749 (1): 001–668 ISSN 1175-5326 (print edition)

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ZOOTAXA
4749

Checklist, Bibliography and Quantitative Data of the Arthropods of Hispaniola

DANIEL E. PEREZ-GELABERT
Integrated Taxonomic Information System (ITIS) and Department of Entomology, U. S. National Museum of Natural History, Smithso-
nian Institution, P. O. Box 37012, Washington, DC 20013-7012, USA. E-mail: [email protected]

Magnolia Press
Auckland, New Zealand

Accepted by E. Bernard: 2 Nov 2019; published: 10 Mar. 2020

DANIEL E. PEREZ-GELABERT
Checklist, Bibliography and Quantitative Data of the Arthropods of Hispaniola
(Zootaxa 4749)
668 pp.; 30 cm.
10 Mar. 2020
ISBN 978-1-77670-895-6 (paperback)
ISBN 978-1-77670-896-3 (Online edition)

FIRST PUBLISHED IN 2020 BY

Magnolia Press
P.O. Box 41-383
Auckland 1346
New Zealand
e-mail: [email protected]
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© 2020 Magnolia Press

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ISSN 1175-5326 (Print edition)

ISSN 1175-5334 (Online edition)

2 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Table of contents

Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Resumen . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
CHECKLIST OF SPECIES. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13
Phylum ONYCHOPHORA . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13
Phylum TARDIGRADA . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13
Phylum ARTHROPODA. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
Subphylum CRUSTACEA . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
Subclass Phyllopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
Class Remipedia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15
Infraclass Cirripedia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15
Subclass Pentastomida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
Subclass Copepoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
Class Ostracoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18
Class Malacostraca . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28
Order Thermosbaenacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 29
Order Mysida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 29
Order Amphipoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 29
Order Isopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31
Order Decapoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35
Subphylum CHELICERATA. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 45
Class Pycnogonida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 45
Class Arachnida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 45
Subclass Acari . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 45
Order Amblypygi . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 54
Order Araneae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 55
Order Opiliones . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77
Order Palpigradi . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 78
Order Pseudoscorpiones . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 78
Order Schizomida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81
Order Scorpiones . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81
Order Solifugae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83
Order Uropygi . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83
Subphylum MYRIAPODA . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84
Class Chilopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84
Class Diplopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86
Class Symphyla . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94
Subphylum HEXAPODA . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94
Class Collembola. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94
Class Protura . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96
Class Insecta . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96
Order Archaeognatha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96
Order Coleoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 97
Order Dermaptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 183
Order Dictyoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 184
Suborder Blattodea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 184
Suborder Mantodea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 184
Order Diptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 190
Order Embiidina . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 234
Order Ephemeroptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 235
Order Hemiptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 235
Suborder Heteroptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 237
Suborder Auchenorryncha. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 238
Suborder Sternorrhyncha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 238
Order Hymenoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 280
Order Lepidoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 318
Order Neuroptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 358
Order Odonata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 361
Order Orthoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 364
Order Phasmatodea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 377
Order Plecoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 378
Order Psocodea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 378

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 3

 Order Siphonaptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 382
Order Strepsiptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 382
Order Thysanoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 383
Order Trichoptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 385
Order Zoraptera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 391
Order Zygentoma . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 97
Literature references . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 391
Index to families, genera and species . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 574

Abstract

An updated and extensively revised checklist of the arthropods of Hispaniola (Dominican Republic and Haiti) is presented 11
years after the publication of the original in 2008. It integrates and quantifies all the terrestrial and surrounding marine
arthropod species (plus those of Tardigrada and Onychophora), reported in the zoological literature for Hispaniola through the
middle of 2019.
A total of 9,920 valid species (8,202 extant and 1,718 fossil) are listed, which represents an increase of 1,683 species
(1,369 extant and 314 fossil) from the original list. The largest component is Insecta (6,784 extant and 1,136 fossil), including
2,206 extant species of Coleoptera, 1,042 species of Hemiptera, 929 species of Diptera, 913 species of Lepidoptera and 774
species of Hymenoptera.
Emphasis is on reviewing and updating the original list, including all newly recorded taxa and all pertinent taxonomic
changes proposed since then. Important corrections have been made, and explanatory notes have been added. For example,
multiple authors have confused the Lesser Antillean island of Dominica with the Dominican Republic. This error is much more
common in the literature than was initially recognized. Erroneous records attributing species from one island to the other have
been identified and corrected. The original spelling of the cricket species Scapsipedus bastardoi Otte & Perez-Gelabert, 2009,
dedicated to Dominican biologist Ruth H. Bastardo, is corrected to Scapsipedus bastardoae nom. emend.
High species endemism is typical of the biota of Caribbean islands. In this checklist, a total of 3,161 arthropod species
(38.6%) are considered endemic or unique to Hispaniola. Among the speciose groups with higher levels of endemism are the
Diplopoda (91.6%), Orthoptera (90.1%), Trichoptera (82.6%), Coleoptera (49.3%) and Araneae (47.5%). Also, a total of 201
arthropod species (174 insects + 27 non-insects) are identified as introduced to the island.
The accompanying bibliography complements the taxonomic information and includes over 5,000 titles.

Key words: Arthropoda, Biodiversity, Quantitative assessment, Dominican Republic, Haiti, Caribbean, Greater Antilles, West Indies

Resumen

Se presenta una lista actualizada y extensamente revisada de los artrópodos de la Hispaniola (República Dominicana y Haití) 11
años después de la publicación original en 2008. Esta integra y cuantifica todas las especies de artrópodos terrestres y de aguas
marinas circundantes (más Tardigrada y Onychophora), reportadas en la literatura zoológica para la Hispaniola hasta mediados
del 2019.
Se lista un total de 9,920 especies válidas (8,202 vivientes y 1,718 fósiles), lo que en relación a la lista original representa
un incremento de 1,683 especies (1,369 especies vivientes y 314 fósiles). El componente más grande es Insecta (6,784 especies
vivientes y 1,135 fosiles), incluyendo 2,206 especies vivientes de Coleoptera, 1,042 especies de Hemiptera, 929 especies de
Diptera, 913 especies de Lepidoptera y 774 especies de Hymenoptera.
Se hizo énfasis en revisar los datos de la lista original, incluir todos los taxones de nuevo registro, así como todos los
cambios taxonómicos pertinentes propuestos desde entonces. Se hicieron correcciones importantes y se agregaron notas
explicativas. Por ejemplo, múltiples autores han confundido la isla de Dominica en las Antillas Menores con la República
Dominicana. Este error es mucho más común en la literatura de lo que se había reconocido inicialmente. Se identificaron y
corrigieron registros erróneos que atribuyen especies de una isla a la otra. La grafía original de la especie de grillo Scapsipedus
bastardoi Otte & Perez-Gelabert, 2009, dedicada a la bióloga dominicana Ruth H. Bastardo, es corregida a Scapsipedus
bastardoae nom. emend.
El alto endemismo de especies es típico de la biota insular caribeña. En esta lista se reconocen un total de 3,161 especies
de artrópodos (38.6%) como endémicas o únicas de la Hispaniola. Entre los grupos especiosos con más altos niveles de
endemismo están los Diplopoda (91.6%), Orthoptera (90.1%), Trichoptera (82.6%), Coleoptera (49.3%) y Araneae (47.5%).
También un total de 201 especies (174 insectos + 27 no insectos) son identificadas como introducidas a la isla.
La bibliografía acompañante complementa la información taxonómica e incluye más de 5,000 títulos.

Palabras clave: Arthropoda, Biodiversidad, Evaluación cuantitativa, Dominican Republic, Haiti, Caribe, Antillas Mayores,
Indias Occidentales.

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Introduction

The Caribbean Islands, also known as the West Indies (Greater Antilles, Lesser Antilles and the Bahamas), are a heterogeneous
group of islands that span a wide area of the tropical Atlantic. Despite their different geological histories, these islands
constitute a biogeographic subunit that has evolved unique biological characteristics. As expected, the largest and most
complex islands support the greatest share of the archipelago’s biodiversity. The Greater Antilles have characteristically high
levels of floristic and faunal endemism while at the same time being depauperate of other zoological groups (especially
mammals) relative to the surrounding continental lands. The Caribbean Islands are considered one of the hotspots of global
biodiversity (Myers et al., 2000), primarily based on the diversity of vascular plants and terrestrial vertebrates. Hotspots are
areas where high concentrations of endemic species are undergoing rapid loss of habitat and as a result are threatened by a high
risk of extinction. By focusing our attention and giving conservation priority to these areas, a greater number of species per
dollar invested can be protected from extinction (Myers et al., 2000).
Hispaniola (Fig. 1) is the second largest island (76,480 km2) in the archipelago of the Greater Antilles. It is centrally
located in the Caribbean basin south of the Tropic of Cancer between 17º40’ and 19º56’ North latitude and 68º20’ and 72º01’
West longitude. This island is shared by two countries, Haiti on the western one third (27,750 km2) and the Dominican Republic
on the eastern two-thirds (48,730 km2). The island is notable for the marked physiographic variability of its terrain. Its
dispersed and rugged mountain systems are the highest and most extensive in the West Indies. Five mountain ranges (Cordillera
Central—Massif du Nord, Cordillera Oriental, Cordillera Septentrional, Sierra de Neiba—Montagnes Noires and the Sierra de
Bahoruco—Massif de la Selle—Massif de la Hotte) traverse the island in a roughly northwest to southeast axis with valleys in-
between. These mountains and valleys include both the highest points in the insular Caribbean (Pico Duarte—3,098 m above
sea level (Orvis, 2003)) and the lowest (Lake Enriquillo basin ~40 m below sea level). High mountains are surrounded by
vegetational gradients that indicate that the peaks themselves constitute islands of habitat. Each mountain mass is a center of
endemism for many plants and animals. Climate in Hispaniola is mainly determined by its subtropical location, the interaction
of trade winds that produce rain, the internal topography of the island, and the adjacent sea. Warm temperatures are prevalent
throughout the year oscillating around 25°C, except in high mountain areas. Rainfall patterns vary across its geography,
creating a mosaic of vegetational life zones. The wettest areas are to the north and northeast, which receive over 2,500 mm of
rain per year. The drier areas to the northwest and southwest receive less than 700 mm per year.

FIGURE 1. Map of the island of Hispaniola.

Biological diversity, and the ecosystems that support it, constitute the primary source of natural goods of countries, their
natural heritage for the future. The species diversity and the structure formed by their interactions are an essential part of

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 5

ecosystem function. Arthropods play a fundamental role in the equilibrium of communities. The lack of ecological and
biological knowledge of the elements making up this diversity frequently limit their productive use and conservation. The
taxonomic characterization of the West Indian biota is still incomplete, especially for many invertebrate groups such as
arthropods, in which the diversity of species is much greater than recognized, and the size of individuals is often orders of
magnitude smaller than that of vertebrates and plants. This is patently obvious on an island like Hispaniola, where the study of
natural history by local researchers has a checkered past. The need for more detailed taxonomic knowledge of the island’s
biodiversity is tied to the need for better conservation of its many unique environments. A demanding and growing human
population puts considerable pressure on its natural resources. Loss of habitat is the main threat to the conservation of the
insular biota of Hispaniola.
This work constitutes an update of the original checklist of the arthropods of Hispaniola by Perez-Gelabert (2008). The
emphasis is on including all taxa newly recorded for the island, but also making all pertinent taxonomic changes proposed in
the zoological literature since then. The overall goal is to summarize and appraise the status of our knowledge of the arthropod
diversity of Hispaniola. Having a comprehensive list will facilitate further biodiversity studies and enhance taxonomic research
and species collections.

FIGURE 2. Known diversity of extant insects in Hispaniola. The preliminary total is 6,784 species.

Format of the checklist

Species records. Taxonomic lists are useful tools for studying biodiversity, providing informed starting points for more
comprehensive research initiatives. This checklist adheres to the International Code of Zoological Nomenclature and includes
all arthropod taxa known by the author to have been recorded in the published zoological literature from the island of
Hispaniola (Dominican Republic and Haiti) through the middle of 2019. Although Tardigrada and Onychophora are not
considered to be arthropods, they are united with Arthropoda in the monophyletic taxon Panarthropoda, and therefore are
included for the sake of completeness. The Collembola and Protura are treated as classes within the subphylum Hexapoda. The
termites are included as Epifamily Termitoidae within the suborder Blattodea, order Dictyoptera (Beccaloni, 2014). Short
taxonomic summaries of the classification used are given at the beginning of larger groups.
Species are listed alphabetically within genera, families, orders, and classes. Valid names of genera and species are listed
together with the author, year of description and at least one reference in which the species is recorded from Hispaniola. All
page number citations refer to a specific record of a species from the island. No attempt was made to include all existing
citations for any species, but the most important are provided. Species names in which the original author and date are not
accompanied by a page number, are those initially described from elsewhere and only later recorded from Hispaniola. Invalid
names (e.g., synonyms) cited for the island are included under the valid species name. In the species index invalid names are

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not italicized to distinguish them from valid names, which are italiziced. Importantly, extant and fossil undetermined species,
identified only to the generic level and reported as “Genus sp.”, are also included in the species counts.
Fossils. Amber is the fossilized resin of ancient trees that after millions of years becomes transformed into a hard
substance through polymerization. Organisms entrapped in the liquid resin are often well preserved. Today they are found in
underground deposits from which they are dug out. The amber pieces containing biological inclusions are effectively time
capsules that allow glimpses of life from the ancient past. The Dominican Republic is one of the world’s best known sources of
fossiliferous amber. Arthropods are among the most common biological inclusions in the Dominican amber. Very often their
small fossilized bodies are found preserved to the last detail. This allows the taxonomic study of these fossil inclusions almost
to the same degree as extant specimens. The age of the Dominican amber has been controversial, the most supported idea is that
it dates from the Burdigalian period of the middle Miocene epoch, some 16 million years ago (Iturralde-Vinent, 2001; Penney,
2008), although a younger age (Pliocene—Lower Pleistocene—3 to 5 million years old) has also been proposed (Braga et al.,
2012). Mineralized fossils of arthropods, especially of Crustacea, are also found in Hispaniola. Most of the crustacean fossils
correspond to shells of Ostracoda, although there are also a large number of fossilized crabs of various families. In this
checklist, fossils are listed together with their extant relatives. Fossils from the Dominican amber are identified with a dagger
symbol (†) and explicitly labelled as from Dominican amber. Mineralized fossils are denoted with a double dagger symbol (‡)
to differentiate them from those in amber.
Endemism. Species unique and/or confined to a specific area are considered endemics. The Caribbean Islands, especially
the larger islands, typically have high levels of endemism in many animal groups. Endemism varies among different arthropod
groups depending on their dispersal abilities. Groups of low vagility and high fidelity to specific environmental conditions will
have higher levels of endemism. In this checklist, endemic species are marked with an asterisk (*), with further indication of
whether the species is endemic to Haiti (*H), Dominican Republic (*DR), or both sides of the island.
Introduced species. The modern world depends on rapid intercommunication and trade between widely separated
geographic areas. This has facilitated the introduction of many species into new environments, often with negative effects on
the invaded ecosystems. Many alien plants and animals have become purposely or inadvertently introduced into the fragile
insular environment of Hispaniola. Among the many introductions, invasive species are one of the main threats to natural
communities, often causing serious damage. In this checklist, an attempt is made to identify those species that have been
introduced to the island. It is often difficult to ascertain whether a species was introduced in recent times or arrived on the
island in the past by its own means.
Type localities. In taxonomic work the “type locality” is the specific place where the type specimen of a species was
collected. It often signifies the region and environmental preferences of the species. A major addition to this list are details
about the type localities (TL:) of all endemic species. In the recording of type localities, I chose not to repeat errors of language
or typographical mistakes.
In the Hispaniolan entomological literature there is mention of the locality “San Francisco Mountains.” This was an
important collecting locality from the early 20th century that however, is not found in any map. Rehn & Hebard (1927: 11)
explained that this somewhat inaccurate term was a designation used in 1904, presumably following a local name used by a few
natives. They explained that all collections labelled San Francisco Mts. came from localities reached by foot or mule on one-
day excursions from a house located near the Blanton copper mine, near a group of hills surrounding Naranjo Dulce on the Rio
Nigua, 13 km north of San Cristóbal.
Another problematic locality is “La Cavite.” This seems to refer to a place in Haiti, perhaps a cave or depression. It is
possibly one of the localities of the herpetological collector R. H. Beck in Haiti. Matching a specific locality to this name in
Haiti has proven elusive.
Erroneous records. All possible efforts have been made to correct erroneous records and citations, often involving
confusion by various authors of the Lesser Antillean island of Dominica with the Dominican Republic. An earlier corrigenda
addressed the beetles included in the original checklist (Perez-Gelabert, 2012). Other errors such as unsupported records, are
explicitly removed from the list of Hispaniolan taxa and some species are marked as dubious records that need confirmation.

Advances

As the result of new information published between 2008 and 2019, the total number of arthropod species added to this
checklist is 1,683 species (16.9%), of which 1,363 are extant species and 314 are fossil species, mostly from Dominican amber.
Thus, on average, the rate of species descriptions and discoveries in these 11 years was about 152 species per year. However,
the year 2009 was notably above this average, with 370 new extant species, 56 new fossil species, and 22 new records of
species.

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 The 2008 checklist was one of the most important advances in the study of the Hispaniolan arthropod fauna. As of this
writing, it has been cited in over 230 zoological publications. Among the most relevant advances made during this period are
the works of Kazantsev & Perez-Gelabert (2009, 2013), providing important additions to the fireflies (Lampyridae) on the
island and doubling the number of known species. Additionally, Perez-Gelabert (2011) examined the distribution of fireflies
and noted that the northern and southern paleoislands seem to have different firefly faunas. A massive, if still introductory,
inventory of the Caribbean crickets (Grylloidea) was published by Otte & Perez-Gelabert (2009), revealing that the Caribbean
islands contain an exceptionally large number (over 500) of cricket species. Skelley (2009) provided a review of the Erotylidae
beetles of the West Indies, significantly increasing the knowledge of the family in the area. Revisionary studies including
molecular data on the Caribbean endemic radiation of Calisto butterflies (Matos-Maraví et al., 2014; Pérez-Asso et al., 2016;
Rayner Nuñez et al., 2017) have refined the knowledge of taxonomy and phylogenetic relationships within this genus. The
publication of a book on Hispaniolan butterflies (Perez-Asso et al., 2017) summarizes for researchers and the public in general
the faunistic knowledge of this best known and most popular insect group. Peck & Cook (2014) contributed a review of the
round fungus beetles (Leiodidae) of the West Indies, adding many new species. Some taxonomic advances were made on the
study of stick insects (Phasmatodea) (Poinar, 2011; Conle et al., 2014; Hennemann et al., 2016), with the descriptions of
several new genera and multiple species. Several works on Entiminae weevils by Franz (2010, 2011, 2012), Girón & Franz
(2010, 2012); Girón (2015), and Girón et al. (2018) expanded the taxonomic knowledge of this group on the island and the
Caribbean. Various genera of tortoise beetles (Cassidinae) were revised (Sekerka & Simões, 2015; Simões, 2017; Simões &
Peterson, 2018; Simões et al., 2018), advancing the knowledge of the subfamily in Hispaniola. A monographic work on bark
beetles (Scolytinae) of the West Indies by Bright (2019) greatly expanded the knowledge of these economically important
beetles. A comprehensive synopsis of the extant ant (Formicidae) fauna of Hispaniola (Lubertazzi, 2019) also broadened our
taxonomic and faunistic understanding of this ecologically important insect group. In the realm of agricultural entomology,
Serra et al. (2011) examined the impacts of exotic invasive species on Dominican agriculture. Valdéz et al. (2016) published an
updated compilation of the arthropod pests and diseases of cultivated plants.
The Museo Nacional de Historia Natural in Santo Domingo (MNHNSD) is the governmental institutional depository of
zoological collections in the Dominican Republic. As such, it maintains one of the two major arthropod collections in the
country. The museum continues to improve its collections and cultivate a new crop of local zoologists who focus their research
on vertebrates and invertebrates of the island and the Caribbean. A most important accomplishment of the museum has been the
stability and high quality attained by its scientific journal Novitates Caribaea since its reestablishment in 2009. This journal is
now published twice a year.
The Instituto de Investigaciones Botánicas y Zoológicas (IIBZ) at the Universidad Autónoma de Santo Domingo (UASD),
has become established as the research center in zoology and botany of the university. The IIBZ is also improving its
entomological, malacological and botanical collections, which in many groups surpass those of the MNHNSD. The IIBZ plays
an important support role providing a research base to students in the biology curriculum. It is currently involved in various
research projects on arthropods, particularly ants and aquatic insects.
In June 2014, I created the Facebook group “Insectos Caribeños” with the purpose of disseminating knowledge about
Caribbean insects using social media. The group unites insect aficionados from different islands with the rest of the world
through the exchange of photographs and information on insects. The group includes nearly 1,800 members, mainly insect
enthusiasts, photographers, and some entomology research professionals.

Status of knowledge

In general, our knowledge of the entomofauna of Hispaniola is still preliminary. Although important advances have been made
in the characterization of multiple groups, a majority of those contributions were made by foreign researchers. Both Haiti and
Dominican Republic need more native specialists to carry out taxonomic, faunistic and biological studies on arthropods. There
are still too few researchers in either country actively publishing on any aspect of arthropod biology. The activities of
productive local researchers would also enrich museum collections.
The total number of arthropod species (extant and fossil) compiled in this checklist now approaches 10,000, while the
number of extant species is 8,202. However, these numbers are actually lower, since they include over 500 species only
determined to the genus level. Tables 1‒4 show that insects comprise over 80% of all the recorded extant species. As expected,
the insect orders Coleoptera, Hemiptera, Lepidoptera, Diptera and Hymenoptera have the largest species richness, with
Coleoptera alone (2,206 species) representing 32.6% of all insects and 27.0% of all arthropods. The only other order that
surpasses a thousand species is Hemiptera with 1,042. Table 4 shows a comparison of the known insect fauna of Hispaniola to
that of some other islands. The recorded number of insect species in Hispaniola should increase substantially with more
exhaustive taxonomic and faunistic research.

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TABLE 1. Numbers of valid extant genera and species, endemic species, % endemism and fossil species of non-hexapod
arthropods recorded from Hispaniola. *Count includes only genera with extant species. ‡Fossil Decapoda and Ostracoda are
mineralized fossils. All others are Dominican amber fossils†.
Taxonomic group Genera* Species Endemic % Introduced Fossil
species Endemism species species†
Tardigrada 10 17 3 17.6% 0 0
Onychophora 2 6 6 100.0% 0 1
Crustacea
Amphipoda 18 26 20 76.9% 0 1
Cirripedia 1 3 0 0.0% 0 5
Copepoda 29 54 4 7.4% 0 4
Decapoda 129 232 9 3.9% 2 40‡
Isopoda 26 38 8 21.1% 3 0
Ostracoda 15 24 6 25.0% 0 297‡
Stomatopoda 8 13 0 0.0% 0 0
Other Crustacea 28 42 7 16.7% 0 0
Pycnogonida 5 9 0 0.0% 0 0
Arachnida
Acari 100 170 38 22.4% 11 19
Amblypygi 2 10 4 40.0% 0 1
Araneae 207 446 212 47.5% 2 174
Opiliones 17 25 21 84.0% 0 4
Palpigradi 1 1 0 0.0% 0 0
Pseudoscorpionida 24 27 8 29.6% 0 9
Schizomida 3 12 11 91.7% 0 2
Scorpiones 8 45 40 88.9% 0 5
Solpugida 3 3 2 66.7% 0 1
Thelyphonida 2 3 2 66.7% 0 0
Myriapoda
Chilopoda 15 24 8 33.3% 4 3
Diplopoda 60 178 163 91.6% 5 15
Symphyla 0 0 0 0.0% 0 1
All non-Hexapoda 713 1,416 570 40.3% 27 582
Hexapoda 3,148 6,784 2,591 38.2% 174 1,136

Total 3,858 8,202 3,163 38.6% 201 1,718

TABLE 2. Numbers of valid extant genera and species, endemic species, % endemism and amber fossil species of Hexapoda
known from Hispaniola. *Count includes only genera with extant species. The termites are included within the Blattodea. In
Hispaniola Plecoptera and Megaloptera are known only as amber fossils. †All known Hispaniolan fossil insects are from the
Dominican amber.
Hexapoda Genera* Species Endemic % Endemism Introduced Fossil
species species species†
Archaeognatha 0 0 0 0.0% 0 1
Blattodea 54 133 48 36.1% 3 47
Coleoptera 848 2,206 1,087 49.3% 65 314
Collembola 17 28 9 32.1% 0 12
Dermaptera 13 18 3 16.7% 0 0
Diptera 408 929 279 30.0% 14 260
Embiidina 2 2 2 100.0% 0 4
Ephemeroptera 7 9 3 100.0% 0 7
Hemiptera 543 1,042 303 29.1% 32 111
Hymenoptera 341 774 224 28.9% 42 246
Lepidoptera 646 913 164 18.0% 9 7
Mantodea 6 11 5 45.5% 0 0
Megaloptera 0 0 0 0.0% 0 1
Neuroptera 30 60 15 25.0% 0 20
Odonata 37 67 7 10.4% 0 2
Orthoptera 85 333 300 90.1% 2 23
Phasmatodea 10 26 24 92.3% 0 1
Plecoptera 0 0 0 0.0% 0 1
Protura 2 2 0 0.0% 0 0
Psocodea 24 47 18 54.5% 0 23
Siphonaptera 3 3 0 0.0% 0 3
Strepsiptera 5 8 3 37.5% 0 10
Thysanoptera 33 55 2 3.64% 7 0
Trichoptera 27 115 95 82.6% 0 33
Zoraptera 1 1 0 0.0% 0 4
Zygentoma 2 2 0 0.0% 0 6

3,148 6,784 2,591 38.2% 174 1,136

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 9

TABLE 3. Diversity of beetles (Coleoptera) reported from Hispaniola by families. *Number of families and genera only
include those with extant species. Belidae, Lucanidae, Micromalthidae, Mycetophagidae, Rhynchitidae and Scraptiidae are
families only known from fossil species.
Families* Genera* Species Endemic % Endemism Introduced Fossil
species species species†
Anobiidae 5 8 1 12.5% 0 3
Anthicidae 9 20 4 20.0% 4 0
Anthribidae 14 73 5 6.8% 1 11
Apionidae 3 11 8 72.7% 0 17
Attelabidae 3 14 7 50.0% 0 1
Belidae 0 0 0 0.0% 0 1
Bostrichidae 10 13 0 0.0% 2 1
Bothrideridae 1 2 0 0.0% 0 0
Brachypsectridae 1 1 1 100.0% 0 1
Brentidae 11 18 2 11.1% 1 1
Buprestidae 25 50 27 54.0% 2 1
Cantharidae 6 52 51 98.1% 0 1
Carabidae 72 194 91 46.9% 1 12
Cerambycidae 80 183 108 59.0% 1 16
Ceratocanthidae 1 1 1 100.0% 0 1
Cerylonidae 6 9 2 22.2% 0 0
Chelonariidae 1 1 0 0.0% 0 0
Chrysomelidae 90 247 164 66.4% 8 18
Ciidae 3 4 0 0.0% 0 0
Cleridae 8 26 20 76.9% 1 1
Coccinellidae 22 30 11 36.7% 2 0
Corylophidae 1 2 2 100.0% 0 0
Cryptophagidae 1 1 1 100.0% 0 0
Cybocephalidae 1 1 0 0.0% 0 0
Curculionidae 119 346 150 43.4% 18 109
Dermestidae 4 15 9 60.0% 3 12
Discolomatidae 1 1 1 100% 0 1
Dryopidae 1 1 0 0.0% 0 0
Dytiscidae 14 25 0 0.0% 0 1
Elateridae 15 45 16 35.6% 0 0
Elmidae 7 10 7 70.0% 0 0
Endomychidae 2 2 1 50.0% 0 1
Erotylidae 5 19 15 78.9% 1 2
Eucnemidae 1 1 0 0.0% 0 1
Geotrupidae 2 6 4 66.7% 0 0
Gyrinidae 3 5 0 0.0% 0 0
Haliplidae 1 4 0 0.0% 0 0
Heteroceridae 3 3 0 0.0% 0 0
Histeridae 11 20 2 11.1% 2 1
Hybosoridae 2 3 1 33.3% 1 3
Hydraenidae 3 3 1 33.3% 0 0
Hydrophilidae 12 32 7 21.9% 0 1
Hydrochidae 1 1 0 0.0% 0 0
Laemophloeidae 5 10 0 0.0% 0 0
Lampyridae 10 69 61 88.4% 0 0
Latridiidae 4 8 6 75.0% 0 0
Leiodidae 9 28 28 100.0 0 1
Limnichidae 4 5 0 0.0% 0 0
Lucanidae 0 0 0 0.0% 0 1
Lutrochidae 1 1 0 0.0% 0 0
Lycidae 5 10 9 90.0% 0 2
Lymexylidae 1 1 0 0.0% 0 1
Meloidae 4 4 0 0.0% 0 1
Melyridae 6 16 15 93.8% 0 0
Micromalthidae 0 0 0 0.0% 0 1
Micropeplidae 1 1 1 100.0% 0 0
Monotomidae 1 1 0 0.0% 0 0
Mordellidae 4 6 4 66.6% 0 0
Mycetophagidae 0 0 0 0.0% 0 1
Mycteridae 2 3 3 100.0% 0 0
......continued on the next page

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 TABLE 3. (Continued)
Families* Genera* Species Endemic % Endemism Introduced Fossil
species species species†
Nitidulidae 7 9 2 22.2% 0 2
Noteridae 5 6 0 0.0% 0 0
Oedemeridae 5 16 2 12.5% 0 0
Passalidae 3 5 4 80.0% 0 0
Passandridae 1 2 1 50.0% 0 0
Phalacridae 3 3 3 100.0% 0 0
Phengodidae 1 1 1 100.0% 0 0
Psephenidae 2 2 2 100.0% 0 0
Rhynchitidae 0 0 0 0.0% 0 2
Rhysodidae 1 4 4 100.0% 0 3
Ripiphoridae 1 1 0 0.0% 0 2
Salpingidae 2 2 2 100.0% 0 0
Scarabaeidae 27 134 87 64.9 % 4 6
Scraptiidae 0 0 0 0.0% 0 1
Silphidae 1 1 1 100.0% 0 0
Silvanidae 8 13 6 46.2% 4 3
Smicripidae 1 1 0 0.0% 0 0
Sphindidae 1 1 1 100.0% 0 0
Staphylinidae 96 217 98 45.2% 5 33
Tenebrionidae 39 104 59 54.8% 3 33
Trogidae 2 2 0 0.0% 0 0
Trogossitidae 4 7 4 57.1% 0 2
Zopheridae 5 12 4 33.3% 0 1
77 families 848 2,206 1,087 49.3% 64 314

TABLE 4. Comparison of reported numbers of extant insect species between Hispaniola and other islands.
Island Size Area No. Insect Species Reference
(km2)
Cuba 110,860 8,316 Genaro & Tejuca (2001)
Hispaniola 76,480 6,784 This work
Puerto Rico 9,104 5,066 Maldonado Capriles (1996)
Galapagos Islands 7,900 1,616 Bensted-Smith (2002)
Canary Islands 7,447 6,042 Izquierdo et al. (2004)
Hawaiian Islands 6,424 8,155 Eldredge & Evenhuis (2003)
Samoa 2,934 2,523 Kami & Miller (1998)
Bermuda 53.3 1,218 Sterrer (1998)

The overall endemicity for extant Hispaniolan arthropods is calculated to be 38.6%, more than one of every three species.
Some speciose groups of exceptionally high endemicity are the Diplopoda (91.6%), Orthoptera (90.1%), Trichoptera (82.6%),
Coleoptera (49.3%), and Araneae (47.5%). When all of the West Indies are considered, more than 200 beetle genera are unique
to the territory. A review of the remarkable level of endemism at the generic level among beetles (Peck & Perez-Gelabert,
2012) emphasized the extraordinary evolutionary richness of this neotropical archipelago. Extrapolating to all insects, as many
as 700 genera of insects may be endemic to the West Indies. This number is far greater than the 269 endemic genera of plants
plus all groups of non-marine vertebrates (Peck & Perez-Gelabert, 2012).
In this list, 201 species of arthropods are identified as introduced to Hispaniola. This is still a conservative listing and
many more arthropod species are likely to be introduced into the island. Among them are many plant pests that have a
significant negative impact on agriculture. Possibly the most publicized news about insects in Hispaniola in the last 11 years
was the accidental introduction of the invasive Mediterranean fruit fly (Ceratitis capitata Wiedemann), reported in March
2015, centered in the Punta Cana area. The rapid intervention of local and international authorities and the release of millions of
sterile females was key to the containment and eradication of this threat (FAO, 2018). Because of the initial ban on fruit exports
like avocado, citrus, papaya, mangoes, and peppers, this introduction resulted in the loss of many millions of dollars to the
horticultural industry of the Dominican Republic. The country was declared free of this pest in July 2017.
Evidence that the arthropod inventory is far from complete for Hispaniola is the fact that there are several groups (classes,
orders and families) of uncommon invertebrates that have not yet been recorded from the island, even though they are almost
certainly present. Examples include the soil-dwelling Diplura (two-pronged bristletails) and Pauropoda (pauropods). Also
awaiting discovery are, the Archaeognatha (jumping bristletails), known from a single fossil in amber. Among aquatic insect
families, Nepidae (water scorpions) are known from a few specimens but have not yet been formally studied. Groups that are

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 11

known from single fossils in the Dominican amber and are now apparently extinct in the island are Plecoptera (stoneflies) and
Megaloptera (dobsonflies and alderflies). Symphyla (garden centipedes) are also known from a single fossil in amber, but are
undoubtedly present and possibly even abundant in the extant fauna.

Conservation

Loss of habitat is the primary threat to the survival of the West Indian biota in general. Contributing factors are the conversion
of forests into agricultural land, charcoal production from wood, logging, mining, urbanization, introduction of exotic species,
over-exploitation of commercial species (e.g., crabs, lobsters), and the effects of climate change. The survival of organisms,
populations, and ultimately species is compromised when their environment is degraded to a point where it no longer supplies
their needs. Many Hispaniolan endemic species have very specific habitat requirements. We can infer that a portion of the
arthropod species in Hispaniola are rare, because they are collected only sporadically or after some perseverance. Those are the
species probably most vulnerable to extinction.
At present, we have insufficient knowledge about the conservation status of arthropod species in Hispaniola, but many
species could be classified as at risk because of negative trends in the quality of their habitats. Some species represent unique
lineages within their families (e.g., the carrion beetle Nicrophorus hispaniola Sikes & Peck, 2000), and as such could be of
extreme importance for understanding the evolution of the entire group (Sikes & Peck, 2000; Perez-Gelabert, 2016). Other
species have not been collected again since their initial discovery. This is the case of the pyrgomorphid grasshopper of Sierra de
Neiba (Jaragua serranus Perez-Gelabert, Dominici & Hierro, 1995). Several periods of intense drought in the southwest during
the last 30 years have apparently affected populations of this insect.
The future conservation of Hispaniolan arthropods will largely depend on preserving the integrity and health of large forest
tracts like the national parks and forest reserves already established in the Dominican Republic and Haiti. Stopping the
degradation of forests throughout the island will require a general adoption of practices like increased environmental education
in school curricula, implementing modern measures to avoid forest fires, protection of all river basins and beaches from the
extraction of sand and gravel, better management of solid waste in cities and rural towns, improved regulation of agricultural
pesticides, and eradication of invasive species. A healthy environment is clearly necessary for a healthy human population. The
task is enormous but with resolve, a harmonious environment can be created for the good of all.

Acknowledgements

Thanks are due to the many authors that have contributed information with species descriptions and taxonomic revisions
including Hispaniolan arthropods. Special gratitude is due to the Smithsonian Libraries, their interlibrary loan service and the
Biodiversity Heritage Library (BHL) project for greatly facilitating access to many publications and essentially making
possible this type of compilations. I am also very grateful to John W. Brown (National Museum of Natural History,
Smithsonian Institution) and Stewart B. Peck (Carleton University, Ottawa, Canada), who contributed language improvements
to the Introduction. Lucrecia H. Rodríguez, provided support through the years and expert advice with the production of the
illustrations. Many colleagues have also supported this initiative by providing reviews of the data in the taxonomic group of
their expertise. Most important among them are:
Alfonso García-Aldrete (Instituto de Biología, Universidad Nacional Autónoma de México—Psocodea), Ruth H. Bastardo
(Universidad Autónoma de Santo Domingo—butterflies), Benoit Vincent (Muséum national d’Histoire naturelle, Paris—
Erebidae), Brett Rattcliffe (University of Nebraska—Scarabaeidae), Carlos Suriel (Museo Nacional de Historia Natural, Santo
Domingo—Diplopoda), Cecilia Waichert (Utah State University—Pompilidae), David Hernández-Martich (Universidad
Autónoma de Santo Domingo—Drosophilidae), David Lubertazzi (Museum of Comparative Zoology, Cambridge—
Formicidae), Elijah Talamas (Florida State Collection of Arthropods, Gainesville—Platygastridae), Gabriel de los Santos
(Museo Nacional de Historia Natural, Santo Domingo—Araneae), Jennifer Girón (University of Kansas—Entiminae,
Hydrophilidae), Julio A. Genaro (Toronto, Canada—Apoidea), Marcoandre Savaris (Universidade de São Paulo, Brazil—
Tephritidae), Marcus Guidoti (Universidade Federal do Rio Grande do Sul, Porto Alegre—Tingidae), Paul H. Freytag
(University of Kentucky, Lexington—Cicadellidae), Paul Skelley (Florida State Collection of Arthropods, Gainesville—
Erotylidae), Rayner Nuñez (Instituto de Ecología y Sistemática, La Habana—Calisto), Robert Anderson (Canadian Museum of
Nature, Ottawa—Curculionidae), Sergio Godinez Cortés (Colegio de Postgraduados, Montecillo, Mexico—Bruchinae),
Stewart B. Peck (Carleton University, Ottawa, Canada—Leiodidae), Solanlly Carrero (Museo Nacional de Historia Natural,
Santo Domingo—Pseudoscorpiones) and Thomas J. Henry (Systematic Entomology Lab, USDA—Miridae).

12 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

CHECKLIST OF SPECIES

Phylum ONYCHOPHORA
Family PERIPATIDAE
Peripatus Guilding, 1826
Peripatus basilensis Brues, 1935: 62. Oliveira et al., 2012: 23 *H > TL: Morne Basile, northwestern part of the island
=Peripatus dominicae basilensis Brues, 1935: 62. Peck, 1975: 348
Peripatus darlingtoni Brues, 1935: 62. Oliveira et al., 2012: 24 *H > TL: Massif de la Hotte, between Camp Perrin and Mafin,
914 asl
=Peripatus dominicae darlingtoni Brues, 1935: 62. Peck, 1975: 348
Peripatus haitiensis Brues, 1913: 519. Oliveira et al., 2012: 25 *H > TL: Massif de la Selle, Furcy
=Peripatus dominicae haitiensis Brues, 1913: 519. Brues, 1935: 61; Peck, 1975: 348
Peripatus lachauxensis Brues, 1935: 61. Oliveira et al., 2012: 26 *H > TL: South eastern foothills of the Massif de la Hotte,
305 m asl
=Peripatus dominicae lachauxensis Brues, 1935: 61. Peck, 1975: 348
Peripatus manni Brues, 1913: 520. Brues, 1935: 61; Peck, 1975: 349; Oliveira et al., 2012: 26 *H > TL: Massif de la Selle
Macroperipatus Clark, 1913
Macroperipatus insularis Clark, 1937. Oliveira et al., 2012: 16 *H > TL: Between Jacmel and Tronin
=Macroperipatus insularis insularis Clark, 1937. Peck, 1975: 347

Family TERTIAPATIDAE†
Tertiapatus† Poinar, 2000
Tertiapatus dominicanus† Poinar, 2000: 105 [Fossil in Dominican amber]

Phylum TARDIGRADA
Family ECHINISCIDAE
Bryodelphax Thulin, 1928
Bryodelphax dominicanus (Schuster & Toftner, 1982: 225) *DR > TL: Constanza
Echiniscus Schultze, 1840
Echiniscus (Echiniscus) cavagnaroi Schuster & Grigarick, 1966. Schuster & Toftner, 1982: 226
Echiniscus (Echiniscus) virginicus Riggin, 1962. Schuster & Toftner, 1982: 226
Pseudechiniscus Mihelcic, 1964
Pseudechiniscus suillus (Ehrenberg, 1853). Schuster & Toftner, 1982: 225

Family HYPSIBIIDAE
Doryphoribius Pilato, 1969
Doryphoribius evelinae (Marcus, 1928). Schuster & Toftner, 1982: 228
Doryphoribius flavus (Iharos, 1966). Schuster & Toftner, 1982: 229
Itaquascon Barros, 1939
Itaquascon pawlowskii Weglarska, 1973. Schuster & Toftner, 1982: 229

Family MACROBIOTIDAE
Macrobiotus Schultze, 1834
Macrobiotus barbarae Kaczmarek et al., 2007: 364 *DR > TL: Puerto Plata Prov., hills near Puerto Plata; moss from tree trunk
Macrobiotus coronatus Barros, 1942. Meyer, 2013: 27
=Macrobiotus harmsworthi coronatus Barros, 1942. Schuster & Toftner, 1982: 227
Macrobiotus hufelandi group—Schultze, 1833. Schuster & Toftner, 1982: 227
Macrobiotus liviae Ramazzotti, 1962. Schuster & Toftner, 1982: 227
Macrobiotus sp.—Schuster & Toftner (1982: 226)
Minibiotus Schuster, 1980
Minibiotus intermedius (Plate, 1888). Schuster & Toftner, 1982: 228
Minilentus Guidetti & Pilato, 2003
Minilentus dubius (Schuster & Toftner, 1982: 228). Meyer, 2013: 32 *DR > TL: Cabrera
Paramacrobiotus Guidetti et al., 2009

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 13

Paramacrobiotus areolatus (Murray, 1907). Schuster & Toftner, 1982: 227; Meyer, 2013: 33
Paramacrobiotus richtersi (Murray, 1911). Schuster & Toftner, 1982: 227: 34

Family MILNESIIDAE
Milnesium Doyère, 1840
Milnesium tardigradum Doyère, 1840. Schuster & Toftner, 1982: 229

Phylum ARTHROPODA
Subphylum CRUSTACEA
Classification of the Crustacea follows Martin & Davis (2001) with modifications.

Class BRANCHIOPODA
Order ANOSTRACA
Family ARTEMIIDAE
Artemia Leach, 1819
Artemia franciscana (Kellog, 1906). Triantaphyllidis et al. (1998: 218)
Artemia sp.—Zapata & Richarson (1993: 83); Triantaphyllidis et al. (1998: 218); Mayer (2002: 29)

Family STREPTOCEPHALIDAE
Streptocephalus Baird, 1852
Streptocephalus similis Baird, 1852b: 22. Moore, 1958: 169; Hartland-Rowe, 1982: 175

Subclass PHYLLOPODA
Order SPINICAUDATA
Family LIMNADIIDAE
Eulimnadia Packard, 1874
Eulimnadia antillarum (Baird, 1852: 30). Daday, 1926: 13; Martin, 1989: 104

Order NOTOSTRACA
Family TRIOPSIDAE
Triops Schrank, 1803
Triops longicaudatus (LeConte, 1846). Hartland-Rowe, 1982: 175

Order DIPLOSTRACA
Suborder CLADOCERA
Infraorder CTENOPODA
Family SIDIDAE
Diaphanosoma Brandt, 1850
Diaphanosoma brachyurum (Liévin, 1848). Collado et al., 1984: 113
Diaphanosoma fluviatile Hansen, 1899. Collado et al., 1984: 113

Infraorder ANOMOPODA
Family CHYDORIDAE
Camptocercus Baird, 1843
Camptocercus uncinatus Smirnov, 1971. Acosta-Mercado et al., 2012: 207
Chydorus Leach, 1816
Chydorus dentifer Daday, 1905. Collado et al., 1984: 113
Chydorus linguilabris Frey, 1982b: 267. Acosta-Mercado, 2012: 207
Chydorus nitidulus (Sars, 1901). Frey, 1982a: 261
Chydorus pubescens Sars, 1901. Collado et al., 1984: 113
Dadaya Sars, 1901
Dadaya macrops (Daday, 1898). Collado et al., 1984: 113
Dunhevedia King, 1853
Dunhevedia americana Rajapaksa & Fernando, 1987: 436

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Dunhevedia crassa King, 1853. Collado et al., 1984: 114; Acosta-Mercado, 2012: 207
Dunhevedia serrata Daday, 1898. Collado et al., 1984: 114
Kurzia Dybowski & Grochowski, 1894
Kurzia latissima (Kurz, 1874). Collado et al., 1984: 114
Leberis Smirnov, 1989
Leberis davidi (Richard, 1895: 192). Collado et al., 1984: 112; Acosta-Mercado, 2012: 208
Oxyurella Dybowski & Grochowski, 1894
Oxyurella ciliata Bergamin, 1939. Collado et al., 1984: 114; Acosta-Mercado, 2012: 207

Family DAPHNIIDAE
Ceriodaphnia Dana, 1853
Ceriodaphnia cornuta Sars, 1885. Collado et al., 1984: 113
Ceriodaphnia dubia Richard, 1894. Collado et al., 1984: 113
Daphnia Müller, 1785
Daphnia magna Straus, 1820. Reyes et al., 2014: 166

Family MACROTHRICIDAE
Echinisca Liévin, 1848
Echinisca rosea (Jurine, 1820). Collado et al., 1984: 114
Macrothrix Baird, 1843
Macrothrix sp.—Richard (1895: 189)

Family MOINIDAE
Moina Baird, 1850
Moina macrocopa (Straus, 1920). Collado et al., 1984: 114
Moina micrura (Kurz, 1874). Collado et al., 1984: 114
Moina wierzejskii Richard, 1895: 195. Collado et al., 1984: 114
Moinodaphnia Herrick, 1887
Moinodaphnia macleayi (King, 1853). Collado et al., 1984: 114
Moinodaphnia sp.—Richard (1895: 189)

Class REMIPEDIA
Family MORLOCKIIDAE
Morlockia García-Valdecasas, 1984
Morlockia emersoni (Lorentzen et al., 2007: 62). Hoenemann et al., 2013: 615 *DR > TL: Cueva Los Jardines Orientales,
south coast near Santo Domingo

Class MAXILLOPODA
Subclass THECOSTRACA
Infraclass CIRRIPEDIA
Superorder THORACICA
Order PEDUNCULATA
Family LEPADIDAE
Lepas Linnaeus, 1758
Lepas anatifera Linnaeus, 1758. CIBIMA, 1994: 328
Lepas anserifera Linnaeus, 1758. CIBIMA, 1994: 328
Lepas pectinata Spengler, 1793. CIBIMA, 1994: 328

Order SESSILIA
Family BALANIDAE
Subfamily BALANINAE
Balanus Da Costa, 1778
Balanus concavus eseptatus‡ Pilsbry, 1924: 1 [Fossil]
Balanus polyporus‡ Pilsbry, 1924: 2 [Fossil]

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Subfamily AMPHIBALANINAE
Amphibalanus Pitombo, 2004
Amphibalanus amphitrite‡ (Darwin, 1854). Pilsbry, 1924: 3 [Fossil]
Amphibalanus nr. eburneus‡ (Gould, 1841). Pilsbry, 1924: 1 [Fossil]

Family CHTHAMALIDAE
Chthamalus Gray, 1825
Chthamalus sp.—Mateo (1991a: 38); CIBIMA (1994: 328)

Family PYRGOMATIDAE
Ceratoconcha Kramberger-Gorjanovic, 1889
Ceratoconcha conicocystata‡ Newman & Ladd, 1974: 391 [Fossil]

Subclass PENTASTOMIDA
Order RAILLIETIELLIDA
Raillietiella Vaney & Sambon, 1910
Raillietiella sp.—Powell et al. (1993: 51); Riley et al., 1991: 53
Note: The inclusion of Porocephalus dominicana Riley & Walters, 1980 in Perez-Gelabert (2008: 18) was erroneous. This
species was found on an individual of the snake Constrictor constrictor nebulosus in the Lesser Antillean island of Dominica.

Subclass COPEPODA
Order CALANOIDA
Family DIAPTOMIDAE
Arctodiaptomus Kiefer, 1932
Arctodiaptomus dorsalis Marsh, 1907. Smith & Fernando, 1978: 2017
Diaptomus Westwood, 1836
Diaptomus proximus Kiefer, 1936: 268
Mastigodiaptomus Light, 1939
Mastigodiaptomus albuquerquensis (Herrick, 1895). Collado et al., 1984: 115
Mastigodiaptomus purpureus (Marsh, 1907). Collado et al., 1984: 115

Order CYCLOPOIDA
Family CYCLOPIDAE
Bryocyclops Kieffer, 1927
Bryocyclops caroli Bjornberg, 1985. Acosta-Mercado et al., 2012: 207
Bryocyclops sp.—Acosta-Mercado et al. (2012: 207)
Ectocyclops Brady, 1904
Ectocyclops phaleratus (Koch, 1838). Collado et al., 1984: 115; Acosta-Mercado et al., 2012: 207
Eucyclops Claus, 1893
Eucyclops bondi Kiefer, 1934: 228. Kiefer, 1936: 291; Smith & Fernando, 1978: 2019; Collado et al., 1984: 115
Eucyclops extensus Kiefer, 1931. Kiefer, 1936: 293; Collado et al., 1984: 115
Eucyclops serrulatus (Fisher, 1851). Collado et al., 1984: 115
Fimbricyclops Reid, 1993
Fimbricyclops cf. jimhensoni Reid, 1993. Acosta-Mercado et al., 2012: 207
Halicyclops Norman, 1903
Halicyclops exiguus Kiefer, 1934: 227. Kiefer, 1936: 282; Collado et al., 1984: 115
Halicyclops similis Kiefer, 1935. Kiefer, 1936: 281; Collado et al., 1984: 115
Macrocyclops Claus, 1893
Macrocyclops albidus (Jurine, 1820). Kiefer, 1936: 291; Collado et al., 1984: 115; Pesce, 1985: 301
Mesocyclops Sars, 1914
Mesocyclops aspericornis (Daday, 1906). Pesce, 1985: 312
Mesocyclops ellipticus Kiefer, 1936. Pesce, 1985: 317
Mesocyclops longisetus (Thiebaud, 1912). Kiefer, 1936: 301; Smith & Fernando, 1978: 2020
Mesocyclops reidae Petkovski, 1986. Gutiérrez-Aguirre et al., 2006: 566

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Metacyclops Kiefer, 1927
Metacyclops (Metacyclops) dianae Pesce, 1985: 304 *H > TL: Dépt. de l’Ouest, Groureau well, at Groureau, hamlet of Cazeau,
Croix-des-Missions
Metacyclops (Metacyclops) mendocinus Wierzejski, 1902. Kiefer, 1936: 310; Pesce, 1985: 305
Microcyclops Claus, 1893
Microcyclops ceibaensis (Marsh, 1919). Kiefer, 1936: 295; Smith & Fernando, 1978: 2020; Pesce, 1985: 311; Reid, 1986: 71;
Acosta-Mercado et al., 2012: 207
Microcyclops dengizicus (Lepeshkin, 1900). Kiefer, 1936: 296; Collado et al., 1984: 115
Microcyclops diversus (Kiefer, 1935). Kiefer, 1936: 295; Collado et al., 1984: 115
Microcyclops dubitabilis (Kiefer, 1934: 229). Kiefer, 1936: 294; Collado et al., 1984: 115; Pesce, 1985: 309
Microcyclops inarmatus Gutiérrez-Aguirre & Cervantes-Martínez, 2016: 38
Microcyclops mendocinus (Wierzejski, 1892). Richard, 1895: 189; Collado et al., 1984: 115
Microcyclops panamensis (Marsh, 1913). Kiefer, 1936: 299; Collado et al., 1984: 115
Microcyclops varicans (Lilljeborg, 1901). Richard, 1895: 189; Kiefer, 1936: 310; Collado et al., 1984: 115; Pesce, 1985: 309
Paracyclops Claus, 1893
Paracyclops fimbriatus chiltoni (Thomson, 1883). Pesce, 1985: 301
Paracyclops sp.—Acosta-Mercado et al. (2012: 207)
Thermocyclops Kiefer, 1927
Thermocyclops decipiens (Kiefer, 1929). Pesce, 1985: 321
Thermocyclops orthonoides (Sars, 1863). Richard, 1895: 189; Collado et al., 1984: 116
Thermocyclops thermocyclopoides (Harada, 1931). Collado et al., 1984: 115
Tropocyclops Kiefer, 1927
Tropocyclops cf. jamaicensis Reid & Janetzky, 1996. Acosta Mercado et al., 2012: 207
Tropocyclops prasinus (Fischer, 1860). Collado et al., 1984: 116; Acosta-Mercado et al., 2012: 207

Family OITHONIDAE
Oithona Baird, 1843
Oithona sp.—Olivares (1984: 7)

Order HARPACTICOIDA
Family AMEIRIDAE
Antillesia Humes, 1958
Antillesia cardisomae Humes, 1958: 82
Cancrincola Wilson, 1913
Cancrincola jamaicensis Wilson, 1913. Humes, 1958: 77
Cancrincola longiseta Humes, 1957. Humes, 1958: 82
Cletocamptus Schmankewitsch, 1875
Cletocamptus deitersi (Richard, 1897). Kiefer, 1936: 301
Cletocamptus dominicanus Kiefer, 1934: 232. Kiefer, 1936: 302 *DR > TL: Lago Enriquillo

Family CANTHOCAMPTIDAE
Attheyella Brady, 1880
Attheyella mervini Janetzki, Martínez Arbizu & Reid, 1996. Acosta-Mercado et al., 2012: 207
Bryocamptus Chapuis, 1929
Bryocamptus sp.—Acosta-Mercado et al. (2012: 207)
Elaphoidella Chappuis, 1929
Elaphoidella bidens (Schmeil, 1894). Acosta-Mercado et al., 2012: 207
Elaphoidella sewelli (Chapuis, 1928). Acosta-Mercado et al., 2012: 207

Family LAOPHONTIDAE
Mexicolaophonte Cottarelli, 1977
Mexicolaophonte arganoi Cottarelli, 1977. Suárez-Morales et al., 2006: 7

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Family MIRACIIDAE
Schizopera Sars, 1905
Schizopera haitiana Kiefer, 1934: 230. Kiefer, 1936: 304; Suárez-Morales et al., 2006: 8 *H > TL: Etang Saumâtre
Schizopera triacantha Kiefer, 1934: 231. Kiefer, 1936: 306 *H > TL: Etang Saumâtre

Family PHYLLOGNATHOPODIDAE
Phyllognathopus Mrázek, 1893
Phyllognathopus vigueri (Maupas, 1892). Acosta-Mercado et al., 2012: 207

Order POECILOMASTOIDA
Family BOMOLOCHIDAE
Holobomolochus Vervoort, 1969
Holobomolochus divaricatus Cressey & Cressey, 1980. Cressey & Cressey, 1980: 4

Family CLAUSIIDAE
Saphirella Scott, 1894
Saphirella sp.—Olivares (1984: 7)

Order SIPHONOMASTOIDA
Family CALIGIDAE
Caligus Mueller, 1785
Caligus atromaculatus Wilson, 1913. Cressey, 1991: 3
Caligus productus Dana, 1852. Cressey & Cressey, 1980: 24

Family PSEUDOCYDNIDAE
Pseudocycnoides Yamaguti, 1963
Pseudocycnoides buccata (Wilson, 1922). Cressey & Cressey, 1980: 4

Class OSTRACODA
Classification follows the World Ostracoda Database found at World Register of Marine Species at: http://
www.marinespecies.org.
Taxonomic summary:
Subclass Myodocopa
Order Myodocopida
Family Sarsiellidae
Subclass Podocopa
Order Podocopida
Family Bairdiidae
Family Bythocytheridae
Family Candonidae
Family Cyclocyprididae
Family Cyprididae
Family Cytherellidae
Family Cushmanideidae
Family Cytheridae
Family Cytherideidae
Family Cytherettidae
Family Cytheromatidae
Family Cytherudidae
Family Darwinulidae
Family Enthocytheridae
Family Hemicytheridae
Family Krithidae
Family Limnocytheridae

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 Family Loxoconchidae
Family Macrocyprididae
Family Microcytheridae
Family Paracytherideidae
Family Paradoxostomatidae
Family Pectocytheridae
Family Pontocyprididae
Family Saididae
Family Thaerocytheridae
Family Trachyleberididae
Family Xestoleberididae

Subclass MYODOCOPA
Order MYODOCOPIDA
Family SARSIELLIDAE
Eusarsiella Cohen & Kornicker, 1975
Eusarsiella dominicana Kornicker, 1986: 158 *DR > TL: Off the Dominican Republic

Subclass PODOCOPA
Order PODOCOPIDA
Family BAIRDIIDAE
Bairdia M’Coy, 1844
Bairdia antillea‡ Bold, 1946. Bold, 1968: 49; Bold, 1981: 58 [Fossil]
Bairdia cassida‡ Bold, 1946. Bold, 1981: 59 [Fossil]
Bairdia condylus‡ Bold, 1965. Bold, 1981: 58 [Fossil]
Bairdia longisetosa‡ Brady, 1902. Bold, 1968: 48; Bold, 1975: 586; Bold, 1988: 22 [Fossil]
Bairdia oarion‡ Bold, 1972. Bold, 1981: 59 [Fossil]
Bairdia sp. aff. formosa‡ Brady, 1868. Bold, 1968: 48; Bold, 1988: 22 [Fossil]
Bairdia sp. aff. fortificata‡ Brady, 1880. Bold, 1968: 48 [Fossil]
Bairdia sp. aff. laevicula‡ Edwards, 1944. Bold, 1975: 577; Bold, 1988: 22 [Fossil]
Bairdia sp. aff. victrix‡ Brady, 1869. Bold, 1981: 58 [Fossil]
Bairdia willisensis‡ (Puri, 1954). Bold, 1975: 586; Bold, 1981: 58
Bairdoppilata Coryell Sample & Jennings, 1935
Bairdoppilata oblongata‡ (Bold, 1946). Bold, 1988: 23 [Fossil]
Bythocypris Brady, 1880
Bythocypris sp.‡—Bold (1981: 59) [Fossil]
Cardobairdia Bold, 1960
Cardobairdia asymmetrica‡ (Bold, 1946). Bold, 1981: 57 [Fossil]
Cardobairdia glabra‡ Bold, 1968: 45. Bold, 1981: 57 [Fossil]
Glyptobairdia Stephenson, 1946
Glyptobairdia sp.‡—Bold (1975: 589) [Fossil]
Macrocyprina Triebel, 1960
Macrocyprina sp.‡—Bold (1981: 58) [Fossil]
Macrocyprina sp. aff. decora‡ (Brady, 1866). Bold, 1975: 600 [Fossil]
Paranesidea Maddocks, 1969
Paranesidea antillea‡ Bold, 1946. Bold, 1988: 23 [Fossil]
Paranesidea dimorpha‡ (Bold, 1963). Bold, 1988: 24 [Fossil]
Paranesidea elegantissima‡ (Bold, 1974). Bold, 1981: 59 [Fossil]
Paranesidea sp. aff. fortificata‡ (Brady, 1880). Bold, 1975: 600; Bold, 1988: 24 [Fossil]
Triebelina Bold, 1946
Triebelina crumena‡ (Stephenson, 1944). Bold, 1981: 59 [Fossil]

Family BYTHOCYTHERIDAE
Bythoceratina Hornibrook, 1952

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Bythoceratina monoceros‡ Bold, 1988: 64 [Fossil]
Bythoceratina sp.‡—Bold (1988: 64) [Fossil]
Sclerochilus Sars, 1866
Sclerochilus sp.‡—Bold (1975: 585). Bold, 1988: 65 [Fossil]
Pseudoceratina Carter, 1885
Pseudoceratina droogeri‡ Bold, 1965. Bold, 1968: 78; Bold, 1975: 600; Bold, 1988: 65 [Fossil]
Pseudocythere Sars, 1866
Pseudocythere sp.‡—Bold (1968: 77) [Fossil]

Family CANDONIDAE
Candona Baird, 1846
Candona ex. gr. angulata‡ G. W. Müller, 1900. Bold, 1990: 225 [Fossil]
Candonopsis Vávra, 1891
Candonopsis cf. kingsleyii (Brady & Robertson, 1870). Acosta-Mercado et al., 2012: 207
Candonopsis hummelincki Broodbaker, 1983a: 289
Caribecandona Broodbaker, 1983
Caribecandona (Caribecandona) ansa Brookbaker, 1983: 314 *H > TL: Well of Gaby Gabriel, at Dessalines, Dépt. de
l’Artibonite
Caribecandona (Caribecandona) auricularia Brookbaker, 1983a: 312 *H > TL: Well of Mrs. Tissé-Coriolan, at Dumonet,
between Caiman and Debas, Dépt. de l’Ouest
Caribecandona (Caribecandona) trapezoidea Brookbaker, 1983a: 307 *H > TL: Round covered well near a house in Gressier,
N. side of road, Dépt. de l’Ouest
Cyclocypris Brady & Norman, 1889
Cyclocypris sp.‡—Bold (1975: 577) [Fossil]

Family CYPRIDIDAE
Cypretta Vávra, 1895
Cypretta sp.‡—Collado et al. (1984: 116) [Fossil]
Cypria Zenker, 1854
Cypria sp.‡—Collado et al. (1984: 116) [Fossil]
Cypridopsis Brady, 1867
Cypridopsis sp.‡—Collado et al. (1984: 116) [Fossil]
Cypridopsis vidua‡ (Müller, 1776). Bold, 1975: 583 [Fossil]
Cypridopsis vidua (Müller, 1876). Acosta-Mercado, 2012: 207
Dolerocypris Kaufmann, 1900
Dolerocypris sp.‡—Bold (1975: 584) [Fossil]
Eucypris Vávra, 1891
Eucypris domingensis Daday, 1910: 189 *DR > TL: St. Domingo
Herpetocypris Brady & Norman, 1889
Herpetocypris reptans (Baird, 1835). Acosta-Mercado et al., 2012: 207
Heterocypris Claus, 1892
Heterocypris sp.‡—Collado et al. (1984: 116) [Fossil]
Paracypris Sars, 1866
Paracypris sp.‡—Bold (1968: 47) [Fossil]
Paracypris sp. B‡—Bold (1988: 21) [Fossil]
Physocypria Vávra, 1897
Physocypria sp.‡—Collado et al. (1984: 116) [Fossil]
Potamocypris Brady, 1870
Potamocypris sp.—Collado et al. (1984: 116)
Potamocypris sp.‡—Bold (1975: 578) [Fossil]
Prionocypris Brady & Norman, 1896
Prionocypris sp.—Acosta-Mercado et al., 2012: 207
Propontocypris Silvester & Bradley, 1947
Propontocypris sp.‡—Bold (1968: 47) [Fossil]

20 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Propontocypris varaderoensis‡ Lubimova & Sánchez-Arango, 1974. Bold, 1988: 19 [Fossil]
Strandesia Stuhlmann, 1888
Strandesia (Acanthocypris) cavernicola Broodbaker, 1983b: 336
Strandesia (Acanthocypris) longula Broodbaker, 1983b: 330
Strandesia (Neocypris) stocki Broodbaker, 1983b: 339
Strandesia pistrix Broodbaker, 1983b: 348 *H > TL: Dépt. du Sud-Est, Karst spring in front of the “Bar de Simon”, along road
from Cayes to Camp-Perrin
Strandesia sp.—Collado et al. (1984: 116)
Strandesia sphaeroidea‡ Broodbaker, 1983. Bold, 1990: 225 [Fossil]
Triangulocypris Teeter, 1975
Triangulocypris pachyconcha‡ (Bold, 1946). Bold, 1988: 21 [Fossil]
Typhlocypris Vejdovsky, 1882
Typhlocypris (Typhlocypris) annae (Mehes, 1914). Karanovic & Datry, 2009: 20
Typhlocypris (Typhlocypris) antilliana (Brookbaker, 1983a: 297). Karanovic, 2005: 377
Typhlocypris (Typhlocypris) caribbeana‡ (Broodbaker, 1983). Bold, 1990: 225; Karanovic & Datry, 2009: 20 [Fossil]

Family CYTHERELLIDAE
Cytherella Jones, 1849
Cytherella caelata‡ Bold, 1963. Bold, 1975: 600 [Fossil]
Cytherella dominicana‡ Bold, 1968: 41 [Fossil]
Cytherella polita Brady, 1869: 161
Cytherella polita‡ Brady, 1869. Bold, 1975: 577 [Fossil]
Cytherella postdenticulata‡ Oertly, 1961. Bold, 1981: 55 [Fossil]
Cytherella sp. aff. pandora‡ Kornicker, 1963. Bold, 1988: 17 [Fossil]
Cytherella sp. aff. pulchra‡ Brady, 1866. Bold, 1975: 577 [Fossil]
Cytherella sp. aff. semitalis‡ Brady, 1868. Bold, 1981: 55 [Fossil]
Cytherella sp. aff. vulgata‡ Ruggieri, 1962. Bold, 1968: 40 [Fossil]
Cytherella sp.‡—Bold (1968: 43) [Fossil]
Cytherella sp. A‡—Bold (1968: 42) [Fossil]
Cytherella sp. B‡—Bold (1968: 43) [Fossil]
Cytherelloidea Alexander, 1931
Cytherelloidea sp.‡—Bold (1975: 585) [Fossil]
Cytherelloidea sp. B‡—Bold (1988: 19) [Fossil]

Family CUSHMANIDEIDAE
Cushmanidea Blake, 1933
Cushmanidea anderseni‡ (Puri, 1952). Bold, 1968: 53; Bold, 1975: 600 [Fossil]
Cushmanidea howei‡ (Bold, 1946). Bold, 1981: 70; Bold, 1988: 28 [Fossil]
Cushmanidea sp.‡—Bold (1975: 580) [Fossil]
Cushmanidea sp. aff. howei‡ (Bold, 1946)‡—Bold (1968: 52) [Fossil]

Family CYTHERIDAE
Gangamocytheridea Bold, 1963
Gangamocytheridea? plicata‡ Bold, 1968: 53. Bold, 1975: 590; Bold, 1988: 29 [Fossil]

Family CYTHERIDEIDAE
Cyprideis Jones, 1857
Cyprideis bensoni‡ Sandberg, 1966. Bold, 1974: 537 [Fossil]
Cyprideis curta‡ (Edwards, 1944). Bold, 1975: 578 [Fossil]
Cyprideis edentata‡ (Klie, 1939). Medley et al., 2008: 411 [Fossil]
Cyprideis maissadensis‡ Bold, 1981: 62 [Fossil]
Cyprideis mexicana‡ (Sandberg, 1964). Bold, 1990: 225; Medley et al., 2008: 411 [Fossil]
Cyprideis n. sp.‡—Bold (1981: 64) [Fossil]
Cyprideis pascagoulaensis‡ (Mincher, 1941). Bold, 1968: 50; Bold, 1975: 578 [Fossil]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 21

Cyprideis portusprospectuensis‡ Bold, 1971. Bold, 1975: 577 [Fossil]
Cyprideis salebrosa Bold, 1963. Meyer et al., 2017: 259
Cyprideis salebrosa‡ Bold, 1963. Bold, 1974: 537; Bold, 1975: 577 [Fossil]
Cyprideis similis‡ (Brady, 1869). Bold, 1975: 577 [Fossil]
Cyprideis subquadraregularis‡ (Brady, 1870). Bold, 1975: 580 [Fossil]
Haplocytheridea Stephenson, 1936
Haplocytheridea karlana‡ (Stephenson, 1938). Bold, 1975: 577 [Fossil]
Haplocytheridea sp.‡—Bold (1975: 577) [Fossil]
Hemicyprideis Malz & Triebel, 1970
Hemicyprideis agoiadiomensis‡ Bold, 1981: 60. Bold, 1988: 25 [Fossil]
Hemicyprideis cubensis cubensis‡ (Bold, 1946). Bold, 1988: 25 [Fossil]
Hemicyprideis stephensoni‡ (Bold, 1946). Bold, 1988: 25 [Fossil]
Messinella Bold, 1969
Messinella guanajayensis‡ (Bold, 1946). Bold, 1981: 70 [Fossil]
Perissocytheridea Stephenson, 1938
Perissocytheridea alata‡ Bold, 1946. Bold, 1981: 76; Bold, 1988: 28 [Fossil]
Perissocytheridea bicelliforma‡ Swain, 1955. Bold, 1975: 577 [Fossil]
Perissocytheridea cahobensis‡ Bold, 1981: 73. Bold, 1988: 28 [Fossil]
Perissocytheridea cytheridellaformis‡ Forester in Bold, 1975. Bold, 1975: 578 [Fossil]
Perissocytheridea? haitensis‡ Bold, 1981: 74 [Fossil]
Perissocytheridea matsoni‡ (Stephenson, 1935). Bold, 1981: 72 [Fossil]
Perissocytheridea plauta‡ Forester in Bold, 1975. Bold, 1975: 578 [Fossil]
Perissocytheridea pumila‡ Bold, 1975. Bold, 1988: 29 [Fossil]
Perissocytheridea rugata‡ Swain, 1955. Bold, 1990: 225 [Fossil]
Perissocytheridea sp. A‡—Bold (1975: 577) [Fossil]
Perissocytheridea sp. B‡—Bold (1975: 577) [Fossil]
Perissocytheridea subrugosa‡ (Brady, 1870). Bold, 1975: 577 [Fossil]
Pumilocytheridea Bold, 1963
Pumilocytheridea sandbergi‡ Bold, 1975. Bold, 1975: 584; Bold, 1988: 29 [Fossil]
Pseudopsammocythere Carbonnel, 1966
Pseudopsammocythere vicksburgensis‡ (Howe & Law, 1936). Bold, 1988: 27 [Fossil]

Family CYTHERETTIDAE
Cytheretta Müller, 1894
Cytheretta dominicana‡ (Bold, 1968). Bold, 1975: 585; Bold, 1988: 46 [Fossil]
=Cytheretta ponceana‡ Bold, 1969. Bold, 1975: 600 [Misidentification]
Cytheretta karlana‡ Howe & Pyeatt, 1935. Bold, 1968: 55; Bold, 1975: 600; Bold, 1988: 46 [Fossil]

Family CYTHEROMATIDAE
Paracytheroma Juday, 1907
Paracytheroma sp.‡—Bold (1975: 578) [Fossil]
Pellucistoma Coryell & Fields, 1937
Pellucistoma magniventra‡ Edwards, 1944. Bold, 1975: 590 [Fossil]
Pellucistoma sp. aff. magniventra‡ Edwards, 1944. Bold, 1981: 85; Bold, 1988: 66 [Fossil]
Pellucistoma? spurium‡ Bold, 1963. Bold, 1988: 66 [Fossil]
Pellucistoma? howei‡ Coryell & Fields, 1937. Bold, 1988: 66 [Fossil]
Pellucistoma sp.‡—Bold (1988: 66) [Fossil]

Family CYTHERURIDAE
Cytheropteron Sars, 1866
Cytheropteron (Aversovalva) bijuduvali‡ Bold, 1988: 61 [Fossil]
Cytheropteron (Cytheropteron) dominicanum‡ Bold, 1988: 59 [Fossil]
Cytheropteron (Cytheropteron) guraboense‡ Bold, 1988: 60 [Fossil]
Cytheropteron (Lobosocytheropteron) palton‡ Bold, 1966. Bold, 1988: 61 [Fossil]

22 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Cytheropteron sp.‡—Bold (1968: 73) [Fossil]
Cytheropteron sp. aff. bichense‡ Bold, 1963. Bold, 1981: 84 [Fossil]
Cytheropteron sp. aff. palton‡ Bold, 1958. Bold, 1981: 84 [Fossil]
Cytheropteron subreticulatum‡ Bold, 1946. Bold, 1975: 589 [Fossil]
Cytheropteron? trinidadensis‡ Bold, 1960. Bold, 1968: 73; Bold, 1981: 85 [Fossil]
Cytherura Sars, 1866
Cytherura cresera‡ Bold, 1968: 71. Bold, 1975: 600; Bold, 1988: 54 [Fossil]
Cytherura ex gr. johnsoni‡ Mincher, 1941. Bold, 1975: 577 [Fossil]
Cytherura sp. aff. swaini‡ Bold, 1963. Bold, 1988: 54 [Fossil]
Cytherura sp. A‡—Bold (1988: 54) [Fossil]
Cytherura sp. C‡—Bold (1988: 54) [Fossil]
Cytherura sp. D‡—Bold (1988: 54) [Fossil]
Cytherura sp. G‡—Bold (1988: 55) [Fossil]
Cytherura sp. H‡—Bold (1988: 55) [Fossil]
Cytherura sp. I‡—Bold (1988: 55) [Fossil]
Eucytherura Mueller, 1893
Eucytherura ex gr. complexa‡ (Brady, 1867). Bold, 1988: 56 [Fossil]
Eucytherura fithianae‡ Bold, 1988: 57 [Fossil]
Eucytherura sp.‡—Bold (1975: 600) [Fossil]
Eucytherura sp. A‡—Bold (1988: 57) [Fossil]
Eucytherura sp. B‡—Bold (1988: 57) [Fossil]
Eucytherura sp. C‡—Bold (1988: 58) [Fossil]
Eucytherura sp. D‡—Bold (1988: 58) [Fossil]
Eucytherura sp. E‡—Bold (1988: 58) [Fossil]
Eucytherura sp. G‡—Bold (1988: 58) [Fossil]
Hemicytherura Elofson, 1941
Hemicytherura bradyi‡ Puri, 1960. Bold, 1988: 56 [Fossil]
Hemicytherura cranekeyensis‡ Puri, 1960. Bold, 1968: 72; Bold, 1975: 590; Bold, 1988: 56 [Fossil]
Hemicytherura sp.‡—Bold (1975: 589) [Fossil]
Kangarina Coryell & Fields, 1937
Kangarina depressa‡ Bold, 1968: 74. Bold, 1975: 600; Bold, 1988: 58 [Fossil]
Kangarina quellita‡ Coryell & Fields, 1937. Bold, 1975: 600; Bold, 1988: 58 [Fossil]
“Kangarina sp. aff. abyssicola‡” (Müller, 1894). Bold, 1988: 59 [Fossil]
Pedicythere‡ Eagar, 1965
Pedicythere variabilis‡ Bold, 1988: 63 [Fossil]
Semicytherura Wagner, 1957
Semicytherura sp. A‡—Bold (1988: 55) [Fossil]

Family DARWINULIDAE
Darwinula Jones, 1885
Darwinula sp.‡—Bold (1975: 577) [Fossil]
Darwinula sp.—Acosta-Mercado et al. (2012: 207)
Darwinula sp. 2—Acosta-Mercado et al. (2012: 207)

Family ENTHOCYTHERIDAE
Entocythere Marshall, 1903
Entocythere ruibali Rioja, 1955. Collado et al., 1984: 116

Family HEMICYTHERIDAE
Aurila Pokorny, 1955
Aurila amigdala‡ (Stephenson, 1944). Bold, 1988: 33 [Fossil]
Aurila auricula‡ (Teeter, 1975). Bold, 1988: 33 [Fossil]
Aurila galerita‡ (Bold, 1965). Bold, 1988: 33 [Fossil]
Aurila laevicula‡ (Edwards, 1944). Bold, 1975: 577 [Fossil]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 23

Aurila macropunctata‡ (Bold, 1946). Bold, 1988: 33 [Fossil]
Aurila sp. aff. cicatricosa‡ (Reuss, 1859). Bold, 1981: 80 [Fossil]
Caudites Coryell & Fields, 1937
Caudites angulatus‡ Puri, 1960. Bold, 1975: 585 [Fossil]
Caudites medialis‡ Coryell & Fields, 1937. Bold, 1975: 600; Bold, 1988: 34 [Fossil]
Caudites nipeensis‡ Bold, 1946. Bold, 1975: 600; Bold, 1988: 34 [Fossil]
Caudites n. sp.‡—Bold (1975); Bold (1975: 600) [Fossil]
Caudites rectangularis‡ (Brady, 1969). Bold, 1988: 34 [Fossil]
Caudites sacer‡ Bold, 1970. Bold, 1988: 34 [Fossil]
Coquimba Ohmert, 1968
Coquimba congestocostata‡ (Bold, 1963). Bold, 1975: 578; Bold, 1981: 81; Bold, 1988: 38 [Fossil]
Coquimba minuta‡ (Bold, 1963). Bold, 1988: 38 [Fossil]
Hermanites Puri, 1955
Hermanites hornibrooki‡ (Puri, 1960). Bold, 1968: 66; Bold, 1975: 600; Bold, 1988: 37 [Fossil]
Hermanites tschoppi‡ (Bold, 1946). Bold, 1988: 37 [Fossil]
Jugosocythereis Puri, 1960
Jugosocythereis pannosa‡ (Brady, 1869). Bold, 1975: 577; Bold, 1988: 36 [Fossil]
Jugosocythereis vicksburgensis‡ (Howe & Law, 1936). Bold, 1988: 36 [Fossil]
Neocaudites Puri, 1960
Neocaudites triplistratus‡ (Edwards, 1944). Bold, 1975: 585 [Fossil]
Neocaudites scottae‡ Teeter, 1975. Bold, 1988: 40 [Fossil]
Orionina Puri, 1954
Orionina serrulata‡ (Brady, 1869). Bold, 1975: 577; Bold, 1988: 35 [Fossil]
Orionina vaughani‡ (Ulrich & Bassler, 1904). Bold, 1988: 35 [Fossil]
Puriana Coryell & Fields, 1953
Puriana gatunensis‡ (Coryell & Fields, 1937). Bold, 1988: 41 [Fossil]
=Puriana floridana‡ Puri, 1960. Bold, 1988: 41 [Fossil]
Puriana krutaki‡ Kontrovitz, 1976. Bold, 1988: 41 [Fossil]
Puriana pustulosa‡ Bold, 1968: 63. Bold, 1975: 600; Bold, 1988: 42 [Fossil]
Puriana scrupulosa‡ Bold, 1968: 61. Bold, 1975: 600; Bold, 1988: 42 [Fossil]
Radimella Pokorny, 1969
Radimella confragosa‡ (Edwards, 1944). Bold, 1988: 31 [Fossil]
Radimella ovata‡ Bold, 1988: 32 [Fossil]

Family KRITHIDAE
Krithe Brady, 1874
Krithe dolichodeira‡ Bold, 1946. Bold, 1968: 51; Bold, 1981: 66; Bold, 1988: 25 [Fossil]
Krithe hiwanneensis‡ Howe & Lea, 1936. Bold, 1981: 66 [Fossil]
Krithe lambi‡ Bold, 1966. Bold, 1981: 67; Bold, 1988: 25 [Fossil]
Krithe morkhoveni‡ Bold, 1960. Bold, 1981: 67 [Fossil]
Krithe prolixa‡ Bold, 1966. Bold, 1981: 68; Bold, 1988: 25 [Fossil]
Krithe reversa‡ Bold, 1958. Bold, 1981: 69 [Fossil]
Krithe trinidadensis‡ Bold, 1958. Bold, 1981: 69; Bold, 1988: 25 [Fossil]
Parakrithe Bold, 1858
Parakrithe alta‡ Bold, 1988: 26 [Fossil]
Parakrithella Hanai, 1959
Parakrithella sp.‡—Bold (1975: 577) [Fossil]

Family LIMNOCYTHERIDAE
Cytheridella Daday, 1807
Cytheridella ilosvayi‡ (Daday, 1905). Bold, 1975: 577 [Fossil]
Elpidium Müller, 1880
Elpidium sp.—Acosta-Mercado et al. (2012: 207)
Elpidium sp. 2—Acosta-Mercado et al. (2012: 207)

24 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Limnocythere Brady, 1868
Limnocythere friabilis‡ (Benson & MacDonald, 1963). Bold, 1975: 583 [Fossil]
Limnocythere opesta Brehm, 1939. Collado et al., 1984: 116
Limnocythere? sp.‡—Bold (1981: 77) [Fossil]
Limnocythere staplini‡ Gutentag & Benson, 1962. Bold, 1975: 577 [Fossil]

Family LOXOCONCHIDAE
Cytheromorpha Hirschmann, 1909
Cytheromorpha apheles‡ Bold, 1963. Bold, 1975: 584 [Fossil]
Cytheromorpha? sp. aff. caudata‡ Bold, 1963: 71 [Fossil]
Heinia Bold, 1985
Heinia sp. aff. howei‡ Bold, 1985. Bold, 1988: 31 [Fossil]
Loxoconcha Sars, 1866
Loxoconcha banesensis‡ Bold, 1946. Bold, 1988: 48 [Fossil]
Loxoconcha dorsotuberculata‡ (Brady, 1866). Bold, 1975: 577 [Fossil]
Loxoconcha fischeri‡ (Brady, 1868). Bold, 1975: 590 [Fossil]
Loxoconcha forda‡ Bold, 1968: 70. Bold, 1975: 600; Bold, 1988: 47 [Fossil]
Loxoconcha levis‡ Brady, 1871. Bold, 1975: 577 [Fossil]
Loxoconcha (Loxoconcha) spinoalata‡ (Bold, 1946). Bold, 1988: 47 [Fossil]
Loxoconcha (Loxoconcha) runa‡ (Bold, 1946). Bold, 1988: 48 [Fossil]
Loxoconcha (Loxocorniculum) antillea‡ Bold, 1946. Bold, 1988: 50 [Fossil]
Loxoconcha (Loxocorniculum) dorsotuberculata‡ (Brady, 1866). Bold, 1988: 49 [Fossil]
Loxoconcha (Loxocorniculum) fischeri‡ (Brady, 1866). Bold, 1988: 49 [Fossil]
Loxoconcha (Loxocorniculum) rugosa‡ Bold, 1946. Bold, 1981: 81; Bold, 1988: 50 [Fossil]
Loxoconcha (Loxocorniculum) tricornata‡ (Krutak, 1971). Bold, 1988: 49 [Fossil]
Loxoconcha (Palmoconcha) banesensis‡ Bold, 1946. Bold, 1988: 48 [Fossil]
Loxoconcha purisubrhomboidea‡ (Edwards, 1953). Bold, 1975: 577 [Fossil]
Loxoconcha sp. A‡—Bold (1981: 83) [Fossil]
Loxoconcha sp. aff. forda‡ Bold, 1970. Bold, 1981: 82 [Fossil]
Loxoconcha sp. aff. woodwardsensis‡ Howe & Law, 1969. Bold, 1981: 82 [Fossil]
Loxoconcha sp. B‡—Bold (1981: 83) [Fossil]
Loxoconcha (Touroconcha) lapidiscola‡ Hartmann, 1959. Bold, 1975: 589; Bold, 1988: 50 [Fossil]
Loxoconcha (Touroconcha) mosquera‡ (Bate et al., 1981). Bold, 1988: 51 [Fossil]
Phlyctocythere Keij, 1958
Phlyctocythere curva‡ Bold, 1988: 52 [Fossil]
Phlyctocythere recta‡ Bold, 1988: 53 [Fossil]
Phlyctocythere sp. 1‡—Bold (1988: 53) [Fossil]
Phlyctocythere sp. 2‡—Bold (1988: 53) [Fossil]
Phlyctocythere stricta‡ Bold, 1988: 52 [Fossil]

Family MACROCYPRIDIDAE
Macrocypris Brady, 1868
Macrocypris sp.‡—Bold (1975: 585) [Fossil]
Macrocypris sp. 1‡—Bold (1988: 21) [Fossil]
Macrocypris sp. 2‡—Bold (1988: 22) [Fossil]

Family MICROCYTHERIDAE
Microcythere Müller, 1894
Microcythere howei‡ (Puri, 1954). Bold, 1988: 67 [Fossil]

Family PARACYTHERIDEIDAE
Paracytheridea Mueller, 1894
Paracytheridea altila‡ Edwards, 1944. Bold, 1968: 76; Bold, 1975: 600 [Fossil]
Paracytheridea calcitrapa‡ Bold, 1988: 62 [Fossil]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 25

Paracytheridea sp.‡—Bold (1968: 76) [Fossil]
Paracytheridea sp. aff. hispida‡ Bold, 1946. Bold, 1968: 76; Bold, 1975: 600; Bold, 1981: 84; Bold, 1988: 63 [Fossil]
Paracytheridea tschoppi‡ Bold, 1946. Bold, 1968: 76; Bold, 1975: 600 [Fossil]

Family PARADOXOSTOMATIDAE
Caribbella Teeter, 1975
Caribbella geisteri‡ Bold, 1988: 45 [Fossil]
Caribbella yoni‡ (Puri, 1960). Bold, 1988: 46 [Fossil]
Paracythere Müller, 1894
Paracythere sp.‡—Bold (1988: 67) [Fossil]
Paradoxostoma Fischer, 1855
Paradoxostoma sp.‡—Bold (1975: 600) [Fossil]
Paradoxostoma sp. 1‡—Bold (1988: 65) [Fossil]
Paradoxostoma sp. 2‡—Bold (1988: 65) [Fossil]
Xiphichilus Brady, 1870
Xiphichilus sp.‡—Bold (1988: 65) [Fossil]

Family PECTOCYTHERIDAE
Munseyella Bold, 1957
Munseyella bermudezi‡ Bold, 1966. Bold, 1975: 600 [Fossil]
Munseyella jungi‡ Bold, 1988: 30 [Fossil]
Munseyella saundersi‡ Bold, 1988: 30 [Fossil]
Munseyella sp.‡—Bold (1981: 71) [Fossil]

Family PONTOCYPRIDIDAE
Abyssocypris Bold, 1974
Abyssocypris pikna‡ (Bold, 1960). Bold, 1981: 57 [Fossil]
Abyssocypris tipica‡ Bold, 1974. Bold, 1981: 57 [Fossil]
Argilloecia Sars, 1866
Argilloecia sp. cf. alexanderi‡ Swain, 1948. Bold, 1981: 57 [Fossil]
Argilloecia hiwanneensis‡ Howe & Lea, 1936. Bold, 1975: 594 [Fossil]
Argilloecia sp.‡—Bold (1975: 594) [Fossil]
Argilloecia sp.‡ cf. vespa Bold, 1988: 20 [Fossil]
Argilloecia sp. 1‡—Bold (1968: 47) [Fossil]
Argilloecia sp. 2‡—Bold (1988: 20) [Fossil]
Argilloecia sp. 3‡—Bold (1988: 21) [Fossil]
Argilloecia vespa‡ Bold, 1988: 20 [Fossil]
Pontocypris Sars, 1866
Pontocypris ex gr. dreikanter‡ (Coryell & Fields, 1958). Bold, 1988: 19 [Fossil]
Pontocypris sp.‡—Bold (1975: 578) [Fossil]

Family SAIDIDAE
Saida Hornibrook, 1952
Saida? kingmai‡ (Bold, 1946). Bold, 1988: 59 [Fossil]

Family THAEROCYTHERIDAE
Bradleya Hornibrook, 1952
Bradleya ex gr. dictyon‡ (Brady, 1880). Bold, 1975: 593; Bold, 1988: 37 [Fossil]
Bradleya hazelae‡ Bold, 1946. Bold, 1968: 66 [Fossil]
Quadracythere Hornibrook, 1952
Quadracythere antillea‡ (Bold, 1946). Bold, 1981: 80 [Fossil]
Quadracythere bichensis‡ (Bold, 1963). Bold, 1975: 589 [Fossil]
Quadracythere brachypygaia‡ Bold, 1965. Bold, 1988: 36 [Fossil]
Quadracythere howei‡ (Puri, 1953). Bold, 1988: 35 [Fossil]

26 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Quadracythere producta‡ (Brady, 1866). Bold, 1968: 57; Bold, 1975: 577; Bold, 1988: 36 [Fossil]

Family TRACHYLEBERIDIDAE
Abyssocythere Benson, 1971
Abyssocythere trinidadensis‡ (Bold, 1957). Bold, 1981: 78 [Fossil]
Acuticythereis Edwards, 1944
Acuticythereis elongata‡ Bold, 1946. Bold, 1968: 68; Bold, 1975: 600 [Fossil]
Agrenocythere Benson, 1972
Agrenocythere hazelae‡ (Bold, 1946). Bold, 1981: 79 [Fossil]
Ambocythere Bold, 1958
Ambocythere exilis‡ Bold, 1965: 13. Bold, 1968: 64; Bold, 1975: 593; Bold, 1988: 42 [Fossil]
Ambocythere subreticulata‡ Bold, 1958. Bold, 1981: 16 [Fossil]
Ambocythere sp.‡—Bold (1965: 11) [Fossil]
Basslerites Howe, 1937
Basslerites minutus‡ Bold, 1958. Bold, 1975: 577 [Fossil]
Bosquetina Brady et al., 1874
Bosquetina sp. aff. tarentina‡ (Baird, 1850). Bold, 1975: 600 [Fossil]
Brachycythere Alexander, 1933
Brachycythere sp. aff. russelli‡ Howe & Lea, 1936. Bold, 1968: 70; Bold, 1988: 43 [Fossil]
Campylocythere Edwards, 1944
Campylocythere? perieri‡ (Brady, 1868). Bold, 1975: 577 [Fossil]
Campylocythere? sp.‡—Bold (1968: 68) [Fossil]
Campylocythere? sp. A‡—Bold (1975: 577) [Fossil]
Cativella Coryell & Fields, 1937
Cativella navis‡ Coryell & Fields, 1937. Bold, 1975: 577; Bold, 1988: 39 [Fossil]
Cativella sp. aff. moriahensis‡ Bold, 1957. Bold, 1988: 39 [Fossil]
Costa Neviani, 1928
Costa bellipulex‡ Levinson, 1974. Bold, 1981: 78; Bold, 1988: 39 [Fossil]
Costa cubana‡ Bold, 1970. Bold, 1981: 78 [Fossil]
Costa dohmi‡ Bold, 1968: 59. Bold, 1981: 78; Bold, 1988: 39 [Fossil]
Costa maquayensis‡ Bold, 1961. Bold, 1981: 78 [Fossil]
Costa praedohmi‡ Bold, 1970. Bold, 1981: 77 [Fossil]
Echinocythereis Puri, 1953
Echinocythereis madremaestrae‡ Bold, 1988: 42 [Fossil]
Henryhowella Puri, 1956
Henryhowella ex gr. asperrima‡ (Reuss, 1850). Bold, 1975: 589 [Fossil]
Occultocythereis Howe, 1951
Occultocythereis angusta‡ Bold, 1963. Bold, 1988: 40 [Fossil]
Procythereis Skogsberg, 1928
Procythereis? deformis‡ (Reuss, 1850). Bold, 1975: 600; Bold, 1988: 32 [Fossil]
Procythereis? howei‡ (Bold, 1975). Bold, 1988: 32 [Fossil]
Procythereis? sp.‡—Bold (1981: 79) [Fossil]
Proteoconcha Plusquellec & Sandberg, 1969
Proteoconcha? evai‡ Bold, 1988: 44 [Fossil]
Pterygocythereis Blake, 1933
Pterygocythereis llinasi‡ Bold, 1988: 40. Bold, 1988: 40 [Fossil]
Pterygocythereis miocenica‡ Bold, 1967. Bold, 1968: 65; Bold, 1975: 600 [Fossil]
Pterygocythereis polita‡ Bold, 1966. Bold, 1968: 64; Bold, 1975: 600; Bold, 1988: 40 [Fossil]
Trachyleberidea Bowen, 1972
Trachyleberidea cubensis‡ (Bold, 1946). Bold, 1968: 58 [Fossil]
Trachyleberidea mammidentata‡ (Bold, 1946). Bold, 1968: 57; Bold, 1975: 589; Bold, 1988: 38 [Fossil]
Trachyleberidea sp. aff. cubensis‡—Bold (1946); Bold (1968: 59) [Fossil]
Trachyleberis Brady, 1898
Trachyleberis? bermudezi crebripustulosa‡ Bold, 1966. Bold, 1988: 38 [Fossil]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 27

Family XESTOLEBERIDIDAE
Microxestoleberis Müller, 1894
Microxestoleberis sp.‡—Bold (1988: 70) [Fossil]
Uroleberis Triebel, 1958
Uroleberis sp.‡—Bold (1975: 583) [Fossil]
Uroleberis torquata‡ Bold, 1968: 81. Bold, 1975: 580; Bold, 1988: 69 [Fossil]
Uroleberis triangula‡ Bold, 1968: 80. Bold, 1975: 600; Bold, 1988: 69 [Fossil]
Uroleberis sp. 1‡—Bold (1988: 69) [Fossil]
Uroleberis sp. aff. Uleberis sp. 1‡—Bold (1988: 70) [Fossil]
Xestoleberis Sars, 1866
Xestoleberis antillea‡ Bold, 1988: 68 [Fossil]
Xestoleberis duo‡ Bold, 1988: 67 [Fossil]
Xestoleberis punctata‡ Tressler, 1949. Bold, 1988: 68 [Fossil]
Xestoleberis sp. 1‡—Bold (1968: 78) [Fossil]
Xestoleberis sp. 2‡—Bold (1968: 78) [Fossil]
Xestoleberis sp. 3‡—Bold (1968: 79) [Fossil]
Xestoleberis sp. 4‡—Bold (1968: 79) [Fossil]
Xestoleberis sp. B‡—Bold (1988: 69) [Fossil]
Xestoleberis sp. E‡—Bold (1988: 69) [Fossil]
Xestoleberis sp. H‡—Bold (1988: 69) [Fossil]

Class MALACOSTRACA
Subclass HOPLOCARIDA
Order STOMATOPODA
Family GONODACTYLIDAE
Neogonodactylus Manning, 1995
Neogonodactylus bredini (Manning, 1969: 315). Herrera-Moreno & Betancourt-Fernández, 2003: 275
Neogonodactylus curacaoensis (Schmitt, 1924). Manning, 1969: 307; Herrera-Moreno & Betancourt-Fernández, 2003: 275
Neogonodactylus oerstedii (Hansen, 1894). Manning, 1969: 325; CIBIMA, 1994: 329; Herrera-Moreno & Betancourt-
Fernández, 2003: 275
Neogonodactylus spinulosus (Manning, 1924). Manning, 1969: 299; Herrera-Moreno & Betancourt-Fernández, 2003: 275
Pseudosquilla Dana, 1852
Pseudosquilla ciliata (Fabricius, 1787). Manning, 1969: 264; CIBIMA, 1994: 329; Herrera-Moreno & Betancourt-Fernández,
2003: 275
Pseudosquilla oculata (Brullé, 1835). Manning, 1969: 271; Herrera-Moreno & Betancourt-Fernández, 2003: 275

Family LYSIOSQUILLIDAE
Lysiosquilla Dana, 1852
Lysiosquilla glabriuscula (Lamarck, 1818). Herrera-Moreno & Betancourt-Fernández, 2003: 276
Lysiosquilla scabricauda (Lamarck, 1818). CIBIMA, 1994: 329; Herrera-Moreno & Betancourt-Fernández, 2003: 276

Family NANNOSQUILLIDAE
Nannosquilla Manning, 1963
Nannosquilla schmitti (Manning, 1962). Perez-Gelabert, 2008: 29

Family SQUILLIDAE
Alima Leach, 1817
Alima hyalina Leach, 1817. Manning, 1969: 128; Herrera-Moreno & Betancourt-Fernández, 2003: 275
Cloridopsis Manning, 1968
Cloridopsis dubia (H. Milne Edwards, 1837). Manning, 1969: 141; Herrera-Moreno & Betancourt-Fernández, 2003: 276
Fennerosquilla Manning & Camp, 1983
Fennerosquilla heptacantha (Chace, 1939). Manning, 1969: 171; Herrera-Moreno & Betancourt-Fernández, 2003: 276
Squilla Gronovius, 1760
Squilla intermedia Bigelow, 1893. Manning, 1969: 228; Herrera-Moreno & Betancourt-Fernández, 2003: 276

28 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Subclass EUMALACOSTRACA
Superorder PERACARIDA
Order TANAIDACEA
Family METAPSEUDIDAE
Pseudoapseudomorpha Gutu, 1991
Pseudoapseudomorpha gomezi Gutu & Ortiz, 2009. Gutu, 2009: 118
Zaraza Gutu, 2006
Zaraza linda Gutu, 2006: 50 *DR > TL: Samaná Peninsula, Playa Colibrí, about 1 km West of Las Terrenas

Order THERMOSBAENACEA
Family MONODELLIDAE
Monodella Ruffo, 1949
Monodella sp.—Stock (1983: 239)
Tethysbaena Wagner, 1993
Tethysbaena gaweini Wagner, 1993: 79 *DR > TL: Prov. Espaillat, Magante, well of Paulino Almonte, near bent in road, 20 m
Tethysbaena haitiensis Wagner, 1993: 86 *H > TL: Dépt. de l’Ouest, E of Croix-des-Missions, from well
Tethysbaena juglandis Wagner, 1993: 95 *H > TL: Dépt. Du Nord, Limbé, N side road D 100, from covered well
Tethysbaena juriaani Wagner, 1993: 68 *DR > TL: Distrito Nacional, Guaricano, well of Ramona Altagracia de Pichardo,
Calle Gregorio Luperón #7, 20 m

Order MYSIDA
Family MYSIDAE
Heteromysis Smith, 1873
Heteromysis actiniae Clarke, 1955. Williams et al., 1983: 45; CIBIMA, 1994: 328
Mysidium Dana, 1852
Mysidium sp.—CIBIMA (1994: 328)
Siriella Dana, 1850
Siriella chierchiae Coifmann, 1937. Tattersall, 1951: 70; Brattegard, 1975: 109; Müller, 1993: 44

Order STYGYOMYSIDA
Family STYGIOMYSIDAE
Stygiomysis Caroli, 1937
Stygiomysis aemete Wagner, 1992: 71. Müller, 1993: 285 *DR > TL: Prov. Pedernales, 2.8 km E of Pedernales, well, half
covered by wood, sea level

Order AMPHIPODA
Suborder GAMMARIDEA
Family BOGIDIELLIDAE
Bogidiella Hertzog, 1933
Bogidiella (Hagidiella) prionura Stock, 1985a: 217 *H > TL: Berquer well at Jérémie, Département de la Grande Anse
Bogidiella (Mexigidiella) hamatula Stock, 1985a: 211 *H > TL: Rivière de la Grande Anse at Fondelain, Département de la
Grande Anse

Family HADZIIDAE
Apoweckelia Stock, 1985
Apoweckelia serrata Stock, 1985b: 418 *H > TL: Dépt. du Sud, spring Dubreuil II at Ducis, some 500 m downstream of a small
dam on the Acul river
Bahadzia Holsinger, 1985
Bahadzia jaraguensis Jaume & Wagner, 1998: 51 *DR > TL: Cueva de Mondesí, near Oviedo, Prov. Pedernales
Bahadzia latipalpus Stock, 1985b: 391 *H > TL: Dépt. de l’Est, Marigot, well of Jacques Simein
Crangoweckelia Stock, 1985
Crangoweckelia mixta Stock, 1985b: 338 *H > TL: Dépt. de l’Artibonite, Dessalines, well Nord Albert
Crangoweckelia spinicauda Stock, 1985b: 345 *H > TL: Dépt. de l’Ouest, two karst springs on the shore of l’Etang Saumâtre

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 29

Metaniphargus Stephensen, 1933
Metaniphargus (Caribdzia) chaetodactylus Stock, 1985b: 370 *H > TL: Dépt. du Sud, banks of the river l’islet, near Cayes
Metaniphargus (Caribdzia) haitianus Stock, 1985b: 360 *H > TL: Dépt. du Nord, open well just N. E. of Cormier Plage
Metaniphargus (Guadzia) crenatus Stock, 1985b: 386 *H > TL: Dépt. de la Grand Anse, Bac well, near Jérémie
Metaniphargus (Hispadzia) longipalpus Stock, 1985b: 365 *H > TL: Dépt. de la Grand Anse, one of the springs of the brook
Tessier
Metaniphargus (Hispadzia) pedunculatus Stock, 1985b: 375 *H > TL: Dépt. de Sud, large karst spring with many outlets on the
road Cayes-Camp Perrin
Metaniphargus (Hispadzia?) plumicauda Stock, 1985b: 380 *H > TL: Dépt. de l’Est, karst spring Ca-Auguste
Pintaweckelia Stock, 1985
Pintaweckelia grandis Stock, 1985b: 350 *H > TL: Dépt. du Nord, Limbé, covered, pumped well of the Presbytère
Radaweckelia Stock, 1985
Radaweckelia brevicauda Stock, 1985b: 409 *H > TL: Dépt. du Centre, karst springs known as Ara, near the waterfall (Saut
d’Eau), hamlet la Selle, Ville Bonheur
Zombiweckelia Stock, 1985
Zombiweckelia parvipalpus Stock, 1985b: 400 *H > TL: Dépt. de l’Ouest, two karst springs E.S.E. of the hamlet of Fond
Parisien

Family HYALELLIDAE
Hyalella Smith, 1874
Hyalella (Parhyalella) cf. platensis—Margalef (1986: 8)
Parhyale Stebbing, 1897
Parhyale hawaiiensis (Dana, 1853). Shoemaker, 1956: 351

Family ISAEIDAE
Photis Krøyer, 1842
Photis trapherus Thomas & Barnard, 1991: 96

Family LEUCOTHOIDAE
Leucothoe Leach, 1814
Leucothoe sp.—Mateo (1991: 7)

Family LYSIANASSIDAE
Ottenwalderia Jaume & Wagner, 1998
Ottenwalderia kymbalion Jaume & Wagner, 1998: 39 *DR > TL: El Agua de Odín, Oviedo, Prov. Pedernales

Family METACRANGONYCTIDAE
Metacrangonyx Chevreux, 1909
Metacrangonyx dominicanus Jaume & Christenson, 2001: 101 *DR > TL: Cueva Seca, near Juan Dolio, San Pedro de Macorís
Prov.
Metacrangonyx samanensis Jaume & Christenson, 2001: 114 *DR > TL: Cueva del Agua de Saturnino, Samaná

Family STENOTHOIDAE
Stenothoe Dana, 1852
Stenothoe sp.—Mateo (1991: 7)

Family TALITRIDAE
Caribitroides Bousfield, 1984
Caribitroides genaroi Ortíz & Lalana, 2009: 26 *DR > TL: Las Abejas, Sierra de Bahoruco, 1314 m
Tethorchestia Bousfield, 1984
Tethorchestia palaeorchestes† Bousfield & Poinar, 1995: 748 [Fossil in Dominican amber]

Family TRISCHIZOSTOMATIDAE
Trischizostoma Boeckh, 1860

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Trischizostoma nicaeense (Costa, 1853). Varela & Bracken-Grissom, 2017: 17

Order ISOPODA
Suborder CYMOTHOIDA
Superfamily ANTHUROIDEA
Family ANTHURIDAE
Cyathura Norman & Stebbing, 1886
Cyathura (Cyathura) tridentata Wagner, 1990a: 152 *DR > TL: La Ciénaga, at beach 100 m northeast of the river Cañada
Baialla, at sea level
Cyathura (Stygocyathura) broodbakkeri Wagner, 1990a: 147 *DR > TL: Majagual, ~1 km S of Cabral, Prov. Barahona, 20 m
Cyathura (Stygocyathura) motasi Botosaneanu & Stock, 1982: 20. Kensley & Schotte, 1989: 36
Cyathura (Stygocyathura) salpiscinalis Botosaneanu & Stock, 1982. Wagner, 1990a: 147; Kensley & Schotte, 1989: 38

Superfamily CYMOTHOIDEA
Family AEGIDAE
Rocinela Leach, 1818
Rocinela signata (Schiödte & Meinert, 1879). Kensley & Schotte, 1989: 120

Family CIROLANIDAE
Anopsilana Paulian & Delamare Deboutteville, 1956
Anopsilana sp.—Notenboom (1981: 323)
Arubolana Botosaneanu & Stock, 1979
Arubolana rotunditelson Botosaneanu & Illife, 2010: 47 *DR > TL: Cueva de los Murciélagos, Cabarete
Booralana Bruce, 1986
Booralana tricarinata Camp & Heard, 1988. Kensley & Schotte, 1989: 275
Calyptolana Bruce, 1985
Calyptolana hancocki Brucke, 1985. Kensley & Schotte, 1989: 132
Cirolana Leach, 1818
Cirolana (Anopsilana) acanthura (Notenboom, 1981: 319). Kensley & Schotte, 1989: 125; Illiffe & Botosaneanu, 2006: 14
Cirolana (Anopsilana) radicicola (Notenboom, 1981: 314). Kensley & Schotte, 1989: 127; Illiffe & Botosaneanu, 2006: 14
Eurydice Leach, 1815
Eurydice personata Kensley, 1987. Kensley & Schotte, 1989: 149

Family CORALLANIDAE
Excorallana Stebbing, 1904
Excorallana tricornis (Hansen, 1890). Williams et al., 1983: 46; CIBIMA, 1994: 328

Family CYMOTHOIDAE
Anilocra Leach, 1818
Anilocra acanthuri Williams & Williams, 1981: 1018. Williams et al., 1983: 45; Kensley & Schotte, 1989: 176; CIBIMA,
1994: 328
Anilocra chaetodontis Williams & Williams, 1981. Williams et al., 1983: 46
Anilocra chromis Williams & Williams, 1981: 1024. Williams et al., 1983: 46; Kensley & Schotte, 1989: 177; CIBIMA, 1994:
328
Anilocra haemuli Williams & Williams, 1981: 1006. Williams et al., 1983: 45; Kensley & Schotte, 1989: 177; CIBIMA, 1994:
328
Anilocra holacanthi Williams & Williams, 1981: 1030. Williams et al., 1983: 46; Kensley & Schotte, 1989: 179
Anilocra myripristis Williams & Williams, 1981: 1016. Williams et al., 1983: 45; Kensley & Schotte, 1989: 180
Ceratothoa Dana, 1852
Ceratothoa deplanata Bovallius, 1885. Kensley & Schotte, 1989: 180 *H > TL: Haiti
Cymothoa Fabricius, 1793
Cymothoa oestrum (Linnaeus, 1758). Kensley & Schotte, 1989: 183
Mothocya Hope, 1851
Mothocya bermudensis Bruce, 1986: 1113

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 31

Renocila Miers, 1880
Renocila bowmani E. Williams & L. Williams, 1980: 585. Williams et al., 1983: 46; Kensley & Schotte, 1989: 191 *DR > TL:
South central coast of Saona Island
Renocila colini E. Williams & L. Williams, 1980. Williams et al., 1983: 46
Renocila waldneri E. Williams & L. Williams, 1980: 578. Williams et al., 1983: 46; Kensley & Schotte, 1989: 193 *DR > TL:
La Caleta, near airport, Santo Domingo

Superfamily BOPYROIDEA
Family BOPYRIDAE
Azygopleon Markham, 1985
Azygopleon schmitti (Pearse, 1932). Markham, 1985: 122; Kensley & Schotte, 1989: 110
Bopyrione Bourdon & Markham, 1980
Bopyrione synalphei Bourdon & Markham, 1980: 222. Markham, 1985: 129; Kensley & Schotte, 1989: 110
Dactylokepon Stebbing, 1910
Dactylokepon caribeus Markham, 1975: 61. Kensley & Schotte, 1989: 111
Eophrixus Caroli, 1930
Eophrixus subcaudalis (Hay, 1917). Markham, 1985: 100; Kensley & Schotte, 1989: 111
Hemiarthrus Giard & Bonnier, 1887
Hemiarthrus synalphei (Pearse, 1950). Markham, 1985: 118; Kensley & Schotte, 1989: 111
Kolourione Markham, 1978
Kolourione premordica Markham, 1978: 483
Probopyrus Giard & Bonnier, 1888
Probopyrus pandalicola (Packard, 1879). Markham, 1985: 29
Synsynella Hay, 1917
Synsynella choprae (Pearse, 1932). Markham, 1985: 129; Kensley & Schotte, 1989: 113
Synsynella deformans Hay, 1917. Bourdon, 1981: 1151

Suborder MICROCERBERIDEA
Family ATLANTASELLIDAE
Atlantasellus Sket, 1980
Atlantasellus dominicanus Jaume, 2001: 222 *DR > TL: Cueva de los Bolos, cenote, at 1.2 km inland from the western shore of
Laguna de Oviedo

Suborder ASELLOTA
Family JANIRIDAE
Jehaia Wagner, 1990
Jehaia stocki Wagner, 1990b: 187 *DR > TL: Prov. Barahona, Los Patos, 250 m E of bridge over cañada

Suborder ONISCIDEA
Family ASELLIDAE
Asellus Geoffroy, 1762
Asellus sp.—Cicero (1981: 121)
Lirceus Agassiz, 1846
Lirceus sp.—Cicero (1981: 121)

Infraorder DIPLOCHAETA
Family LIGIIDAE
Ligia Weber, 1795
Ligia (Megaligia) exotica Roux, 1828. Van Name, 1936: 48

Superfamily ONISCOIDEA
Family ONISCIDAE
Cubaris Billberg, 1820
Cubaris murina Brandt, 1833. Van Name, 1936: 48 [Introduced]

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Exosphaeroma Stebbing, 1900
Exosphaeroma bondi Van Name, 1936: 512
Oniscus Latreille, 1802
Oniscus asellus Linnaeus, 1758. Van Name, 1942: 326 [Introduced]
Phalloniscus Budde-Lund, 1909
Phalloniscus avrilensis (Van Name, 1940: 114). Leistikow & Wägele, 1999: 25
Porcellio Latreille, 1804
Porcellio laevis Latreille, 1804. Van Name, 1936: 34 [Introduced]
Venezillo Verhoeff, 1928
Venezillo hendersoni (Boone, 1934: 595). Van Name, 1936: 337; Leistikow & Wägele, 1999: 48
Venezillo phylax (Van Name, 1936: 353). Leistikow & Wägele, 1999: 50
Venezillo tuberosus (Budde-Lund, 1904). Van Name, 1936: 377; Leistikow & Wägele, 1999: 51

Family PLATYARTHRIDAE
Trichorhina Budde-Lund, 1908
Trichorhina sp.†—Schmalfuss (1984: 13) [Fossil in Dominican amber]
Trichorhina termophila (Dollfus, 1896). Van Name, 1936: 192; Leistikow & Wägele, 1999: 30

Family PSEUDOARMADILLIDAE
Pseudarmadillo Saussure, 1857
Pseudarmadillo cristatus† Schmalfuss, 1980: 4. Leistikow & Wägele, 1999: 52 [Fossil in Dominican amber]
Pseudarmadillo tuberculatus† Schmalfuss, 1984: 10. Leistikow & Wägele, 1999: 52 [Fossil in Dominican amber]

Family RHYSCOTIDAE
Rhyscotoides Arcangeli, 1947
Rhyscotoides ciferrii (Arcangeli, 1930: 35). Van Name, 1936: 268; Leistikow & Wägele, 1999: 32 *DR > TL: Los Hermanos
islands near Santo Domingo
Rhyscotus Budde-Lund, 1885
Rhyscotus jacksoni Arcangeli, 1930: 32. Van Name, 1936: 273; Leistikow & Wägele, 1999: 32

Family SCLEROPACTIDAE
Protosphaeroniscus† Schmalfuss, 1980
Protosphaeroniscus tertiarius† Schmalfuss, 1980: 4. Leistikow & Wägele, 1999: 39; Schmidt, 2007: 74 [Fossil in Dominican
amber]

Suborder SPHAEROMATIDEA
Family SPHAEROMATIDAE
Dynamenella Hansen, 1905
Dynamenella perforata (Moore, 1901). Kensley & Schotte, 1989: 215; CIBIMA, 1994: 328
Dynameniscus Richardson, 1905
Dynameniscus carinatus (Richardson, 1900). Kensley & Bruce, 2001: 191
Paracerceis Hansen, 1905
Paracerceis caudata (Say, 1818). Kensley & Schotte, 1989: 219
Paracerceis edithae Boone, 1930: 51. Kensley & Schotte, 1989: 221 *H > TL: Gonave Bay
Paradella Harrison & Holdich, 1982
Paradella quadripunctata (Menzies & Glynn, 1968). Kensley & Schotte, 1989: 224
Sphaeroma Latreille, 1802
Sphaeroma terebrans Bates, 1866. CIBIMA, 1994: 328

Superorder EUCARIDA

Taxonomic summary:
Order Decapoda
Suborder Dendrobranchiata

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 33

Superfamily Penaeoidea
Family Aristeidae
Family Penaeidae
Family Solenoceridae
Suborder Pleocyemata
Infraorder Stenopodidea
Family Stenopodidae
Infraorder Caridea
Superfamily Alpheoidea
Family Alpheidae
Family Hippolytidae
Family Ogyridae
Superfamily Atyoidea
Family Atyidae
Family Xiphocarididae
Superfamily Palaemonoidea
Family Gnathophyllidae
Family Palaemonidae
Infraorder Pasiphaeoidea
Family Pasiphaeidae
Infraorder Stylodactyloidea
Family Stylodactylidae
Infraorder Astacidea
Superfamily Enoplometopoidea
Family Enoplometopidae
Superfamily Nephropoidea
Family Nephropidae
Superfamily Astacoidea
Family Cambaridae
Infraorder Axiidea
Family Callianassidae
Infraorder Gebiidea
Family Laomediidae
Family Upogebiidae
Infraorder Achelata
Family Palinuridae
Family Scyllaridae
Infraorder Anomura
Family Albuneidae
Family Coenobitidae
Family Diogenidae
Family Galatheidae
Family Hippidae
Family Pylochelidae
Family Porcellanidae
Infraorder Brachyura
Family Aethridae
Family Calappidae
Family Cancridae
Family Dromiidae
Family Gecarcinidae
Family Goneplacidae
Family Grapsidae
Family Homolidae

34 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

 Family Leucosiidae
Family Majidae
Family Ocypodidae
Family Parthenopidae
Family Pinnotheridae
Family Portunidae
Family Pseudothelphusidae
Family Rhynchocinetidae
Family Xanthidae

Order DECAPODA
Suborder DENDROBRANCHIATA
Superfamily PENAEOIDEA
Family ARISTEIDAE
Hepomadus Bate, 1881
Hepomadus tener Smith, 1894. Pérez Farfante, 1973: 442; D’Incao, 1998: 1511

Family PENAEIDAE
Eusicyonia Stebbing, 1914
Eusicyonia sp.—Mateo & Rodríguez (1986: 153) [As Sicyonia]
Farfantepenaeus Burukovskii, 1971
Farfantepenaeus aztecus (Ives, 1891). CIBIMA, 1994: 329
Farfantepenaeus duorarum notialis Pérez Farfante, 1967. Pérez Farfante, 1969: 509
Farfantepenaeus subtilis (Pérez Farfante, 1967: 87)
Litopenaeus Pérez Farfante, 1969
Litopenaeus schmittii (Burkenroad, 1936). Pérez Farfante, 1969: 488; CIBIMA, 1994: 329
Litopenaeus vannamei (Boone, 1931). SEMARENA, 2004: 138
Metapenaeopsis Bouvier, 1905
Metapenaeopsis martinella Pérez Farfante, 1971: 16
Metapenaeopsis smithi Pérez Farfante, 1971: 29
Penaeopsis Bate, 1881
Penaeopsis serrata Bate, 1881. Pérez Farfante, 1979: 748
Xiphopeneus Smith, 1869
Xiphopeneus kroyeri (Heller, 1862). CIBIMA, 1994: 329

Family SOLENOCERIDAE
Hadropenaeus Pérez Farfante, 1977
Hadropenaeus affinis (Bouvier, 1906). Pérez Farfante, 1977: 317
Mesopenaeus Pérez Farfante, 1977
Mesopenaeus tropicalis (Bouvier, 1905). Pérez Farfante, 1977: 332
Pleoticus Bate, 1888
Pleoticus robustus (Smith, 1885). Pérez Farfante, 1977: 297
Solenocera Lucas, 1849
Solenocera acuminata Pérez Farfante & Bullis, 1973: 8
Solenocera atlantidis Burkenroad, 1939. Pérez Farfante & Bullis, 1973: 20

Suborder PLEOCYEMATA
Infraorder STENOPODIDEA
Family STENOPODIDAE
Stenopus Latreille, 1819
Stenopus hispidus (Olivier, 1811). Williams et al., 1983: 46
Stenopus scutellatus (Rankin, 1898). Williams et al., 1983: 46

Infraorder CARIDEA

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 35

Superfamily ALPHEOIDEA
Family ALPHEIDAE
Alpheopsis Coutière, 1896
Alpheopsis labis Chace, 1972: 55
Alpheus Bosc, 1802
Alpheus angulatus McClure, 1995. Nizinski, 2003: 105
Alpheus armatus Rathbun, 1901. Williams et al., 1983: 46; CIBIMA, 1994: 330
Alpheus estuariensis Christoffersen, 1984. Nizinski, 2003: 105
Alpheus immaculatus Knowlton & Keller, 1983: 354
Alpheus platycheirus Boone, 1930: 49
Alpheus polystictus Knowlton & Keller, 1985: 894
Alpheus rathbunae (Schmitt, 1924). Armstrong, 1949: 13; CIBIMA, 1994: 330
Alpheus viridari (Armstrong, 1949: 8). Chace, 1972: 73; CIBIMA, 1994: 330; Anker, 2012: 85
Parabetaeus Coutière, 1896
Parabetaeus hummelincki (Schmitt, 1936). Anker, 2007: 34
Salmoneus Holthuis, 1955
Salmoneus hispaniolensis Anker, 2010: 178 *DR > TL: Boca Chica
Synalpheus Bate, 1888
Synalpheus antillensis Coutière, 1909. Anker et al., 2012: 16
Synalpheus barahonensis Armstrong, 1949: 20. Chace, 1972: 86 *DR > TL: From live colony of Agaricia agaricites behind
Piedra Prieta reef, Barahona Harbor
Synalpheus bousfieldi Chace, 1972. Markham, 1985: 129
Synalpheus cf. africanus Crosnier & Forest, 1965. Anker et al., 2012: 9
Synalpheus disparodigitus Armstrong, 1949: 15. Chace, 1972: 92
Synalpheus dominicensis Armstrong, 1949: 23. Chace, 1972: 92; Anker et al., 2012: 37 *DR > TL: Quiet water inside Piedra
Prieta Reef, Barahona Harbor
Synalpheus filidigitus Armstrong, 1949: 15. Chace, 1972: 92; Anker et al., 2012: 40
Synalpheus fritzmuelleri Coutière, 1909. Anker et al., 2012: 41
Synalpheus hemphilli Coutière, 1909. Markham, 1985: 129
Synalpheus kensleyi (Ríos & Duffy, 2007). Anker et al., 2012: 59
Synalpheus longicarpus (Herrick, 1891). Markham, 1985: 130; CIBIMA, 1994: 330
Synalpheus ul (Ríos & Duffy, 2007). Anker et al., 2012: 72

Family HIPPOLYTIDAE
Calliasmata Holthuis, 1973
Calliasmata rimolii Chace, 1975: 37 *DR > TL: Cave 4 km from town of Estero Hondo, Prov. Puerto Plata
Lysmata Risso, 1816
Lysmata amboinensis (De Man, 1888). Williams et al., 1983: 46; CIBIMA, 1994: 329
Lysmata ankeri Rhyne & Lin, 2006: 179
Lysmata grabhami Gordon, 1935. Williams et al., 1983: 46; CIBIMA, 1994: 329
Lysmata rathbunae Chace, 1970: 59

Family OGYRIDIDAE
Ogyrides Stebbing, 1914
Ogyrides yaquiensis Armstrong, 1949: 3. Chace, 1972: 106 *DR > TL: Taken on grass banks 1–3 feet deep near mouth of Rio
Yaque del Sur

Superfamily ATYOIDEA
Family ATYIDAE
Atya Leach, 1816
Atya innocuous (Herbst, 1872). CIBIMA, 1994: 329
Atya lanipes Holthuis, 1963. CIBIMA, 1994: 329
Atya scabra (Leach, 1815). CIBIMA, 1994: 329
=Atya punctata Kingsley, 1878. Chace & Hobbs, 1969: 63

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Jonga Hart, 1961
Jonga serrei (Bouvier, 1909). CIBIMA, 1994: 329
Micratya Bouvier, 1913
Micratya poeyi (Guérin-Méneville, 1855). CIBIMA, 1994: 329
Potimirin Holthuis, 1954
Potimirin mexicana (Saussure, 1857). CIBIMA, 1994: 325
Typhlatya Creaser, 1936
Typhlatya monae Chace, 1954. Chace, 1975: 29

Family XIPHOCARIDIDAE
Xiphocaris Martens, 1872
Xiphocaris elongata (Guérin-Méneville, 1855). CIBIMA, 1994: 329

Superfamily PALAEMONOIDEA
Family GNATHOPHYLLIDAE
Gnathophyllum Desmarest, 1823
Gnathophyllum americanum Guérin-Méneville, 1855. Williams et al., 1983: 46; CIBIMA, 1994: 330

Family PALAEMONIDAE
Brachycarpus Bate, 1888
Brachycarpus biunguiculatus (Lucas, 1846). Rodríguez & Mateo, 1983: 19; Mateo & Rodríguez, 1986: 153; CIBIMA, 1994:
252
Brachycarpus sp.—CIBIMA (1994: 329)
Macrobrachium Bate, 1888
Macrobrachium acanthurus (Wiegmann, 1836). CIBIMA, 1994: 329
Macrobrachium carcinus (Linnaeus, 1758). CIBIMA, 1994: 329
Macrobrachium crenulatum Holthuis, 1950. CIBIMA, 1994: 329
Macrobrachium faustinum (Saussure, 1857). Chace, 1972: 20; CIBIMA, 1994: 329
Macrobrachium heterochirus (Wiegmann, 1836). CIBIMA, 1994: 329
Macrobrachium lucifugum Holthuis, 1974. Hobbs et al., 1977: 150
=Macrobrachium crybelum Chace, 1975: 30
Macrobrachium olfersi (Wiegmann, 1836). Markhan, 1985: 128
Macrobrachium rosenbergii (De Man, 1879). Cicero, 1981: 106 [Purposely introduced]
Periclimenes Costa, 1844
Periclimenes pedersoni Chace, 1958. Williams et al., 1983: 46; CIBIMA, 1994: 325
Periclimenes perlatus (Boone, 1930: 45). Chace, 1972: 29 *H > TL: Gonave Bay
Periclimenes yucatanicus (Ives, 1891). CIBIMA, 1994: 325; Williams et al., 1983: 46

Infraorder PASIPHAEOIDEA
Family PASIPHAEIDAE
Pasiphaea Savigny, 1816
Pasiphaea merriami Schmitt, 1931. Hayashi, 2004: 344
=Pasiphaea nishei Iwasaki, 1990. Hayashi, 2004: 344

Infraorder STYLODACTYLOIDEA
Family STYLODACTYLIDAE
Stylodactylus A. Milne Edwards, 1881
Stylodactylus rectirostris A. Milne Edwards, 1883. Cleva, 1990: 168

Infraorder ASTACIDEA
Superfamily ENOPLOMETOPOIDEA
Family ENOPLOMETOPIDAE
Enoplometopus A. Milne Edwards, 1862
Enoplometopus (Hoplometopus) antillensis Lütken, 1865. Borrell, 1981: 93

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 37

Superfamily NEPHROPOIDEA
Family NEPHROPIDAE
Nephrops Leach, 1814
Nephrops aequus‡ Rathbun, 1920: 382 [Fossil]
Nephrops maoensis‡ Rathbun, 1920: 381 [Fossil]
Nephropsis Wood-Mason, 1873
Nephropsis agassizii A. Milne Edwards, 1880. Holthuis, 1974: 796
Nephropsis neglecta Holthuis, 1974: 792

Superfamily ASTACOIDEA
Family CAMBARIDAE
Procambarus Ortmann, 1905
Procambarus (Scapulicambarus) clarkii (Girard, 1852). Cicero, 1981: 106; Hobbs, 1989: 76 [Purposely introduced]

Infraorder AXIIDEA
Family CALLIANASSIDAE
Callianassa Leach, 1814
Callianassa branneri Rathbun, 1900. CIBIMA, 1994: 330
Callianassa latigidita‡ Rathbun, 1918: 165 [Fossil]
Callianassa miocenica‡ Rathbun, 1918: 166 [Fossil]
Callichirus Stimpson, 1866
Callichirus placida (De Man, 1928). CIBIMA, 1994: 330

Infraorder GEBIIDEA
Family LAOMEDIIDAE
Naushonia Kingsley, 1897
Naushonia portoricensis (Rathbun, 1901). Dworschak et al., 2006: 2; Komai & Anker, 2015: 348

Family UPOGEBIIDAE
Upogebia Leach, 1814
Upogebia casis Williams, 1993: 27
Upogebia omissa Gomez-Corrêa, 1968. Williams, 1993: 54

Infraorder ACHELATA
Family PALINURIDAE
Palinurellus Martens, 1878
Palinurellus gundlachii Martens, 1878. CIBIMA, 1994: 330
Panulirus Fabricius, 1798
Panulirus argus (Latreille, 1804). Williams et al., 1983: 46; CIBIMA, 1994: 330
Panulirus guttatus (Latreille, 1804). Williams et al., 1983: 46; CIBIMA, 1994: 330
Panulirus laevicauda (Latreille, 1817). CIBIMA, 1994: 330

Family SCYLLARIDAE
Parribacus Dana, 1852
Parribacus antarcticus (Lund, 1793). CIBIMA, 1994: 330
Scylla de Haan, 1833
Scylla costata‡ Rathbun, 1918: 170. Luque et al., 2017: 60 [Fossil]
Scyllarides Gill, 1898
Scyllarides aequinoctialis (Lund, 1793). CIBIMA, 1994: 330

Infraorder ANOMURA
Family ALBUNEIDAE
Albunea Weber, 1795

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Albunea gibbesi Stimpson, 1859. Boyko, 2002: 359
Albunea paretii Guérin-Méneville, 1853. Boyko, 2002: 328
Lepidopa Stimpson, 1858
Lepidopa richmondi Benedict, 1903. Boyko, 2002: 151
Lepidopa sp.—Williams et al. (1983: 46); CIBIMA, 1994: 332
Paraleucolepidopa Calado, 1996
Paraleucolepidopa distincta (Gomes Corrêa, 1968). Boyko, 2002: 86

Family COENOBITIDAE
Coenobita Latreille, 1829
Coenobita clypeatus (Fabricius, 1787). CIBIMA, 1994: 330

Family DIOGENIDAE
Allodardanus Haig & Provenzano, 1965
Allodardanus bredini Haig & Provenzano, 1965. CIBIMA, 1994: 330
Calcinus Dana, 1851
Calcinus tibicen (Herbst, 1791). Mateo, 1993: 76; CIBIMA, 1994: 330
Clibanarius Dana, 1852
Clibanarius antillensis Stimpson, 1862. Mateo, 1993: 76; CIBIMA, 1994: 330
Clibanarius cubensis (Saussure, 1858). CIBIMA, 1994: 330
Clibanarius tricolor (Gibbes, 1850). Mateo, 1993: 76; CIBIMA, 1994: 330
Clibanarius vittatus (Bosc, 1802). CIBIMA, 1994: 330
Dardanus Paulson, 1875
Dardanus venosus (H. Milne Edwards, 1848). Mateo, 1993: 76; CIBIMA, 1994: 330
Dardanus squamatus‡ Collins in Collins, Portell & Donovan, 2009: 91. Luque et al., 2017: 59 [Fossil]
Iridopagurus de St. L.-Dechance, 1966
Iridopagurus margaritensis García-Gómez, 1983: 28.
Iridopagurus reticulatus García-Gómez, 1983: 37. Nizinski, 2003: 119
Paguristes Dana, 1851
Paguristes cadenati Forest, 1954. CIBIMA, 1994: 330
Paguristes grayi Benedict, 1901. Provenzano, 1959: 387; Mateo, 1993: 76; CIBIMA, 1994: 330
Paguristes sericeus A. Milne Edwards, 1880. Mateo, 1986: 4
Paguristes tortugae Schmitt, 1933. Mateo, 1988: 7
Pagurus Fabricius, 1775
Pagurus annulipes (Stimpson, 1862). Mateo, 1985b: unpaginated
Pagurus brevidactylus (Stimpson, 1859). Mateo, 1993: 76
Pagurus criniticornis (Dana, 1852). Mateo, 1993: 76
Pagurus longimanus Wass, 1963. Mateo, 1993: 76
Pagurus provenzanoi Forest & de Saint Laurent, 1967. Mateo, 1985b: unpaginated
Petrochirus Stimpson, 1858
Petrochirus bahamensis (Herbst, 1796). Alvarez, 1981: 71
Petrochirus diogenes (Linnaeus, 1758). Williams et al., 1983: 46; Mateo, 1993: 76; CIBIMA, 1994: 330
Petrochirus inaequalis‡ Rathbun, 1918: 167. Luque et al., 2017: 59 [Fossil]
Pylopagurus A. Milne Edwards & Bouvier, 1893
Pylopagurus operculatus (Stimpson, 1862). Mateo, 1985b: unpaginated

Family GALATHEIDAE
Munidopsis Whiteaves, 1874
Munidopsis bradleyi Pequegnat & Pequegnat, 1971. Baba et al. 2008: 135
Munidopsis platirostris (A. Milne Edwards & Bouvier, 1894). Navas et al., 2003: 209
Munidopsis riveroi Chace, 1939. Pequegnat & Pequegnat, 1971: 21

Family HIPPIDAE
Emerita Gronovius, 1764

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 39

Emerita portoricensis Schmitt, 1935. Efford, 1976: 178

Family PYLOCHELIDAE
Pylocheles A. Milne Edwards, 1880
Pylocheles (Bathycheles) cubensis Ortíz & Gómez, 1986. McLaughlin et al., 2007: 42
=Pylocheles (Bathycheles) chacei Forest, 1987: 66

Family PORCELLANIDAE
Pachycheles Stimpson, 1858
Pachycheles ackleianus A. Milne Edwards, 1880. Gore, 1974: 705
Pachycheles rugimanus A. Milne Edwards, 1880. CIBIMA, 1994: 330
Petrolisthes Stimpson, 1858
Petrolisthes armatus (Gibbes, 1850). CIBIMA, 1994: 331
Petrolisthes hispaniolensis Werding & Hiller, 2005: 1186 *DR > TL: Las Terrenas, Samaná Peninsula, rocky shore, west of
Playa Las Ballenas
Petrolisthes politus (Gray, 1831). CIBIMA, 1994: 330
Petrolisthes quadratus Benedict, 1901. CIBIMA, 1994: 331
Porcellana Mueller, 1766
Porcellana sayana (Leach, 1820). Gore, 1974: 715; CIBIMA, 1994: 331
Porcellana sigsbeiana A. Milne Edwards, 1880. CIBIMA, 1994: 331

Infraorder BRACHYURA
Family AETHRIDAE
Hepatus Latreille, 1802
Hepatus ephelipticus (Linnaeus, 1763). Williams, 1984: 279; CIBIMA, 1994: 331; Nizinski, 2003: 126
Hepatus guraboensis‡ Collins in Collins, Portell & Donovan, 2009: 94. Luque et al., 2017: 59 [Fossil]
Hepatus pudibundus (Herbst, 1785). CIBIMA, 1994: 331

Family CALAPPIDAE
Calappa Weber, 1795
Calappa flammea‡ (Herbst, 1794). Rathbun, 1918: 169; Luque et al., 2017: 59 [Fossil]
Calappa flammea (Herbst, 1794). CIBIMA, 1994: 331
Calappa gallus (Herbst, 1803). CIBIMA, 1994: 331
Calappa ocellatus Holthuis, 1958: 158. CIBIMA, 1994: 331
Calappa sp.‡—Luque et al. (2017: 59) [Fossil]
Calappella Rathbun, 1918
Calappella (?) sp.‡—Rathbun (1918: 169) [Fossil]
Cryptosoma Brullé, 1839
Cryptosoma bairdii‡ (Stimpson, 1862). Rathbun, 1918: 169 Luque et al., 2017: 59 [Fossil]
Cryptosoma bairdii (Stimpson, 1862). CIBIMA, 1994: 331 (as Cycloes)
Osachila Stimpson, 1870
Osachila tuberosa Stimpson, 1871. CIBIMA, 1994: 331

Family CANCRIDAE
Cancer Linnaeus, 1758
Cancer irroratus Say, 1817. Drew, 2011: 241

Family DROMIIDAE
Hypoconcha Guérin-Méneville, 1854
Hypoconcha sabulosa (Herbst, 1799). Holthuis & Manning, 1987: 1022

Family GECARCINIDAE
Cardisoma Latreille, 1824
Cardisoma guanhumi Latreille, 1825. Chace, 1969: 195; CIBIMA, 1994: 333

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Gecarcinus Leach, 1814
Gecarcinus lateralis (Freminville, 1835). CIBIMA, 1994: 333
Gecarcinus ruricola (Linnaeus, 1758). CIBIMA, 1994: 333

Family GONEPLACIDAE
Frevillea A. Milne Edwards, 1881
Frevillea hirsuta (Borradaile, 1916). CIBIMA, 1994: 333

Family GRAPSIDAE
Aratus H. Milne Edwards, 1853
Aratus pisonii (H. Milne Edwards, 1837). CIBIMA, 1994: 333
Armases Abele, 1992
Armases miersii (Rathbun, 1897). Abele, 1992: 43; CIBIMA, 1994: 333
Armases ricordi (H. Milne Edwards, 1853). Abele, 1992: 32; CIBIMA, 1994: 333
Armases roberti (H. Milne Edwards, 1853). Abele, 1992: 39; CIBIMA, 1994: 333
Cyclograpsus H. Milne Edwards, 1837
Cyclograpsus integer H. Milne Edwards, 1837. CIBIMA, 1994: 333
Euchirograpsus H. Milne Edwards, 1853
Euchirograpsus americanus A. Milne Edwards, 1880. CIBIMA, 1994: 333
Geograpsus Stimpson, 1858
Geograpsus lividus (H. Milne Edwards, 1837). CIBIMA, 1994: 333
Goniopsis de Haan, 1835
Goniopsis cruentata (Latreille, 1802). CIBIMA, 1994: 333
Grapsus Lamarck, 1801
Grapsus grapsus (Linnaeus, 1758). CIBIMA, 1994: 333
Metasesarma H. Milne Edwards, 1853
Metasesarma rubripes? Rathbun, 1897. CIBIMA, 1994: 333
Metopaulias Rathbun, 1896
Metopaulias depressus Rathbun, 1896. CIBIMA, 1994: 333
Pachygrapsus Randall, 1840
Pachygrapsus gracilis (Saussure, 1858). CIBIMA, 1994: 333
Pachygrapsus transversus (Gibbes, 1850). CIBIMA, 1994: 333
Percnon Gistl, 1848
Percnon gibbesi (H. Milne Edwards, 1853). CIBIMA, 1994: 333
Plagusia Latreille, 1804
Plagusia depressa (Fabricius, 1775). Chace & Hobbs, 1969: 192; CIBIMA, 1994: 333
Planes Bowdich, 1825
Planes minutus (Linnaeus, 1758). Chace, 1951: 86

Family HOMOLIDAE
Homola Leach, 1815
Homola barbata (Fabricius, 1793). CIBIMA, 1994: 331

Family LEUCOSIIDAE
Iliacantha Stimpson, 1871
Iliacantha sp.‡—Luque et al. (2017: 60) [Fossil]
Iliacantha subglobosa Stimpson, 1871. CIBIMA, 1994: 331
Myropsis Stimpson, 1871
Myropsis quinquespinosa Stimpson, 1871. García, 1980: 2; CIBIMA, 1994: 331
Persephona Leach, 1817
Persephona finneganae Rathbun, 1933. CIBIMA, 1994: 331
Persephona prepunctata‡ Rathbun, 1919: 170. Luque et al., 2017: 60 [Fossil]
Persephona punctata (Linnaeus, 1758). CIBIMA, 1994: 331

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 41

Family MAJIDAE
Acanthonyx Latreille, 1827
Acanthonyx petiverii H. Milne Edwards, 1834. García, 1983: 434; CIBIMA, 1994: 331
Collodes Stimpson, 1860
Collodes trispinosus (Stimpson, 1851). CIBIMA, 1994: 331
Epialtus Edwards, 1834
Epialtus bituberculatus (H. Milne Edwards, 1834). CIBIMA, 1994: 331
Macrocoeloma Miers, 1879
Macrocoeloma diplacanthum (Stimpson, 1860). CIBIMA, 1994: 331
Microphrys H. Milne Edwards, 1851
Microphrys antillensis Rathbun, 1920. CIBIMA, 1994: 331
Microphrys bicornutus (Latreille, 1825). CIBIMA, 1994: 331
Mithrax Edwards, 1834
Mithrax (Mithraculus) cinctimanus (Stimpson, 1860). CIBIMA, 1994: 331
Mithrax (Mithraculus) coryphe (Herbst, 1801). CIBIMA, 1994: 331
Mithrax (Mithraculus) forceps A. Milne Edwards, 1875. CIBIMA, 1994: 331
Mithrax (Mithraculus) ruber (Stimpson, 1871). CIBIMA, 1994: 332
Mithrax (Mithraculus) sculptus (Lamarck, 1818). CIBIMA, 1994: 332
Mithrax (Mithrax) acuticornis Stimpson, 1870. CIBIMA, 1994: 331
Mithrax (Mithrax) caribbeus Rathbun, 1920. CIBIMA, 1994: 331
Mithrax (Mithrax) pleuracanthus Stimpson, 1871. CIBIMA, 1994: 332
Mithrax (Mithrax) spinossisimus (Lamarck, 1818). CIBIMA, 1994: 332
Mithrax (Mithrax) verrucosus H. Milne Edwards, 1832. CIBIMA, 1994: 332
Mithrax sp.‡—Luque et al. (2017: 61) [Fossil]
Pelia Bell, 1836
Pelia mutica (Gibbes, 1850). CIBIMA, 1994: 332
Pitho Bell, 1836
Pitho aculeata (Gibbes, 1850). CIBIMA, 1994: 332
Pitho therminieri (Schramm, 1867). CIBIMA, 1994: 332
Podochela Stimpson, 1860
Podochela gracilipes Stimpson, 1871. CIBIMA, 1994: 332
Stenocionops Leach, 1823
Stenocionops furcata (Oliver, 1791). CIBIMA, 1994: 332
Stenocionops spinosissima (Saussure, 1857). Nizinski, 2003: 1132
Stenorhynchus Lamarck, 1818
Stenorhynchus seticornis (Herbst, 1788). Williams et al., 1983: 46; García, 1983: 434; CIBIMA, 1994: 332
Teleophrys Stimpson, 1860
Teleophrys beebei Boone, 1930: 42

Family OCYPODIDAE
Ocypode Weber, 1795
Ocypode quadrata (Fabricius, 1787). CIBIMA, 1994: 334
Sandomingia‡ Rathbun, 1919
Sandomingia yaquiensis‡ Rathbun, 1919: 180. Luque et al., 2017: 60 [Fossil]
Uca Leach, 1814
Uca (Leptuca) leptodactyla Rathbun, 1898. CIBIMA, 1994: 334
Uca (Leptuca) pugilator (Bosc, 1802). Williams, 1984: 475
Uca (Minuca) burguersi Holthuis, 1967. CIBIMA, 1994: 333; Arruda Bezerra, 2012: 216
Uca (Minuca) pugnax (Smith, 1870). CIBIMA, 1994: 334
Uca (Minuca) rapax (Smith, 1870). CIBIMA, 1994: 334
Uca (Minuca) thayeri Rathbun, 1900. CIBIMA, 1994: 334
Uca (Minuca) vocator (Herbst, 1782). CIBIMA, 1994: 334
Ucides Rathbun, 1897
Ucides cordatus (Linnaeus, 1763). CIBIMA, 1994: 334

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Family PARTHENOPIDAE
Mesorhoea Stimpson, 1870
Mesorhoea mauryae‡ Rathbun, 1919: 181. Luque et al., 2017: 60 [Fossil]
Parthenope Weber, 1795
Parthenope agona Stimpson, 1871. CIBIMA, 1994: 332
Parthenope granulata (Kingsley, 1879). CIBIMA, 1994: 332
Parthenope serrata (H. Milne Edwards, 1834). CIBIMA, 1994: 332
Parthenope sp. indet.‡—Rathbun (1918: 181); Rathbun (1923: 5) [Fossil]
Platylambrus Stimpson, 1871
Platylambrus obscura‡ Rathbun, 1919: 180. Luque et al., 2017: 60 [Fossil]

Family PINNOTHERIDAE
Dissodactylus Smith, 1870
Dissodactylus mellitae (Rathbun, 1900). Mateo, 1981b: 37/81
Zaops Rathbun, 1900
Zaops ostreum (Say, 1817). CIBIMA, 1994: 333

Family PORTUNIDAE
Achelous de Haan, 1833
Achelous sebae (H. Milne Edwards, 1834). CIBIMA, 1994: 332
Arenaeus Dana, 1851
Arenaeus cribrarius (Lamarck, 1818). CIBIMA, 1994: 332
Callinectes Stimpson, 1860
Callinectes danae Smith, 1869. CIBIMA, 1994: 332
Callinectes declivis‡ Rathbun, 1918: 174. Luque et al., 2017: 60 [Fossil]
Callinectes exasperatus (Gerstaecker, 1856). CIBIMA, 1994: 312
=Callinectes tumidus Ordway, 1863. Rathbun, 1896: 359
Callinectes larvatus Ordway, 1863. Williams, 1984: 371; CIBIMA, 1994: 332
Callinectes ornatus Ordway, 1863. Rathbun, 1896: 356; Williams, 1984: 373; CIBIMA, 1994: 332
Callinectes sapidus Rathbun, 1896. CIBIMA, 1994: 332
Callinectes sp. indet.‡—Rathbun (1918: 174) [Fossil]
Cronius Stimpson, 1860
Cronius tumidulus Stimpson, 1871. CIBIMA, 1994: 332
Euphylax Stimpson, 1860
Euphylax domingensis‡ (Rathbun, 1919: 175). Schweizer et al., 2006: 126. Luque et al., 2017: 60 [Fossil]
Lupella Rathbun, 1897
Lupella forceps (Fabricius, 1793). CIBIMA, 1994: 332
Neocallichirus Sakai, 1988
Neocallichirus? quisquellanus‡ Schweitzer et al., 2006: 115 [Fossil]
Portunus Weber, 1795
Portunus anceps (Saussure, 1858). CIBIMA, 1994: 332
Portunus depressifrons (Stimpson, 1859). CIBIMA, 1994: 332
Portunus gabbi‡ Rathbun, 1919: 172. Luque et al., 2017: 60 [Fossil]
Portunus haitensis‡ Rathbun, 1923: 1. Luque et al., 2017: 62 [Fossil]
Portunus oblongus‡ Rathbun, 1920: 383. Luque et al., 2017: 60 [Fossil]
Portunus ordwayi (Stimpson, 1860). CIBIMA, 1994: 332
Portunus sayi (Gibbes, 1850). CIBIMA, 1994: 332
Portunus spinicarpus (Stimpson, 1871). CIBIMA, 1994: 332
Portunus spinimanus Latreille, 1819. CIBIMA, 1994: 332
Portunus sp. indet.‡—Rathbun (1918: 174) [Fossil]
Portunus sp. 1‡—Schweizer et al. (2006: 121) [Fossil]
Portunus sp. 3‡—Schweizer et al. (2006: 121) [Fossil]
Portunus tenuis‡ Rathbun, 1919: 173. Luque et al., 2017: 60 [Fossil]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 43

Psygmophthalmus‡ Schweitzer, Iturralde-Vinent, Hetler & Velez-Juarbe, 2006
Psygmophthalmus bifurcatus‡ Collins in Collins, Portell & Donovan, 2009: 103. Luque et al., 2017: 60
Rathbunites‡ Schweitzer, Dworschak & Martin, 2011
=Rathbunella‡ Collins in Collins, Portell & Donovan, 2009 [Preoccupied]
Rathbunites pentaspinosa‡ (Collins in Collins, Portell & Donovan, 2009: 96). Luque et al., 2017: 60 [Fossil]

Family PSEUDOTHELPHUSIDAE
Epilobocera Stimpson, 1860
Epilobocera haytensis Rathbun, 1893: 658. Rodríguez, 1982: 27; CIBIMA, 1994: 333 *H, *DR > TL: Hayti
Epilobocera wetherbeei Rodríguez & Williams, 1995: 77 *DR > TL: Prov. Santiago, Rio Magua, tributary of Rio Mao, Sierra
Platicos, northern slopes of the Cordillera Central, 2300 m

Family RHYNCHOCINETIDAE
Cinetorhynchus Holthuis, 1995
Cinetorhynchus rigens (Gordon, 1936). Williams et al., 1983: 46; CIBIMA, 1994: 329

Family XANTHIDAE
Carpilius Desmarest, 1823
Carpilius corallinus Herbst, 1783. Williams et al., 1983: 46; CIBIMA, 1994: 332
Eriphia Latreille, 1817
Eriphia gonagra (Fabricius, 1781). CIBIMA, 1994: 332
Eurypanopeus A. Milne Edwards, 1880
Eurypanopeus abbreviatus (Stimpson, 1860). García, 1983: 438
Eurypanopeus sp. aff. E. crenatus‡ H. Milne-Edwards, 1834. Luque et al., 2017: 61 [Fossil]
Eurytium Stimpson, 1859
Eurytium limosum (Say, 1818). CIBIMA, 1994: 332
Glyptoxanthus A. Milne Edwards, 1879
Glyptoxanthus erosus (Stimpson, 1859). CIBIMA, 1994: 332
Lobonotus Uhler, 1877
Lobonotus sculptus‡ (A. Milne Edwards, 1864: 279). Luque et al., 2017: 60 [Fossil]
=Archaeopilumnus caelatus‡ Rathbun, 1919: 177 [Fossil]
Menippe de Haan, 1833
Menippe nodifrons Stimpson, 1859. CIBIMA, 1994: 332
Micropanope Stimpson, 1870
Micropanope nuttingi (Rathbun, 1898). CIBIMA, 1994: 333
Neopanope A. Milne Edwards, 1880
Neopanope? sp.—CIBIMA (1992: 333)
Ozius Desmarest, 1825
Ozius reticulatus (Desbonne & Schramm, 1867). García, 1983: 438; CIBIMA, 1994: 333
Panopeus H. Milne Edwards, 1834
Panopeus herbsti H. Milne Edwards, 1834. CIBIMA, 1994: 333
Panopeus sp.‡—Rathbun (1918: 179); Rathbun (1923: 2); Luque et al. (2017: 61) [Fossil]
Pilumnus Dejean, 1821
Pilumnus caribbeus Desbonne & Schramm, 1867. CIBIMA, 1994: 333
Pilumnus floridanus Stimpson, 1871. CIBIMA, 1994: 333
Pilumnus levimanus? Rathbun, 1933. CIBIMA, 1994: 333
Pilumnus sayi Rathbun, 1897. CIBIMA, 1994: 333
Pilumnus sp.‡—Collins, Portell & Donovan (2009: 102); Luque et al. (2017: 61) [Fossil]
Pilumnus subequus‡ Rathbun, 1919: 177. Luque et al., 2017: 61 [Fossil]
Xantho Leach, 1815
Xantho denticulata White, 1847. CIBIMA, 1994: 333
Zanthopsis M’Coy, 1849
Zanthopsis sp.‡—Rathbun (1923: 3) [Fossil]

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Subphylum CHELICERATA
Class PYCNOGONIDA
Family AMMOTHEIDAE
Ascorhynchus Sars, 1877
Ascorhynchus colei Hedgpeth, 1943. Hedgpeth, 1948: 257; Stock, 1986: 412
Ascorhynchus latipes (Cole, 1906). Hedgpeth, 1948: 256
Eurycyde Schiödte, 1857
Eurycyde raphiaster Loman, 1912. Hedgpeth, 1948: 260; Stock, 1986: 409

Family COLOSSENDEIDAE
Colossendeis Jarschinsky, 1870
Colossendeis colossea Wilson, 1881. Stock, 1986: 418

Family PHOXICHILIDIIDAE
Anoplodactylus Wilson, 1878
Anoplodactylus batangensis (Helfer, 1938). Perez-Gelabert, 2008: 42
Pallenopsis Wilson, 1881
Pallenopsis (Bathypallenopsis) mollissima atlantica Stock, 1986: 434
Pallenopsis (Pallenopsis) childi Stock, 1986: 429
Pallenopsis (Pallenopsis) forficifera Wilson, 1881. Stock, 1986: 425
Pallenopsis (Pallenopsis) schmitti Hedgpeth, 1943. Stock, 1986: 425

Class ARACHNIDA
Subclass ACARI

Taxonomic summary:
Superorder Acariformes
Order Oribatida
Family Carabodidae
Family Eremaeozetidae
Family Fortuyniidae
Family Hermanniellidae
Family Mochlozetidae
Family Neoliodidae
Family Oppiidae
Family Oribatidae
Family Oribotritiidae
Family Oripodidae
Family Otocepheiidae
Family Scutoverticidae
Family Selenoribatidae
Family Trhypochthonidae
Order Trombidiformes
Suborder Prostigmata
Family Bdellidae
Family Caeculidae
Family Eriophyidae
Family Erythraeidae
Family Halacaridae
Family Hydromidae
Family Limnesiidae
Family Pygmephoridae
Family Smarididae
Family Tarsonemidae

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 45

 Family Tenuipalpidae
Family Tetranychidae
Family Trombiculidae
Order Sarcoptiformes
Family Acaridae
Family Atopomelidae
Family Chaetodactylidae
Family Knemidokoptidae
Family Macrodinychidae
Family Psoroptidae
Family Pterolichidae
Superorder Parasitiformes
Order Metastigmata
Family Argasidae
Family Ixodidae
Order Mesostigmata
Family Ascidae
Family Blattisocidae
Family Macronyssidae
Family Melicharidae
Family Opilioacaridae
Family Otopheidomenidae
Family Phytoseiidae
Family Spelaeorhynchidae
Family Trigonuropodidae
Family Uropodidae
Family Varroidae
Order Holothyrida
Family Neothyridae

Superorder ACARIFORMES
Order ORIBATIDA
Family CARABODIDAE
Carabodes Koch, 1835
Carabodes? sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]
Klapperiches Mahunka, 1978
Klapperiches nigrosetosus Mahunka, 1978: 555 *DR > TL: Loma Casabito, 1250 m

Family EREMAEOZETIDAE
Eremaeozetes Berlese, 1913
Eremaeozetes sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family FORTUYNIIDAE
Alismobates Luxton, 1992
Alismobates cf. inexpectatus Pfingst & Schuster, 2012. Pfingstl et al., 2016: 42

Family HERMANNIELLIDAE
Sacculobates Grandjean, 1962
Sacculobates sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family MOCHLOZETIDAE
Mochloribatula Mahunka, 1978
Mochloribatula grandjeani Mahunka, 1978: 563 *DR > TL: Baní, 65 m
Mochloribatula multiporosa Mahunka, 1978: 561 *DR > TL: Loma Casabito, 1250 m

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Mochlozetes Grandjean, 1930
Mochlozetes sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family NEOLIODIDAE
Neoliodes Berlese, 1888
Neoliodes dominicus† Heethoff, Helfen & Norton, 2009: 154 [Fossil in Dominican amber]
=Liodes sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]
Teleoliodes Grandjean, 1934
Teleoliodes sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family OPPIIDAE
Aeroppia Hammer, 1961
Aeroppia insularis Higgins, 1966: 106 *DR > TL: From Artocarpus communis at San Cristóbal
Oppia Koch, 1935
Oppia? sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family ORIBATIDAE
Arphthicarus Niedbala, 1994
Arphthicarus cataulacus Niedbala, 2004: 103
Arphthicarus pedanos Niedbala, 2004: 110 *DR > TL: Jarabacoa, 5 km S of Jarabacoa on Highway 28
Arphthicarus prolixus Niedbala, 2004: 111 *DR > TL: Jarabacoa, 5 km S of Jarabacoa on Highway 28
Atropacarus Ewing, 1917
Atropacarus (Hoplophorella) hamatus (Ewing, 1909). Niedbala, 2004: 135; Niedbala, 2008: 809
Austrophthiracarus Balogh & Mahunka, 1978
Austrophthiracarus valdiviaensis Niedbala, 2008: 730
Heterobelba Berlese, 1913
Heterobelba furcata Mahunka, 1978: 553 *DR > TL: Colonia Ramfis, ca. 1000 m
Hoplophthiracarus Jacot, 1933
Hoplophthiracarus illinoisensis (Ewing, 1909). Niedbala, 2008: 790
Indotritia Jacot, 1928
Indotritia bellingeri Niedbala & Schatz, 1996. Niedbala, 2004: 177
Indotritia krakatauensis (Sellnick, 1923). Niedbala, 2004: 177; Niedbala, 2008: 764
Meristacarus Grandjean, 1934
Meristacarus longisetosus Mahunka, 1978: 551 *DR > TL: Loma Casabito, 1250 m
Mesoplophora Berlese, 1905
Mesoplophora (Mesoplophora) pusilla Schuster, 1962. Niedbala, 2008: 758
Mesoplophora (Parplophora) bacula Niedbala, 2003. Niedbala, 2004: 177
Protophthiracarus Balogh, 1972
Protophthiracarus kyphos Niedbala, 2004. Niedbala, 2008: 805
Rhysotritia Markel & Meyer, 1959
Rhysotritia clavata Markel, 1964. Niedbala, 2004: 177
Rhysotritia comtea Mahunka, 1983. Niedbala, 2004: 53
Rhysotritia parallelos Niedbala, 2003. Niedbala, 2004: 177
Rhysotritia peruensis (Hammer, 1961). Niedbala, 2004: 59
Zygoribatula Berlese, 1916
Zygoribatula heterotricha Mahunka, 1978: 558 *DR > TL: Baní, 65 m

Family ORIBOTRITIIDAE
Oribotritia Jacot, 1924
Oribotritia sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family ORIPODIDAE
Oripoda Banks & Pergande, 1904
Oripoda sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 47

Family OTOCEPHEIIDAE
Dolicheremaeus Jacot, 1938
Dolicheremaeus sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family SCUTOVERTICIDAE
Arthrovertex Balogh, 1970
Arthrovertex baloghi Mahunka, 1978: 556 *DR > TL: Loma Casabito, 1250 m
Arthrovertex sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Family SELENORIBATIDAE
Carinozetes Pfingstl & Schuster, 2012
Carinozetes cf. mangrovi Pfingstl, Lienhard & Jagersbacher-Baumann, 2014. Pfingstl et al., 2016: 42
Thalassozetes Schuster, 1963
Thalassozetes cf. barbara Pfingstl, 2013. Pfingstl et al., 2016: 42

Family TRHYPOCHTHONIDAE
Allonothrus Hammen, 1953
Allonothrus sp.†—Norton & Poinar (1993: 64) [Fossil in Dominican amber]

Order TROMBIDIFORMES
Suborder PROSTIGMATA
Family BDELLIDAE
Bdella Latreille, 1775
Bdella distincta Baker & Balock, 1944. Sánchez Martínez et al., 2014: 548
Probably Bdella sp.†—Wu (1997: 115) [Fossil in Dominican amber]
Hexabdella Van der Schyff et al., 2004
Hexabdella aff. cinquaginta Hernandez, Daud & Feres, 2007. Sánchez Martínez et al., 2014: 548

Family CAECULIDAE
Caeculus Dufour, 1832
Probably Caeculus sp.†—Wu (1997: 115) [Fossil in Dominican amber]
Procaeculus Jacot, 1936
Procaeculus dominicensis† Coineau & Poinar, 2000: 143 [Fossil in Dominican amber]

Family ERIOPHYIDAE
Acalitus Keifer, 1965
Acalitus gossypii (Banks, 1904). Schmutterer, 1990: 301
Aceria Keifer, 1944
Aceria guerreronis (Keifer, 1965). Domínguez, 1981: 110; Schmutterer, 1990: 111; Sánchez Martínez et al., 2014: 548
[Introduced]
Aceria hibisci Nalepa, 1906. Serra et al., 2003: 108 [Introduced]
Aceria sheldoni (Ewing, 1937). Zuluaga & Mesa, 2000: 127 [Introduced]
Aceria tulipae (Keifer, 1938). Reyes, 1994: 8/94; Serra, 2005: 121 [Introduced]
Aculops Keifer, 1966
Aculops lycopersici (Massee, 1937). SEA, 1999: 118
Anthocoptes Nalepa, 1892
Anthocoptes bakeri (Keifer, 1959). Taveras et al., 2000: 111
Calepitrimerus Kiefer, 1938
Calepitrimerus muesebecki Keifer, 1940. SEA, 1999: 66 [Introduced]
Colomerus Newkirk & Keifer, 1971
Colomerus bucidae (Nalepa, 1904). Sánchez Martínez et al., 2014: 548
Eriophyes von Siebold, 1851
Eriophyes annonae Keifer, 1973. Serra et al., 2003: 108 [Introduced]

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Phyllocoptruta Keifer, 1938
Phyllocoptruta oleivora (Ashmead, 1879). Schmutterer, 1990: 176; Zuluaga & Mesa, 2000: 127

Family ERYTHRAEIDAE
Genus near Fallopia Oudemans, 1905
Genus nr. Fallopia sp.†—Poinar (1992: 226) [Fossil in Dominican amber]
Leptus Latreille, 1796
Leptus sp.—Armas & Prieto Trueba (2003: 133); Botosaneanu & Bolland (1997: 71)

Family HALACARIDAE
Rhombognathus Trouessart, 1888
Rhombognathus caribaeus Bartsch, 2007: 18 *DR > TL: Samaná Peninsula, Playa Colibrí, about 1 km west of Las Terrenas,
from dead corals covered with algae

Family HYDROMIDAE
Hydroma Koch, 1837
Hydroma despiciens (O. F. Müller, 1776). Lundblad, 1936: 1

Family LIMNESIIDAE
Arrenurus Dugès, 1834
Arrenurus birgei Marshall, 1903. Lundblad, 1936: 16
Arrenurus megalurus Marshall, 1903. Lundblad, 1936: 21
Arrenurus sp.—Perez-Gelabert & Alba Suriel (2016: 98)
Arrenurus valencius Marshall, 1919. Lundblad, 1936: 24
Atax Fabricius, 1805
Atax gracilipalpis tenuis Lundblad, 1936: 8 *DR > TL: Laguna Rincón
Centrolimnesia Lundblad, 1936
Centrolimnesia bondi Lundblad, 1936: 5 *H > TL: Trou Caiman
Koenikea Wolcott, 1900
Koenikea haitiana Lundblad, 1936: 12 *DR > TL: Laguna Rincón
Limnesia Koch in Panzer, 1835
Limnesia cordifera Daday, 1905. Lundblad, 1936: 2
Neomamersa Lundblad, 1954
Neomamersa haitiana Cook, 1981: 137 *H > TL: Taken by the Karaman-Chappuis method, Rivière Voldrogue at Marché León,
80 m
Pentatax Viets, 1926
Pentatax furcula Lundblad, 1936: 11 *DR > TL: Laguna Rincón
Tubomixdea Cook, 1981
Tubomixdea botosaneanui (Cook, 1981: 142) *H > TL: Taken by the Karaman-Chappuis method, Rivière Voldrogue at Marché
León, 80 m
Tubomixdea voldroguea (Cook, 1981: 140) *H > TL: Taken by the Karaman-Chappuis method, Rivière Voldrogue at Marché
León, 80 m
Voldroguella Cook, 1981
Voldroguella hispaniolae (Cook, 1981: 138) *H > TL: Taken by the Karaman-Chappuis method, Rivière Voldrogue at Marché
León, 80 m

Family PYGMEPHORIDAE
Pygmephorus Kramerr, 1877
Pygmephorus klapperichi Mahunka, 1982: 81 *DR > TL: Colonia Ramfis, ca. 1000 m

Family SMARIDIDAE
Smaris Latreille, 1796
Smaris sp.†—Wu (1997: 115) [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 49

Family TARSONEMIDAE
Phytonemus Lindquist, 1986
Phytonemus pallidus (Banks, 1901). SEA, 1999: 103
Polyphagotarsonemus Beer & Nucifora, 1965
Polyphagotarsonemus latus (Banks, 1904). Schmutterer, 1990: 389; Zuluaga & Mesa, 2000: 197; Sánchez Martínez et al.,
2014: 548 [Introduced]
Steneotarsonemus Beer, 1954
Steneotarsonemus comosus Ochoa in Ochoa et al., 1991. SEA, 1999: 29
Steneotarsonemus spinki Smiley, 1967. Taveras et al., 2000: 111; Zuluaga & Mesa, 2000: 197; Ramos et al., 2001: 6; Hummel
et al., 2009: 551; Sánchez Martínez et al., 2014: 548 [Introduced]

Family TENUIPALPIDAE
Brevipalpus Donnadieu, 1876
Brevipalpus californicus (Banks, 1904). De Leon, 1965: 518
Brevipalpus lewisi McGregor, 1949. SEA, 1999: 111
Brevipalpus mori De Leon, 1960. De Leon, 1965: 518
Brevipalpus phoenicis (Geijskes, 1939). De Leon, 1965: 518; Sánchez Martínez et al., 2014: 548
Brevipalpus sp.—Baker (1979: 355)
Brevipalpus trinidadensis Baker, 1949. Evans et al., 1993: 136; Sánchez Martínez et al., 2014: 549
Brevipalpus yothersi Baker, 1949. Beard et al., 2015: 51
Raoiella Hirst, 1924
Raoiella indica Hirst, 1924. Serra, 2006: 37; Pons & Bliss, 2007: 4; Perez-Gelabert, 2008: 46; Serra et al., 2011: 147; Sánchez
Martínez et al., 2014: 549 [Introduced]
Tenuipalpus Donnadieu, 1876
Tenuipalpus annonae De Leon, 1957. Sánchez Martínez et al., 2014: 549
Tenuipalpus anoplus Baker & Pritchard, 1953. De Leon, 1965: 518
Tenuipalpus bakeri McGregor, 1949. Evans et al., 1993: 146
Tenuipalpus domingensis De Leon, 1965: 521 *DR > TL: San Cristóbal, on Catalpa longissima
Tenuipalpus vexus De Leon, 1965: 520 *DR > TL: Santo Domingo, on Tabebuia sp.

Family TETRANYCHIDAE
Dolichotetranychus Sayed, 1938
Dolichotetranychus floridanus (Banks, 1900). SEA, 1999: 28
Eutetranychus Banks, 1917
Eutetranychus banksi (McGregor, 1950). Zuluaga & Mesa, 2000: 127; Sánchez Martínez et al., 2014: 549
Mononychellus Wainstein, 1971
Mononychellus caribbeanae (McGregor, 1950: 283). Rowland & De la Cruz, 2001: 107; Sánchez Martínez et al., 2014: 549
Panonychus Yokohama, 1929
Panonychus citri (McGregor, 1916). Schmutterer, 1990: 179; Zuluaga & Mesa, 2000: 127
Petrobia Murray, 1877
Petrobia (Tetranychina) hispaniola Sánchez Martínez & Fletchmann in Sánchez Martínez et al., 2014: 550 *DR > TL: Río
Seco, La Vega, from orange (Citrus sinensis)
Tetranychus Dufour, 1832
Tetranychus bimaculatus Harvey, 1892. Wolcott, 1927: 17; Santoro, 1960: 27
Tetranychus cinnabarinus (Boisduval, 1867). SEA, 1999: 119
Tetranychus evansi Baker & Pritchard, 1960. Zuluaga & Mesa, 2000: 197
Tetranychus gloveri Banks, 1900. Schmutterer, 1990: 277; Sánchez Martínez et al., 2014: 549
Tetranychus ludeni Zacher, 1913. SEA, 1999: 123
Tetranychus telarius (Linnaeus, 1758). SEA, 1999: 146
Tetranychus urticae Koch, 1836. Schmutterer, 1990: 388; Sánchez Martínez et al., 2014: 549

Family TROMBICULIDAE
Trombicula Berlese, 1905
Trombicula (Eutrombicula) alfreddugesi (Oudemans, 1920). Jenkins, 1949: 305; Zippel et al., 1996: 43

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Trombicula insularis Ewing, 1925. Ewing, 1931: 8

Order SARCOPTIFORMES
Family ACARIDAE
Acarus Linnaeus, 1758
Acarus siro Linnaeus, 1758. Abud, 1981: 31/81; SEA, 1999: 128
Rhizoglyphus Claperède, 1869
Rhizoglyphus robini Claperède, 1869. Schmutterer, 1990: 475 [Introduced]

Family ATOPOMELIDAE
Cubanochirus Fain, 1970
Cubanochirus sp.—De la Cruz (1980: 11)
Domingoecius Fain & Lukoschus in Fain, 1979
Domingoecius cruzi Fain & Lukoschus in Fain, 1979: 146 *DR > TL: Sur Plagiodontia aedium, Gurabo
Plagiodontochirus Fain, 1976
Plagiodontochirus nudus Fain, 1976: 63. Fain, 1979: 140

Family CHAETODACTYLIDAE
Chaetodactylus Rondani, 1866
Chaetodactylus antillarum Klimov & O’Connor, 2008: 113

Family KNEMIDOKOPTIDAE
Knemidokoptes Fuerstenburg, 1870
Knemidokoptes jamaicensis Turk, 1950. Latta & O’Connor, 2001: 437; Latta, 2003: 730

Family MACRODINYCHIDAE
Macrodinychus Berlese, 1917
Macrodinychus (Macrodinychus) sellnicki Hirschmann & Zirngiebl-Nicol, 1975. Kontschán, 2011: 1623

Family PSOROPTIDAE
Protalges Trouessart & Mégnin, 1883
Protalges sp.—Brea Tió (1985: 7)

Family PTEROLICHIDAE
Aralichus Gaud, 1966
Aralichus nobilis Atyeo, 1988. Atyeo & Perez, 1990: 11

Family SARCOPTIDAE
Chirnyssoides Fain, 1959
Chirnyssoides caparti Fain, 1959. Fain, 1960: 291
Sarcoptes Latreille, 1802
Sarcoptes scabiei (Linnaeus, 1758). Rojas et al. 1984a: 34

Superorder PARASITIFORMES
Order METASTIGMATA
Family ARGASIDAE
Argas Latreille, 1795
Argas (Argas) dulus Keirans et al., 1971: 1411 *DR > TL: From nest of Dulus dominicus, Nisibón, El Cedro
Argas (Persicargas) persicus (Oken, 1818). De la Cruz, 2001: 87
Carios Latreille, 1796
Carios elongatus (Kohls et al., 1965). Morel, 1967: 349; De la Cruz, 2001: 88
Carios hasei (Schulze, 1935). De la Cruz, 2001: 89
Carios marginatus (Banks, 1910). De la Cruz, 2001: 91
Carios puertoricensis (Fox, 1947). Morel, 1967: 342; Endris et al., 1989: 146; De la Cruz, 2001: 89

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 51

Carios viguerasi (Cooley & Kohls, 1941). De la Cruz, 2001: 90
Ornithodorus Agassiz, 1845
Ornithodorus (Alectorobius) sp.†—Estrada-Peña & de la Fuente (2018: 255) [Fossil in Dominican amber]
Ornithodorus antiquus† Poinar, 1995: 385 [Fossil in Dominican amber]

Family IXODIDAE
Amblyomma Koch, 1844
Amblyomma albopictum Neumann, 1899. Morel, 1967: 342
Amblyomma cajennense (Fabricius, 1787). Thompson, 1950: 227
Amblyomma cruciferum Neumann, 1901. Thompson, 1950: 221; Morel, 1967: 342; De la Cruz, 2001: 93
Amblyomma dissimile Koch, 1844. De la Cruz, 2001: 92
Amblyomma quadricavum (Schulze, 1941). Keirans & Klompen, 1996: 164; De la Cruz, 2001: 92
Amblyomma sp.†—Lane & Poinar (1986: 75); Keirans et al. (2002: 61) [Fossil in Dominican amber]
Dermacentor Koch, 1844
Dermacentor nitens (Neumann, 1897). Wolcott, 1927: 22; Thompson, 1950: 221; De la Cruz, 2001: 93
Rhipicephalus Koch, 1844
Rhipicephalus microplus (Canestrini, 1887). Thompson, 1950: 227
Rhipicephalus sanguineus (Latreille 1806). Wolcott, 1927: 22; Thompson, 1950: 227

Order MESOSTIGMATA
Family ASCIDAE
Asca Von Heyden, 1826
Asca elongatula De Leon, 1967. Abo-Shnaf et al., 2016: 614
Asca foxi De Leon, 1967. Abo-Shnaf et al., 2016: 616

Family BLATTISOCIDAE
Lasioseius Berlese, 1916
Lasioseius chaudhrii (Wu & Wang, 1982). De Moraes et al., 2015: 6
Lasioseius dominicensis Abo-Shnaf et al., 2016: 619 *DR > TL: La Palmita, Prov. María Trinidad Sánchez, from L. camara
Lasioseius oryzae Abo-Shnaf et al., 2016: 622 *DR > TL: El Pozo, Prov. María Trinidad Sánchez, from Oryza sativa
Lasioseius prorsoperitrematus Abo-Shnaf et al., 2016: 627 *DR > TL: El Aguacate, Prov. María Trinidad Sánchez, from fruits
of C. nucifera
Lasioseius sanchezensis Abo-Shnaf et al., 2016: 631 *DR > TL: Las Cejas, Prov. María Trinidad Sánchez, from Sida sp.

Family MACRONYSSIDAE
Steatonyssus Kolenati, 1858
Steatonyssus spinosus Willmann, 1936: 152

Family MELICHARIDAE
Proctolaelaps Berlese, 1923
Proctolaelaps bickleyi (Bram, 1956). Abo-Shnaf et al., 2016: 636
Proctolaelaps aff. fiseri Samsinak, 1960. Abo-Shnaf et al., 2016: 637
Proctolaelaps lobatus De Leon, 1963. Abo-Shnaf et al., 2016: 637

Family OPILIOACARIDAE
Caribeacarus Vázquez & Klompen, 2009
Caribeacarus armasi Vázquez & Klompen, 2009: 36

Family OTOPHEIDOMENIDAE
Dicrocheles Krantz & Khot, 1962
Dicrocheles phalaenodectes (Treat, 1954). Treat, 1969: 654
Hemipteroseius Evans, 1963
Hemipteroseius ageneius Treat, 1965: 7
Hemipteroseius antilleus Treat, 1965: 3

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Hemipteroseius parvulus Treat, 1965: 6

Family PHYTOSEIIDAE
Subfamily AMBLYSEIINAE
Amblydromalus Chant & McMurtry, 2005
Amblydromalus congeae (De Leon, 1965). Ferragut et al., 2011: 38
Amblydromalus rapax (De Leon, 1965). Ferragut et al., 2011: 38; Abo-Shnaf et al., 2016: 608
Amblyseius Berlese, 1914
Amblyseius aerialis (Muma, 1955). Ferragut et al., 2011: 39; Abo-Shnaf et al., 2016: 609
Amblyseius chiapensis De Leon, 1961. Ferragut et al., 2011: 39
Amblyseius fernandezi Chant & Baker, 1965. Abo-Shnaf et al., 2016: 609
Amblyseius herbicolus (Chant, 1959). Ferragut et al., 2011: 39
Amblyseius largoensis (Muma, 1955). Ferragut et al., 2011: 40; Abo-Shnaf et al., 2016: 610
Amblyseius obtusus (Koch, 1839). Abo-Shnaf et al., 2016: 607
=Amblyseiopsis musae Garman, 1958: 73 [Misidentification]
Amblyseius tamatavensis Blommers, 1974. Abo-Shnaf et al., 2016: 610
Euseius Wainstein, 1962
Euseius naindaimei (Chant & Baker, 1965). Ferragut et al., 2011: 40
Euseius sibelius (De Leon, 1962). Ferragut et al., 2011: 40
Iphiseiodes De Leon, 1966
Iphiseiodes quadripilis (Banks, 1904). Ferragut et al., 2011: 40
Iphiseiodes zuluagai Denmark & Muma, 1972. Abo-Shnaf et al., 2016: 611
Neoseiulus Hughes, 1948
Neoseiulus longispinosus (Evans, 1952). Ferragut et al., 2011: 41; Abo-Shnaf et al., 2016: 611
Neoseiulus plumosus Denmark & Muma, 1975. Ferragut et al., 2011: 41
Phytoscutus Muma, 1961
Phytoscutus sexpilis Muma, 1961. Abo-Shnaf et al., 2016: 610
Phytoseiulus Evans, 1952
Phytoseiulus macropilis (Banks, 1904). Abo-Shnaf et al., 2016: 612
Proprioseiopsis Muma, 1961
Proprioseiopsis sandersi (Chant, 1959: 102). De Moraes et al., 2004: 192; Abo-Shnaf et al., 2016: 607
=Amblyseius perlongisetus Berlese, 1916. De Moraes et al., 2004: 47 [Misidentification]
Transeius Chant & McMurtry, 2004
Transeius aciculus (De Leon, 1967). Ferragut et al., 2011: 41
Typhlodromalus Muma, 1961
Typhlodromalus peregrinus (Muma, 1955). Ferragut et al., 2011: 42
Typhlodromips De Leon, 1965
Typhlodromips dentilis (De Leon, 1959). Ferragut et al., 2011: 42. Abo-Shnaf et al., 2016: 613
Subfamily PHYTOSEIINAE
Phytoseius Ribaga, 1904
Phytoseius averrhoae De Leon, 1965. Ferragut et al., 2011: 45
Phytoseius benneti De Leon, 1965. Ferragut et al., 2011: 45
Phytoseius dominicensis Ferragut & De Moraes in Ferragut et al., 2011: 43 *DR > TL: Boca del Diablo, Las Galeras (Peninsula
de Samaná)
Phytoseius rex De Leon, 1967. Ferragut et al., 2011: 45; Abo-Shnaf et al., 2016: 612
Phytoseius woodburyi De Leon, 1965. Ferragut et al., 2011: 46
Subfamily TYPHLODROMINAE
Galendromus Muma, 1961
Galendromus (Galendromus) annectens (De Leon, 1958). Ferragut et al., 2011: 46
Typhlodromina Muma, 1961
Typhlodromina subtropica Muma & Denmark, 1969. Ferragut et al., 2011: 46; Abo-Shnaf et al., 2016: 613
Typhlodromus Scheuten, 1857
Typhlodromus (Anthoseius) evectus (Schuster, 1966). Ferragut et al., 2011: 47
Typhloseiopsis De Leon, 1959

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Typhloseiopsis adventitius Ferragut & Moraes in Ferragut et al., 2011: 48 *DR > TL: Jarabacoa, on Cecropia peltata, 640 m

Family SPELAEORHYNCHIDAE
Spelaeorhynchus Neumann, 1902
Spelaeorhynchus praecursor Neumann, 1902. Fain et al., 1967: 542
=Spelaeorhynchus latus Banks, 1917. Fain et al., 1967: 547

Family TRIGONUROPODIDAE
Trigonuropoda Trägårdh, 1952
Trigonuropoda (Baloghiatrigon) dominicana Konstchán, 2008: 56 *DR > TL: Monseñor Nouel, 10 km NNW of Bonao W of
Jima/Pringamosa village

Family UROPODIDAE
Rotundabaloghia Hirschmann, 1975
Rotundabaloghia arragensis Kontschán, 2005: 245 *DR > TL: S of Arraga
Rotundabaloghia dominicana Kontschán, 2005: 242 *DR > TL: Near Constanza
Rotundabaloghia pocsiana Kontschán, 2005: 247 *DR > TL: Near Jima village
Rotundabaloghia punctata Kontschán, 2005: 245 *DR > TL: S of Arraga
Rotundabaloghia szuetsi Kontschán, 2005: 243 *DR > TL: SW edge of Constanza

Family VARROIDAE
Varroa Oudemans, 1904
Varroa destructor Anderson & Trueman, 2000. Zuluaga & Mesa, 2000: 197; Perez-Gelabert, 2008: 44 [Introduced]

Order HOLOTHYRIDA
Family NEOTHYRIDAE
Caribothyrus Kontschán & Mahunka, 2004
Caribothyrus barbatus Kontschán & Mahunka, 2004: 343 *DR > TL: Along a tributary of Arrago streamlet, south of Arraga
village in valley, 1120-1160 m

Order AMBLYPYGI
Key reference: Armas (2006), Teruel (2016).

Family CHARINIDAE
Charinus Simon, 1892
Charinus bahoruco Teruel, 2016: 7 *DR > TL: Rio Mulito, Banano, Pedernales Prov.
Charinus dominicanus Armas & Pérez-Gonzalez, 2001: 50. Armas, 2006: 228. Teruel, 2016: 3 *DR > TL: Los Charcos, San
Rafael, Sierra de Bahoruco
Charinus magua Seiter, Schramm & Schwaha, 2018: 350 *DR > TL: Subida a Casabito, Monseñor Nouel Prov., 382 m

Family PHRYNIDAE
Phrynus Lamarck, 1801
Phrynus alejandroi Armas & Teruel, 2010. Armas, 2013: 91
Phrynus decoratus Teruel & Armas, 2005: 129. Armas, 2006: 233
Phrynus eucharis Armas & Pérez-Gonzalez, 2001: 49. Armas, 2006: 234
Phrynus hispaniolae Armas & Pérez-Gonzalez, 2001: 48. Armas, 2006: 236
Phrynus kennidae Armas & Pérez-Gonzalez, 2001: 50. Armas, 2006: 236 *DR > TL: Playa del Coco, NW Isla Beata, Prov.
Pedernales
Phrynus longipes (Pocock, 1894: 536). Armas, 2006: 238
Phrynus marginemaculatus C. L. Koch, 1840. Armas, 2006: 239
Phrynus resinae† (Schawaller, 1979: 4). Schawaller, 1982: 2 [Fossil in Dominican amber]

Order ARANEAE
Key references: Penney and Perez-Gelabert (2002), Penney (2006)

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Taxonomic summary:
Suborder Opisthothelae
Infraorder Mygalomorphae
Family Dipluridae
Family Cyrtauchenidae
Family Microstigmatidae
Family Barychelidae
Family Theraphosidae
Infraorder Araneomorpha
Family Filistatidae
Family Sicariidae
Family Scytodidae
Family Drymusidae
Family Ochyroceratidae
Family Pholcidae
Family Caponiidae
Family Tetrablemmidae
Family Segestridae
Family Oonopidae
Family Plectreuridae
Family Mimetidae
Family Oecobidae
Family Hersiliidae
Family Deinopidae
Family Uloboridae
Family Nesticidae
Family Theridiidae
Family Theridiosomatidae
Family Symphytognathidae
Family Anapidae
Family Mysmenidae
Family Linyphiidae
Family Tetragnathidae
Family Araneidae
Family Lycosidae
Family Pisauridae
Family Oxyopidae
Family Ctenidae
Family Agelenidae
Family Desidae
Family Hahniidae
Family Dictynidae
Family Amaurobiidae
Family Eutichuridae
Family Miturgidae
Family Anyphaenidae
Family Liocranidae
Family Phrurolithidae
Family Clubionidae
Family Corinnidae
Family Trachelidae
Family Trochanteridae
Family Prodidomidae
Family Gnaphosidae

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 55

 Family Selenopidae
Family Sparassidae
Family Philodromidae
Family Thomisidae
Family Salticidae

Suborder OPISTHOTHELAE
Infraorder MYGALOMORPHAE
Family DIPLURIDAE
Ischnothele Ausserer, 1875
Ischnothele garcia Coyle, 1995: 50. Penney & Perez-Gelabert, 2002: 205 *DR > TL: Sierra Martín García, Barahona Prov.
Ischnothele jeremie Coyle, 1995: 48. Penney & Perez-Gelabert, 2002: 205 *H > TL: Formond, elev. 975 m
Ischnothele sp. indet.†—Wunderlich (1988a: 33); Penney (2006a: 26) [Fossil in Dominican amber]
Masteria Koch, 1873
Masteria galipote Passanha & Brescovit, 2018: 50 *DR > TL: La Pirámide
Masteria sexoculata† (Wunderlich, 1988a: 47). Penney, 2004: 184 [Fossil in Dominican amber]
Masteria sp.—Sánchez-Ruíz (2009: 24)

Family CYRTAUCHENIIDAE
Bolostromus Ausserer, 1875
Bolostromus destructus† Wunderlich, 1988a: 42 [Fossil in Dominican amber]
Bolostromus sp.—Alayón García & De los Santos (2009: 45)

Family MICROSTIGMATIDAE
Parvomygale† Wunderlich, 2004
Parvomygale distincta† Wunderlich, 2004: 617 [Fossil in Dominican amber]

Family BARYCHELIDAE
Psalistops Simon, 1889
Psalistops fulvus Bryant, 1948: 335. Penney & Perez-Gelabert, 2002: 205 *H > TL: Diquini
Psalistops hispaniolensis† Wunderlich, 1988a: 50 [Fossil in Dominican amber]
Psalistops maculosus Bryant, 1948: 336. Penney & Perez-Gelabert, 2002: 205 *H > TL: Milot

Family THERAPHOSIDAE
Antillena Bertani, Huff & Fukushima in Fukushima & Bertani, 2017
Antillena rickwesti (Bertani & Huff, 2013: 333). Raderka, 2016: 121; Fukushima & Bertani, 2017: 153 *DR > TL: Track into
park between Manuel Goya and Oviedo, Parque Nacional Jaragua
Citharacanthus Pock, 1901
Citharacanthus spinicrus (Latreille, 1819). Penney & Perez-Gelabert, 2002: 205
Cyrtopholis Simon, 1892
Cyrtopholis agilis Pocock, 1903: 97. Bryant, 1948: 337; Penney & Perez-Gelabert, 2002: 205 *DR > TL: San Domingo
Cyrtopholis cursor (Ausserer, 1875: 176). Bryant, 1948: 334; Penney & Perez-Gelabert, 2002: 205 *H, *DR > TL: St.
Domingo
Scopelobates Simon, 1903
Scopelobates sericeus Simon, 1903: 928. Bryant, 1948: 334; Penney & Perez-Gelabert, 2002: 205; Guadanucci et al., 2017:
270 *DR > TL: Santo Domingo
Ischnocolinopsis Wunderlich, 1988
Ischnocolinopsis acutus† Wunderlich, 1988a: 49 [Fossil in Dominican amber]
Phormictopus Pocock, 1901
Phormictopus atrichomatus Schmidt, 1991: 7. Rudloff, 2008: 8 *DR > TL: By error originally said to be from “Honduras”
without additional data, but this species is endemic to Hispaniola.
Phormictopus cancerides (Latreille, 1806). Bryant, 1948: 339; Penney & Perez-Gelabert, 2002: 205; Rudloff, 2008: 20
Phormictopus melodermus Chamberlin, 1917: 62. Rudloff, 2008: 41 *H > TL: “West Indies?”
Trichopelma Simon, 1888

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Trichopelma maddeni Esposito & Agnarsson in Bloom et al., 2014: 151 *DR > TL: Cueva Seibo, Parque Nacional del Este
Trichopelma nitidum Simon, 1888. Bryant, 1948: 334; Penney & Perez-Gelabert, 2002: 205

Infraorder ARANEOMORPHAE
Family FILISTATIDAE
Antilloides Brescovit et al., 2016
Antilloides didicostae† (Penney, 2005d: 95). Magalhães, 2018: 247 [Fossil in Dominican amber]
Antilloides haitises Brescovit et al., 2016: 418 *DR > TL: Cueva Lines, Parque Nacional Los Haitises
Kukulcania Lehtinen, 1967
Kukulcania hibernalis (Hentz, 1842). Bryant, 1948: 339; Penney & Perez-Gelabert, 2002: 206; Brescovit et al., 2016: 429

Family SICARIIDAE
Loxosceles Heineken & Lowe, 1832
Loxosceles aculicaput† Wunderlich, 2004: 703 [Fossil in Dominican amber]
Loxosceles caribbaea Gertsch, 1958: 17. Gertsch & Ennik, 1983: 340; Penney & Perez-Gelabert, 2002: 206
Loxosceles cubana Gerstch, 1958. Gertsch & Ennik, 1983: 344; Penney & Perez-Gelabert, 2002: 206
Loxosceles defecta† Wunderlich, 1988a: 69 [Fossil in Dominican amber]
Loxosceles deformis† Wunderlich, 1988a: 68 [Fossil in Dominican amber]
Loxosceles taino Gerstch & Ennik, 1983. Penney & Perez-Gelabert, 2002: 206; Sánchez Ruiz & Brescovit, 2013: 221

Family SCYTODIDAE
Scytodes Latreille, 1804
Scytodes fusca Walckenaer, 1837. Bryant, 1948: 365; Penney & Perez-Gelabert, 2002: 206
Scytodes longipes Lucas, 1844. Bryant, 1948: 365; Penney & Perez-Gelabert, 2002: 206
Scytodes piliformis† Wunderlich, 1988a: 71 [Fossil in Dominican amber]
Scytodes planithorax† Wunderlich, 1988a: 72 [Fossil in Dominican amber]
Scytodes stridulans† Wunderlich, 1988a: 72 [Fossil in Dominican amber]

Family DRYMUSIDAE
Drymusa Simon, 1892
Drymusa simoni Bryant, 1948: 363. Valerio, 1971: 196; Penney & Perez-Gelabert, 2002: 206 *H > TL: La Hotte, 5,000-7,800
ft.

Family OCHYROCERATIDAE
Arachnolithulus† Wunderlich, 1988
Arachnolithulus longipes† Wunderlich, 2004f: 733 [Fossil in Dominican amber]
Arachnolithulus pygmaeus† Wunderlich, 1988a: 63 [Fossil in Dominican amber]
Fageicera Dumitrescu & Georgescu, 1992
Fageicera sp.—De los Santos & Carrero (2015: 135)
Ochyrocera Simon, 1892
Ochyrocera cachote Hormiga et al., 2007: 3 *DR > TL: Reserva Natural Cachote, Barahona Prov., 1220 m

Family PHOLCIDAE
Artema Walckenaer, 1833
Artema atlanta Walckenaer, 1837. Bryant, 1948: 366; Penney & Perez-Gelabert, 2002: 206
Coryssocnemis Simon, 1893
Coryssocnemis velteni† Wunderlich, 2004f: 743 [Fossil in Dominican amber]
Micropholcus Deeleman-Reinhold & Prinsen, 1987
Micropholcus baoruco (Huber in Huber & Wunderlich, 2006: 2342). Huber et al., 2014: 435 *DR > TL: Near Polo, Sierra de
Baoruco, 850 m
Micropholcus fauroti (Simon, 1887). Penney, 2004: 184
Micropholcus hispaniola (Huber, 2000: 77). Huber & Wunderlich, 2006: 2347; Penney & Perez-Gelabert, 2002: 206; Huber et
al., 2014: 435 *DR > TL: La Ciénaga, Parque Nacional A. Bermúdez, 1100 m
Micropholcus kiskeya† (Huber & Wunderlich, 2006: 2355); Huber et al., 2014: 435 [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 57

Micropholcus toma (Huber in Huber & Wunderlich, 2006: 2353). Huber et al., 2014: 435 *DR > TL: Near La Toma, San
Cristóbal Prov., 70 m
Modisimus Simon, 1893
Modisimus angulatus Huber & Fischer in Huber et al., 2010: 264 *DR > TL: Near La Toma, San Cristóbal Prov., 70 m
Modisimus bachata Huber & Fischer in Huber et al., 2010: 282 *DR > TL: Near La Ciénaga, La Vega Prov., ~1100 m
Modisimus berac Huber & Fischer in Huber et al., 2010: 288 *H > TL: Berac, La Visite N. P., 1580 m
Modisimus calcar† Wunderlich, 1988a: 78 [Fossil in Dominican amber]
Modisimus calcaroides† Wunderlich, 1988a: 79 [Fossil in Dominican amber]
Modisimus cienaga Huber & Fischer in Huber et al., 2010: 286 *DR > TL: Near La Ciénaga, La Vega Prov., ~1000 m
Modisimus crassifemoralis† Wunderlich, 1988a: 80 [Fossil in Dominican amber]
Modisimus cuadro Huber & Fischer in Huber et al., 2010: 283 *DR > TL: Near La Toma, San Cristóbal Prov., 70 m
Modisimus culicinus (Simon, 1893). Huber & Fischer in Huber et al., 2010: 287
Modisimus enriquillo Huber & Fischer in Huber et al., 2010: 276 *DR > TL: North-west of Paraiso, Barahona Prov., 180 m
Modisimus epepye Huber & Fischer in Huber et al., 2010: 273 *H > TL: Near Seguin, Dept Sud-Est, ~990 m
Modisimus femoratus Bryant, 1948: 307. Bryant, 1948: 367; Penney & Perez-Gelabert, 2002: 206; Huber et al., 2010: 256 *DR
> TL: Samaná Prov., Sánchez
Modisimus fuscus Bryant, 1948: 368. Penney & Perez-Gelabert, 2002: 206; Huber et al., 2010: 272 *H > TL: Kenscoff, 4500
feet
Modisimus glaucus Simon, 1893: 322. Bryant, 1948: 369; Penney & Perez-Gelabert, 2002: 206
Modisimus jima Huber & Fischer in Huber et al., 2010: 263 *DR > TL: Near Jima, Monseñor Nouel Prov., ~700 m
Modisimus kiskeya Huber & Fischer in Huber et al., 2010: 264 *DR > TL: Near La Ciénaga, La Vega Prov., ~1100 m
Modisimus leprete Huber & Fischer in Huber et al., 2010: 278 *H > TL: Near Le Prete, Dept du Sud, 170 m
Modisimus macaya Huber & Fischer in Huber et al., 2010: 280 *H > TL: Macaya B. R., 1290 m
Modisimus makandal Huber & Fischer in Huber et al., 2010: 271 *DR > TL: Near Cana, Santiago Rodríguez Prov., 150 m
Modisimus mango Huber & Fischer in Huber et al., 2010: 285 *H > TL: Le Prete, Dept du Sud, ~170 m
Modisimus mariposas Huber & Fischer in Huber et al., 2010: 280 *DR > TL: Near Jima, Monseñor Nouel Prov., ~700 m
Modisimus miri Huber & Fischer in Huber et al., 2010: 289 *DR > TL: Reserva Científica Loma Quita Espuela, near summit
Modisimus oculatus† Wunderlich, 1988a: 77 [Fossil in Dominican amber]
Modisimus palvet Huber & Fischer in Huber et al., 2010: 287 *H > TL: Macaya B.R., 1350 m
Modisimus paraiso Huber in Huber et al., 2010: 291 *DR, *H > TL: ~7 km north-west of Paraíso, Barahona Prov., ~180 m
Modisimus pelejil Huber & Fischer in Huber et al., 2010: 284 *DR > TL: East of San Juan de la Maguana, San Juan Prov., 410
m
Modisimus roumaini Huber in Huber et al., 2010: 274 *H > TL: Between Léogâne and Jacmel, 730 m
Modisimus seguin Huber & Fischer in Huber et al., 2010: 274 *H > TL: Near Seguin, Dept Sud-Est, 990 m
Modisimus tiburon Huber & Fischer in Huber et al., 2010: 277 *H > TL: Case Dent, La Visite, N. P., 1880 m
Modisimus toma Huber & Fischer in Huber et al., 2010: 268 *DR > TL: Near La Toma, San Cristóbal Prov., 70 m
Modisimus tuberosus† Wunderlich, 1988a: 81 [Fossil in Dominican amber]
Modisimus vittatus Bryant, 1948: 370. Penney & Perez-Gelabert, 2002: 206; Huber et al., 2010: 269 *H > TL: Port-au-Prince
Pholcophora Banks, 1896
Pholcophora brevipes† Wunderlich, 1988a: 83 [Fossil in Dominican amber]
Pholcophora gracilis† Wunderlich, 1988a: 82 [Fossil in Dominican amber]
Pholcophora longicornis† Wunderlich, 1988a: 83 [Fossil in Dominican amber]
Physocyclus Simon, 1893
Physocyclus globosus (Taczanowski, 1874). Bryant, 1948: 372; Penney & Perez-Gelabert, 2002: 206; Valdez-Mondragón,
2010: 25
Serratochorus Wunderlich, 1988
Serratochorus pygmaeus† Wunderlich, 1988a: 85 [Fossil in Dominican amber]
Tainonia Huber, 2000
Tainonia bayahibe Huber & Astrin, 2009: 299 *DR > TL: Near Bahahibe
Tainonia cienaga Huber & Astrin, 2009: 290 *DR > TL: Near La Ciénaga
Tainonia samana Huber & Astrin, 2009: 293 *DR > TL: S of Las Galeras, Samaná Prov.
Tainonia serripes (Simon, 1893). Bryant, 1948: 366; Huber, 2000: 146; Penney & Perez-Gelabert, 2002: 206 *DR > TL: St.
Dom. [Santo Domingo]
Tainonia visite Huber & Astrin, 2009: 293 *H > TL: La Visite National Park, Case Dent

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Family CAPONIIDAE
Cubanops Sánchez-Ruiz, Platnick & Dupérré, 2010
Cubanops darlingtoni (Bryant, 1948: 342). Penney & Perez-Gelabert, 2002: 206; Sánchez-Ruiz, Platnick & Dupérré, 2010: 17
*DR > TL: San José de las Matas, Santiago
Cubanops vega Sánchez-Ruiz, Platnick & Dupérré, 2010: 17 *DR > TL: 10 km NE of Jarabacoa, La Vega, 550 m
Nops MacLeay, 1838
Nops blandus (Bryant, 1942). Chickering, 1967: 4; Penney & Perez-Gelabert, 2002: 206; Sánchez-Ruiz et al., 2015: 44;
Sánchez-Ruiz & Brescovit, 2018: 22
Nops ernestoi Sánchez-Ruiz, 2005: 24. Sánchez-Ruiz & Brescovit, 2018: 28 *DR > TL: Prov. La Vega: Hotel Montaña, 550
msnm, 10 km NE de Jarabacoa
Nops gertschi Chickering, 1967: 9. Penney & Perez-Gelabert, 2002: 206; Sánchez-Ruiz & Brescovit, 2018: 32 *DR > TL:
Loma Cibao, La Vega, 600-1600 m
Nops hispaniola Sánchez-Ruiz et al., 2015: 55; Sánchez-Ruiz & Brescovit, 2018: 43 *DR, *H > TL: Puerto Escondido,
Independencia Prov.
Nops jaragua Sánchez-Ruiz & Brescovit, 2018: 45 *DR > TL: El Cajuil, course between Oviedo and La Laguna, Jaragua
National Park, Oviedo, Pedernales
Nops lobatus† Wunderlich, 1988a: 65. Penney, 2001: 988 [Fossil in Dominican amber]
=Nops segmentatus† Wunderlich, 1988a: 67 [Fossil in Dominican amber]

Family TETRABLEMMIDAE
Monoblemma Gertsch, 1941
Monoblemma? spinosum† Wunderlich, 1988a: 61 [Fossil in Dominican amber]

Family SEGESTRIIDAE
Ariadna Audouin, 1826
Ariadna multispinosa Bryant, 1948: 339. Beatty, 1970: 474; Penney & Perez-Gelabert, 2002: 207 *DR > TL: Loma Rucilla
Mountains, Pico del Yaque, 8,000-10,000 ft.
Ariadna paucispinosa† Wunderlich, 1988a: 53 [Fossil in Dominican amber]

Family OONOPIDAE
Fossilopaea Wunderlich, 1988
Fossilopaea sulci† Wunderlich, 1988a: 59 [Fossil in Dominican amber]
Heteroonops Dalmas, 1916
Heteroonops castelloides Platnick & Dupérré, 2009b: 48 *DR > TL: Ciénaga, Parque Nacional A. Bermúdez, 1100 m
Heteroonops iviei Platnick & Dupérré, 2009b: 41 *DR > TL: Las Abejas, ca. 35 km N of Cabo Rojo, 1250 m
Heteroonops validus (Bryant, 1948: 341). Chickering, 1972a: 114; Penney & Perez-Gelabert, 2002: 207; Platnick & Dupérré,
2009b: 45 *DR > TL: Loma Rucilla mountains
Heteroonops vega Platnick & Dupérré, 2009b: 41 *DR > TL: Salto Jimenoa, 7 km SE Jarabacoa, 800 m
Hortoonops Platnick & Dupérré, 2012
Hortoonops excavatus Platnick & Dupérre, 2012: 61 *DR > TL: Salto Jimenoa, 7 km SE Jarabacoa, 800 m
Ischnothyreus Simon, 1890
Ischnothyreus peltifer (Simon, 1891). Chickering, 1968b: 80; Penney & Perez-Gelabert, 2002: 207
=Dysderina antillana Bryant, 1942. Bryant, 1948: 340
Opopaea Simon, 1891
Opopaea deserticola Simon, 1891. Platnick & Dupérré, 2009a: 4
Orchestina Simon, 1882
Orchestina dentifera Simon, 1893. Izquierdo & Ramírez, 2017: 26 [Introduced]
Orchestina dominicana† Wunderlich, 1981: 104. Penney, 2000b: 348 [Fossil in Dominican amber]
Orchestina sp. indet.†—Penney (2000b: 348) [Fossil in Dominican amber]
Orchestina tibialis† Wunderlich, 1988a: 57 [Fossil in Dominican amber]
Ponsoonops Bolzern, 2014
Ponsoonops lavega Bolzern, 2014: 52 *DR > TL: Berlese sample, near Concepción de la Vega, Parque Nacional José Armando
Bermúdez, 1124 m
Scaphioides Bryant, 1942

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 59

Scaphioides miches Platnick & Dupérre, 2012: 32 *DR > TL: Cacao farm near Miches, El Seibo
Stenoonops Simon 1892
Stenoonops incertus† (Wunderlich, 1988a: 60). Penney, 2000b: 348 [Fossil in Dominican amber]
Stenoonops jara Platnick & Dupérré, 2010: 45 *DR > TL: Hotel Montaña, 10 km NE Jarabacoa, 550 m
Stenoonops seldeni† Penney, 2000b: 345. Marusik & Wunderlich, 2008: 75 [Fossil in Dominican amber]
Triaeris Simon, 1892
Triaeris stenaspis Simon, 1891. Bryant, 1948: 342; Chickering, 1968c: 358; Penney & Perez-Gelabert, 2002: 207; Platnick et
al., 2012: 22

Family PLECTREURIDAE
Plectreurys Simon, 1893
Plectreurys pittfieldi† Penney, 2009: 66 [Fossil in Dominican amber]

Family MIMETIDAE
Mimetus Hentz, 1831
Mimetus? bituberculatus† Wunderlich, 1988a: 176 [Fossil in Dominican amber]
Mimetus hispaniolae Bryant, 1948: 395. Penney & Perez-Gelabert, 2002: 207 *DR > TL: Cordillera Central, rain forest near
Valle Nuevo, 6,000 ft.+
Mimetus notius Chamberlin, 1923. De los Santos & Alayón, 2012: 26
Mimetus syllepsicus Hentz, 1832. Penney & Perez-Gelabert, 2002: 207

Family OECOBIIDAE
Oecobius Lucas, 1846
Oecobius concinnus Simon, 1893. Penney, 2004: 184
=Oecobius audanti Bryant, 1948: 344
Oecobius piliformis† Wunderlich, 1988a: 86 [Fossil in Dominican amber]

Family HERSILIIDAE
Fictotama Petrunkevitch, 1963
Fictotama maculosa† Wunderlich, 2011: 462 [Fossil in Dominican amber]
Prototama Petrunkevitch, 1971
Prototama maior† (Wunderlich, 1988a: 90). Penney, 2006b: 901 [Fossil in Dominican amber]
Prototama media† (Wunderlich, 1988a: 89). Penney, 2006b: 901 [Fossil in Dominican amber]
Prototama minor† (Wunderlich, 1987: 93). Penney, 2006b: 902 [Fossil in Dominican amber]
Yabisi Rheims & Brescovit, 2004
Yabisi guaba Rheims & Brescovit, 2004: 234; Rheims et al., 2014: 405 *DR > TL: Pedernales (4 km north of Oviedo, 10 mts.)
Yabisi habanensis Rheims & Brescovit, 2004. De los Santos & Carrero-Jimenez, 2015: 136

Family DEINOPIDAE
Deinopis MacLeay, 1838
Deinopis lamia MacLeay, 1839. Bryant, 1948: 394; Penney & Perez-Gelabert, 2002: 207

Family ULOBORIDAE
Miagrammopes Cambridge, 1870
Miagrammopes dominicanus† Wunderlich, 2004g: 868 [Fossil in Dominican amber]
Miagrammopes latens Bryant, 1936. Bryant, 1948: 393; Penney & Perez-Gelabert, 2002: 207
Miagrammopes sp. indet.†—Penney (2001: 989) [Fossil in Dominican amber]
Philoponella Mello-Leitão, 1917
Philoponella semiplumosa (Simon, 1893). Penney & Perez-Gelabert, 2002: 207
Uloborus Latreille, 1806
Uloborus trilineatus Keyserling, 1883. Penney & Perez-Gelabert, 2002: 207
Zosis Walckenaer, 1833
Zosis geniculata (Olivier, 1791). Bryant, 1948: 394; Penney & Perez-Gelabert, 2002: 207

60 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Family NESTICIDAE
Eidmannella Roewer, 1935
Eidmannella pallida (Emerton, 1875). Penney & Perez-Gelabert, 2002: 207
Hispanonesticus† Wunderlich, 1986
Hispanonesticus latopalpus† Wunderlich, 1986: 128 [Fossil in Dominican amber]
Nesticus Thorell, 1869
Nesticus maculatus Bryant, 1948: 384. Penney & Perez-Gelabert, 2002: 207 *DR > TL: Cordillera Central, Loma Vieja, south
of Constanza, 6,000 ft.

Family THERIDIIDAE
Achaearanea Strand, 1929
Achaearanea extincta† Wunderlich, 1988a: 166 [Fossil in Dominican amber]
Achaearanea tesselata (Keyserling, 1884). Alayón García, 2007: 43
Anelosimus Simon, 1891
Anelosimus fraternus Bryant, 1948: 381. Agnarsson, 2006: 511
Anelosimus studiosus (Hentz, 1850). Bryant, 1948: 382; Penney & Perez-Gelabert, 2002: 207
Argyrodes Simon, 1894
Argyrodes elevatus Taczanowski, 1873. Exline & Levi, 1962: 134; Penney & Perez-Gelabert, 2002: 207
Argyrodes furcatus (O. P.-Cambridge, 1894). Exline & Levi, 1962: 116; Penney & Perez-Gelabert, 2002: 207
Argyrodes nephilae Taczanowski, 1873. Bryant, 1948: 380; Exline & Levi, 1962: 139; Penney & Perez-Gelabert, 2002: 207
Argyrodes parvipatellaris† Wunderlich, 1988a: 130 [Fossil in Dominican amber]
Ariamnes Thorell, 1869
Ariamnes haitensis (Exline & Levi, 1962: 129). Penney & Perez-Gelabert, 2002: 207 *H > TL: 32 km from Aux Cayes, 330 m
elev.
Chrosiothes Simon, 1894
Chrosiothes biconigerus† Wunderlich, 1988a: 140 [Fossil in Dominican amber]
Chrosiothes curvispinosus† Wunderlich, 1988a: 141 [Fossil in Dominican amber]
Chrosiothes emulgatus† Wunderlich, 1988a: 139 [Fossil in Dominican amber]
Chrosiothes longispinosus† Wunderlich, 1988a: 142 [Fossil in Dominican amber]
Chrosiothes monoceros† Wunderlich, 1988a: 141 [Fossil in Dominican amber]
Chrosiothes tumulus† Wunderlich, 1988a: 143 [Fossil in Dominican amber]
Chrosiothes unicornis† Wunderlich, 1988a: 144 [Fossil in Dominican amber]
Chrysso Cambridge, 1882
Chrysso albomaculata O. P.-Cambridge, 1882. Bryant, 1948: 382; Levi, 1955b: 61; Levi, 1962b: 215; Penney & Perez-
Gelabert, 2002: 208
Chrysso? conspicua† Wunderlich, 1988a: 162 [Fossil in Dominican amber]
Chrysso? dubia† Wunderlich, 1988a: 163 [Fossil in Dominican amber]
Chrysso pulcherrima (Mello-Leitão, 1917). Penney & Perez-Gelabert, 2002: 208
Coleosoma Cambridge, 1882
Coleosoma floridanum Banks, 1900. Bryant, 1948: 373; Penney & Perez-Gelabert, 2002: 208
Cornutidion† Wunderlich, 1988
Cornutidion elongatum† Wunderlich, 1988a: 174 [Fossil in Dominican amber]
Craspedisia Simon, 1894
Craspedisia spatulata Bryant, 1948: 373. Levi, 1963b: 178; Penney & Perez-Gelabert, 2002: 208 *DR > TL: Rainforest near
Valle Nuevo, 1800 m
Craspedisia yapchoontecki† Penney et al., 2012: 585 [Fossil in Dominican amber]
Dipoena Thorell, 1869
Dipoena dominicana Wunderlich, 1986: 73. Penney & Perez-Gelabert, 2002: 208 *DR > TL: Nordgebirge, nördlich von
Santiago [Mountains north of Santiago]
Dipoenata Wunderlich, 1988
Dipoenata altioculata† Wunderlich, 1988a: 156 [Fossil in Dominican amber]
Dipoenata cala† Wunderlich, 1988a: 159 [Fossil in Dominican amber]
Dipoenata clypeata† Wunderlich, 1988a: 158 [Fossil in Dominican amber]
Dipoenata globulus† Wunderlich, 1988a: 160 [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 61

Dipoenata morosa (Bryant, 1948: 376). Penney & Perez-Gelabert, 2002: 208
Dipoenata praedominicana† (Wunderlich, 1986: 74) [Fossil in Dominican amber] Nomen dubium
Dipoenata stipes† Wunderlich, 1988a: 155 [Fossil in Dominican amber]
Dipoenata yolandae† Wunderlich, 1988a: 160. Penney, 2004: 184 [Fossil in Dominican amber]
Episinus Latreille, 1809
Episinus antecognatus† Wunderlich, 1986: 77 [Fossil in Dominican amber] Nomen dubium
Episinus brevipalpus† Wunderlich, 1988a: 134 [Fossil in Dominican amber]
Episinus cornutus† Wunderlich, 1988a: 135 [Fossil in Dominican amber]
Episinus dominicus Levi, 1955a: 77. Levi, 1964b: 15; Penney & Perez-Gelabert, 2002: 208 *DR > TL: Valle de Polo, 2000-
3000 m
Episinus praecognatus† Wunderlich, 1982: 34 [Fossil in Dominican amber]
Episinus tibiaseta† Wunderlich, 2011: 463 [Fossil in Dominican amber]
Episinus tuberosus† Wunderlich, 1988a: 133 [Fossil in Dominican amber]
Faiditus Simon, 1894
Faiditus americanus (Taczanowski, 1874). Exline & Levi, 1962: 161; Penney & Perez-Gelabert, 2002: 207
Faiditus caudatus (Taczanowski, 1874). Exline & Levi, 1962: 176; Penney & Perez-Gelabert, 2002: 207
Faiditus crassipatellaris† (Wunderlich, 1988a: 129). Penney, 2001: 989 [Fossil in Dominican amber]
Faiditus darlingtoni (Exline & Levi, 1962: 184). Penney & Perez-Gelabert, 2002: 207 *DR > TL: Rain forest near Valle
Nuevo, 2000 m elev.
Femurraptor Wunderlich, 2011
Femurraptor dominicanus† Wunderlich, 2011: 465 [Fossil in Dominican amber]
Genus Incertae Sedis
Genus indet. clypeatus† (Wunderlich, 1988a: 165). Penney, 2001: 995 [Fossil in Dominican amber]
Lasaeola Simon, 1881
Lasaeola puta† Wunderlich, 1988a: 153 [Fossil in Dominican amber]
Lasaeola pristina† (Wunderlich, 1988a: 152) [Fossil in Dominican amber]
Lasaeola vicina† (Wunderlich, 1982: 36) [Fossil in Dominican amber]
Lasaeola vicinoides† Wunderlich, 1988a: 151 [Fossil in Dominican amber]
Latrodectus Walckenaer, 1805
Latrodectus geometricus C. L. Koch, 1841. Bryant, 1948: 377; Penney & Perez-Gelabert, 2002: 208
Latrodectus mactans (Fabricius, 1775). Bryant, 1948: 377; Penney & Perez-Gelabert, 2002: 208
Neopisinus Marques, Buckup & Rodrigues, 2011
Neopisinus gratiosus (Bryant, 1940: 313). Bryant, 1948: 382; Levi, 1964: 13; Penney & Perez-Gelabert, 2002: 208
Neospintharus Exline, 1950
Neospintharus furcatus (O.P.-Cambridge, 1894). Penney & Perez-Gelabert, 2002: 208
Nesticodes Archer, 1950
Nesticodes rufipes (Lucas, 1846). Penney & Perez-Gelabert, 2002: 208
Pholcomma Thorell, 1869
Pholcomma sp.—De los Santos & Alayón (2012: 27)
Romphaea L. Koch, 1872
Rhomphaea fictilium (Hentz, 1850). De los Santos & Alayón, 2015: 136
Spintharus Hentz, 1850
Spintharus frosti Van Patten & Agnarsson in Agnarsson et al., 2018: 777 *DR > TL: San Juan, Parque Nacional Armando
Bermúdez, Compartición Camp, 2476 m
Spintharus leonardodicaprioi Van Patten & Agnarsson in Agnarsson et al., 2018: 772 *DR > TL: Loma Quita Espuela, 200 m
Spintharus longisoma† Wunderlich, 1988a: 136 [Fossil in Dominican amber]
Spintharus skelly Van Patten & Agnarsson in Agnarsson et al., 2018: 779 *DR > TL: La Vega, Armando Bermúdez National
Park, Los Tablones, 1304 m
Steatoda Sundevall, 1833
Steatoda erigoniformis (O.P.-Cambridge, 1872). Levi, 1962a: 25; Penney & Perez-Gelabert, 2002: 208
Steatoda grossa (C. L. Koch, 1838). Levi, 1957: 404; Levi, 1962a: 26; Penney & Perez-Gelabert, 2002: 208
=Teutona modesta Bryant, 1948: 375
Stemmops Cambridge, 1894
Stemmops incertus† Wunderlich, 1988a: 146 [Fossil in Dominican amber]

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Stemmops prominens† Wunderlich, 1988a: 145 [Fossil in Dominican amber]
Styposis Simon, 1894
Styposis pholcoides† Wunderlich, 1988a: 146. Penney, 2001: 990; Wunderlich, 2011: 466 [Fossil in Dominican amber]
Theridion Walckenaer, 1805
Theridion antillanum Simon, 1894. Levi, 1959a: 116; Penney & Perez-Gelabert, 2002: 209
Theridion australe Banks, 1899. Levi, 1959a: 82; Penney & Perez-Gelabert, 2002: 209
Theridion contrarium† Wunderlich, 1988a: 168 [Fossil in Dominican amber]
Theridion dilucidum Simon, 1897. Levi, 1959a: 84; Penney & Perez-Gelabert, 2002: 209
Theridion erectoides† Wunderlich, 1988a: 173 [Fossil in Dominican amber]
Theridion erectum† Wunderlich, 1988a: 172 [Fossil in Dominican amber]
Theridion fuesslini Simon, 1894. Penney & Perez-Gelabert, 2002: 209 [Nomen dubium]
Theridion hassleri Levi, 1963a: 557. Penney & Perez-Gelabert, 2002: 209
Theridion inversum† Wunderlich, 1988a: 171 [Fossil in Dominican amber]
Theridion melanostictum O. P. Cambridge, 1876. Penney, 2004: 184
Theridion positivum Chamberlin, 1924. Levi, 1959a: 114; Levi, 1963a: 665; Penney & Perez-Gelabert, 2002: 209
Theridion variosoma† Wunderlich, 1988a: 170 [Fossil in Dominican amber]
Theridion wunderlichi† Penney, 2001: 994 [Fossil in Dominican amber]
=Theridion ovale† Wunderlich, 1988a: 169 [Preoccupied name]
Theridula Emerton, 1882
Theridula gonygaster (Simon, 1873). Levi, 1966: 127; Penney & Perez-Gelabert, 2002: 209
Thymoites Keyserling, 1884
Thymoites banksi (Bryant, 1948: 378). Penney & Perez-Gelabert, 2002: 209 *DR > TL: Cordillera Central, Loma Rucilla
Mountains, 5,000 ft.
Thymoites guanicae (Petrunkevitch, 1930). Penney & Perez-Gelabert, 2002: 209
Thymoites pallidus (Emerton, 1913). Bryant, 1948: 385; Levi, 1964a: 470; Penney & Perez-Gelabert, 2002: 209
Tidarren Chamberlin & Ivie, 1934
Tidarren sisyphoides (Walckenaer, 1842). Levi, 1955b: 70; Penney & Perez-Gelabert, 2002: 209

Family THERIDIOSOMATIDAE
Allototua Bryant, 1945
Allototua guttata Bryant, 1945: 411. Penney & Perez-Gelabert, 2002: 209 *DR > TL: Loma Vieja, Cordillera Central, south of
Constanza, 6,000 ft. [Nomen dubium]
Ogulnius Cambridge, 1882
Ogulnius fulvus Bryant, 1945: 412. Penney & Perez-Gelabert, 2002: 209 *DR > TL: Foot hills of Cordillera Central, south of
Santiago, 2,000-5,000 ft.
Ogulnius latus Bryant, 1948: 394. Penney & Perez-Gelabert, 2002: 209 *DR > TL: Cordillera Central, Loma Rucilla, 5,000-
8,000 ft.
Palaeoepeirotypus† Wunderlich, 1988
Palaeoepeirotypus iuvenis† Wunderlich, 1988a: 115 [Fossil in Dominican amber]
Palaeoepeirotypus iuvenoides† Wunderlich, 1988a: 117 [Fossil in Dominican amber]
Theridiosoma Cambridge, 1879
Theridiosoma incompletum† Wunderlich, 1988a: 114 [Fossil in Dominican amber]
Wendilgarda Keyserling, 1886
Wendilgarda clara Keyserling, 1886. Coddington, 1986: 88; Penney & Perez-Gelabert, 2002: 209

Family SYMPHYTOGNATHIDAE
Patu Marples, 1951
Patu sp.—Hormiga et al. (2007: 13)

Family ANAPIDAE
Palaeoanapis† Wunderlich, 1988
Palaeoanapis nana† Wunderlich, 1988a: 119 [Fossil in Dominican amber]

Family MYSMENIDAE

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 63

Dominicanopsis† Wunderlich, 2004
Dominicanopsis grimaldii† Wunderlich, 2004: 1078 [Fossil in Dominican amber]
Microdipoena Banks, 1895
Microdipoena sp.—Hormiga et al. (2007: 13)
Mysmenopsis Simon, 1898
Mysmenopsis lissycoleyae† Penney, 2000b: 349 [Fossil in Dominican amber]

Family LINYPHIIDAE
Ceraticelus Simon, 1884
Ceraticelus paludigenus Crosby & Bishop, 1925. Penney, 2004: 184
=Ceraticelus solitarius Bryant, 1948: 387. Penney, 2004: 184
Floricomus Crosby & Bishop, 1928
Floricomus fossilis† Penney, 2005a: 60 [Fossil in Dominican amber]
Florinda Cambridge, 1896
Florinda coccinea (Hentz, 1850). Bryant, 1948: 387; Penney & Perez-Gelabert, 2002: 209
Frontinella Cambridge, 1902
Frontinella bella Bryant, 1948: 386. Penney & Perez-Gelabert, 2002: 209 *DR > TL: Cordillera Central, Constanza, 3,000-
4,000 ft.
Frontinella communis (Hentz, 1850). Bryant, 1948: 387; Penney & Perez-Gelabert, 2002: 209
Grammonota Emerton, 1882
Grammonota calcarata Bryant, 1948: 390. Penney & Perez-Gelabert, 2002: 209 *DR > TL: Cordillera Central, Loma Rucilla
Mountains, 5,000-8,000 ft.
Lomaita Bryant, 1948
Lomaita darlingtoni Bryant, 1948: 392. Penney & Perez-Gelabert, 2002: 209; Da Silva-Moreira & Hormiga, 2015: 252 *DR >
TL: Cordillera Central, Loma Vieja, south of Constanza, 6,000 ft.
Meioneta Hull, 1920
Meioneta bigibber† (Wunderlich, 1988a: 123). Penney, 2001: 997 [Fossil in Dominican amber]
Meioneta fastigata† (Wunderlich, 1988a: 122). Penney, 2001: 997 [Fossil in Dominican amber]
Meioneta separata† (Wunderlich, 1988a: 125). Penney, 2001: 997 [Fossil in Dominican amber]
Meioneta sp. indet.†—Penney (2001: 997) [Fossil in Dominican amber]
=Agyneta sp.†—Wunderlich (1988a: 124) [Fossil in Dominican amber]
Mermessus O. Pickard-Cambridge, 1899
Mermessus dentiger O. Pickard-Cambridge, 1899. Miller, 2007: 124
=Eperigone antillana Bryant, 1948: 388
=Eperigone serrata Ivie & Barrows, 1935. Millidge, 1987: 18; Penney & Perez-Gelabert, 2002: 209
Selenyphantes Gertsch & Davis, 1946
Selenyphantes flagellifera† (Wunderlich, 1986: 83) [Fossil in Dominican amber]
Tapinopa Westring, 1851
Tapinopa bilineata Banks, 1893. De los Santos & Alayón, 2012: 27
Tutaibo Chamberlin, 1917
Tutaibo anglicanus (Hentz, 1850). Bryant, 1948: 388; Penney & Perez-Gelabert, 2002: 210

Family TETRAGNATHIDAE
Agriognatha Cambridge, 1896
Agriognatha argyra Bryant, 1945: 397. Penney & Perez-Gelabert, 2002: 210 *DR > TL: Cordillera Central, rain forest north of
Valle Nuevo, 6,000 ft.+
Alcimosphenus Simon, 1895
Alcimosphenus licinus Simon, 1895. Bryant, 1945: 385; Penney & Perez-Gelabert, 2002: 211
Antillognatha Bryant, 1945
Antillognatha lucida Bryant, 1945: 403. Penney & Perez-Gelabert, 2002: 210
Azilia Keyserling, 1881
Azilia hispaniolensis† Wunderlich, 1988a: 102. Penney, 2001: 998 [Fossil in Dominican amber]
=Azilia muellenmeisteri† Wunderlich, 1988a: 103. Penney, 2001: 998 [Fossil in Dominican amber]
Azilia montana Bryant, 1940. Penney & Perez-Gelabert, 2002: 210

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Chrysometa Simon, 1894
Chrysometa bigibbosa (Keyserling, 1864). Penney & Perez-Gelabert, 2002: 210
Chrysometa conspersa (Bryant, 1945: 391). Penney & Perez-Gelabert, 2002: 210
Chrysometa cornuta (Bryant, 1945: 392). Levi, 1986: 120; Penney & Perez-Gelabert, 2002: 210 *DR > TL: Cordillera Central,
Loma Vieja, south of Constanza, 6,000 ft.
Chrysometa maculata (Bryant, 1945: 388). Levi, 1986: 126; Penney & Perez-Gelabert, 2002: 210 *H, *DR > TL: Ennery
Chrysometa obscura (Bryant, 1945: 394). Levi, 1986: 145; Penney & Perez-Gelabert, 2002: 210 *DR > TL: Mt. Diego de
Ocampo, north range, 3,000-4,000 ft.
Chrysometa sabana Levi, 1986: 122. Penney & Perez-Gelabert, 2002: 210 *DR > TL: Forest between Hato Mayor and Sabana
de la Mar
Cyrtognatha Keyserling, 1881
Cyrtognatha espanola (Bryant, 1945: 399). Penney & Perez-Gelabert, 2002: 210; Dimitrov & Hormiga, 2009: 62 *H > TL:
Kenscoff, 1310 m
Cyrtognatha rucilla (Bryant, 1945: 401). Dimitrov & Hormiga, 2009: 73 *DR > TL: Loma Rucilla mountains, 1500-2400 m
Cyrtognatha weitschati† Wunderlich, 1988a: 97 [Fossil in Dominican amber]
Glenognatha Simon, 1888
Glenognatha argyrostilba (O. Pickard-Cambridge, 1876). Cabra-García & Brescovit, 2016: 45
=Glenognatha mira Bryant, 1945: 405. Penney & Perez-Gelabert, 2002: 210
Glenognatha foxi (McCook, 1894). Cabra-García & Brescovit, 2016: 109
Glenognatha minuta Banks, 1898. Cabra-García & Brescovit, 2016: 148
Hispanognatha Bryant, 1945
Hispanognatha guttata Bryant, 1945: 406. Penney & Perez-Gelabert, 2002: 210 *DR > TL: Cordillera Central, Valle Nuevo,
south west of Constanza, 7,000 ft.
Homalometa Simon, 1898
Homalometa fossilis† Wunderlich, 1988a: 100 [Fossil in Dominican amber]
Leucauge White, 1841
Leucauge argyra (Walckenaer, 1842). Bryant, 1945: 390; Penney & Perez-Gelabert, 2002: 210
Leucauge moerens (O.-P. Cambridge, 1896). De los Santos & Alayón, 2012: 27
Leucauge regnyi (Simon, 1897). Bryant, 1945: 386; Penney & Perez-Gelabert, 2002: 210
Leucauge venusta (Walckenaer, 1842). Penney, 2004: 184
Leucauge mariana (Taczanowski, 1881). Levi, 2008: 170
=Leucauge venustella Strand, 1916. Perez-Gelabert, 2008: 56; Levi, 2008: 170
Metabus O.P. Cambridge, 1899
Metabus ebanoverde Alvarez-Padilla, 2007: 306
Tetragnatha Walckenaer, 1805
Tetragnatha elongata Walckenaer, 1842. Bryant, 1945: 407; Penney & Perez-Gelabert, 2002: 210
Tetragnatha nitens Audouin, 1826. Penney & Perez-Gelabert, 2002: 210
=Tetragnatha haitiensis Bryant, 1945: 408
Tetragnatha orizaba (Banks, 1898). Bryant, 1945: 409; Penney & Perez-Gelabert, 2002: 210
Tetragnatha pallescens F.O.P.-Cambridge, 1903. Bryant, 1945: 410; Penney & Perez-Gelabert, 2002: 210; Penney, 2004: 184
Tetragnatha pristina† Schawaller, 1982: 3 [Fossil in Dominican amber]
Tetragnatha tenuissima O.P.-Cambridge, 1889. Bryant, 1945: 410; Penney & Perez-Gelabert, 2002: 210
=Tetragnatha festina Bryant, 1945: 407

Family ARANEIDAE
Acacesia Simon, 1895
Acacesia hamata (Hentz, 1847). Bryant, 1945: 364; Penney & Perez-Gelabert, 2002: 211
Acanthepeira Marx, 1883
Acanthepeira stellata (Walckenaer, 1805). Bryant, 1945: 376; Penney & Perez-Gelabert, 2002: 211
Acanthepeira venusta (Banks, 1896). De los Santos & Carrero, 2011: 55
Aculepeira Chamberlin & Ivie, 1942
Aculepeira busu Levi, 1991a: 308. Penney & Perez-Gelabert, 2002: 211 *DR > TL: Mt. Busú, Sierra Martín García, 1000-1300
m
Aculepeira visite Levi, 1991a: 307. Penney & Perez-Gelabert, 2002: 211 *H > TL: La Visite, 1800-2100 m

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 65

 =Parawixia darlingtoni Bryant, 1945: 382 (part). Levi, 1991a: 307
Allocyclosa Levi, 1999
Allocyclosa bifurca (McCook, 1887). Bryant, 1945: 367; Levi, 1999: 304; Penney & Perez-Gelabert, 2002: 211
Araneometa Wunderlich, 1988
Araneometa excelsa† Wunderlich, 1988a: 111 [Fossil in Dominican amber]
Araneometa herrlingi† Wunderlich, 1988a: 109 [Fossil in Dominican amber]
Araneometa spirembolus† Wunderlich, 1988a: 105 [Fossil in Dominican amber]
Araneus Clerck, 1757
Araneus elizabethae Levi, 1991: 263. Penney & Perez-Gelabert, 2002: 211 *H > TL: Kenscoff, Ouest, 1400 m
=Meta blanda Bryant, 1945: 386. Levi, 1991a: 263
=Araneus bryantae Brignoli, 1983: 262. Levi, 1991a: 262
=Neosconella parva Bryant, 1945: 381. Levi, 1991a: 262
Araneus hispaniola (Bryant, 1945: 366). Levi, 1991a: 263; Penney & Perez-Gelabert, 2002: 211 *H > TL: Kenscoff, 4,300 ft
[1300 m]
Araneus hotteiensis (Bryant, 1945: 387). Levi, 1991a: 263; Penney & Perez-Gelabert, 2002: 211 *H > TL: Foothills NE of
Massif de la Hotte, 1000-1100 m
Araneus? nanus† Wunderlich, 1988a: 110 [Fossil in Dominican amber]
Araneus perplexus (Walckenaer, 1842). Bryant, 1948: 440; Penney & Perez-Gelabert, 2002: 211 [Nomen dubium]
Argiope Audouin, 1827
Argiope argentata (Fabricius, 1775). Bryant, 1945: 361; Levi, 2004: 58; Penney & Perez-Gelabert, 2002: 211
Argiope trifasciata (Forskål, 1775). Bryant, 1945: 361; Levi, 2004: 54; Penney & Perez-Gelabert, 2002: 211
Cyclosa Menge, 1866
Cyclosa berlandi Levi, 1999. Penney & Perez-Gelabert, 2002: 211
Cyclosa bifurcata (Walckenaer, 1842). Levi, 1999: 365; Penney & Perez-Gelabert, 2002: 211
Cyclosa caroli (Hentz, 1850). Levi, 1999: 336; Penney & Perez-Gelabert, 2002: 211
Cyclosa haiti Levi, 1999: 365. Penney & Perez-Gelabert, 2002: 211
Cyclosa turbinata (Walckenaer, 1842). Levi, 1999: 356; Penney & Perez-Gelabert, 2002: 211
Cyclosa walckenaeri (O.P.-Cambridge, 1889). Levi, 1977: 84; Levi, 1999: 360; Penney & Perez-Gelabert, 2002: 211
=Cyclosa oculata (Walckenaer, 1802). Bryant, 1945: 367 [Misidentification]
Cyrtophora Simon, 1864
Cyrtophora citricola (Forskål, 1775). Alayón García et al., 2001: 9; Penney & Perez-Gelabert, 2002: 211 [Introduced]
Enacrosoma Mello-Leitão, 1932
Enacrosoma verrucosa† (Wunderlich, 1988a: 107). Penney, 2001: 998 [Fossil in Dominican amber]
Eriophora Simon, 1864
Eriophora ravilla (C. L. Koch, 1844). Penney & Perez-Gelabert, 2002: 211
Eustala Simon, 1895
Eustala anastera (Walckenaer, 1842). Genaro, 2007: 4
Eustala bisetosa Bryant, 1945: 370. Penney & Perez-Gelabert, 2002: 211 *DR, *H > TL: Foot hills of Cordillera Central, south
of Santiago, 1,000-3,000 ft.
Eustala delasmata Bryant, 1945: 372. Penney & Perez-Gelabert, 2002: 211 *DR > TL: San José de las Matas, 1,500+ ft.
Eustala devia (Gerstch & Mulaik, 1936). Penney & Perez-Gelabert, 2002: 211
Eustala fuscovittata (Keyserling, 1864). Bryant, 1945: 372; Penney & Perez-Gelabert, 2002: 211
Eustala perdita Bryant, 1945: 373. Penney & Perez-Gelabert, 2002: 211
Eustala vegeta (Keyserling, 1865). Bryant, 1945: 374; Penney & Perez-Gelabert, 2002: 211
Fossilaraneus† Wunderlich, 1988
Fossilaraneus incertus† Wunderlich, 1988a: 113 [Fossil in Dominican amber]
Gasteracantha Sundevall, 1833
Gasteracantha cancriformis (Linnaeus, 1758). Bryant, 1945: 414; Levi, 1996: 140; Penney & Perez-Gelabert, 2002: 211
=Plectana atlantica Walckenaer, 1841: 167. Levi, 1996: 142
=Gasteracantha rubiginosa C. L. Koch, 1845: 55. Levi, 1996: 142
=Gasteracantha mascula Strand, 1916: 119. Levi, 1996: 142
=Gasteracantha tetracantha (Linnaeus, 1767). Bryant, 1945: 413; Levi, 1996: 142
Gea Koch, 1843
Gea heptagon (Hentz, 1850). Bryant, 1945: 361; Levi, 2004: 49; Penney & Perez-Gelabert, 2002: 211

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Kapogea Levi, 1997
Kapogea alayoi (Archer, 1958). De los Santos & Alayón, 2012: 27
Kapogea sellata (Simon, 1895: 155). Levi, 1997: 242; Penney & Perez-Gelabert, 2002: 211
Larinia Simon, 1874
Larinia minor (Bryant, 1945: 367). Harrod et al., 1990: 254; Penney & Perez-Gelabert, 2002: 211 *H > TL: Port-au-Prince
Mangora Cambridge, 1889
Mangora fascialata Franganillo, 1936. Levi, 2005: 162; Penney & Perez-Gelabert, 2002: 211
=Mangora striatipes Bryant, 1945: 375
Mecynogea Simon, 1903
Mecynogea martiana (Archer, 1958). Levi, 1997: 232; Penney & Perez-Gelabert, 2002: 211
Metazygia Cambridge, 1903
Metazygia cienaga Levi, 1995. Penney & Perez-Gelabert, 2002: 211
Metazygia crewi (Banks, 1903). Bryant, 1945: 364; Penney & Perez-Gelabert, 2002: 211
=Singa crewii Banks, 1903: 342
Metazygia dubia (Keyserling, 1864). Penney & Perez-Gelabert, 2002: 211
Matazygia genialis (Keyserling, 1892). De los Santos & Carrero, 2011: 55
Metazygia gregalis (O. P.-Cambridge, 1889). Penney & Perez-Gelabert, 2002: 211
=Metazygia manni Bryant, 1945: 377
Metepeira Cambridge, 1903
Metepeira compsa (Chamberlin, 1916). Piel, 2001: 48; Penney, 2004: 185
Metepeira datona Chamberlin & Ivie, 1942. Piel, 2001: 20; Penney & Perez-Gelabert, 2002: 211
=Metepeira inerma Bryant, 1945: 378
Metepeira jamaicensis Archer, 1958. Penney & Perez-Gelabert, 2002: 211
Metepeira triangularis (Franganillo, 1930). Piel, 2001: 63; Penney & Perez-Gelabert, 2002: 212
=Metepeira bani Archer, 1965: 132
Metepeira vigilax (Keyserling, 1893). Piel, 2001: 30; Penney & Perez-Gelabert, 2002: 212
=Metepeira dominicana Archer, 1965: 132
Micrathena Sundevall, 1833
Micrathena forcipata (Thorell, 1859). Chickering, 1964: 257; Penney & Perez-Gelabert, 2002: 212
=Acrosoma flavomaculata Keyserling, 1864. Chickering, 1964: 257
Micrathena militaris (Fabricius, 1775). Bryant, 1945: 414; Chickering, 1964: 269; Penney & Perez-Gelabert, 2002: 212
Micrathena similis Bryant, 1945: 415. Chickering, 1964: 279; Penney & Perez-Gelabert, 2002: 212
Molinaranea Mello-Leitão, 1940
Molinaranea mitnickii† Saupe, Selden & Penney, 2010: 715 [Fossil in Dominican amber]
Neoscona Simon, 1864
Neoscona arabesca (Walckenaer, 1842). Berman & Levi, 1971: 474; Levi, 1993: 230; Penney & Perez-Gelabert, 2002: 212
Neoscona marcanoi Levi, 1993: 226. Penney & Perez-Gelabert, 2002: 212; Alayón, 2011: 24 *DR > TL: Terreno Salado
between Neiba and Duvergé
Neoscona moreli (Vinson, 1863). Levi, 1993: 232; Penney & Perez-Gelabert, 2002: 212
=Neoscona neotheis (Petrunkevitch, 1911). Berman & Levi, 1971: 490; Levi, 1993: 232
Neoscona nautica (L. Koch, 1875). Berman & Levi, 1971: 498; Levi, 1993: 228; Penney & Perez-Gelabert, 2002: 212
=Epeira volucripes Keyserling, 1885: 528. Levi, 1993: 228
Neoscona oaxacensis (Keyserling, 1864). Bryant, 1945: 380; Penney & Perez-Gelabert, 2002: 212
Nephila Leach, 1815
Nephila breviembolus† Wunderlich, 1986: 79 [Fossil in Dominican amber]
Nephila clavipes (Linnaeus, 1767). Bryant, 1945: 395; Penney & Perez-Gelabert, 2002: 210
Nephila dommeli† Wunderlich, 1982: 32 [Fossil in Dominican amber]
Nephila furca† Wunderlich, 1986: 81 [Fossil in Dominican amber]
Nephila longembolus† Wunderlich, 1986: 82 [Fossil in Dominican amber]
Nephila sp.†—Wunderlich (1988: 100) [Fossil in Dominican amber]
Nephila tenuis† Wunderlich, 1986: 80 [Fossil in Dominican amber]
Ocrepeira Marx, 1883
Ocrepeira darlingtoni (Bryant, 1945: 382). Penney & Perez-Gelabert, 2002: 212
Ocrepeira serrallesi (Bryant, 1947). Penney & Perez-Gelabert, 2002: 212

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 67

Parawixia Cambridge, 1903
Parawixia tredecimnotata F. O. P.-Cambridge, 1904. Penney & Perez-Gelabert, 2002: 212
Pozonia Schenkel, 1953
Pozonia andujari Alayón García, 2007: 42 *DR > TL: Honduras-Matadero, provincia Peravia
Pulchellaranea† Poinar, 2014
Pulchellaranea pedunculata† Poinar, 2014: 106 [Fossil in Dominican amber]
Pycnosinga† Wunderlich, 1988
Pycnosinga fossilis† Wunderlich, 1988a: 106 [Fossil in Dominican amber]
Verrucosa Mello-Leitão, 1939
Verrucosa arenata (Walckenaer, 1842). Bryant, 1945: 384; Penney & Perez-Gelabert, 2002: 212
Wagneriana Cambridge, 1904
Wagneriana tauricornis (O. P.-Cambridge, 1889). Bryant, 1945: 385; Levi, 1991b: 411; Penney & Perez-Gelabert, 2002: 212
Wagneriana undecimtuberculata (Keyserling, 1865). Bryant, 1948: 440; Penney & Perez-Gelabert, 2002: 212
Wagneriana vallenuevo Alayón, 2011: 23 *DR > TL: Parque Nacional R. P. Rancier (Valle Nuevo)
Wagneriana vegas Levi, 1991b: 403. Penney & Perez-Gelabert, 2002: 212
Witica Cambridge, 1895
Witica crassicaudus (Keyserling, 1865). Bryant, 1945: 368; Penney & Perez-Gelabert, 2002: 212

Family LYCOSIDAE
Agalenocosa Mello-Leitão, 1944
Agalenocosa bryantae Roewer, 1951. Penney & Perez-Gelabert, 2002: 212
=Lycosa festina Bryant, 1948: 354
Arctosa Koch, 1847
Arctosa inconspicua (Bryant, 1948: 408). Penney & Perez-Gelabert, 2002: 212
Hogna Simon, 1885
Hogna reducta (Bryant, 1942). Bryant, 1948: 356; Penney & Perez-Gelabert, 2002: 212
Hogna tantilla (Bryant, 1948: 352). Penney & Perez-Gelabert, 2002: 212
Pardosa Koch, 1847
Pardosa hamifera F.O.P.-Cambridge, 1902. Penney & Perez-Gelabert, 2002: 212
Pardosa portoricensis Banks, 1902. Bryant, 1948: 356; Penney & Perez-Gelabert, 2002: 212
Pirata Sundevall, 1833
Pirata sedentarius Montgomery, 1904. Wallace & Exline, 1978: 72
Trochosa Koch, 1847
Trochosa reichardtiana (Strand, 1916: 141). Penney & Perez-Gelabert, 2002: 212

Family PISAURIDAE
Thaumasia Perty, 1833
Thaumasia velox Simon, 1898. Da Silva & Carico, 2012: 13
=Thaumasia annecta Bryant, 1948: 349. Penney & Perez-Gelabert, 2002: 212
Thaumasia marginella (C. L. Koch, 1847). Bryant, 1948: 439; Penney & Perez-Gelabert, 2002: 212
Tinus Cambridge, 1903
Tinus connexus (Bryant, 1940). Bryant, 1948: 350; Penney & Perez-Gelabert, 2002: 212

Family OXYOPIDAE
Hamataliwa Keyserling, 1887
Hamataliwa communicans (Chamberlin, 1925: 128). Bryant, 1948: 358; Penney & Perez-Gelabert, 2002: 212 *H > TL: Hayti
Hamataliwa haytiana (Chamberlin, 1925: 127). Bryant, 1948: 358; Penney & Perez-Gelabert, 2002: 212 *H > TL: Hayti
Hamataliwa nigritarsa Bryant, 1948: 360. Penney & Perez-Gelabert, 2002: 212 *DR > TL: Cordillera Central, Loma Rucilla
Mountains, 5,000-8,000 ft.
Hamataliwa rana (Simon, 1897). Penney & Perez-Gelabert, 2002: 212
Oxyopes Latreille, 1806
Oxyopes crewi Bryant, 1948: 361. Penney & Perez-Gelabert, 2002: 212; Santos, 2017: 471
Oxyopes defectus† Wunderlich, 1988a: 198 [Fossil in Dominican amber]
Peucetia Thorell, 1869

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Peucetia viridans (Hentz, 1832). Bryant, 1948: 363; Penney & Perez-Gelabert, 2002: 212

Family CTENIDAE
Ciba Bloom et al., 2014
Ciba seibo Alayón García & Agnarsson in Bloom et al., 2014: 150 *DR > TL: Cueva Seibo, Parque Nacional del Este
Ctenus Walckenaer, 1805
Ctenus avidus Bryant, 1948: 397. Penney & Perez-Gelabert, 2002: 212 Alayón García, 2004: 282 *H > TL: Pétionville
Ctenus darlingtoni Bryant, 1948: 398. Penney & Perez-Gelabert, 2002: 212
Ctenus haina Alayón García, 2004: 278 *DR > TL: Cajuilito, Haina, Prov. San Cristóbal
Ctenus haitiensis Strand, 1909: 323. Bryant, 1948: 440; Penney & Perez-Gelabert, 2002: 212; Alayón García, 2004: 281
Ctenus hiemalis Bryant, 1948: 399. Penney & Perez-Gelabert, 2002: 212
Ctenus insulanus Bryant, 1948: 400. Penney & Perez-Gelabert, 2002: 212 *DR > TL: San José de las Matas, 1,500 ft.
Ctenus jaragua Alayón García, 2004: 280 *DR > TL: Fondo Paradí, Parque Nacional Jaragua, Oviedo, Prov. Pedernales
Ctenus manni Bryant, 1948: 401. Penney & Perez-Gelabert, 2002: 212
Ctenus monticola Bryant, 1948: 401. Penney & Perez-Gelabert, 2002: 212 *H > TL: Furcy, Mt. Cabaio, 7,000 ft.
Ctenus naranjo Alayón García, 2004: 278 *DR > TL: El Naranjo, Puerto Escondido, Prov. Independencia
Ctenus ottleyi (Petrunkevitch, 1930). De los Santos & Alayón, 2012: 27
Cupiennius Simon, 1891
Cupiennius salei (Keyserling, 1877). Bryant, 1948: 402; Penney & Perez-Gelabert, 2002: 212
Cupiennius vodou Brescovit & Polotow, 2005: 772 *H > TL: Miragoâne, Centre
Nanoctenus† Wunderlich, 1988
Nanoctenus longipes† Wunderlich, 1988a: 199 [Fossil in Dominican amber]
Trujillina Bryant, 1948
Trujillina hursti (Bryant, 1948: 403). Penney & Perez-Gelabert, 2002: 212 *DR > TL: Puerto Plata
Trujillina spinipes Bryant, 1948: 405. Alayón García, 1995b: 2; Penney & Perez-Gelabert, 2002: 212 *DR > TL: Cordillera
Central, Loma Rucilla Mountains, 5,000-8,000 ft.

Family AGELENIDAE
Barronopsis Chamberlin & Ivie, 1941
Barronopsis barrowsi (Gertsch, 1934). Alayón, 2012: 21
Barronopsis pelempito Alayón, 2012: 18 *DR > TL: Hoyo de Pelempito, Parque Nacional Sierra de Bahoruco
Neowadotes Alayón García, 1995
Neowadotes casabito Alayón García, 1995a: 3. Penney & Perez-Gelabert, 2002: 213 *DR > TL: 11 km del entronque de
Abanico, Casabito, Cordillera Central

Family DESIDAE
Paratheuma Bryant, 1940
Paratheuma insulana (Banks, 1902). Banks, 1904: 270; Penney & Perez-Gelabert, 2002: 213

Family HAHNIIDAE
Neohannia Mello-Leitao, 1917
Neohannia sp.—De los Santos & Alayón (2012: 28)

Family DICTYNIDAE
Emblyna Chamberlin, 1948
Emblyna altamira (Gertsch & Davis, 1942). Penney, 2004: 185
=Phantyna guanica (Gertsch, 1946). Bryant, 1948: 372; Penney, 2004: 185
Hispaniolyna† Wunderlich, 1988
Hispaniolyna hirsuta† Wunderlich, 1988a: 194 [Fossil in Dominican amber]
Hispaniolyna magna† Wunderlich, 1988a: 193 [Fossil in Dominican amber]
Palaeodyctina† Wunderlich, 1988
Palaeodyctina intermedia† Wunderlich, 1988a: 185 [Fossil in Dominican amber]
Palaeodyctina longispina† Wunderlich, 1988a: 187 [Fossil in Dominican amber]
Palaeodyctina? singularis† Wunderlich, 1988a: 189 [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 69

Palaeodyctina spiculum† Wunderlich, 1988a: 186 [Fossil in Dominican amber]
Palaeodyctina termitophila† Wunderlich, 1988a: 184 [Fossil in Dominican amber]
Palaeodyctina unispina† Wunderlich, 1988a: 188 [Fossil in Dominican amber]
Palaeolathys† Wunderlich, 1986
Palaeolathys? circumductus† Wunderlich, 1988a: 183 [Fossil in Dominican amber]
Palaeolathys copalis† Wunderlich, 1986: 66 [Fossil in Dominican amber]
Palaeolathys quadruplex† Wunderlich, 1988a: 181 [Fossil in Dominican amber]
Palaeolathys similis† Wunderlich, 1988a: 182 [Fossil in Dominican amber]
Palaeolathys spinosa† Wunderlich, 1986: 66 [Fossil in Dominican amber]
Succinya† Wunderlich, 1988
Succinya longembolus† Wunderlich, 1988a: 191 [Fossil in Dominican amber]
Succinya pulcher† Wunderlich, 1988a: 190 [Fossil in Dominican amber]
Succinya spinipalpus† Wunderlich, 1988a: 192 [Fossil in Dominican amber]

Family AMAUROBIIDAE
Retiro Mello-Leitão, 1915
Retiro gratus (Bryant, 1948: 348). Penney & Perez-Gelabert, 2002: 213
Tugana Chamberlin, 1948
Tugana cavatica (Bryant, 1940). Alayón García, 1992a: 6
=Alauximus infumatus Bryant, 1948: 347. Alayón García, 1992a: 6
Tugana crassa (Bryant, 1948: 346). Alayón García, 1992a: 7; Penney & Perez-Gelabert, 2002: 213 *DR > TL: North
Cordillera Central, Loma Rucilla Mountains, 5,000-8,000 ft.

Family CHEIRACANTHIIDAE
Cheiracanthium Koch, 1839
Cheiracanthium inclusum (Hentz, 1847). Bryant, 1948: 420; Penney & Perez-Gelabert, 2002: 213
Strotarchus Simon, 1888
Strotarchus heidti† Wunderlich, 1988a: 224 [Fossil in Dominican amber]
Strotarchus sp.—Sánchez-Ruíz (2009: 24)

Family MITURGIDAE
Odo Keyserling, 1887
Odo abudi Alayón García, 2002c: 30. Penney, 2004: 186 *DR > TL: Loma La Vigia, Azua
Syspira Simon, 1895
Syspira agujas Brescovit, Sánchez-Ruiz & Bonaldo, 2018: 63 *DR > TL: Via Agujas, Prov. San José de Ocoa
Syspira cimitarra Brescovit, Sánchez-Ruiz & Bonaldo, 2018: 58 *DR > TL: Parque Nacional Valle Nuevo, 15 km of
Constanza, Prov. La Vega
Syspira eclectica Chamberlin, 1924. Ramírez, 2014: 371
Syspira jimmyi Brescovit, Sánchez-Ruiz & Bonaldo, 2018: 59 *DR > TL: Parque Nacional Valle Nuevo, 15 km of Constanza,
Prov. La Vega
Syspira medialuna Brescovit, Sánchez-Ruiz & Bonaldo, 2018: 62 *DR > TL: Reserva Científica Ebano Verde, 1029 m,
Constanza, Prov. La Vega
Teminius Keyserling, 1887
Teminius hirsutus (Petrunkevitch, 1925). Penney & Perez-Gelabert, 2002: 213
Teminius insularis (Lucas, 1857). Bryant, 1948: 409; Penney & Perez-Gelabert, 2002: 213
Teminius monticola Bryant, 1948: 411. Penney & Perez-Gelabert, 2002: 213 *DR > TL: Cordillera Central, Loma Rucilla
Mountains, 5,000-8,000 ft.

Family ANYPHAENIDAE
Anyphaena Sundevall, 1833
Anyphaena dominicana Roewer, 1951: 445. Penney & Perez-Gelabert, 2002: 213 *DR > TL: Cordillera Central, Loma Rucilla
Mountains, 5,000-8,000 ft.
=Anyphaena lauta Bryant, 1948: 421 [Preoccupied name]
Anyphaena modesta Bryant, 1948: 422. Penney & Perez-Gelabert, 2002: 213 *H > TL: Foot hills north east of Massif de la

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Hotte, 3,000-4,000 ft.
Anyphaena pusilla Bryant, 1948: 423. Penney & Perez-Gelabert, 2002: 213
Anyphaenoides Berland, 1913
Anyphaenoides bulla† (Wunderlich, 1988a: 220). Penney, 2000a: 223 [Fossil in Dominican amber]
Hibana Brescovit, 1991
Hibana tenuis (L. Koch, 1866). Bryant, 1948: 430; Penney & Perez-Gelabert, 2002: 213
=Aysha ravida Simon, 1897: 508
Hibana velox (Becker, 1879). Bryant, 1948: 430; Penney & Perez-Gelabert, 2002: 213
Lupettiana Brescovit, 1996
Lupettiana levii Brescovit, 1999: 72. Penney & Perez-Gelabert, 2002: 213 *DR > TL: Jardin Botánico, Santo Domingo
Lupettiana ligula† (Wunderlich, 1988a: 221). Penney, 2000a: 223 [Fossil in Dominican amber]
Lupettiana parvula (Banks, 1903: 342). Bryant, 1948: 430; Brescovit, 1999: 67; Penney & Perez-Gelabert, 2002: 214
Lupettiana spinosa (Bryant, 1948: 432). Brescovit, 1999: 70; Penney & Perez-Gelabert, 2002: 214
Thaloe Brescovit, 1993
Thaloe ebano Brescovit & Oliveira, 2019: 262 *DR > TL: Reserva Científica Ebano Verde, 1029 m
Thaloe ennery Brescovit, 1993. Penney & Perez-Gelabert, 2002: 214
Thaloe leboulet Brescovit & Oliveira, 2019: 259 *H > TL: Le Boulet
Thaloe remotus (Bryant, 1948: 424). Penney & Perez-Gelabert, 2002: 214
Wulfila Cambridge, 1895
Wulfila fasciculus (Bryant, 1948: 425). Penney & Perez-Gelabert, 2002: 214 *DR > TL: Cordillera Central, Loma Vieja, south
of Constanza, 6,000 ft.
Wulfila fragilis (Bryant, 1948: 427). Penney & Perez-Gelabert, 2002: 214 *H > TL: Foot hills northeast of Massif de la Hotte,
3,000-4,000 ft.
Wulfila gracilipes (Banks, 1903: 342). Bryant, 1948: 428. Penney & Perez-Gelabert, 2002: 214 *H > TL: Hayti
Wulfila spinipes† Wunderlich, 1988a: 218. Penney, 2000a: 224 [Fossil in Dominican amber]

Family LIOCRANIDAE
Lausus Simon, 1898
Lausus pulchellus Bryant, 1948: 437. Penney & Perez-Gelabert, 2002: 214 *DR > TL: Rain forest, north of Valle Nuevo,
Cordillera Central, 6000 feet+ [Incertae sedis]

Family PHRUROLITHIDAE
Phrurolithus Koch, 1839
Phrurolithus spinosus Bryant, 1948: 433. Penney & Perez-Gelabert, 2002: 213 *H > TL: Étang Lachaux

Family CLUBIONIDAE
Elaver Cambridge, 1898
Elaver arawakan Saturnino & Bonaldo, 2015: 25 *H > TL: La Boule
Elaver implicata (Gertsch, 1941: 18). Penney & Perez-Gelabert, 2002: 214; Saturnino & Bonaldo, 2015: 22
Elaver nutua† (Wunderlich, 1988a: 222). Penney & Perez-Gelabert, 2002: 214 [Fossil in Dominican amber]

Family CORINNIDAE
Castianeira Keyserling, 1880
Castianeira tenebricosa† Wunderlich, 1988a: 204 [Fossil in Dominican amber]
Chemmisomma Wunderlich, 1988
Chemmisomma dubia† Wunderlich, 1988a: 216 [Fossil in Dominican amber]
Corinna Koch, 1841
Corinna colombo Bonaldo, 2000. Alayón García, 2007: 43
Corinna flageliformis† Wunderlich, 1988a: 203 [Fossil in Dominican amber]
Corinna parvula Bryant, 1940. Bryant, 1948: 434; Penney & Perez-Gelabert, 2002: 214
Corinna pulchella (Bryant, 1948: 437). Penney & Perez-Gelabert, 2002: 214 *DR > TL: Rain forest north of Valle Nuevo,
Cordillera Central, 6,000 ft.+
Corinna sp.—Alayón García (2007: 43)
Corinna toussainti Bryant, 1948: 435. Penney & Perez-Gelabert, 2002: 214 *H > TL: La Visite, La Selle Range, 6,000-7,000 ft.

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 71

Creugas Thorell, 1878
Creugas gulosus Thorell, 1878. Bonaldo, 2000: 99; Penney & Perez-Gelabert, 2002: 214
Mazax O.-P. Cambridge, 1898
Mazax sp.—De los Santos & Alayón (2012: 27)
Megalostrata Karsch, 1880
Megalostrata grandis† Wunderlich, 1988a: 201 [Fossil in Dominican amber]
Megalostrata raptor (L. Koch, 1866). De los Santos & Alayón, 2012: 28
Xeropigo Cambridge, 1882
Xeropigo tridentiger (O. P.-Cambridge, 1869). Bonaldo, 2000: 56; Penney & Perez-Gelabert, 2002: 214

Family TRACHELIDAE
Trachelas Koch, 1866
Trachelas bicolor Keyserling, 1887. Bryant, 1948: 439; Chickering, 1972b: 216; Penney & Perez-Gelabert, 2002: 214
Trachelas dilatus Platnick & Shadab, 1974: 18. Penney & Perez-Gelabert, 2002: 214 *DR > TL: San José de las Matas, elev.
1500 ft.
Trachelas erectus Platnick & Shadab, 1974: 19. Penney & Perez-Gelabert, 2002: 214 *H > TL: La Visite, elev. 6000 to 7000 ft.
Trachelas poinari† Penney, 2001: 1000 [Fossil in Dominican amber]
Trachelas tomaculus Platnick & Shadab, 1974: 19. Penney & Perez-Gelabert, 2002: 214

Family TROCHANTERIIDAE
Veterator Petrunkevitch, 1963
Veterator angustus† Wunderlich, 1988a: 211 [Fossil in Dominican amber]
Veterator ascutum† Wunderlich, 1988a: 208 [Fossil in Dominican amber]
Veterator incompletus† Wunderlich, 1982: 40 [Fossil in Dominican amber]
Veterator longipes† Wunderlich, 1988a: 209 [Fossil in Dominican amber]
Veterator loricatus† Wunderlich, 1988a: 212 [Fossil in Dominican amber]
Veterator porrectus† Wunderlich, 1988a: 206 [Fossil in Dominican amber]
Veterator viduus† Wunderlich, 1988a: 214 [Fossil in Dominican amber]

Family PRODIDOMIDAE
Lygromma Simon, 1893
Lygromma chamberlini Gertsch, 1941. Sánchez-Ruíz, 2009: 24
Zimiris Simon, 1882
Zimiris doriai Simon, 1882. Platnick & Penney, 2004: 8; Penney, 2004: 186

Family GNAPHOSIDAE
Camillina Berland, 1919
Camillina sp.—De los Santos & Alayón (2012: 28)
Cesonia Simon, 1893
Cesonia nadleri Platnick & Shadab, 1980: 382. Penney & Perez-Gelabert, 2002: 214 *DR > TL: Boca Chica, National District
Drassyllinus† Wunderlich, 1988
Drassyllinus aliter† Wunderlich, 1988a: 227 [Fossil in Dominican amber]
Gnaphosa Latreille, 1804
Gnaphosa sericata (L. Koch, 1866). Penney & Perez-Gelabert, 2002: 215
Sergiolus Simon, 1892
Sergiolus magnus (Bryant, 1948: 406). Penney & Perez-Gelabert, 2002: 215 *H > TL: Miragoâne
Scopoides Platnick, 1989
Scopoides dominicanus† Wunderlich, 2011: 467 [Fossil in Dominican amber]
Zelotes Gistel, 1848
Zelotes sp.—De los Santos & Alayón (2012: 28)

Family SELENOPIDAE
Selenops Latreille, 1819
Selenops anacaona Crews, 2018: 129 *DR> TL: Unmarked track into Parque Nacional Jaragua between Manuel Goya and

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Oviedo, Pedernales, 83 m
Selenops bani Alayón García, 1992b: 5. Penney & Perez-Gelabert, 2002: 215; Crews, 2011: 118 *DR > TL: La Laguna,
carretera Baní-Manaclar
Selenops beynai† Schawaller, 1984: 3; Crews, 2011: 95 [Fossil in Dominican amber]
Selenops bocacanadensis Crews, 2011: 103; Crews, 2018: 133 *DR > TL: Km 13 Carretera Alcoa, Pedernales
Selenops caonabo Crews, 2018: 130 *DR > TL: Los Tablones, La Vega
Selenops denia Crews, 2011: 108 *DR > TL: Mata Grande, Santiago, 1009 m
Selenops dominicanus† Wunderlich, 2004: 1734 [Fossil in Dominican amber]
Selenops duan Crews, 2011: 107 *DR > TL: Cañada Miguel, Elías Piña
Selenops enriquillo Crews, 2011: 122 *DR > TL: La Azufrada, La Descubierta, Lago Enriquillo, -15 m
Selenops guerrero Crews, 2011: 110 *DR > TL: Loma de Isabel, Puerto Plata, ~96 m
Selenops insularis Keyserling, 1881. Bryant, 1948: 414; Muma, 1953: 24; Alayón García, 1992b: 2; Penney & Perez-Gelabert,
2002: 215; Crews, 2011: 96
Selenops lindborgi Petrunkevitch, 1926. Crews & Gillespie, 2010: 297; Crews, 2011: 77
Selenops marcanoi Alayón García, 1992b: 3. Penney & Perez-Gelabert, 2002: 215; Crews, 2011: 101 *DR > TL: El Aguacate,
Neyba
Selenops morro Crews, 2011: 126; Crews, 2018: 134 *DR > TL: El Morro, Monte Cristi, 20-40 m
Selenops oviedo Crews, 2011: 104 *DR > TL: El Cajuil, Laguna Oviedo, Pedernales, ~86 m
Selenops pensilis Muma, 1953: 42. Alayón García, 1992b: 2; Penney & Perez-Gelabert, 2002: 215; Crews, 2011: 120; Crews,
2018: 128 *H > TL: Grand’Anse, Haiti
Selenops phaselus Muma, 1953: 42. Penney & Perez-Gelabert, 2002: 215; Crews, 2011: 124 *H, *DR > TL: Kenscoff, Haiti,
4500-5500 feet
Selenops sp.†—Penney (2001: 1002) [Fossil in Dominican amber]

Family SPARASSIDAE
Heteropoda Nuttall, 1834
Heteropoda venatoria (Linnaeus, 1767). Bryant, 1948: 414; Penney & Perez-Gelabert, 2002: 215
Neostasina Rheims & Alayón, 2016
Neostasina antiguensis (Bryant, 1923). Bryant, 1948: 414; Rheims & Alayón, 2016: 310
Neostasina baoruco Rheims & Alayón, 2016: 310 *DR > TL: Pedernales, Rabo de Gato Lagoon
Neostasina bermudezi Rheims & Alayón, 2016: 312 *DR > TL: La Ciénaga, Parque Nacional A. Bermúdez
Neostasina bicolor (Banks, 1914). Rheims & Alayón, 2016: 313
Neostasina cachote Rheims & Alayón, 2016: 316 *DR > TL: Cachote, Parque Nacional Sierra de Baoruco
Neostasina setosa (Bryant, 1948: 413). Rheims & Alayón, 2016: 336
Neostasina taino Rheims & Alayón, 2016: 338 *DR > TL: San Rafael, Parque Nacional del Este
Pseudosparianthis Simon, 1887
Pseudosparianthis pfeifferi† (Wunderlich, 1988a: 233). Penney, 2001: 1004 [Fossil in Dominican amber]

Family PHILODROMIDAE
Apollophanes Cambridge, 1898
Apollophanes punctatus (Bryant, 1948: 419). Penney & Perez-Gelabert, 2002: 215 *DR > TL: Cordillera Central, Valle Nuevo,
south east of Constanza, 7,000 ft.
Tibellus Simon, 1875
Tibellus sp.—De los Santos & Alayón (2012: 28)

Family THOMISIDAE
Heterotmarus Wunderlich, 1988
Heterotmarus altus† Wunderlich, 1988a: 229 [Fossil in Dominican amber]
Isaloides Cambridge, 1900
Isaloides toussaintii Banks, 1903: 343. Bryant, 1948: 416; Penney & Perez-Gelabert, 2002: 215
Komisumena Ono, 1981
Komisumena rosae† Ono, 1981: 4 [Fossil in Dominican amber]
Mecaphesa Simon, 1900
Mecaphesa asperata (Hentz, 1847). Penney & Perez-Gelabert, 2002: 215

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Mecaphesa californica (Banks, 1896). Penney, 2004: 187
Mecaphesa celer (Hentz, 1847). Bryant, 1948: 420; Penney & Perez-Gelabert, 2002: 215
Misumenops Cambridge, 1900
Misumenops bellulus (Banks, 1896). Bryant, 1948: 420; Penney & Perez-Gelabert, 2002: 215
Rejanellus Lise Arno, 2005
Rejanellus pallescens (Bryant, 1940). Lise, 2005: 155
Rejanellus venustus (Bryant, 1948: 416). Lise, 2005: 153; Penney & Perez-Gelabert, 2002: 215 *DR > TL: Loma Rucilla
Mountains, Cordillera Central
Tobias Simon, 1895
Tobias taczanowskii Roewer, 1951. Penney & Perez-Gelabert, 2002: 215
Xysticus Koch in Panzer, 1834
Xysticus pellax O.-P. Cambridge, 1894. De los Santos & Alayón, 2012: 28

Family SALTICIDAE
Subfamily EUOPHRYINAE
Agobardus Keyserling, 1885
Agobardus anormalis Keyserling, 1885. Bryant, 1943: 450; Penney & Perez-Gelabert, 2002: 215
Agobardus anormalis montanus Bryant, 1943: 452. Penney & Perez-Gelabert, 2002: 215 *H, *DR > TL: Foot hills northeast of
La Hotte, 3000–4000 m
Agobardus bahoruco Zhang & Maddison, 2012: 3 *DR > TL: Pedernales, Parque Nacional Sierra de Bahoruco, 1340 m
Agobardus brevitarsus Bryant, 1943: 453. Penney & Perez-Gelabert, 2002: 215 *H > TL: La Visite, 6000–7000 m
Agobardus cordiformis Zhang & Maddison, 2012: 5 *DR > TL: East of Pedernales, 30 m
Agobardus gramineus Zhang & Maddison, 2012: 7 *DR > TL: Near Pedernales, 8 m
Agobardus modestus (Bryant, 1943: 466). Penney & Perez-Gelabert, 2002: 215; Zhang & Maddison, 2015: 18 *DR > TL: Valle
Nuevo, southeast of Constanza, 7000 feet
Agobardus obscurus Bryant, 1943: 455. Penney & Perez-Gelabert, 2002: 215 *H > TL: Swamp north of Dessalines
Agobardus oviedo Zhang & Maddison, 2012: 10 *DR > TL: Laguna de Oviedo, Pedernales, 5 m
Agobardus perpilosus Bryant, 1943: 456. Penney & Perez-Gelabert, 2002: 215 *DR > TL: Loma Rucilla, Pico del Yaque,
8000-10000 feet
Agobardus phylladiphilus Zhang & Maddison, 2012: 11 *DR > TL: Pedernales, Parque Nacional Sierra de Bahoruco, 1340 m
Amphidraus Simon, 1900
=Nebridia Simon, 1902
Amphidraus manni (Bryant, 1943: 492); Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 22 *H > TL: Foot
hills of La Hotte, 3000-4000 feet
Amphidraus mendicus (Bryant, 1943: 494). Penney & Perez-Gelabert, 2002: 216 *DR > TL: Rain forest near Valle Nuevo,
6000 feet
Anasaitis Bryant, 1950
Anasaitis adorabilis Zhang & Maddison, 2012: 13 *DR > TL: Laguna de Oviedo, Pedernales, 5 m
Anasaitis banksi (Roewer, 1951). Penney & Perez-Gelabert, 2002: 215; Zhang & Maddison, 2015: 15
Anasaitis brunnea Zhang & Maddison, 2012: 15 *DR > TL: Highway 44, south of Barahona, 50 m
Anasaitis elegantissima (Simon, 1888). Bryant, 1943: 471; Penney & Perez-Gelabert, 2002: 215; Zhang & Maddison, 2015: 15
*H, *DR > TL: San Domingo
Anasaitis hebetata Zhang & Maddison, 2012: 17 *DR > TL: Parque Nacional Sierra Martín García, 170 m
Anasaitis laxa Zhang & Maddison, 2012: 19 *DR > TL: Parque Nacional Sierra Martín García, 170 m
Anasaitis locuples (Simon, 1888). Bryant, 1943: 474; Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 15 *H,
*DR > TL: San Domingo
Anasaitis perplexa (Peckham & Peckham, 1901). Banks, 1903: 341; Proszynski, 2016: 3198
Antillattus Bryant, 1943
Antillatus applanatus Zhang & Maddison, 2012: 22 *DR > TL: Road Constanza to Ocoa, Valle Nuevo, 2070 m
Antillatus cambridgei (Bryant, 1943: 457). Penney & Perez-Gelabert, 2002: 215; Zhang & Maddison, 2015: 17 *DR > TL:
Constanza, 3000-4000 feet
Antillatus darlingtoni (Bryant, 1943: 504). Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 17 *DR > TL:
Loma Rucilla Mountains, 5000-8000 feet
Antillatus electus (Bryant, 1943: 506). Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 17 *DR > TL: Valle

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Nuevo, southeast of Constanza, 7000 feet
Antillattus gracilis Bryant, 1943: 461. Penney & Perez-Gelabert, 2002: 215 *H > TL: Dame-Marie
Antillatus maxillosus (Bryant, 1943: 508). Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 17 *DR > TL:
Loma Vieja, south of Constanza, 6000 feet
Antillatus montanus (Bryant, 1943: 510). Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 17 *DR > TL: Loma
Rucilla, Pico del Yaque, 3000-10000 feet
Antillatus peckhami (Bryant, 1943: 512). Penney & Perez-Gelabert, 2002: 217; Zhang & Maddison, 2015: 17 *DR > TL: Loma
Rucilla Mountains, 5000-8000 feet
Antillattus placidus Bryant, 1943: 462. Penney & Perez-Gelabert, 2002: 215 *H > TL: Ennery
Antillatus scutiformis Zhang & Maddison, 2015: 17 *DR > TL: Rain forest near Valle Nuevo, 6000 feet
=Siloca electa Bryant, 1943: 516
Bythocrotus Simon, 1903
Bythocrotus cephalotes (Simon, 1888). Bryant, 1943: 464; Penney & Perez-Gelabert, 2002: 215 *H, *DR > TL: San Domingo
Bythocrotus crypticus Zhang & Maddison, 2012: 23 *DR > TL: Parque Nacional Sierra Martín García, 170 m
Cobanus Cambridge, 1900
Cobanus sp.—De los Santos & Alayón (2012: 30)
Compsodecta Simon, 1903
Compsodecta haytiensis (Banks, 1903: 344). Bryant, 1943: 468; Penney & Perez-Gelabert, 2002: 215 *H > TL: Hayti
Compsodecta peckhami Bryant, 1943: 470. Penney & Perez-Gelabert, 2002: 215 *H > TL: Foot hills of Massif de la Hotte,
3000-4000 feet
Corticatus Zhang & Maddison, 2012
Corticatus latus Zhang & Maddison, 2012: 29 *DR > TL: Laguna de Oviedo, Pedernales, 5 m
Corythalia Koch, 1850
=Dinattus Bryant, 1943
Corythalia broccai Zhang & Maddison, 2012: 30 *DR > TL: Pedro Sánchez - Miches road, 157 m
Corythalia bromelicola Zhang & Maddison, 2012: 32 *DR > TL: Cachote - La Ciénaga road, Barahona, 730 m
Corythalia coronai Zhang & Maddison, 2012: 36 *DR > TL: Rio Mulito, Pedernales, 270 m
Corythalia decora (Bryant, 1943: 519). Penney & Perez-Gelabert, 2002: 217; Zhang & Maddison, 2015: 16 *H > TL:
Kenscoff, 5,000-7,000 ft.
Corythalia ocululiter† Wunderlich, 1988a: 241 [Fossil in Dominican amber]
Corythalia erebus (Bryant, 1943: 485. Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 15 *H > TL: La Visite,
6000-7000 feet
Corythalia heros (Bryant, 1943: 482). Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 15 *H > TL: Foot hills
northeast of La Hotte, 4000 feet
Corythalia minor (Bryant, 1943: 485). Penney & Perez-Gelabert, 2002: 216; Zhang & Maddison, 2015: 15 *H > TL: Foot hills
northeast of La Hotte, 4000 feet
Corythalia peblique Zhang & Maddison, 2012: 38 *DR > TL: Peblique, Pedernales, 270 m
Corythalia pilosa† Wunderlich, 1982: 43 [Fossil in Dominican amber]
Corythalia scissa† Wunderlich, 1988a: 240 [Fossil in Dominican amber]
Descangeles† Wunderlich, 1988
Descangeles pygmaeus† Wunderlich, 1988a: 248 [Fossil in Dominican amber]
Descanso Peckham, 1892
Descanso formosus Bryant, 1943: 476. Penney & Perez-Gelabert, 2002: 216 *H > TL: Trou Caiman
Descanso magnus Bryant, 1943: 479. Penney & Perez-Gelabert, 2002: 216 *DR > TL: Jarabacoa
Descanso montanus Bryant, 1943: 480. Penney & Perez-Gelabert, 2002: 216 *DR > TL: Constanza to Jarabacoa, 2000-4000
feet
Eris Koch, 1846
Eris flava (Peckham & Peckham, 1888). Maddison, 1986: 145; Penney & Perez-Gelabert, 2002: 216
Habronattus Cambridge, 1901
Habronattus brunneus (Peckham & Peckham, 1901). Bryant, 1943: 487; Penney & Perez-Gelabert, 2002: 216
Hasarius Simon, 1872
Hasarius adansoni (Audouin, 1826). Penney & Perez-Gelabert, 2002: 216
Hentzia McCook, 1894
Hentzia antillana Bryant, 1940. Bryant, 1943: 487; Penney & Perez-Gelabert, 2002: 216

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Hentzia mandibularis (Bryant, 1943: 500). Penney & Perez-Gelabert, 2002: 216 *DR, *H > TL: Foot hills of Cordillera
Central, south of Santiago, 1000-3000 feet
Hentzia vittata (Keyserling, 1885). Penney & Perez-Gelabert, 2002: 216
Hentzia zombia Richman, 1989. Penney & Perez-Gelabert, 2002: 216
Icius Simon, 1876
Icius separatus Banks, 1903: 344. Bryant, 1948: 440; Penney & Perez-Gelabert, 2002: 216
Jollas Simon, 1901
Jollas armatus (Bryant, 1943: 495). Penney & Perez-Gelabert, 2002: 216 *DR > TL: Loma Rucilla Mountains, Cordillera
Central, 5000-8000 feet
Jollas crassus (Bryant, 1943: 497). Penney & Perez-Gelabert, 2002: 216 *DR > TL: Sánchez
Lyssomanes Hentz, 1845
Lyssomanes antillanus Peckham et al., 1889: 226. Bryant, 1943: 489; Penney & Perez-Gelabert, 2002: 216
Lyssomanes pristinus† Wunderlich, 1986: 86. Penney, 2001: 1005 [Fossil in Dominican amber]
=Lyssomanes galianoae† Reiskind, 1989: 281 [Fossil in Dominican amber]
Lyssomanes pulcher† Wunderlich, 1988a: 238 [Fossil in Dominican amber]
Lyssomanes viridis (Walckenaer, 1837). Penney & Perez-Gelabert, 2002: 216
Menemerus Simon, 1869
Menemerus bivittatus (Dufour, 1831). Bryant, 1943: 489; Penney & Perez-Gelabert, 2002: 216
Metacyrba Cambridge, 1901
Metacyrba pictipes Banks, 1903: 343. Bryant, 1943: 489; Penney & Perez-Gelabert, 2002: 216 *H > TL: Hayti
Metacyrba taeniola (Hentz, 1846). Bryant, 1943: 490; Penney & Perez-Gelabert, 2002: 216
Myrmarachne MacLeay, 1838
Myrmarachne parallela (Fabricius, 1794). Penney & Perez-Gelabert, 2002: 216
Nilakantha Peckham & Peckham, 1901
Nilakantha beugelorum† (Wolff, 1990: 406). Bustamante et al., 2015: 188 [Fossil in Dominican amber]
Nilakantha cockerelli Peckham & Peckham, 1901. Banks, 1903: 341; Proszynski, 2016: 3947
Paraphidippus Cambridge, 1901
Paraphidippus aurantius (Lucas, 1833). Penney & Perez-Gelabert, 2002: 216
Parathiodina Bryant, 1943
Parathiodina compta Bryant, 1943: 502. Penney & Perez-Gelabert, 2002: 216 *H > TL: Camp Perrin
Peckhamia Simon, 1901
Peckhamia americana (Peckham & Peckham, 1892). Sánchez-Ruíz, 2009: 24
Pelegrina Franganillo, 1930
Pelegrina proxima (Peckham & Peckham, 1901). Maddison, 1996: 265; Penney & Perez-Gelabert, 2002: 216
Pensacolatus† Wunderlich, 1988
Pensacolatus coxalis† Wunderlich, 1988a: 243 [Fossil in Dominican amber]
Pensacolatus spinipes† Wunderlich, 1988a: 244 [Fossil in Dominican amber]
Pensacolatus? tibialis† Wunderlich, 2004: 1802 [Fossil in Dominican amber]
Phidippus Koch, 1846
Phidippus regius C. L. Koch, 1846. Bryant, 1943: 514; Penney & Perez-Gelabert, 2002: 217
Phlegrata Wunderlich, 1988
Phlegrata pala† Wunderlich, 1988a: 246 [Fossil in Dominican amber]
Plexippus Koch, 1846
Plexippus paykulli (Audouin, 1826). Bryant, 1943: 516; Penney & Perez-Gelabert, 2002: 217
Popcornella Zhang & Maddison, 2012
Popcornella furcata Zhang & Maddison, 2012: 40 *DR > TL: Reserva Científica Ébano Verde (antenna), 1460 m
Popcornella spiniformis Zhang & Maddison, 2012: 44 *DR > TL: Cachote, Barahona, 1220 m
Sarinda Peckham, 1892
Sarinda sp. indet.†—Wunderlich (2004ee: 1803) [Fossil in Dominican amber]
Truncattus Zhang & Maddison, 2012
Truncattus cachotensis Zhang & Maddison, 2012: 48 *DR > TL: Cachote, Barahona, 1220 m
Truncattus dominicanus Zhang & Maddison, 2012: 50 *DR > TL: P. N. Armando Bermúdez, 1120—1250 m
Truncattus flavus Zhang & Maddison, 2012: 52 *DR > TL: P. N. Armando Bermúdez, 1120—1250 m

76 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Order OPILIONES

Taxonomic summary:
Suborder Laniatores
Infraorder Grassatores
Superfamily Samooidea
Family Biantidae
Family Kimulidae
Family Samoidae
Superfamily Gonyleptoidea
Family Agoristenidae
Family Cosmetidae
Superfamily Phalangodoidea
Family Phalangodidae
Infraorder Grassatores Incertae Sedis
Suborder Eupnoi
Superfamily Phalangioidea
Family Sclerosomatidae

Suborder LANIATORES
Infraorder GRASSATORES
Superfamily SAMOOIDEA
Family BIANTIDAE
Bidoma Šilhavý, 1973
Bidoma indivisa Šilhavý, 1973b: 143 *H > TL: Grande Rivière
Vestitecola Šilhavý, 1973
Vestitecola haitensis Šilhavý, 1973b: 142 *H > TL: La Vestite [La Visite]

Family KIMULIDAE
Kimula Goodnight & Goodnight, 1942
Kimula cokendolpheri Pérez González & Armas, 2000: 257 *DR > TL: Casabito, Constanza
Kimula? sp.†—Cokendolpher & Poinar (1992: 55) [Fossil in Dominican amber]

Family SAMOIDAE
Hummelinckiolus Šilhavý, 1979
Hummelinckiolus silhavyi† Cokendolpher & Poinar, 1998: 10 [Fossil in Dominican amber]

Superfamily GONYLEPTOIDEA
Family AGORISTENIDAE
Agoristenus Šilhavý, 1973
Agoristenus haitensis Šilhavý, 1973b: 116 *DR > TL: Loma Vieja, Cord. Centr., S. of Constanza, 6000 ft.
Ahotta Šilhavý, 1973
Ahotta hispaniolica Šilhavý, 1973b: 119 *H > TL: Foothills NE of La Hotte, 3–4000 ft.
Haitimera Šilhavý, 1973
Haitimera peninsularis Šilhavý, 1973b: 119 *H > TL: La Visite, 6–7000 ft.
Meriosfera Šilhavý, 1973
Meriosfera gertschi Šilhavý, 1973b: 124 *H > TL: La Visite, 6–7000 ft.
Meriosfera lineata Šilhavý, 1973b: 125 *H > TL: La Hotte, 5–7000 ft.

Family COSMETIDAE
Arucillus Šilhavý, 1971
Arucillus armasi Pérez González & Vasconcelos, 2003: 135 *DR > TL: La Nevera, Valle Nuevo, Prov. La Vega
Arucillus hispaniolicus Šilhavý, 1971: 139 *DR > TL: Loma Rucilla, 8–10,000 ft.
Cynorta Koch, 1839

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Cynorta hassleri Goodnight & Goodnight, 1942: 8 *H > TL: Aux Cayes
Cynortoides Roewer, 1912
Cynortoides caraibica (Sørensen, 1932: 408). Kury, 2003: 50; Kury & Alonso-Zarazaga, 2011: 50
Cynortoides haitiensis Roewer, 1912: 62. Goodnight & Goodnight, 1942: 10 *H, *DR > TL: Sánchez, Samaná Bay
Cynortoides marginata Goodnight & Goodnight, 1942: 10; Kury & Alonso-Zarazaga, 2011: 50 *DR > TL: Dominican
Republic
Cynortoides v-album (Simon, 1879: 198)
Metacynortoides Roewer, 1912
Metacynortoides bilineatus Goodnight & Goodnight, 1942: 14 *DR > TL: Sánchez
Metacynortoides obscurus (Banks, 1903). Kury, 2003: 68
Metacynortoides romanus Goodnight & Goodnight, 1942: 15 *DR > TL: La Romana
Metacynortoides transversalis Goodnight & Goodnight, 1942: 15 *DR > TL: La Romana
Proerginus Roewer, 1917
Proerginus lineatus Roewer, 1917: 99 *DR > Santo Domingo: around the city

Superfamily PHALANGODOIDEA
Family PHALANGODIDAE
Arganotus Šilhavý, 1971
Arganotus robustus Šilhavý, 1979: 13 *H > TL: La Visite, 6–7000 ft.
Orsa Šilhavý, 1979
Orsa daphne Šilhavý, 1979: 4 *H > TL: La Visite, 6–7000 ft.
Pellobunus Banks, 1905
Pellobunus haitiensis (Šilhavý, 1979: 25) *H > TL: Manneville
Pellobunus proavus† Cokendolpher, 1986: 206 [Fossil in Dominican amber]

Infraorder GRASSATORES Incertae Sedis

Mirda Šilhavý, 1973a
Mirda insulanus (Banks, 1901). Banks, 1903: 342; Šilhavý, 1973a: 146; Kury, 2003: 25
Philacarus Sorensen, 1932
Philacarus hispaniolensis† Cokendolpher & Poinar, 1992: 54 [Fossil in Dominican amber]

Suborder EUPNOI
Superfamily PHALANGIOIDEA
Family SCLEROSOMATIDAE
Geaya Roewer, 1910
Geaya haitiensis Goodnight & Goodnight, 1942: 3 *H > TL: Kenscoff

Order PALPIGRADI
Family EUKOENENIIDAE
Eukoenenia Boerner, 1901
Eukoenenia sp.—Armas (2004: 29)

Order PSEUDOSCORPIONES
Key reference: Harvey (1990); Cosgrove et al. (2016).

Taxonomic summary:
Superfamily Chthonioidea
Family Chthoniidae
Superfamily Neobisioidea
Family Bochicidae
Family Syariniidae
Superfamily Garypoidea
Family Olpiidae
Superfamily Cheiridioidea

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 Family Cheiridiidae
Family Sternophoridae
Superfamily Cheliferoidea
Family Atemnidae
Family Chernetidae
Family Pseudochiridiidae
Family Withiidae
Family Cheliferidae

Superfamily CHTHONIOIDEA
Family CHTHONIIDAE
Lechytia Balzan, 1892
Lechytia tertiaria† Schawaller, 1980: 9 [Fossil in Dominican amber]
Lechytia trinitatis Beier, 1970. Beier, 1976: 46
Pseudochthonius Balzan, 1892
Pseudochthonius squamosus† Schawaller, 1980: 3 [Fossil in Dominican amber]
Tyrannochthonius Chamberlin, 1929
Tyrannochthonius imitatus Hoff, 1959. Beier, 1976: 45; Harvey, 1990: 209; Cosgrove et al., 2016: 264
Lagynochthonius Beier, 1951
Lagynochthonius proximus (Hoff, 1959). Beier, 1976: 45; Harvey, 1990: 212; Cosgrove et al., 2016: 264
Lagynochthonius dominicanus (Muchmore, 1996: 104). Cosgrove et al., 2016: 264

Superfamily NEOBISIOIDEA
Family BOCHICIDAE
Mexobisium Muchmore, 1972
Mexobisium dominicanum Muchmore, 1998: 126 *DR > TL: Cuevas Pomier, Borbón, San Cristóbal Prov.

Family SYARINIIDAE
Ideobisium Balzan, 1892
Ideobisium puertoricense Muchmore, 1982: 201

Superfamily GARYPOIDEA
Family OLPIIDAE
Antillolpium Muchmore, 1991
Antillolpium hummelincki Muchmore, 1991. Cosgrove et al., 2016: 264
Novohorus Hoff, 1945
Novohorus suffuscus Hoff, 1945. Cosgrove et al., 2016: 264
Pachyolpium Beier, 1931
Pachyolpium medium Hoff, 1945. Cosgrove et al., 2016: 264
=Olpiolum monae Hoff, 1964. Beier, 1976: 46; Harvey, 1990: 290
Planctolpium Hoff, 1964
Planctolpium arboreum Hoff, 1964. Beier, 1976: 46; Harvey, 1990: 297; Cosgrove et al., 2016: 264

Superfamily CHEIRIDIOIDEA
Family CHEIRIDIIDAE
Cryptocheiridium Chamberlin, 1931
Cryptocheiridium antiquum† Schawaller, 1981: 3 [Fossil in Dominican amber]

Family STERNOPHORIDAE
Garyops Banks, 1909
Garyops depressus Banks, 1909. Beier, 1976: 46; Harvey, 1990: 449
Idiogaryops Hoff, 1963
Idiogaryops pumilus† Judson, 1998: 419 [Fossil in Dominican amber]

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Superfamily CHELIFEROIDEA
Family ATEMNIDAE
Paratemnoides Harvey, 1990
Paratemnoides elongatus (Banks, 1895). Beier, 1976: 46; Harvey, 1990: 470 [Dubious record in need of confirmation]

Family CHERNETIDAE
Americhernes Muchmore, 1976
Americhernes sp.†—Schawaller (1981: 14) [Fossil in Dominican amber]
Byrsochernes Beier, 1959
Byrsochernes caribicus Beier, 1976: 50. Harvey, 1990: 548
Caribochernes Beier, 1976
Caribochernes pumilus Beier, 1976: 49 *DR > TL: Colonia, Cord. Central, 1000 m
Chernes Menge, 1855
Chernes hispaniolicus Beier, 1976: 52 *DR > TL: San Cristóbal
Gomphochernes Beier, 1932
Gomphochernes communis (Balzan, 1888). Beier, 1976: 46; Harvey, 1990: 578 [Dubious record in need of confirmation]
Hesperochernes Chamberlin, 1924
Hesperochernes vespertilionis Beier, 1976: 53 *DR > TL: Sabaneta
Pachychernes Beier, 1932
Pachychernes effossus† Schawaller, 1980: 5 [Fossil in Dominican amber]
Pachychernes sp.†—Schawaller (1980: 12) [Fossil in Dominican amber]
Parachernes Chamberlin, 1931
Parachernes (Parachernes) dominicanus Beier, 1976: 47 *DR > TL: Boca Chica, 10 m
Parachernes (Scapanochernes) compressus (Tullgren, 1907). Beier, 1976: 48
Parazaona Beier, 1932
Parazaona klapperichi Beier, 1976: 51 *DR > TL: Sabaneta

Family PSEUDOCHIRIDIIDAE
Pseudochiridium With, 1906
Pseudochiridium insulae Hoff, 1964. Judson, 2007: 49
Pseudochiridium lindae† Judson, 2007: 46 [Fossil in Dominican amber]

Family WITHIIDAE
Cacodemonius Chamberlin, 1931
Cacodemonius segmentidentatus (Balzan, 1887). Harvey, 1990: 642 [Dubious record in need of confirmation]
Dolichowithius Chamberlin, 1931
Dolichowithius sp.—Muchmore, 1997: 279
=Dolichowithius simplex Beier, 1932. Beier, 1976: 55; Harvey, 1990: 644

Family CHELIFERIDAE
Cubachelifer Hoff, 1946
Cubachelifer strator Hoff, 1946. Beier, 1976: 57 [Dubious record in need of confirmation]
Parachelifer Chamberlin, 1932
Parachelifer dominicanus Beier, 1976: 55 *DR > TL: Boca Chica, 10 m
Parawithius Chamberlin, 1931
Parawithius? sp.†—Schawaller (1981: 5) [Fossil in Dominican amber]
Tyrannochelifer Chamberlin, 1932
Tyrannochelifer floridanus (Banks, 1891). Beier, 1976: 58; Harvey, 1990: 533
Tyrannochelifer macropalpus (Tullgren, 1907: 26) *H > TL: St. Marc

Order SCHIZOMIDA
Key reference: Armas (2004).

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Family HUBBARDIIDAE
Antillostenochrus Armas & Teruel, 2002
Antillostenochrus subcerdoso (Armas & Abud, 1990: 11). Armas, 2004: 14 *DR > TL: La Laguna, carretera Baní a Manaclar
km 6, Baní, Prov. Peravia
Antillostenochrus pseudoannulatus† (Krüger & Dunlop, 2010: 51). Armas & Teruel, 2011: 336 [Fossil in Dominican amber]
Rowlandius Reddell & Cokendolpher, 1995
Rowlandius anasilviae (Armas & Abud, 1990: 21). Armas, 2004: 16 *DR > TL: La Laguna, sección Montería, Baní, Prov.
Peravia
Rowlandius casabito (Armas & Abud, 1990: 6). Armas, 2004: 16 *DR > TL: Carretera Bonao a Casabito, km 10.5, aprox. 1100
m
Rowlandius ducoudrayi (Armas & Abud, 1990: 3). Armas, 2004: 16 *DR > TL: Rincón Naranjo (Los Haitises), Bayaguana,
Prov. Monte Plata
Rowlandius engombe Armas & Abud, 2002: 15. Armas, 2004: 17 *DR > TL: Engombe, Santo Domingo
Rowlandius isabel Armas & Abud, 2002: 13. Armas, 2004: 17 *DR > TL: Ladera S Loma Isabel de Torres, Prov. Puerto Plata
Rowlandius jarmilae Armas & Cokendolpher, 2001: 4. Armas, 2004: 17 *DR > TL: Cueva El Pomier No. 2, cerca de Borbón,
Prov. San Cristóbal. Original spelling jarmillae corrected to jarmilae by Fet 2002: 90
Rowlandius lantiguai (Armas & Abud, 1990: 7). Armas, 2004: 17 *DR > TL: Los Conucos, Guayacanes, Prov. San Pedro de
Macorís
Rowlandius longipalpus (Rowland & Reddell, 1979: 161). Armas, 2004: 18 *H > TL: La Visite
Rowlandius naranjo (Armas & Abud, 1990: 9). Armas, 2004: 18 *DR > TL: El Naranjo (Los Haitises), Sánchez, Prov. Samaná
Rowlandius velteni† (Krüger & Dunlop, 2010: 49). Armas & Teruel, 2011: 335 [Fossil in Dominican amber]
Rowlandius virginiae Armas & Abud, 2002: 12. Armas, 2004: 18 *DR > TL: 3.5 km N Guayabal, Postrer Rio, Prov.
Independencia
Stenochrus Chamberlin, 1922
Stenochrus portoricensis (Chamberlin, 1922). Armas, 2004: 20; Monjaraz-Ruedas et al., 2019: 69

Order SCORPIONES

Taxonomic summary:
Superfamily Buthoidea
Family Buthidae
Superfamily Scorpionoidea
Family Scorpionidae
Family Hemiscorpiidae

Superfamily BUTHOIDEA
Family BUTHIDAE
Centruroides Marx, 1890
Centruroides alayoni Armas, 1999c: 122 *DR > TL: Playa del Coco, NW Isla Beata, Prov. Pedernales
=Centruroides haitiensis Lourenço, 2016: 18
Centruroides altagraciae Teruel, Armas & Kovarik, 2015a: 14. Armas & Teruel, 2016: 153 *DR > TL: ~4 km N de Bayahibe,
48 m
Centruroides bani Armas & Marcano, 1987: 5
Centruroides jaragua Armas, 1999c: 124 *DR > TL: 400 m S Destacamento de la Marina (playa de Punta Beata), N Isla Beata,
Prov. Pedernales
Centruroides lucidus Teruel, Armas & Kovarik, 2015: 2 *DR > TL: Los Tres Charcos, Oviedo, 71 m
Centruroides marcanoi Armas, 1981b: 10 *DR > TL: Río Mulito, Banano, Prov. Pedernales
Centruroides margaritatus (Gervais, 1841). Armas, 1982b: 6
Centruroides tenuis (Thorell, 1876: 153). Santiago-Blay, 1993: 6
=Centruroides nitidus (Thorell, 1876: 152). Armas & Marcano, 1987: 12
=Centruroides nitidus taino Armas & Marcano, 1987: 14
=Centruroides beynai† Schawaller, 1979: 6. Armas, 1982a: 25 [Fossil in Dominican amber]
Centruroides platnicki Armas, 1981. Teruel & Seiter, 2016: 76
Isometrus Ehrenberg, 1831

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Isometrus maculatus (De Geer, 1775). Santiago-Blay, 1993: 7
Microtityus Kjellesvig-Waering, 1966
Microtityus (Parvabsonus) ambarensis† (Schawaller, 1982: 3). Armas, 1988: 1; Santiago-Blay et al., 1990: 115 [Fossil in
Dominican amber]
Microtityus (Parvabsonus) barahona Armas & Teruel, 2012: 73 *DR > TL: Playa Azul, sección Juan Esteban, Barahona
Microtityus (Parvabsonus) consuelo Armas & Marcano, 1987: 16; Armas & Teruel, 2012: 78 *DR > TL: Cumayasa, Prov. San
Pedro de Macorís
Microtityus (Parvabsonus) dominicanensis Santiago-Blay, 1985: 1. Armas, 2002: 99; Armas & Teruel, 2012: 80 *DR > TL:
Cajuelito
Microtityus (Parvabsonus) ivei Armas, 1999c: 104. Armas & Teruel, 2012: 81 *DR > TL: 24 km N Cabo Rojo, Prov.
Pedernales, 365 m
Microtityus (Parvabsonus) lantiguai Armas & Marcano, 1987: 7 *DR > TL: 8 km N de Pedernales, Prov. Pedernales
Microtityus (Parvabsonus) lourencoi Armas & Teruel, 2012: 77 *DR > TL: Carretera a Guaraguao, 1 km después de la entrada
a Bayahibe, San Rafael del Yuma, Prov. La Altagracia
Microtityus (Parvabsonus) minimus Kovarik & Teruel, 2014: 1 *DR > TL: El Número, Prov. Azua, 167 m
Microtityus (Parvabsonus) paucidentatus Armas & Marcano, 1992: 11. Armas & Teruel, 2012: 82 *DR > TL: Segundo Paso,
sección Apolinar Perdomo, Neiba, Provincia Baoruco
Microtityus (Parvabsonus) prendinii Armas & Teruel, 2012: 83 *DR > TL: Palmarito, sección Las Galeras, Samaná
Microtityus (Parvabsonus) reini Armas & Teruel, 2012: 74 *DR > TL: La Laguna, carretera a Manaclar Km 6, Baní, Prov.
Peravia
Microtityus (Parvabsonus) solegladi Armas & Teruel, 2012: 71 *DR > TL: 3.5 km al N de Guayabal, Postrer Río, Prov.
Independencia
Microtityus (Parvabsonus) virginiae Armas, 1999c: 102. Armas & Teruel, 2012: 86 *DR > TL: Los Bolos, Sierra de Neiba,
Postrer Río, prov. Independencia, 1100 m
Heteroctenus Pocock, 1893
Heteroctenus abudi (Armas & Marcano, 1988: 19). Teruel, 2006: 50; Prendini et al., 2009: 207; Esposito et al., 2017: 25
Heteroctenus bonettii (Armas, 1999c: 126) *DR > TL: Playa del Coco, NW Isla Beata
Heteroctenus princeps (Karsch, 1879). Armas & Marcano, 1987: 20; Esposito et al., 2017: 51 *H, *DR > TL: Port-au-Prince
Heteroctenus renelauerae† (Lourenço in Lourenço & Velten, 2016: 61) [Fossil in Dominican amber]
Tityus C. L. Koch, 1836
Tityus abudi Armas, 1999c: 105 *DR > TL: Loma Alto de la Bandera, 1690 m, NE Parque Nacional Armando Bermúdez,
sección Juncalito, Jánico, Prov. Santiago
Tityus altithronus Armas, 1999c: 115 *DR > TL: Pico Duarte (sobre 3,000 m), Prov. San Juan
Tityus bellulus Armas, 1999c: 112 *DR > TL: Parque Nacional “Armando Bermúdez”, Sierra Atravesada (1,964 m), sección
Rincón de Piedra, San José de las Matas, Prov. Santiago
Tityus crassimanus (Thorell, 1876). Armas & Abud, 2004: 59 *H, *DR > TL: Erroneously “Mexico”, endemic to Haiti and
Dominican Republic
=Tityus bahoruco Teruel & Armas, 2006: 140; Teruel, 2017a: 2
Tityus elii Armas & Marcano, 1992: 22 *DR > TL: Loma La Sal, Jarabacoa, Prov. La Vega
=Tityus ebanoverde Armas, 1999c: 118; Teruel, 2017a: 3
Tityus geratus† Santiago-Blay & Poinar, 1998: 346 [Fossil in Dominican amber]
=Tityus azari† Lourenco, 2013: 1; Teruel, 2017a: 5 [Fossil in Dominican amber]
=Tityus hartkorni† Lourenco, 2009: 4; Teruel, 2017a: 5 [Fossil in Dominican amber]
Tityus haetianus Teruel & Santos, 2018: 1 *DR > TL: Département du Sud, Tiburon Peninsula, Massif de la Hotte, Pic Macaya
National Park, Formond, 2,300 m a.s.l.
Tityus kindli Kovarik & Teruel, 2014: 10 *DR > TL: 2 km East of La Nevera, Valle Nuevo, 2239 m
Tityus neibae Armas, 1999c: 110 *DR > Los Bolos (1,100 m), Sierra de Neiba, Postrer Río, Prov. Independencia
Tityus ottenwalderi Armas, 1999c: 108 *DR > TL: Loma Alto de la Bandera (1,597 m), Parque Nacional “Armando
Bermúdez”, Jánico, Prov. Santiago
=Tityus anasilviae Armas & Abud, 2004: 54. Armas & Abud, 2015: 136; Teruel, 2017a: 1
Tityus portoplatensis Armas & Marcano, 1992: 30. Armas & Abud, 2004: 60 *DR > TL: Pico Isabel de Torres, Prov. Puerto
Plata
=Tityus septentrionalis Armas & Abud, 2004: 56; Teruel, 2017a: 4
Tityus quisqueyanus Armas, 1982a: 13 *DR > TL: Valle Nuevo, Constanza

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Tityus schrammi Teruel & Santos, 2018: 8 *DR > TL: Elias Piña Prov., Cordillera Central, Nalga de Maco National Park, Rio
Limpio, 1,643 m a.s.l.

Superfamily SCORPIONOIDEA
Family SCORPIONIDAE
Cazierius Francke, 1978
Cazierius cicero (Armas & Marcano, 1987: 3) *DR > TL: Conuco, Guayacanes, Prov. San Pedro de Macorís
Cazierius neibae Kovarik & Teruel, 2014: 16 *DR > TL: Segundo Paso, Apolinar Perdomo, Sierra de Neiba, 150 m
Cazierius politus (Pocock, 1898). Armas, 1981a: 10; Armas & Marcano, 1987: 2 *H, *DR > TL: Erroneously given as
“Brazil”.
Heteronebo Pocock, 1899
Heteronebo barahonae Teruel, Armas & Kovarik, 2015b: 24 *DR > TL: Barahona Prov., Cabral Municipality: Sierra de
Bahoruco: km 7–9 of road from Cabral to Polo, 221 m asl
Heteronebo dominicus Armas, 1981a: 5. Teruel, 2005: 166 *DR > TL: Nueve km al N de Pedernales, Prov. Pedernales
Heteronebo monticola (Armas, 1999c: 131). Teruel, 2005: 167 *DR > TL: Casi 35 km N Cabo Rojo, El Aceitillar (4,500 pies),
Prov. Pedernales
Heteronebo oviedo (Armas, 1999c: 133). Teruel, 2005: 167 *DR > TL: Sección Los Tres Charcos, (carretera Oviedo-
Pedernales), Oviedo, Prov. Pedernales
Heteronebo pumilus Armas, 1981a: 2 *H > TL: Pedregal frente a Cayo La Vache, Les Cayes
Heteronebo sp.†—Santiago-Blay & Craig (1998: 89) [Fossil in Dominican amber]

Family HEMISCORPIIDAE
Opistacanthus Peters, 1861
Opistacanthus lepturus (Palisot de Beauvois, 1805). Armas, 1982b: 5; Armas & Marcano, 1987: 2; Lourenço, 1987: 924;
Armas & Marcano, 1992: 5 *H, *DR > TL: Erroneously South America.
=Opistacanthus laevicauda Thorell, 1876. Armas & Marcano, 1992: 2

Order SOLIFUGAE
Family AMMOTRECHIDAE
Ammotrechella Roewer, 1934
Ammotrechella hispaniolana Armas & Alegre, 2001: 5. Armas, 2004: 32 *DR > TL: Cabo Rojo, Prov. Pedernales
Ammotrechinus Roewer, 1934
Ammotrechinus gryllipes (Gervais, 1842). Muma, 1970: 50
Antillotrecha Armas, 1994
Antillotrecha fraterna Armas, 1994: 3. Armas, 2004: 34 *DR > TL: Isla Cabrito, Prov. Independencia
Happlodontus† Poinar & Santiago-Blay, 1989
Happlodontus proterus† Poinar & Santiago-Blay, 1989: 126 [Fossil in Dominican amber]

Order UROPYGI
Key reference: Armas (2004).

Family TELYPHONIDAE
Mastigoproctus Pocock, 1894
Mastigoproctus proscorpio (Latreille, 1806). Armas, 2004: 24
Ravilops Viquez & Armas, 2005
Ravilops kovariki Teruel, 2017b: 16 *DR > TL: La Ceiba, Apolinar Perdomo, Sierra de Neiba, 808 m
Ravilops wetherbeei (Armas, 2002: 39). Armas, 2004: 24 *DR > TL: Límite N del Parque Nacional Armando Bermúdez (1381
m), sección Juncalito, Jánico, Prov. Santiago

Subphylum MYRIAPODA
Class CHILOPODA
Key reference: Martínez-Muñoz & Perez-Gelabert (2018).

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 83

Taxonomic summary:
Order Scutigeromorpha
Family Pselliodidae
Family Scutigeridae
Order Scolopendromorpha
Family Cryptopidae
Family Scolopendridae
Family Scolopocryptidae
Order Geophilomorpha
Family Geophilidae
Family Mecistocephalidae
Family Oryidae
Family Schendylidae

Order SCUTIGEROMORPHA
Family PSELLIODIDAE
Sphendononema Verhoeff, 1904
Sphendononema guildingii (Newport, 1844). Perez-Gelabert & Edgecombe, 2013: 39; Martínez-Muñoz & Perez-Gelabert,
2018: 80
=Pselliodes guildinguii (Newport, 1845). Wurmli, 1978: 137; Perez-Gelabert, 2008: 71
=Pselliophora haitiensis Chamberlin, 1918: 170
=Pselliophora pulchritarsis Verhoeff, 1904. Chamberlin, 1918: 167

Family SCUTIGERIDAE
Dendrothereua Verhoeff, 1944
Dendrothereua linceci (Wood, 1867). Perez-Gelabert & Edgecombe, 2013: 39; Martínez-Muñoz & Perez-Gelabert, 2018: 78
Note: The invalid synonym Scutigera forceps Rafinesque, 1820 is cited by Wolcott, 1927: 12 for Haiti. Also the name
Scutigera coleoptrata Linnaeus, 1758 was used for some fossilized specimens in Dominican amber by Wu, 1997: 129.
However, living S. coleoptrata are thought to originate in the Mediterranean area and have never been found in Hispaniola.
Fossil Miocene scutigeromorphs in Dominican amber are not likely to represent this species (Perez-Gelabert & Edgecombe,
2013).

Order SCOLOPENDROMORPHA
Family CRYPTOPIDAE
Cryptops Leach, 1814
Cryptops manni Chamberlin, 1915: 501. Martínez-Muñoz & Perez-Gelabert, 2018: 85 *H > TL: Milot
Cryptops sp.†—Poinar (1992: 92); Martínez-Muñoz & Perez-Gelabert, 2018: 85 [Fossil in Dominican amber]

Family SCOLOPENDRIDAE
Subfamily OTOSTIGMINAE
Otostigmus Porat, 1878
Otostigmus (Parotostigmus) occidentalis Meinert, 1886: 185. Chamberlin, 1918: 155; Perez-Gelabert, 2008: 70; Martínez-
Muñoz & Perez-Gelabert, 2018: 83 *H > TL: Grande Anse
Rhysida Wood, 1878
Rhysida celeris (Humbert & Saussure, 1870). Schileyko, 2014: 183; Martínez-Muñoz & Perez-Gelabert, 2018: 84
Rhysida longipes longipes (Newport, 1845). Chao, 2002: 28; Martínez-Muñoz & Perez-Gelabert, 2018: 84 [Introduced]
Rhysida nuda (Newport, 1845). Chamberlin, 1918: 156; Martínez-Muñoz & Perez-Gelabert, 2018: 84 [Introduced]
Subfamily SCOLOPENDRINAE
Cormocephalus Newport, 1844
Cormocephalus guildingi (Newport, 1845). Chamberlin, 1918: 156; Chamberlin, 1944: 182; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 80
=Cormocephalus impressus Porat, 1876: 15
Scolopendra Linnaeus, 1758
Scolopendra alternans Leach, 1816. Arthaud, 1787: 427; Shelley, 2006: 6; Mercurio, 2016: 12; Perez-Gelabert, 2008: 70;

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Martínez-Muñoz & Perez-Gelabert, 2018: 80
=Scolopendra sagrae Gervais, 1837. Brandt, 1840: 157
Scolopendra morsitans Linnaeus, 1758. Chamberlin, 1918: 158; Shelley, 2006: 5; Perez-Gelabert, 2008: 70; Martínez-Muñoz
& Perez-Gelabert, 2018: 80 [Introduced]
=Scolopendra platypus Brandt, 1840. Shelley, 2006: 8; Perez-Gelabert, 2008: 70
Scolopendra subspinipes Leach, 1816. Kraepelin, 1904: 322; Martínez-Muñoz & Perez-Gelabert, 2018: 82; Chamberlin, 1918:
158 [Introduced]
Note: Scolopendra gigantea Linnaeus, 1758 was cited for Hispaniola based on one specimen from Haiti (Shelley & Kiser,
2000: 161; Shelley, 2006: 5). This appears to be a labeling error.

Family SCOLOPOCRYPTOPIDAE
Subfamily NEWPORTIINAE
Newportia Gervais, 1847
Newportia cubana Chamberlin, 1915. Martínez-Muñoz & Perez-Gelabert, 2018: 88
Newportia ernsti ernsti Pocock, 1891. Chamberlin, 1918: 153; Schileyko & Minelli, 1999: 274; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 89
Newportia longitarsis (Newport, 1844). Chamberlin, 1918: 153; Martínez-Muñoz & Perez-Gelabert, 2018: 88
Newportia longitarsis stechowi Verhoeff, 1938. Schileyko, 2014: 162; Martínez-Muñoz & Perez-Gelabert, 2018: 88
Newportia sp.†—Wu (1996: 126); Martínez-Muñoz & Perez-Gelabert (2018: 88) [Fossil in Dominican amber]
Subfamily SCOLOPOCRYPTOPINAE
Scolopocryptops Newport, 1844
Scolopocryptops ferrugineus ferrugineus (Linnaeus, 1767). Pocock, 1893: 463; Chamberlin, 1918: 152; Perez-Gelabert, 2008:
70; Martínez-Muñoz & Perez-Gelabert, 2018: 86
Scolopocryptops melanostoma Newport, 1845. Kraepelin, 1904: 245; Chamberlin, 1918: 152; Chamberlin, 1944: 177;
Schileyko, 2014: 154; Martínez-Muñoz & Perez-Gelabert, 2018: 86
Scolopocryptops sp.†—Poinar & Poinar (1999: 82); Martínez-Muñoz & Perez-Gelabert (2018: 85) [Fossil in Dominican
amber]

Order GEOPHILOMORPHA
Family GEOPHILIDAE
Nesidiphilus Chamberlin, 1915
Nesidiphilus juvenis Chamberlin, 1918: 161. Foddai et al., 2000: 79; Perez-Gelabert, 2008: 70; Martínez-Muñoz & Perez-
Gelabert, 2018: 90
Polycricus Saussure & Humbert, 1872
Polycricus haitiensis (Chamberlin, 1915: 526). Chamberlin, 1918: 163; Foddai et al., 2000: 88; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 90 *H > TL: Furcy
Polycricus nesiotes (Chamberlin, 1915: 527). Chamberlin, 1918: 164; Foddai et al., 2000: 89; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 90 *H > TL: Pétionville
Telocricus Chamberlin, 1915
Telocricus multipes Chamberlin, 1915: 521. Chamberlin, 1918: 162; Foddai et al., 2000: 102; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 90 *H > TL: Manneville

Family MECISTOCEPHALIDAE
Mecistocephalus Newport, 1843
Mecistocephalus guildingii Newport, 1843. Chamberlin, 1918: 164; Foddai et al., 2000: 63; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 90

Family ORYIDAE
Titanophilus Chamberlin, 1915
Titanophilus fratrellus Chamberlin, 1915: 505. Chamberlin, 1918: 161; Foddai et al., 2000: 118; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 91 *H > TL: Pétionville
Titanophilus maximus Chamberlin, 1915: 503. Chamberlin, 1918: 160; Foddai et al., 2000: 118; Perez-Gelabert, 2008: 70;
Martínez-Muñoz & Perez-Gelabert, 2018: 91 *H > TL: Grand Rivière

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 85

Family SCHENDYLIDAE
Ctenophilus Cook, 1896
Ctenophilus nesiotes (Chamberlin, 1918: 159). Attems, 1929: 87. Pereira, 1981: 180. Pereira & Demange, 1991: 93. Pereira &
Demange, 1997: 310. Foddai et al., 2000: 121. Perez-Gelabert, 2008: 70; Martínez-Muñoz & Perez-Gelabert, 2018: 91 *H >
TL: Diquini

Class DIPLOPODA
Key reference: Pérez-Asso and Perez-Gelabert (2001).

Taxonomic summary:
Subclass Penicillata
Order Polyxenida
Family Lophoproctidae
Order Glomeridesmida
Family Glomeridesmidae
Order Polyzoniida
Family Siphonotidae
Order Polydesmida
Family Chelodesmidae
Family Furhmannodesmidae
Family Haplodesmidae
Family Platyrrhacidae
Family Paradoxosomatidae
Family Pyrgodesmidae
Family Sphaerodesmidae
Order Siphonophorida
Family Siphonophoridae
Order Spirobolida
Family Spirobolellidae
Family Rhinocricidae
Family Pseudospirobolellidae
Family Trigoniulidae
Order Spirostreptida
Family Pseudonannolenidae
Family Spirostreptidae
Order Stemmiulida
Family Stemmiulidae

Subclass PENICILLATA
Order POLYXENIDA
Family LOPHOPROCTIDAE
Lophoturus Broleman, 1931
Lophoturus aequatus (Loomis, 1936: 7). Pérez-Asso & Perez-Gelabert, 2001: 68 *H > TL: Petite Rivière de Artibonite
Lophoturus niveus (Loomis, 1934: 5). Pérez-Asso & Perez-Gelabert, 2001: 68 *DR > TL: Beata Island
Lophoturus? sp.†—Santiago-Blay & Poinar (1992: 363); Pérez-Asso & Perez-Gelabert (2001: 68) [Fossil in Dominican amber]

Order GLOMERIDESMIDA
Family GLOMERIDESMIDAE
Glomeridesmus Gervais, 1844
Glomeridesmus concolor Chamberlin, 1918. Pérez-Asso & Perez-Gelabert, 2001: 68 *H > TL: Fond des Negres
Glomeridesmus jenkinsi Loomis, 1936. Pérez-Asso & Perez-Gelabert, 2001: 68 *H > TL: Le Borgne
Glomeridesmus sp.†—Santiago-Blay & Poinar (1992: 366); Pérez-Asso & Perez-Gelabert (2001: 68) [Fossil in Dominican
amber]

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Order POLYZONIIDA
Family SIPHONOTIDAE
Rhinotus Cook, 1896
Rhinotus purpureus (Pocock, 1894). Loomis, 1936: 14; Pérez-Asso & Perez-Gelabert, 2001: 68
Rhinotus sp.†—Santiago-Blay & Poinar (1992: 366); Pérez-Asso & Perez-Gelabert (2001: 68) [Fossil in Dominican amber]

Order POLYDESMIDA
Family CHELODESMIDAE
Achromoporus Loomis, 1936
=Biaporus Loomis, 1941
=Craterodesmus Loomis, 1941
=Lasiomazus Loomis, 1941
=Synecheporus Loomis, 1941
Achromoporus andujari Pérez-Asso, 2009: 50 *DR > TL: Prov. La Vega, alrededores de La Pirámide (2,380 msnm), Parque
Nacional Valle Nuevo
Achromoporus atrechoensis Suriel, 2010: 14 *DR > TL: Mancha de bosque latifoliado entre pinares a pocos metros de la loma
El Atrecho, camino de Alto de la Rosa a Sabana Vieja, prov. San Juan, 1722 m
Achromoporus bastardoae Suriel, 2011: 22 *DR > TL: Las Yayitas, al final del caserío y antes de cruzar el río, El Recodo, prov.
Peravia
Achromoporus bayaguana Pérez-Asso, 2005a: 55 *DR > TL: Bayaguana, Prov. Monte Plata
Achromoporus cachote Pérez-Asso, 2009: 74 *DR > TL: Prov. Barahona, Cachote (1,200 msnm), camino de Ciénaga a Polo, a
15 km NW de Ciénaga
Achromoporus coloratus Loomis, 1936: 113. Pérez-Asso & Perez-Gelabert, 2001: 73; Pérez-Asso, 2005a: 57; Pérez-Asso,
2009: 36 *H > TL: Thor, near Port-au-Prince
=Achromoporus coloratus neiba Pérez-Asso, 2005a: 57
Achromoporus concolor (Loomis, 1941a: 52). Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2009: 39 *DR > TL: Loma
Vieja, cerca de Constanza
Achromoporus chocoensis Pérez-Asso, 2009: 63 *DR > TL: Provincia Puerto Plata, área protegida El Choco, cerca del poblado
de Cabarete
Achromoporus ebanoverde Pérez-Asso, 2005a: 53; Pérez-Asso, 2009: 48 *DR > TL: Prov. La Vega, Loma de La Sal (1,000
msnm), La Guira, Reserva Científica Ebano Verde
Achromoporus elegans Suriel, 2010: 18 *DR > TL: Mancha del bosque latifoliado entre pinares a pocos metros de la loma El
Atrecho, camino de Alto de la Rosa a Sabana Vieja, 1722 m
Achromoporus enneryensis Loomis, 1936: 115. Pérez-Asso & Perez-Gelabert, 2001: 73 *H > TL: Ennery
Achromoporus fractus Suriel, 2011: 26 *DR > TL: Loma Los Guayuyos, Manaclar, Prov. Peravia, 1,359 m
Achromoporus furcipes Loomis, 1936: 117. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Near Plaisance
Achromoporus heteromus Loomis, 1941a: 49. Pérez-Asso & Perez-Gelabert, 2001: 74 *DR > TL: Sánchez and vicinity
Achromoporus isabelensis Pérez-Asso, 2009: 68 *DR > TL: Prov. Puerto Plata, Los Charcos del Damajagua
Achromoporus jaragua Pérez-Asso, 2009: 65 *DR > TL: Prov. Pedernales, La Curva del Guano, carretera de Oviedo a
Pedernales, km 22–23, Parque Nacional Jaragua
Achromoporus magnus Pérez-Asso, 2009: 70 *DR > TL: Prov. Ocoa, alrededores del poblado de La Nuez (1,700 msnm),
carretera de San José de Ocoa a Valle Nuevo
Achromoporus martingarcia Pérez-Asso, 2009: 61 *DR > TL: Prov. Barahona, cerca de Puerto Alejandro, Sierra Martín García
Achromoporus montanus (Loomis, 1941a: 62). Pérez-Asso & Perez-Gelabert, 2001: 74; Pérez-Asso, 2009: 47 *DR > TL: Pico
del Yaque y Loma Rucilla
Achromoporus occultus Pérez-Asso, 2009: 56 *DR > TL: Prov. La Vega, La Pirámide (2,380 msnm), Parque Nacional Valle
Nuevo
Achromoporus ovatus (Loomis, 1941a: 60). Pérez-Asso & Perez-Gelabert, 2001: 74; Pérez-Asso, 2009: 41 *DR > TL: Between
3,000 and 4,000 ft. elev., Mt. Diego de Ocampo, Northern Range
Achromoporus pallidus Pérez-Asso, 2009: 54 *DR > TL: Prov. La Vega, alrededores de Las Pirámides, carretera entre La Nuez
y la caseta del Parque Nacional Valle Nuevo
Achromoporus platyurus (Loomis, 1941a: 64). Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2009: 43 *DR > TL: Loma
Rucilla (2,400—3,000 msnm)
Achromoporus robustus Loomis, 1936: 118. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Near Cancoque

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Achromoporus surieli Pérez-Asso, 2009: 58 *DR > TL: Prov. La Vega, La Ciénaga, sendero de Los Tablones al Alto de La
Cotorra (1,400—1,500 msnm), Parque Nacional Armando Bermúdez
Achromoporus vallenuevo Pérez-Asso, 2009: 52 *DR > TL: Entre el poblado de La Nuez y la caseta del Parque Nacional Valle
Nuevo
Beatadesmus Loomis, 1934
=Dendrodesmus Pérez-Asso, 2003
=Pogonodesmus Loomis, 1936
=Quisquicia Loomis, 1936
=Aplopododesmus Loomis, 1936
Beatadesmus bahorucoensis Pérez-Asso, 2010: 8 *DR > TL: Las Abejas, Sierra de Bahoruco, 1200 m
Beatadesmus engonatus (Loomis, 1936: 132). Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2010: 11 *H > TL: Roche
Croix, Morne La Hotte, 5000 ft.
Beatadesmus insignis (Chamberlin, 1918: 234). Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2010: 11 *H > TL: In
rotting Yucca stumps above Kenscoff
Beatadesmus ivonneae Pérez-Asso, 2008: 16 *DR > TL: Curva del Guano, carretera Oviedo a Pedernales, km 22–23, Parque
Nacional Jaragua
Beatadesmus longipes (Loomis, 1936: 118). Pérez-Asso & Perez-Gelabert, 2001: 74; Pérez-Asso, 2010: 11 *H > TL: Morne La
Selle, 7,500 ft.
Beatadesmus pedernales Pérez-Asso, 2008: 19 *DR > TL: Carretera de Pedernales a Las Mercedes a 3 km del entroque a
Pedernales
Beatadesmus pullus (Loomis, 1936: 138). Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2010: 11 *H > TL: Kenscoff,
between 3000 and 6000 ft.
Beatadesmus rubens (Loomis, 1936: 135). Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2010: 11 *H > TL: Morne La
Selle, 7,500 ft.
Beatadesmus sallei (Saussure, 1860). Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2010: 11
Beatadesmus scitulus Loomis, 1936: 130. Pérez-Asso & Perez-Gelabert, 2001: 75; Pérez-Asso, 2010: 11 *H > TL: Near Trouin
Beatadesmus utowani Loomis, 1934: 31. Pérez-Asso & Perez-Gelabert, 2001: 74 *DR > TL: Beata Island
Beatadesmus yuma (Pérez-Asso, 2003: 58). Pérez-Asso, 2010: 11 *DR > TL: Sendero a la Cueva de Bienvo, Boca de Yuma,
Parque Nacional del Este, Prov. La Altagracia
Chondrotropis Loomis, 1936
Chondrotropis niger Loomis, 1936: 121. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Base of a cliff near the road below
Ennery
Chondrotropis pictus Loomis, 1936: 123. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Morne Pilboreau
Chondrotropis venustus Loomis, 1936: 126. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Morne Brigand, near Bayeaux
Cyrtaphe Loomis, 1936
Cyrtaphe alternata Loomis, 1936: 143. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Morne Pilboreau, above Ennery
Cyrtaphe continuata Loomis, 1936: 147. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Near Trouin
Cyrtaphe domingensis Loomis, 1941a: 55. Pérez-Asso & Perez-Gelabert, 2001: 74 *DR > TL: Between 3,000 and 4,000 ft.
elev., Mt. Diego de Ocampo, Northern Range
Cyrtaphe lobipes Loomis, 1936: 145. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Ennery
Cyrtaphe neiba Pérez-Asso, 2010: 1 *DR > TL: Entre Sabana Real y Pirámide 204, Sierra de Neiba
Ebanodesmus Pérez-Asso, 2005
Ebanodesmus victori Pérez-Asso, 2005b: 62 *DR > TL: El Arroyazo (900 msnm), camino de El Arroyazo a Casabito, Reserva
Científica Ébano Verde, Prov. La Vega
Ellipodesmus Chamberlin, 1918
Ellipodesmus simplex Chamberlin, 1918: 243. Pérez-Asso & Perez-Gelabert, 2001: 74 *H > TL: Jacmel
Hypselodesmus Loomis, 1941
Hypselodesmus assoi Suriel, 2009: 2 *DR > TL: Sabana Vieja, Prov. San Juan, 1,944 m
Hypselodesmus bicolor Loomis, 1941a: 54. Pérez-Asso & Perez-Gelabert, 2001: 74 *DR > TL: Loma Quita Espuela
Hypselodesmus subtilissimus Suriel, 2012: 50 *DR > TL: Manaclar de bosque nublado a 1,227 m en la Loma Barbacoa
Podiscodesmus Loomis, 1941
Podiscodesmus carinatus Loomis, 1941a: 58. Pérez-Asso & Perez-Gelabert, 2001: 75 *DR > TL: Sánchez and vicinity
Podiscodesmus neiba Pérez-Asso, 2009: 8 *DR > TL: Entre Sabana Real y Pirámide 204, a 4–5 km de Sabana Real
Pterygiodesmus Loomis, 1936

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Pterygiodesmus strumosus Loomis, 1936: 150. Pérez-Asso & Perez-Gelabert, 2001: 75 *H > TL: La Visite, 6000 to 7000 ft.
Chelodesmidae of uncertain generic position
Amphelictogon Chamberlin, 1918
Amphelictogon? manni Chamberlin, 1918: 231. Pérez-Asso & Perez-Gelabert, 2001: 75 *H > TL: Diquini, near Port-au-Prince

Family FUHRMANNODESMIDAE
Agenodesmus Loomis, 1934
Agenodesmus reticulatus Loomis, 1934: 40. Loomis, 1936: 156; Pérez-Asso & Perez-Gelabert, 2001: 77
Chilaphrodesmus Loomis, 1934
Chilaphrodesmus darlingtoni (Loomis, 1941a: 65). Pérez-Asso & Perez-Gelabert, 2001: 77 *DR > TL: Loma Vieja, south of
Constanza, elev. about 6,000 ft.
Chilaphrodesmus rubellus Loomis, 1934: 43. Pérez-Asso & Perez-Gelabert, 2001: 77 *H > TL: Near the road on the summit of
Morne Pilboreau, above Ennery
Dasyodontus Loomis, 1934
Dasyodontus hispaniolus Loomis, 1936: 158. Pérez-Asso & Perez-Gelabert, 2001: 77 *H > TL: Le Borgne
Dasyodontus sp.†—Santiago-Blay & Poinar (1992: 367); Pérez-Asso & Perez-Gelabert (2001: 77) [Fossil in Dominican
amber]
Hexadesmus Loomis, 1933
Hexadesmus lateridens Loomis, 1933: 363. Pérez-Asso & Perez-Gelabert, 2001: 77
Trematodesmus Loomis, 1936
Trematodesmus setiger Loomis, 1936: 155. Pérez-Asso & Perez-Gelabert, 2001: 77 *H > TL: Morne La Hotte, between 3000
and 7800 ft.

Family HAPLODESMIDAE
Prosopodesmus Silvestri, 1910
Prosopodesmus jacobsoni Silvestri, 1910. Pérez-Asso & Perez-Gelabert, 2001: 77
=Homodesmus parvus Chamberlin, 1918: 223. Loomis, 1936: 180

Family PLATYRHACIDAE
Proaspis Loomis, 1941
Proaspis aitia Loomis, 1941b: 194. Pérez-Asso & Perez-Gelabert, 2001: 76; Shelley & Martínez-Torres, 2013: 481 *H > TL:
Between Camp Perrin and Rivière Glace

Family PARADOXOSOMATIDAE
Asiomorpha Verhoeff, 1936
Asiomorpha coarctata (Saussure, 1860). Pérez-Asso & Perez-Gelabert, 2001: 77; Suriel, 2012: 28 [Introduced]
Chondromorpha Silvestri, 1897
Chondromorpha xantotricha (Attems, 1898). Jeekel, 1999: 23; Shelley & Cupul-Magaña, 2007: 213; Suriel, 2012: 26
[Introduced]
Oxidus Cook, 1911
Oxidus gracilis (C. L. Koch, 1847). Suriel, 2012: 25 [Introduced]

Family PYRGODESMIDAE
Augesmus Chamberlin, 1951
Augesmus varilobatus (Loomis, 1936: 184). Pérez-Asso & Perez-Gelabert, 2001: 78 *H > TL: Hill east of Petit Goave from
Leogane
Coccoelasma Loomis, 1936
Coccoelasma incisura Loomis, 1936: 167. Pérez-Asso & Perez-Gelabert, 2001: 78 *H > TL: Ile de Cabrit, near Bayeaux
Cyphotylus Loomis, 1936
Cyphotylus prolatus Loomis, 1936: 170. Pérez-Asso & Perez-Gelabert, 2001: 78 *H > TL: Morne Brigand, near Bayeaux
Dilophops Loomis, 1934
Dilophops bullatus Loomis, 1934: 59. Pérez-Asso & Perez-Gelabert, 2001: 78
Docodesmus Cook, 1896
Docodesmus alifer Loomis, 1941a: 68. Pérez-Asso & Perez-Gelabert, 2001: 78 *DR > TL: Pico del Yaque, 8,000 to 10,000 ft.

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elev.
Docodesmus angustus Loomis, 1941a: 71. Pérez-Asso & Perez-Gelabert, 2001: 78 *DR > TL: Valle Nuevo, southeast of
Constanza, elev. about 7,000 ft.
Docodesmus brodzinskyi† Shear, 1981: 53. Pérez-Asso & Perez-Gelabert, 2001: 78 [Fossil in Dominican amber]
Docodesmus griseus Loomis, 1941a: 69. Pérez-Asso & Perez-Gelabert, 2001: 78 *DR > TL: Sánchez and vicinity
Docodesmus haitiensis Chamberlin, 1918: 216. Pérez-Asso & Perez-Gelabert, 2001: 78 *H, *DR > TL: Diquini
Docodesmus parvior Chamberlin, 1918: 218. Pérez-Asso & Perez-Gelabert, 2001: 78 *H > TL: Furcy
Docodesmus semiseptus Loomis, 1936: 163. Pérez-Asso & Perez-Gelabert, 2001: 78 *H > TL: Morne Pilboreau
Docodesmus sp.†—Santiago-Blay & Poinar (1992: 367); Pérez-Asso & Perez-Gelabert (2001: 78) [Fossil in Dominican
amber]
Eoromus Loomis, 1936
Eoromus aberrans Loomis, 1936: 176. Pérez-Asso & Perez-Gelabert, 2001: 78 *H > TL: Base of limestone cliff, south of the
divide near Trouin, on the road to Jacmel
Fennellia Loomis, 1941
Fennellia ovipes Loomis, 1941b: 194. Pérez-Asso & Perez-Gelabert, 2001: 78 *H > TL: Jérémie
Henicomus Loomis, 1941
Henicomus septiporus Loomis, 1941a: 79. Pérez-Asso & Perez-Gelabert, 2001: 79 *DR > TL: About 6,000 ft. elevation, Loma
Vieja, south of Constanza
Iomoides Loomis, 1934
Iomoides conjuncta Loomis, 1941a: 76. Pérez-Asso & Perez-Gelabert, 2001: 79 *DR > TL: Villa Altagracia
Iomoides glabra Loomis, 1934: 53. Pérez-Asso & Perez-Gelabert, 2001: 79 *H > TL: Citadel
Iomoides hispidus Loomis, 1934: 51. Pérez-Asso & Perez-Gelabert, 2001: 79 *H > TL: Lower slope of Morne Brigand, facing
Bayeaux
Iomoides parallela Loomis, 1941a: 75. Pérez-Asso & Perez-Gelabert, 2001: 79 *DR > TL: Sánchez and vicinity
Iomus Cook, 1911
Iomus? sp.†—Santiago-Blay & Poinar (1992: 367); Pérez-Asso & Perez-Gelabert (2001: 79) [Fossil in Dominican amber]
Jeekelia Loomis, 1950
Jeekelia granulosa (Loomis, 1941a: 74). Pérez-Asso & Perez-Gelabert, 2001: 79 *DR > TL: Between 5,000 and 8,000 ft. elev.,
Loma Rucilla and mountains north
Lobodesmus Loomis, 1936
Lobodesmus granosus Loomis, 1936: 164. Pérez-Asso & Perez-Gelabert, 2001: 79 *H > TL: Morne La Hotte, between 3000
and 7800 ft.
Lophodesmus Pocock, 1894
Lophodesmus? sp.†—Santiago-Blay & Poinar (1992: 367); Pérez-Asso & Perez-Gelabert (2001: 79) [Fossil in Dominican
amber]
Penteporus Loomis, 1936
Penteporus crenellatus Loomis, 1936: 180. Pérez-Asso & Perez-Gelabert, 2001: 79 *H > TL: Fond des Negres
Poratia Cook, 1894
Poratia granulofrons (Chamberlin, 1918: 221). Pérez-Asso & Perez-Gelabert, 2001: 79
=Dominicodesmus geophilus Chamberlin, 1923: 189
Psochodesmus Cook, 1896
Psochodesmus sp.†—Santiago-Blay & Poinar (1992: 367); Pérez-Asso & Perez-Gelabert (2001: 79) [Fossil in Dominican
amber]
Styraxodesmus Chamberlin, 1918
Styraxodesmus ater Loomis, 1936: 188. Pérez-Asso & Perez-Gelabert, 2001: 79 *H > TL: Ile de Cabrit, near Bayeaux
Styraxodesmus furcatus Chamberlin, 1918: 222. Pérez-Asso & Perez-Gelabert, 2001: 79 *H > TL: Jacmel
Pyrgodesmidae of uncertain generic position
Treseolobus Attems, 1907
Treseolobus? caraibianus Chamberlin, 1918: 220. Pérez-Asso & Perez-Gelabert, 2001: 80
Polydesmidea of uncertain taxonomic status
Inodesmus Cook, 1896
Inodesmus? sp.†—Santiago-Blay & Poinar (1992: 367); Pérez-Asso & Perez-Gelabert (2001: 80) [Fossil in Dominican amber]
Mesethodesmus Chamberlin, 1918
Mesethodesmus haitianus Chamberlin, 1918: 248. Pérez-Asso & Perez-Gelabert, 2001: 80 *H > TL: Furcy

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Family SPHAERIODESMIDAE
Haplocyclodesmus Attems, 1940
Haplocyclodesmus angustipes (Loomis, 1936: 86). Pérez-Asso & Perez-Gelabert, 2001: 75 *H > TL: Near Trouin
Haplocyclodesmus crassartus (Loomis, 1936: 81). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Fond des Negres
Haplocyclodesmus enneryensis (Loomis, 1936: 92). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Ennery
Haplocyclodesmus falcarius (Loomis, 1936: 98). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Morne Pilboreau, above
Ennery
Haplocyclodesmus globulus (Loomis, 1936: 90). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Between Leogane and Petit
Goave
Haplocyclodesmus haitianus (Chamberlin, 1918: 215). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Diquini
Haplocyclodesmus incisus (Loomis, 1936: 88). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Port-au-Prince
Haplocyclodesmus insulanus (Loomis, 1936: 93). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Ile de Cabrit, near
Bayeaux
Haplocyclodesmus montanus (Loomis, 1936: 95). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Morne Pilboreau, above
Ennery
Haplocyclodesmus nudatus (Loomis, 1936: 85). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: L’Archaie
Haplocyclodesmus obesus (Loomis, 1936: 83). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Fond des Negres
Haplocyclodesmus rubellus (Loomis, 1936: 97). Pérez-Asso & Perez-Gelabert, 2001: 76 (H > TL: Le Borgne
Haplocyclodesmus setosus (Loomis, 1936: 101). Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: La Visite, between 6000
and 7000 ft.
Lophocyclus Loomis, 1936
Lophocyclus laxatus Loomis, 1936: 106. Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: North of Trouin
Lophocyclus munitus Loomis, 1936: 105. Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Roche Croix, 5000 ft.
Lophocyclus passus Loomis, 1936: 103. Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Roche Croix, 5000 ft.
Lophocyclus pumilus Loomis, 1936: 108. Pérez-Asso & Perez-Gelabert, 2001: 76 *H > TL: Morne La Hotte, between 3000 and
7800 ft.

Order SIPHONOPHORIDA
Family SIPHONOPHORIDAE
Siphonocybe Pocock, 1903
Siphonocybe alba Loomis, 1936: 13. Pérez-Asso & Perez-Gelabert, 2001: 73 *H > TL: Near Trouin
Siphonocybe sp.†—Shear (1981: 54); Pérez-Asso & Perez-Gelabert (2001: 73) [Fossil in Dominican amber]
Siphonophora Brandt, 1837
Siphonophora gracilior Chamberlin, 1918: 173. Pérez-Asso & Perez-Gelabert, 2001: 73 *H > TL: Pétionville
Siphonophora hoffmani† Santiago-Blay & Poinar, 1992: 367. Pérez-Asso & Perez-Gelabert, 2001: 73 [Fossil in Dominican
amber]
Siphonophora manni Chamberlin, 1918: 173. Pérez-Asso & Perez-Gelabert, 2001: 73 *H > TL: Grande Rivière
Siphonophora platops Loomis, 1941a: 19. Pérez-Asso & Perez-Gelabert, 2001: 73 *DR > TL: Rain forest near Valle Nuevo,
6,000 ft. elevation
Siphonophora proxima Chamberlin, 1918: 173. Pérez-Asso & Perez-Gelabert, 2001: 73 *H > TL: Grand Rivière
Siphonophora sp.†—Santiago-Blay & Poinar (1992: 366); Pérez-Asso & Perez-Gelabert (2001: 73) [Fossil in Dominican
amber]
Siphonophora velezi† Santiago-Blay & Poinar, 1992: 366. Pérez-Asso & Perez-Gelabert, 2001: 73 [Fossil in Dominican
amber]

Order SPIROBOLIDA
Family SPIROBOLELLIDAE
Spirobolellus Pocock, 1894
Spirobolellus concinnus (Loomis, 1936: 45). Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Morne Brigand, near Bayeaux
Spirobolellus erythrotermus (Loomis, 1936: 43). Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Le Brande
Spirobolellus esulcatus (Loomis, 1936: 41). Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Morne Brigand, near Bayeaux
Spirobolellus furcianus (Loomis, 1936: 48). Pérez-Asso & Perez-Gelabert, 2001: 70 *H > TL: Near Furcy on Mt. Noir, 6,100
ft.
Spirobolellus instratus (Loomis, 1941a: 47). Pérez-Asso & Perez-Gelabert, 2001: 70 *DR > TL: Between 1,000 and 3,000 ft.

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elevation, Mt. Quita Espuela
Spirobolellus lineatus (Chamberlin, 1918: 209). Pérez-Asso & Perez-Gelabert, 2001: 70 *H > TL: Carrefour, Rivière Froid
Spirobolellus pullus (Loomis, 1941a: 46). Pérez-Asso & Perez-Gelabert, 2001: 70 *DR > TL: Between 1,000 and 3,000 ft.
elevation, Mt. Quita Espuela
Spirobolellus sigillatus (Loomis, 1934: 20). Pérez-Asso & Perez-Gelabert, 2001: 70 *H > TL: Morne Pilboreau
Spirobolellus signatus (Loomis, 1941a: 42). Pérez-Asso & Perez-Gelabert, 2001: 70 *DR > TL: Mt. Diego de Ocampo,
Northern Range, 3,000 to 4,000 ft. elevation
Spirobolellus tenuipes (Loomis, 1941a: 44). Pérez-Asso & Perez-Gelabert, 2001: 70 *DR > TL: Between 5,000 and 8,000 ft.
elevation, Loma Rucilla and mountains north

Family RHINOCRICIDAE
Alcimobolus Loomis, 1936
Alcimobolus domingensis (Saussure & Humbert, 1860: 363). Pérez-Asso & Perez-Gelabert, 2001: 70 *DR > TL: San Lorenzo,
Samaná Bay
Bahorucocricus Pérez-Asso, 2004
Bahorucocricus crassus Pérez-Asso, 2004: 16 *DR > TL: Carretera 44, km 22, entre San Rafael y La Ciénaga, Sierra de
Bahoruco Oriental, Provincia Barahona
Dibothrocricus Pérez-Asso, 2004
Dibothrocricus pedernales Pérez-Asso, 2004: 14 *DR > TL: Carretera de Pedernales a Los Arroyos, alrededor del km 13,
Provincia Pedernales
Haitobolus Mauries & Hoffman, 1998
Haitobolus haitensis (Gervais in Walckenaer & Gervais, 1847: 191). Pérez-Asso & Perez-Gelabert, 2001: 70 *H > TL: Ile St.
Domingue (Haïti)
=Rhinocricus latespargor Loomis, 1941b: 190
Haitobolus lethifer (Loomis, 1936: 67). Pérez-Asso & Perez-Gelabert, 2001: 70 *H > TL: Between Petit Goave and Miragoâne
Jobocricus Pérez-Asso, 2004
Jobocricus maltzani (Pocock, 1894: 495). Hoffman, 1998: 108; Hoffman, 1999: 92; Pérez-Asso & Perez-Gelabert, 2001: 70 *H
> TL: Cape Haitien in St. Domingo
Leiocricus Loomis, 1936
Leiocricus diversipes Loomis, 1936: 51. Pérez-Asso & Perez-Gelabert, 2001: 71 *H > TL: Top of Morne Pilboreau, between
Ennery and Plaisance
Nesobolus Chamberlin, 1918
Nesobolus loomisi Hoffman 1998: 105. Pérez-Asso & Perez-Gelabert, 2001: 71 *H > TL: No type material known, a neotype
needs to be selected from material at USNM (Hoffman, 1998: 105)
Quisquellacricus Pérez-Asso, 2004
Quisquellacricus hispaniolus (Loomis, 1941a: 39). Pérez-Asso, 2004: 17 *DR > TL: Jarabacoa
Quisquellacricus jamey Pérez-Asso, 2004: 20 *DR > TL: Orillas del río Jamey, El Caobal, provincia San Cristóbal
Quisquellacricus quitaespuela Pérez-Asso, 2004: 19 *DR > TL: Loma Quita Espuela (800—985 msnm), provincia Duarte
Rhinocricidae of uncertain generic position
Anadenobolus Silvestri, 1897
Anadenobolus albolatus (Loomis, 1936: 71). Pérez-Asso & Perez-Gelabert, 2001: 71; Pérez-Asso, 2004: 12 *H > TL: Kenscoff
Anadenobolus cinctus (Loomis, 1936: 72). Pérez-Asso & Perez-Gelabert, 2001: 71; Pérez-Asso, 2004: 12 *H > TL: Morne La
Hotte
Anadenobolus? curtior (Chamberlin, 1918: 194). Pérez-Asso & Perez-Gelabert, 2001: 71 *H > TL: Grande Rivière
Anadenobolus furcianus (Chamberlin, 1918: 192). Pérez-Asso & Perez-Gelabert, 2001: 71; Pérez-Asso, 2004: 12 *H > TL:
Furcy
Anadenobolus? mediator (Chamberlin, 1918: 189). Pérez-Asso & Perez-Gelabert, 2001: 71 *H > TL: Furcy
Anadenobolus? modestior (Silvestri, 1908). Pérez-Asso & Perez-Gelabert, 2001: 71
Anadenobolus monilicornis (Porat, 1876). Pérez-Asso & Perez-Gelabert, 2001: 71; Shelley, 2014: 2
Anadenobolus nigrescens (Chamberlin, 1918: 195). Pérez-Asso & Perez-Gelabert, 2001: 72; Pérez-Asso, 2004: 12 *H > TL:
Furcy
Anadenobolus pedernales Pérez-Asso, 2004: 12 *DR > TL: Carretera de Las Mercedes al Hoyo de Pelempito, Provincia
Pedernales
Anadenobolus? ramulus (Loomis, 1936: 76). Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Roche Croix on Morne La

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Hotte, 5000 ft.
Anadenobolus? rarior (Chamberlin, 1918: 187). Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Grande Rivière, near Jacmel
Anadenobolus? scobinellus (Loomis, 1936: 63). Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Foothills of Morne La Hotte,
4000 ft.
Cubobolus Chamberlin, 1918
Cubobolus? consutus Loomis, 1936: 53. Pérez-Asso & Perez-Gelabert, 2001: 71 *H > TL: Le Brande

Family PSEUDOSPIROBOLELLIDAE
Pseudospirobolellus Carl, 1912
Pseudospirobolellus avernus (Butler, 1876). Pérez-Asso & Perez-Gelabert, 2001: 72 [Introduced]

Family TRIGONIULIDAE
Trigoniulus Pocock, 1894
Trigoniulus corallinus (Gervais, 1842). Loomis, 1936: 50; Pérez-Asso & Perez-Gelabert, 2001: 72 [Introduced]

Order SPIROSTREPTIDA
Family PSEUDONNANOLENIDAE
Cambalomma Loomis, 1941
Cambalomma laeve (Loomis, 1941b: 189). Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Jérémie
Epinannolene Brolemann, 1903
Epinannolene convexus Loomis, 1936: 32. Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Morne La Hotte, between 3,000
and 7,800 ft.
Epinannolene haitensis Chamberlin, 1918: 179. Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Grande Rivière du Nord,
near Cap Haitien
Epinannolene ornata Chamberlin, 1918: 180. Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Pétionville
Epinannolene sp.†—Santiago-Blay & Poinar (1992: 366); Pérez-Asso & Perez-Gelabert (2001: 72) [Fossil in Dominican
amber]
Epinannolene virgata Loomis, 1936: 34. Pérez-Asso & Perez-Gelabert, 2001: 72 *H > TL: Near Trouin

Family SPIROSTREPTIDAE
Orthoporus Silvestri, 1897
Orthoporus caelatus Loomis, 1936: 38. Pérez-Asso & Perez-Gelabert, 2001: 73 *H > TL: Between Port Margo and Limbe
Orthoporus haitiensis Chamberlin, 1918: 183. Pérez-Asso & Perez-Gelabert, 2001: 73 *H > TL: St. Marc
Orthoporus triquetrus Loomis, 1936: 37. Pérez-Asso & Perez-Gelabert, 2001: 73 *DR > TL: San Lorenzo, Samana Bay

Order STEMMIULIDA
Family STEMMIULIDAE
Prostemmiulus Silvestri, 1916
Prostemmiulus abditus Loomis, 1936: 18. Pérez-Asso & Perez-Gelabert, 2001: 68 *H > TL: Morne Pilboreau, above Ennery
Prostemmiulus affinis Loomis, 1936: 28. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Between Petit Goave and Leogane
Prostemmiulus clarus Chamberlin, 1918: 176. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Jacmel
Prostemmiulus clavipes Loomis, 1936: 26. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Near Trouin
Prostemmiulus cognatus Loomis, 1936: 25. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Near Trouin
Prostemmiulus gracilipes Loomis, 1941a: 26. Pérez-Asso & Perez-Gelabert, 2001: 69 *DR > TL: Rain forest near Valle Nuevo,
about 6,000 ft. elev.
Prostemmiulus interruptus Loomis, 1936: 24. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: La Visite, La Selle Range,
6000‒7000 ft.
Prostemmiulus iuloides Loomis, 1941a: 22. Pérez-Asso & Perez-Gelabert, 2001: 69 *DR > TL: Pico del Yaque, Loma Rucilla,
8,000 to 10,000 ft. elev.
Prostemmiulus quadristriatus Loomis, 1936: 22. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Morne Pilboreau, above
Ennery
Prostemmiulus scaurus Loomis, 1941a: 24. Pérez-Asso & Perez-Gelabert, 2001: 69 *DR > TL: Valle Nuevo, southeast of
Constanza, elev. about 7,000 ft.
Prostemmiulus setosus Loomis, 1941a: 31. Pérez-Asso & Perez-Gelabert, 2001: 69 *DR > TL: Valle Nuevo, southeast of

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Constanza, elev. about 7,000 ft.
Prostemmiulus subulatus Loomis, 1936: 20. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Morne Brigand, near Bayeaux
Prostemmiulus sulcatus Loomis, 1941a: 30. Pérez-Asso & Perez-Gelabert, 2001: 69 *DR > TL: Between 1,000 and 3,000 ft.
elev., Mt. Quita Espuela
Prostemmiulus tridigitatus Loomis, 1941a: 28. Pérez-Asso & Perez-Gelabert, 2001: 69 *DR > TL: Between 1,000 and 3,000 ft.
elev., Mt. Quita Espuela
Prostemmiulus venustus Loomis, 1936: 16. Pérez-Asso & Perez-Gelabert, 2001: 69 *H > TL: Morne Brigand, near Bayeaux
Prostemmiulus xenops (Loomis, 1950: 164). Pérez-Asso & Perez-Gelabert, 2001: 69
*H > TL: Pétionville
=Prostemmiulus heterops Loomis, 1936: 29 [Preoccupied name]

Class SYMPHYLA
Family SCUTIGERELLIDAE
Scutigerella Ryder, 1882
Scutigerella dominicana† Poinar & Edwards, 1995: 391 [Fossil in Dominican amber]

Subphylum HEXAPODA
Class COLLEMBOLA
Classification follows: Checklist of the Collembola http://www.collembola.org/taxa/collembo.htm - by Frans Janssens.
Key reference: Mari Mutt and Bellinger (1990).

Taxonomic summary:
Order Poduromorpha
Superfamily Neanuroidea
Family Brachystomellidae
Superfamily Hypogastruroidea
Family Hypogastruridae
Order Entomobryomorpha
Superfamily Isotomoidea
Family Isotomidae
Superfamily Entomobryoidea
Family Entomobryidae
Family Paronellidae
Family Seiridae
Order Symphypleona
Superfamily Dicyrtomoidea
Family Dicyrtomidae
Superfamily Sminthuroidea
Family Sminthuridae

Order PODUROMORPHA
Superfamily NEANUROIDEA
Family BRACHYSTOMELLIDAE
Brachystomella Ågren, 1903
Brachystomella parvula (Schäffer, 1896). Mari Mutt, 1977: 282; Mari Mutt & Bellinger, 1990: 25

Superfamily HYPOGASTRUROIDEA
Family HYPOGASTRURIDAE
Ceratophysella Börner in Bromer, 1932
Ceratophysella armata (Nicolet, 1842). Mari Mutt, 1977: 281; Mari Mutt & Bellinger, 1990: 6
Xenylla Tullberg, 1869
Xenylla sp.—Palacios Vargas (2008: 46)

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Order ENTOMOBRYOMORPHA
Superfamily ISOTOMOIDEA
Family ISOTOMIDAE
Subfamily ANUROPHORINAE
Cryptopygus Willem, 1901
Cryptopygus sp.—Mari Mutt & Gruia (1977: 282)
Cryptopygus sp.†—Mari Mutt (1983: 578) [Fossil in Dominican amber]
Subfamily PROISOTOMINAE
Folsomides Stach, 1922
Folsomides parvulus Stach, 1922. By synonymy of species below.
=Folsomides americanus Denis, 1931. Mari Mutt, 1977: 282; Mari Mutt & Bellinger, 1990: 63
Subfamily ISOTOMINAE
Isotoma Bourlet, 1839
Isotoma (Desoria) sp. 1†—Mari Mutt (1983: 578) [Fossil in Dominican amber]
Isotoma (Desoria) sp. 2†—Mari Mutt (1983: 578) [Fossil in Dominican amber]
Isotomiella Bagnall, 1939
Isotomiella minor (Schäffer, 1896). Mari Mutt, 1977: 282; Mari Mutt & Bellinger, 1990: 82
Isotomurus Börner, 1903
Isotomurus sp.—Mari Mutt & Gruia (1977: 282)

Superfamily ENTOMOBRYOIDEA
Family ENTOMOBRYIDAE
Subfamily CYPHODERINAE
Campylothorax Schött, 1893
Campylothorax cubanus Gruia, 1983. Soto-Adames, 2016: 23
Campylothorax dominicanus Soto-Adames, 2016: 20 *DR > TL: Loma Casabito, 15.8 km NW Bonao, 1455 m
Campylothorax hexosetosus Soto-Adames, 2016: 16 *DR > TL: Loma Casabito, 15.8 km NW Bonao, 1487 m
Campylothorax notidanus Soto-Adames, 2016: 11 *DR > TL: Loma Casabito, 15.8 km NW Bonao, 1455 m
Campylothorax sp.—Mari Mutt & Gruia (1977: 282)
Cyphoderus Nicolet, 1842
Cyphoderus similis Folsom, 1927. Mari Mutt, 1977: 282
Cyphoderus sp. 1†—Mari Mutt (1983: 581) [Fossil in Dominican amber]
Cyphoderus sp. 2†—Mari Mutt (1983: 581) [Fossil in Dominican amber]
Dicranocentrus Schött, 1893
Dicranocentrus antillensis Mari Mutt, 1979: 57. Mari Mutt & Bellinger, 1990: 87
Dicranocentrus biseriatus Mari Mutt, 1981: 97. Mari Mutt & Bellinger, 1990: 88 *H > TL: 3 km above SHADA headquarters,
2,200 m
Dicranocentrus celatus Mari Mutt, 1985: 105. Mari Mutt & Bellinger, 1990: 88
Dicranocentrus millsi millsi Mari Mutt, 1979: 61. Mari Mutt & Bellinger, 1990: 89 *H > TL: La Salle, above Fond Verrettes,
1000 m
Dicranocentrus zebratus Mari Mutt, 1985: 106. Mari Mutt & Bellinger, 1990: 91 *DR > TL: Prov. La Vega, El Río-Bonao, km
24, 600-800 m

Family LEPIDOCYRTIDAE
Subfamily LEPIDOCYRTINAE
Lepidocyrtus Bourlet, 1839
Lepidocyrtus sp.—Mari Mutt & Gruia (1977: 282)
Lepidocyrtus sp.†—Mari Mutt (1983: 579) [Fossil in Dominican amber]
Metasinella Denis, 1929
Metasinella (Sulcuncus) millsi Mari Mutt & Gruia, 1983. Mari Mutt & Bellinger, 1990: 118
Metasinella (Sulcuncus) rapoporti Massoud & Gruia, 1973. Mari Mutt, 1977: 282; Mari Mutt & Bellinger, 1990: 119
Pseudosinella Schäffer, 1897
Pseudosinella sp.†—Mari Mutt (1983: 581) [Fossil in Dominican amber]

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Family PARONELLIDAE
Subfamily CYPHODERINAE
Paronella Schött, 1893
Paronella sp.—Mari Mutt & Gruia (1977: 282)
Paronella sp.†—Mari Mutt (1983: 581) [Fossil in Dominican amber]
Trogolaphysa Mills, 1938
Trogolaphysa haitica (Palacios Vargas et al., 1986: 16). Mari Mutt & Bellinger, 1990: 121 *H > TL: Depto. du Sud: Gruta
Counoubais, 1 km SW Camp Perrin
Trogolaphysa maya (Mills, 1938). Mari Mutt & Bellinger, 1990: 122
=Troglopedetes delamarei Massoud & Gruia, 1973. Mari Mutt, 1977: 282
Trogolaphysa quisqueyana Soto-Adames, Jordana & Baquero, 2014: 10 *DR > TL: Loma del Toro, Sierra de Bahoruco, 2,316
m
Subfamily PARONELLINAE
Salina MacGillivray, 1894
Salina sp.†—Mari Mutt (1983: 581) [Fossil in Dominican amber]

Family SEIRIDAE
Seira Lubbock, 1870
Seira dowlingi (Wray, 1953). Wray, 1959: 67; Mari Mutt & Bellinger, 1990: 104
Seira sp.†—Mari Mutt (1983: 581) [Fossil in Dominican amber]

Order SYMPHYPLEONA
Superfamily DICYRTOMOIDEA
Family DICYRTOMIDAE
Subfamily PTENOTHRICINAE
Ptenothrix Börner, 1906
Ptenothrix dominicana Mari Mutt, 1977: 285 *DR > TL: La Vega Prov., El Río, Bonao, 600–800 m

Superfamily SMINTHURIDOIDEA
Family SMINTHURIDAE
Subfamily SPHYROTHECINAE
Sphyrotheca Börner, 1906
Sphyrotheca sp.—Mari Mutt (1977: 285)
Sphyrotheca sp.†—Mari Mutt (1983: 585) [Fossil in Dominican amber]

Family SMINTHURIDIDAE
Electrosminthuridia† Robin, D’Haese & Barden, 2019
Electrosminthuridia helibionta† Robin, D’Haese & Barden, 2019: 5 [Fossil in Dominican amber]

Class PROTURA
Family ACERENTOMIDAE
Bolivaridia Bonet, 1942
Bolivaridia perissochaeta Bonet, 1942. Nosek & Mari Mutt, 1978: 305

Family EOSENTOMIDAE
Eosentomon Berlese, 1908
Eosentomon puertoricoense Nosek, 1978. Nosek & Mari Mutt, 1978: 305

Class INSECTA
Subclass MONOCONDYLIA
Order ARCHAEOGNATHA (=MICROCORYPHIA)
Family MEINERTELLIDAE
Neomachilellus Wygodzinsky, 1953
Neomachilellus (Praeneomachilellus) dominicanus† Sturm & Poinar, 1997: 162 [Fossil in Dominican amber]

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Subclass APTERYGOTA
Order ZYGENTOMA
Family LEPISMATIDAE
Ctenolepisma Escherich, 1905
Ctenolepisma (Ctenolepisma) electrans† Mendes, 1998: 899 [Fossil in Dominican amber]
Ctenolepisma (Ctenolepisma) rothschildi Silvestri, 1907. Irish, 1995: 562
Lepisma Linnaeus, 1758
Lepisma saccharina Linnaeus, 1758. Wolcott, 1927: 135
Protolepisma† Mendes & Poinar, 2013
Protolepisma tainicum† Mendes & Poinar, 2013: 3 [Fossil in Dominican amber]

Family NICOLETIIDAE
Archeatelura† Mendes, 1997
Archeatelura sturmi† Mendes, 1997: 41. Mendes, 1997: 804 [Fossil in Dominican amber]
Hemitrinemura Mendes, 1994
Hemitrinemura exstincta† Mendes & Poinar, 2004: 103 [Fossil in Dominican amber]
Trinemurodes Silvestri, 1921
Trinemurodes antiquus† Sturm & Mendes, 1998: 2 [Fossil in Dominican amber]
Trinemurodes miocenicus† Sturm & Mendes, 1998: 3 [Fossil in Dominican amber]

Order COLEOPTERA

Taxonomic summary:
Suborder Archostemata
Family Micromalthidae
Suborder Adephaga
Family Gyrinidae
Family Haliplidae
Family Noteridae
Family Dytiscidae
Family Carabidae
Suborder Polyphaga
Infraorder Staphyliniformia
Superfamily Hydrophiloidea
Family Hydrochidae
Family Hydrophilidae
Family Histeridae
Superfamily Staphylinoidea
Family Hydraenidae
Family Leiodidae
Family Silphidae
Family Staphylinidae
Infraorder Scarabaeiformia
Superfamily Scarabaeoidea
Family Passalidae
Family Trogidae
Family Geotrupidae
Family Ceratocanthidae
Family Hybosoridae
Family Scarabaeidae
Family Lucanidae
Infraorder Elateriformia
Superfamily Buprestoidea
Family Buprestidae

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Superfamily Byrrhoidea
Family Elmidae
Family Dryopidae
Family Limnichidae
Family Lutrochidae
Family Heteroceridae
Family Psephenidae
Family Chelonariidae
Superfamily Elateroidea
Family Brachypsectridae
Family Eucnemidae
Family Elateridae
Family Lycidae
Family Phengodidae
Family Lampyridae
Family Cantharidae
Infraorder Bostrichiformia
Superfamily Bostrichoidea
Family Dermestidae
Family Bostrichidae
Family Anobiidae
Infraorder Cucujiformia
Superfamily Lymexyloidea
Family Lymexylidae
Superfamily Cleroidea
Family Trogossitidae
Family Cleridae
Family Melyridae
Superfamily Cucujoidea
Family Sphindidae
Family Nitidulidae
Family Smicripidae
Family Monotomidae
Family Silvanidae
Family Passandridae
Family Laemophloeidae
Family Phalacridae
Family Cryptophagidae
Family Erotylidae
Superfamily Coccinelloidea
Family Bothrideridae
Family Cerylonidae
Family Coccinellidae
Family Corylophidae
Family Discolomatidae
Family Endomychidae
Family Latridiidae
Superfamily Tenebrionoidea
Family Ciidae
Family Mordellidae
Family Scraptiidae
Family Ripiphoridae
Family Zopheridae
Family Mycetophagidae

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 Family Tenebrionidae
Family Oedemeridae
Family Meloidae
Family Mycteridae
Family Salpingidae
Family Anthicidae
Superfamily Chrysomeloidea
Family Cerambycidae
Family Chrysomelidae
Superfamily Curculionoidea
Family Anthribidae
Family Attelabidae
Family Apionidae
Family Brentidae
Family Belidae
Family Curculionidae
Family Rhynchitidae

Suborder ARCHOSTEMATA
Family MICROMALTHIDAE
Micromalthus LeConte, 1878
Micromalthus debilis† LeConte, 1878. Hörnschemeyer, Wedmann & Poinar, 2010: 302 [Fossil in Dominican amber]
=Micromalthus anansi† Perkovsky, 2007: 626 [Fossil in Dominican amber]

Suborder ADEPHAGA
Family GYRINIDAE
Subfamily GYRININAE
Tribe GYRININI
Dineutus MacLeay, 1825
Dineutus americanus (Linnaeus, 1767). Ochs, 1938: 86; Gustafson & Miller, 2015: 23
Dineutus longimanus longimanus (Olivier, 1792). Chevrolat, 1863: 203; Ochs, 1930: 126; Ochs, 1938: 86; Blackwelder, 1944:
81; Gustafson & Miller, 2015: 69 *DR, *H > TL: Saint Domingue
Tribe OREOTOCHILINI
Gyretes Brullé, 1834
Gyretes nigrilabris Ochs, 1938: 92. Blackwelder, 1944: 82
Gyretes vulnerata Aubé, 1838. Chevrolat, 1863: 203; Ochs, 1938: 89; Blackwelder, 1944: 82
Gyrinus Geoffroy, 1862
Gyrinus rugifer Régimbart, 1883. Ochs, 1924: 7; Ochs, 1938: 85

Family HALIPLIDAE
Haliplus Latreille, 1802
Haliplus (Liaphlus) cubensis Chapin, 1930. Vondel in Nilsson & Vondel, 2005: 41
Haliplus (Liaphlus) gravidus Aubé, 1838. Vondel in Nilsson & Vondel, 2005: 46
=Haliplus brandeni Wehncke, 1883: 146. Blackwelder, 1944: 72; Vondel in Nilsson & Vondel, 2005: 46
Haliplus (Liaphlus) havaniensis Wehncke, 1880. Vondel in Nilsson & Vondel, 2005: 47
Haliplus youngi Vondel & Spangler, 2008: 124

Family NOTERIDAE
Subfamily NOTERINAE
Hydrocanthus Say, 1825
Hydrocanthus (Hydrocanthus) advena Sharp, 1882. Peck, 2005: 43; Nilsson in Nilsson & Vondel, 2005: 110
Mesonoterus Sharp, 1882
Mesonoterus laevicollis Sharp, 1882. Peck, 2005: 43
Notomicrus Sharp, 1882

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Notomicrus sharpi J. Balfour-Browne, 1939. Peck, 2005: 43; Nilsson in Nilsson & Vondel, 2005: 139
Suphis Aubé, 1836
Suphis inflatus (LeConte, 1863). Peck, 2005: 43
Suphisellus Crotch, 1873
Suphisellus binotatus (Fleutiaux & Sallé, 1890). Peck, 2005: 43; Nilsson in Nilsson & Vondel, 2005: 127
Suphisellus insularis (Sharp, 1882). Blackwelder, 1944: 73; Nilsson in Nilsson & Vondel, 2005: 130

Family DYTISCIDAE
Subfamily COLYMBETINAE
Rhantus Dejean, 1833
Rhantus binotatus (Harris, 1828). Blackwelder, 1944: 79
Rhantus calidus (Fabricius, 1792). Blackwelder, 1944: 79
Rhantus gutticollis (Say, 1830). Nilsson, 2001: 50
=Colymbetes binotatus Aubé, 1838. Nilsson, 2001: 50
=Rhantus domingensis Gemminger & Harold, 1868. Nilsson, 2001: 50
Subfamily COPELATINAE
Copelatus Erichson, 1832
Copelatus caelatipennis Chevrolat, 1863. Manuel et al., 2018: 379
Copelatus insolitus Chevrolat, 1863. Blackwelder, 1944: 77; Manuel et al., 2018: 380
Copelatus martini Manuël et al., 2018: 373 *DR > TL: La Vega, Reserva Científica Ebano Verde, 1,070 m
Copelatus posticatus (Fabricius, 1801). Blackwelder, 1944: 78; Manuel et al., 2018: 380
Copelatus predaveterus† Miller in Miller & Balke, 2003: 812 [Fossil in Dominican amber]
Copelatus vitraci Legros, 1948. Manuel et al., 2018: 380
Subfamily DYTISCINAE
Acilius Leach, 1817
Acilius semisulcatus Aubé, 1838. Blackwelder, 1944: 80
Hydaticus Leach, 1817
Hydaticus rimosus Aubé, 1838. Peck, 2005: 46
Megadytes Sharp, 1882
Megadytes fraternus Sharp, 1882. Blackwelder, 1944: 80
Megadytes giganteus (Laporte, 1835). Peck, 2005: 47
Thermonectus Dejean, 1833
Thermonectus basillaris (Harris, 1829). Peck, 2005: 47
Thermonectus circumscriptus (Latreille, 1812). García Avila & Peña, 1993: 214
Thermonectus margineguttatus (Aubé, 1838). Peck, 2005: 47
Subfamily HYDROPORINAE
Bidessonotus Régimbart, 1895
Bidessonotus browneanus J. Balfour-Browne, 1947. Young, 1990: 376
Brachyvatus Zimmermann, 1919
Brachyvatus apicatus (Clark, 1862). Blackwelder, 1944: 76
Derovatellus Sharp, 1882
Derovatellus lentus (Wehncke, 1876). K. Miller, 2005: 448
Desmopachria Babington, 1841
Desmopachria (Desmopachria) darlingtoni Young, 1989: 307
Hydrovatus Motschulsky, 1853
Hydrovatus caraibus Sharp, 1882. Trémouilles et al., 2005: 63
Neobidessus Young, 1967
Neobidessus pullus floridanus (Fall, 1917). Peck, 2005: 46
Subfamily LACCOPHILINAE
Laccomimus Toledo & Michat, 2015
Laccomimus bordoni Toledo & Michat, 2015: 320
Laccophilus Leach, 1815
Laccophilus americanus Aubé in Jacquelin du Val, 1856. Peck, 2005: 44
Laccophilus bifasciatus Chevrolat, 1863. Blackwelder, 1944: 74

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Laccophilus gentilis suavis Sharp, 1882. Peck, 2005: 44
Laccophilus proximus Say, 1823. Young, 1954: 45
Laccophilus quadrivittatus Aubé, 1838. Peck, 2005: 45
Laccophilus vacaensis Young, 1953. Peck, 2005: 45

Family CARABIDAE
Subfamily BRACHININAE
Brachinus Weber, 1801
Brachinus brunneus Laporte, 1836. Erwin & Sims, 1984: 428
Brachinus sp.†—Wu (1997: 172) [Misspelled Brachynus] [Fossil in Dominican amber]
Pheropsophus Solier, 1834
Pheropsophus aequinoctialis (Linnaeus, 1763). Blackwelder, 1944: 70
Subfamily CARABINAE
Tribe CARABINI
Calosoma Weber, 1801
Calosoma (Calosoma) splendidum Dejean, 1831. Chevrolat, 1863: 186; Erwin & Sims, 1984: 423; Erwin, 2007: 104;
Bousquet, 2012: 232
Calosoma (Castrida) sayi Dejean, 1826. Gidaspow, 1963: 300; Erwin, 2007: 103; Bousquet, 2012: 229
Subfamily CICINDELINAE
Tribe MEGACEPHALINI
Tetracha Westwood, 1838
Tetracha (Tetracha) sobrina infuscata (Mannerheim, 1837). Hummelinck, 1983: 109; Freitag, 1992: 154; Naviaux, 2007: 51
Tetracha (Neotetracha) affinis (Dejean, 1825). Blackwelder, 1944: 12; Hummelinck, 1983: 106
Tetracha (Neotetracha) acutipennis (Dejean, 1825: 13). Wolcott, 1927: 215; Hummelinck, 1983: 106; Erwin & Sims, 1984:
423; Freitag, 1992: 154; Naviaux, 2007: 103
Tetracha (Neotetracha) misella Naviaux, 2007: 104 *H, *DR > TL: Haiti
Tetracha (Neotetracha) rutilans Thomson, 1857. Erwin & Sims, 1984: 424
Tribe CICINDELINI
Brasiella Rivalier, 1954
Brasiella bellorum Acciavati, 2011: 111 *DR > TL: La Vega, 4.1 km SW El Convento, 1730 m
Brasiella darlingtoniana Acciavati, 2011: 117 *H > TL: Kenscoff to La Visite & vic, La Selle Range, 5000 ft
Brasiella davidsoni Acciavati, 2011: 124 *H > TL: Departement du Sud, Ville Formon, 1405 m
Brasiella dominicana (Mandl, 1983: 117). Freitag, 1992: 157 *H, *DR > TL: Baní
Brasiella iviei Acciavati, 2011: 138 *DR > TL: Prov. Pedernales, ca. 35 km N Cabo Rojo, Las Abejas, 1250 m
Brasiella ocoa Acciavati, 2011: 145 *DR > TL: Peravia Prov., Río Ocoa, 130 m
Brasiella philipi Acciavati, 2011: 149 *DR > TL: Santiago Prov., Sierrecita above Mata Grande, 1290 m
Brasiella rawlinsi Acciavati, 2011: 155 *DR > TL: Pedernales, 37 km N Cabo Rojo, 1500 m
Brasiella youngi Acciavati, 2011: 163 *DR > TL: Prov. Pedernales, 13.5 km N Cabo Rojo, 140 m
Cicindela Linnaeus, 1758
Cicindela argentata Fabricius, 1801. Blackwelder, 1944: 17; Erwin & Sims, 1984: 424
Cicindela marginata Fabricius, 1775. Choate, 2003: 115
Cicindela olivacea Chaudoir, 1854. Choate, 2003: 101
Cicindela (Cicindelidia) trifasciata Fabricius, 1781. Wolcott, 1927: 215; Hummelinck, 1983: 74; Erwin & Sims, 1984: 425;
Freitag, 1992: 154
Cicindela (Cicindelidia) viridiflavescens Horn, 1923. Erwin & Sims, 1984: 423; Freitag, 1992: 154
Cicindela (Plectographa) schaefferi Horn, 1903. Erwin & Sims, 1984: 424; Freitag, 1992: 157
Cylindera Westwood, 1831
Cylindera (Plectographa) suturalis suturalis (Fabricius, 1798). Wolcott, 1927: 215; Hummelinck, 1983: 76; Erwin & Sims,
1984: 424
Cylindera (Plectographa) suturalis hebraea Klug, 1834. Hummelinck, 1983: 78; Freitag, 1992: 157
Habroscelimorpha Dokhturov, 1883
Habroscelimorpha boops (Dejean, 1831). Leng & Mutchler, 1914: 393; Wolcott, 1927: 215; Hummelinck, 1983: 81; Erwin &
Sims, 1984: 424
Note: Cicindela (Cicindelidia) rufiventris rufiventris Dejean, 1825 is cited for Hispaniola by Blackwelder, 1944: 19 and Erwin

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& Sims, 1984: 424. Bousquet, 2012: 323 postulates that this is incorrect and gives records for this species only from USA.
Subfamily HARPALINAE
Tribe CHLAENINI
Chlaenius Bonelli, 1810
Chlaenius circumcinctus Say, 1830. Peck, 2005: 34; Bousquet, 2012: 977
=Chlaenius perplexus Dejean, 1831. Peck, 2005: 34
Tribe CTENODACTYLINI
Leptotrachelus Latreille, 1829
Leptotrachelus dorsalis (Fabricius, 1801). Peck & Thomas, 1998: 23; Bousquet, 2012: 1272
Tribe CYCLOSOMINI
Anaulacus Macleay, 1825
Anaulacus (Aephnidius) ciliatus (Mutchler, 1934). Ball & Shpeley, 2002: 288
Anaulacus (Aephnidius) sericatus (Chaudoir, 1846). Peck, 2005: 37
Anaulacus (Aephnidius) sp.†—Ball & Shpeley (2002: 312) [Fossil in Dominican amber]
Tetragonoderus Dejean, 1821
Tetragonoderus (Peronoscelis) quadriguttatus Dejean, 1829. Shpeley & Ball, 2008: 12
Tribe GALERITINI
Galerita Fabricius, 1801
Galerita lecontei tenebricosa (Klug, 1834). Peck, 2005: 39
Galerita montana Darlington, 1935: 211 *H > TL: Vicinity of La Visite, La Selle Range, 5,000–6,000 ft.
Galerita striata Klug, 1834: 66. Reichardt, 1976: 110 *H > TL: Port-au-Prince
Galerita tristis Reiche, 1842. Peck, 2009: 14
=Galerita unicolor Latreille & Dejean, 1824. Leng & Mutchler, 1914: 397
Pseudaptinus Laporte, 1835
Pseudaptinus cubanus (Chaudoir, 1877). Bousquet, 2012: 1359
Pseudaptinus dorsalis (Brullé, 1834). Bousquet, 2012: 1360
Pseudaptinus marginicollis fumipes Darlington, 1935: 212
Tribe HARPALINI
Amblygnathus Dejean, 1829
Amblygnathus puncticollis Putzeys, 1878a: 34. Ball & Maddison, 1987: 223 *DR > TL: St. Domingue
Arthrostictus Csiki, 1932
Arthrostictus sp.†—Poinar & Poinar (1999: 203) [Fossil in Dominican amber]
Bradycellus Erichson, 1837
Bradycellus selleanus Darlington, 1935: 204. Erwin & Sims, 1984: 439 *H > TL: La Visite and vicinity, La Selle Range,
5,000–7,000 ft.
Discoderus LeConte, 1835
Discoderus beauvoisii (Dejean, 1829). Peck, 2005: 32; Shpeley et al., 2017: 117
Discoderus cyaneopacus (Darlington, 1934: 107). Erwin & Sims, 1984: 440; Shpeley et al., 2017: 121 *H > TL: Cap Haitien
Discoderus thoracicus Putzeys, 1878a: 59. Erwin & Sims, 1984: 441; Shpeley et al., 2017: 124 *H > TL: Haïti
Pogonodaptus Horn, 1881
Pogonodaptus rostratus Darlington, 1935: 204. Erwin & Sims, 1984: 439 *H > TL: Swamps north of Dessalines
Neodiachipteryx Noonan, 1985
Neodiachipteryx cariniger (Putzeys, 1878a: 44). Blackwelder, 1944: 49; Shpeley et al., 2017: 33 *DR > TL: St. Domingue
Neodiachipteryx davidsoni Shpeley et al., 2017: 36 *DR > TL: Sabaneta, Prov. Santiago Rodríguez, Zamba
Selenophorus Dejean, 1829
Selenophorus alternans Dejean, 1829. Putzeys, 1878a: 13; Peck, 2005: 32; Shpeley et al., 2017: 96
Selenophorus chalybaeus Dejean, 1829. Erwin & Sims, 1984: 440
Selenophorus clypealis Ball & Shpeley in Ball, 1992: 101 *H > TL: Source Matelas
Selenophorus cubanus ubancus Ball & Shpeley in Ball, 1992: 103
Selenophorus dessalinesi Ball & Shpeley in Ball, 1992: 102; Shpeley et al., 2017: 58 *H, *DR > TL: Just north of Dessalines,
Artibonite Department
Selenophorus discopunctatus Dejean, 1829. Erwin & Sims, 1984: 440; Bousquet, 2012: 1137
Selenophorus fabricii Shpeley et al., 2017: 79
Selenophorus flavilabris ubancus Ball & Shpeley, 1992: 103. Shpeley et al., 2017: 86

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Selenophorus integer (Fabricius, 1801). Erwin & Sims, 1984: 440; Bousquet, 2012: 1143; Shpeley et al., 2017: 88
Selenophorus latior Darlington, 1934: 109. Erwin & Sims, 1984: 440: Shpeley et al., 2017: 48
Selenophorus mundus Putzeys, 1878 [By synonymy of species below]
=Selenophorus haitianus Darlington, 1934: 107. Erwin & Sims, 1984: 440; Ball, 1992: 96
Selenophorus nonseriatus Darlington, 1934: 109. Erwin & Sims, 1984: 440; Shpeley et al., 2017: 77
Selenophorus pyritosus Dejean, 1829. Peck, 2005: 33; Shpeley et al., 2017: 97
Selenophorus parumpunctatus Dejean, 1829. Bousquet, 2012: 1145; Shpeley et al., 2017: 106
Selenophorus pseudomundus Ball & Shpeley in Ball, 1992: 99. Shpeley et al., 2017: 69 *DR > TL: Las Mercedes, Pedernales
Province
Selenophorus sinuatus Gyllenhal, 1806. Peck, 2005: 33
Selenophorus sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Selenophorus striatopunctatus Putzeys, 1878. Peck, 2005: 33; Bousquet, 2012: 1147; Shpeley et al., 2017: 109
Selenophorus subquadratus (Putzeys, 1878b: 293). Blackwelder, 1944: 48; Erwin & Sims, 1984: 441
Stenolophus Dejean, 1821
Stenolophus ochropezus (Say, 1823). Bousquet, 2012: 1050
Stenomorphus Dejean, 1831
Stenomorphus californicus manni Darlington, 1934: 102. Erwin & Sims, 1984: 441 *H, *DR > TL: Manneville, Ouest
Department
Tribe LACHNOPHORINI
Anchonoderus Reiche, 1843
Anchonoderus darlingtoni Liebherr, 1988: 25
Eucaerus LeConte, 1853
Eucaerus haitianus Darlington, 1935: 210 *H, *DR > TL: Etang Lachaux
Euphorticus Horn, 1881
Euphorticus pubescens (Dejean, 1831). Peck & Thomas, 1998: 23
Lachnophorus Dejean, 1831
Lachnophorus leucopterus Chevrolat, 1863: 198. Peck, 2005: 36
Tribe LEBIINI
Apenes LeConte, 1851
Apenes davidsoni Ball & Shpeley, 2009: 146 *H > TL: Along Río Mulito, 13 km N Pedernales
Apenes fasciata Chaudoir, 1875. Ball, 1992: 106; Ball & Shpeley, 2009: 147 *DR > TL: 4 km southeast of Río Limpio
Apenes lachauxi Ball & Shpeley, 2009: 167 *H > TL: Etang Lachaux, Departement du Sud
Apenes laevicincta Darlington, 1934: 119. Ball & Shpeley, 2009: 165 *H > TL: Port-au-Prince, about 3000 ft.
Apenes lucidula dulculia Ball & Shpeley, 1992: 116. Ball & Shpeley, 2009: 138
Apenes marginalis (Dejean, 1831). Ball & Shpeley, 2009: 106
Apenes morio (Dejean, 1825). Ball & Shpeley, 2009: 119
Apenes ovalis Darlington, 1935: 210. Ball & Shpeley, 2009: 132 *H > TL: La Visite and vicinity, La Selle Range, 5,000–7,000
ft.
Apenes philipi Ball & Shpeley, 2009: 151 *DR > TL: 2 km N Bayahibe, 10 m
Apenes rawlinsi Ball & Shpeley, 2009: 168 *DR > TL: Parque Los Haitises, 3 km W Cueva de Arena, 20 m
Apenes scobifera scobifera Darlington, 1953. Ball & Shpeley, 2009: 170
Apenes toussainti Ball & Shpeley, 2009: 163 *H > TL: Swamps N of Dessalines
Apenes youngi Ball & Shpeley, 2009: 134 *DR > TL: 13.5 km SE Puerto Escondido, Sierra de Bahoruco, 1789 m
Callida Lepeletier & Dejean, 1824
Callida pretiosa Chaudoir, 1872: 124. Erwin & Sims, 1984: 441 *DR > TL: Dans la partie espagnole de Haïti
Coptodera Dejean, 1825
Coptodera festiva Dejean, 1825. Shpeley & Ball, 1993: 82; Bousquet, 2012: 1282
Coptodera picea Dejean, 1826. Shpeley & Ball, 2000: 148; Bousquet, 2012: 1282
Cymindis Latreille, 1805
Cymindis morio Dejean, 1825. Bousquet, 2012: 1620
Lebia Latreille, 1802
Lebia gibba Darlington, 1935: 207. Erwin & Sims, 1984: 447 *H > TL: Hinche
Lebia haitiana Darlington, 1935: 205 *H > TL: Vicinity of Port-au-Prince
Lebia nigrita Darlington, 1935: 206. Erwin & Sims, 1984: 447 *H > TL: Hills south of Port-au-Prince (edge of Massif de la

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Selle), about 2,000 ft.
Lebia nubicola Darlington, 1939: 98. Erwin & Sims, 1984: 447 *DR > TL: Mountains to the north of Loma Rucilla, about
7,000 ft.
Lebia terricola Darlington, 1939: 98. Erwin & Sims, 1984: 447 *DR > TL: Loma del Toro, south of Santiago (north of Loma
Rucilla), about 5,000 ft. altitude
Lebia viridis Say, 1823. Bousquet, 2012: 1333
Phloeoxena Chaudoir, 1869
Phloeoxena montana Darlington, 1935: 208. Erwin & Sims, 1984: 446 *H > TL: Mt. La Hotte, on and near summit (probably
not below 7,000 ft.)
Phloeoxena (Phloeoxena) schwarzi Darlington, 1934. Shpeley & Ball, 2000: 103
Phloeoxena sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Stenognathus Chaudoir, 1843
Stenognathus sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Tribe LICININI
Badister Clairville, 1806
Badister reflexus LeConte, 1880. Peck, 2005: 33; Bousquet, 2012: 1010
Tribe MORIONINI
Morion Latreille, 1810
Morion monilicornis (Latreille, 1806) [By synonymy of species below]
=Morion georgiae Palisot de Beauvois, 1805. Blackwelder, 1944: 33
Tribe ODACANTHINI
Colliuris DeGeer, 1874
Colliuris caymanensis Darlington, 1947. Bousquet, 2010: 22; Bousquet, 2012: 1270
Colliuris portoricensis Liebke, 1930. Blackwelder, 1944: 66
Tribe OODINI
Oodinus Motschulsky, 1865
Oodinus pseudopiceus Bousquet, 1996: 462. Peck, 2005: 34; Bousquet, 2012: 954
Stenocrepis Chaudoir, 1857
Stenocrepis duodecimstriata (Chevrolat, 1835). Bousquet, 1996: 485; Peck, 2005: 34; Bousquet, 2012: 958
Stenocrepis olivacea (Bates, 1878). Bousquet, 1996: 489; Peck, 2005: 34
Stenocrepis palustris Darlington, 1935: 202; Bousquet, 1996: 483 *H, *DR > TL: Swamps north of Dessalines
Stenocrepis subdepressa Darlington, 1934: 101. Erwin & Sims, 1984: 438; Bousquet, 1996: 484 *H, *DR > TL: Grande
Rivière
Stenocrepis tibialis (Chevrolat, 1834). Bousquet, 1996: 493; Peck, 2005: 34; Bousquet, 2012: 959
Tribe PANAGAEINI
Hologaeus Ogueta, 1966
Hologaeus asuai (Ogueta, 1966: 8). Erwin & Sims, 1984: 437; Häckel & Farkac, 2012: 89 *DR > TL: Santo Domingo
Tribe PENTAGONICINI
Pentagonica Schmidt-Goebel, 1846
Pentagonica flavipes flavipes (LeConte, 1853). Bousquet, 2012: 1266
Pentagonica flavipes picipes Darlington, 1935: 211. R. Bell, 1985: 326
Pentagonica maculicornis Bates, 1883. R. Bell, 1985: 322
Pentagonica vittula Darlington, 1939: 100. R. Bell, 1985: 322 *DR > TL: Los Limones, s. of Santiago, 3000 ft.
Tribe PERIGONINI
Perigona Laporte, 1835
Perigona nigriceps (Dejean, 1831). Peck & Thomas, 1998: 23 [Introduced]
Perigona picea Darlington, 1934: 98
Tribe PLATYNINI
Agonum Bonelli, 1810
Agonum decorum (Say, 1823). Liebherr, 1986: 124; Peck, 2005: 35; Bousquet, 2012: 1226
Platynus Bonelli, 1810
Platynus aequinoctialis (Chaudoir, 1850). Darlington, 1935: 198; Liebherr, 1987: 354
Platynus agonellus (Darlington, 1935: 187). Erwin & Sims, 1984: 433; Liebherr, 1987: 345 *H > TL: La Visite and vicinity,
5,000 m (on Mt. Bourette), La Selle Range

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Platynus altifluminis (Darlington, 1935: 198). Darlington, 1935: 198; Erwin & Sims, 1984: 433; Liebherr, 1987: 340 *H > TL:
Vicinity of La Visite, main La Selle Range, about 6,000 ft.
Platynus amone (Darlington, 1935: 190). Liebherr, 1987: 335 *H > TL: Near Summit La Hotte, about 7,000 ft.
Platynus baorucensis Liebherr, 1992: 34 *DR > TL: Pedernales, La Abeja, 38 km NNW Cabo Rojo, 1250 m
Platynus biramosus (Darlington, 1939: 89). Erwin & Sims, 1984: 434; Liebherr, 1987: 327 *DR > TL: Loma Rucilla and
mountains to the north, 5,000–8,000 ft.
Platynus calathinus (Darlington, 1939: 92). Erwin & Sims, 1984: 434; Liebherr, 1987: 328 *DR > TL: Mountains north of
Loma Rucilla, Cordillera Central, about 7,000 ft. altitude
Platynus christophe (Darlington, 1935: 191). Erwin & Sims, 1984: 434; Liebherr, 1987: 329 *H > TL: Summit plateau of Mt.
Basil, northern Haiti, 4,700 ft.
Platynus contricticeps (Darlington, 1935: 194). Erwin & Sims, 1984: 434; Liebherr, 1987: 329 *H > TL: Summit Mt. La Hotte,
about 7,800 ft.
Platynus cychrinus (Darlington, 1935: 192). Erwin & Sims, 1984: 434; Liebherr, 1987: 330 *H > TL: Haiti
Platynus fractilinea (Darlington, 1934: 96). Erwin & Sims, 1984: 434; Liebherr, 1987: 342 *H > TL: Diquini
Platynus fractilateralis Liebherr, 1987: 340 *DR > TL: Baoruco Prov.
Platynus jaegeri (Dejean, 1831: 728). Darlington, 1935: 192; Erwin & Sims, 1984: 434; Liebherr, 1987: 331 *H, *DR > TL:
Saint-Domingue
Platynus laetificus (Darlington, 1935: 200). Erwin & Sims, 1984: 433; Liebherr, 1987: 346 *H > TL: Summit plateau Mt. Basil,
4,700 ft.
Platynus laeviceps (Darlington, 1939: 91). Erwin & Sims, 1984: 434; Liebherr, 1987: 335 *DR > TL: North slope of Loma
Rucilla, Cordillera Central, about 8,000 ft. altitude
Platynus marcus (Darlington, 1935: 180). Erwin & Sims, 1984: 434; Liebherr, 1987: 337 *H > TL: Mt. La Hotte, 5,000–7,800
ft.
Platynus metallosomus Liebherr, 1987: 346. Liebherr, 1987: 346 *DR > TL: Vic. Valle Nuevo, 6000’ elev.
Platynus pavens (Darlington, 1935: 188). Erwin & Sims, 1984: 434; Liebherr, 1987: 348 *H > TL: Haiti
Platynus ramoni (Darlington, 1939: 92). Erwin & Sims, 1984: 434; Liebherr, 1987: 338 *DR > TL: Valle Nuevo and vicinity,
Cordillera Central, 6,000–7,000 ft. altitude
Platynus scriptellus (Darlington, 1939: 94). Liebherr, 1987: 343 *DR > TL: Valle Nuevo, Cordillera Central, about 7,000 ft.
altitude
Platynus scriptus (Darlington, 1939: 93). Erwin & Sims, 1984: 434; Liebherr, 1987: 343 *DR > TL: Valle Nuevo, southeast of
Constanza, about 7,000 ft.
Platynus sellensis (Darlington, 1937b: 122). Liebherr, 1987: 350 *H > TL: La Visite and vicinity, Massif de la Selle, 5,000–
7,000 ft. altitude
Platynus subcordens (Darlington, 1935: 192). Erwin & Sims, 1984: 435; Liebherr, 1987: 333 *H > TL: La Visite and vicinity,
main La Selle Range, 5,000–7,000 ft.
Platynus tipoto (Darlington, 1935: 193). Erwin & Sims, 1984: 435; Liebherr, 1987: 334 *H > TL: Vicinity La Visite, main
range La Selle, about 6,500 ft.
Platynus transcibao (Darlington, 1939: 91). Liebherr, 1987: 334 *DR > TL: Summit dome of Mt. Diego de Ocampo, about
4,000 ft. altitude
=Colpodes biramosus transcibao Darlington, 1939: 91
Platynus visitor (Darlington, 1935: 195). Erwin & Sims, 1984: 435; Liebherr, 1987: 344 *H > TL: La Visite and vicinity, La
Selle Range, about 6,000 ft.
Platynus wolla (Darlington, 1935: 189). Erwin & Sims, 1984: 435; Liebherr, 1987: 338 *H > TL: Ridge just above Roche
Croix, on a spur of Mt. La Hotte, 5,000 ft.
Tribe PTEROSTICHINI
Barylaus Liebherr, 1986
Barylaus puncticeps (Darlington, 1939: 94) *DR > TL: Loma Rucilla and mountains to the north, Cordillera Central, 6,000–
8,000 ft.
Colpodes MacLeay, 1825
Colpodes cuprascens (Motschulsky, 1864: 305). Darlington, 1935: 182 *H, *DR > TL: St. Domingue
Colpodes mannerheimi Chaudoir, 1859: 360. Chaudoir, 1878: 339; Leng & Mutchler, 1914: 394 *H, *DR > TL: Haiti [Nomen
Dubium]
Euchroa Brullé, 1834
Euchroa cupripennis (Chaudoir, 1874). Erwin & Sims, 1984: 435 Frania & Ball, 2007: 31 *H > TL: Haiti

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 =Dyschromus cupripennis (Chaudoir, 1874). Perez-Gelabert, 2008: 82
Euchroa centralis (Darlington, 1939: 88). Erwin & Sims, 1984: 435 *DR > TL: Constanza, at foot of Loma Vieja, about 3,000
ft. altitude
Euchroa independencia Frania & Ball, 2007: 29 *DR > TL: Independencia, Sierra de Neiba near crest, 5.5 km NNW Angel
Feliz, 1750 m
Euchroa opaca (Chaudoir, 1835). Frania & Ball, 2007: 25; Perez-Gelabert, 2008: 82
Euchroa pedernales Frania & Ball, 2007: 33 *DR > TL: Pedernales, 1 km S Los Arroyos, 1125 m
Euchroa perezi (Darlington, 1939: 88). Erwin & Sims, 1984: 435 *DR > TL: Mt. Diego de Ocampo, about 3,500 ft. altitude
Euchroa tiburonica (Darlington, 1935: 179). Erwin & Sims, 1984: 434; Frania & Ball, 207: 24 *H > TL: Northeastern foothills
La Hotte, 2,000–4,000 ft.
Loxandrus LeConte, 1852
Loxandrus cubanus Tschitschérine, 1901. Allen, 1972: 45; Erwin & Sims, 1984: 437
Loxandrus infimus Bates, 1882. Erwin & Sims, 1984: 436; Bousquet, 2012: 763
=Loxandrus mutans Darlington, 1935: 180.
Loxandrus rectangulus LeConte, 1878. Peck, 2005: 31
Tribe ZUPHIINI
Thalpius LeConte, 1851
Thalpius marginicollis (Darlington, 1934: 212). Erwin & Sims, 1984: 441
Zuphium Latreille, 1806
Zuphium haitianum Darlington, 1935: 213 *H > TL: Etang Lachaux
Subfamily OMOPHRONINAE
Omophron Latreille, 1802
Omophron dominicense Chaudoir, 1868: 58. Blackwelder, 1944: 22; Erwin, 2007: 65 *H, *DR > TL: Saint-Domingue
Subfamily PAUSSINAE
Eohomopterus Wasmann, 1920
Eohomopterus paulmuelleri† Nagel, 1987: 140; Carvalho, 1987: 995 [Fossil in Dominican amber]
Eohomopterus poinari† Nagel, 1997: 350 [Fossil in Dominican amber]
Homopterus Westwood, 1841
Homopterus (Homopterus) hispaniolensis† Nagel, 1987: 147 [Fossil in Dominican amber]
Protopaussus Gestro, 1892
Protopaussus pristinus† Nagel, 1997: 347 [Fossil in Dominican amber]
Subfamily SCARITINAE
Tribe CLIVININI
Ardistomis Putzeys, 1846
Ardistomis alticolus Darlington, 1935: 173. Erwin & Sims, 1984: 427; Nichols, 1988b: 119 *H > TL: Mt. Bourette, La Selle
massif, 5,000 ft.
Ardistomis hispaniolensis Nichols, 1988a. Nichols, 1988b: 120
Ardistomis nigroclarus Darlington, 1939: 83. Erwin & Sims, 1984: 427; Nichols, 1988b: 120 *DR > TL: Valle Nuevo, SE of
Constanza, about 6,000 ft. altitude
Ardistomis rufoclarus Darlington, 1939: 82. Erwin & Sims, 1984: 427; Nichols, 1988b: 120 *DR > TL: Mountains to the north
of Loma Rucilla, about 6,000 ft. altitude
Aspidoglossa Putzeys, 1846
Aspidoglossa schach (Fabricius, 1792). Nichols, 1988b: 119
Clivina Latreille, 1802
Clivina (Isoclivina) limbipennis Jacquelin du Val, 1857. Peck, 2005: 28
Dyschirius Bonelli, 1813
Dyschirius larochellei Bousquet, 1988. Bousquet, 2012: 445
=Dyschirius (Dyschirius) nr. erythrocerus LeConte, 1857. Nichols, 1988b: 116
=Dyschiriodes larochellei (Bousquet, 1988). Peck, 2005: 28
Halocoryza Alluaud, 1919
Halocoryza arenaria (Darlington, 1939: 84). Erwin & Sims, 1984: 427; Bousquet, 2012: 426
Nyctosyles Putzeys, 1866
Nyctosyles planicollis (Reiche, 1842). Nichols, 1988b: 118
Oxydrepanus Putzeys, 1866

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Oxydrepanus micans Putzeys, 1866. Nichols, 1988b: 119
Oxydrepanus reicheoides Darlington, 1939: 83. Erwin & Sims, 1984: 427 *DR > TL: Loma Rucilla, Cordillera Central, about
6,000 ft. altitude
Oxydrepanus rufus (Putzeys, 1846). Bousquet, 2012: 427
Pachyteles Perty, 1830
Pachyteles sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Paraclivina Kult, 1947
Paraclivina bipustulata (Fabricius, 1798). Nichols, 1988b: 118; Peck, 2005: 28; Bousquet, 2012: 413
Paraclivina fasciata (Putzeys, 1846). Nichols, 1988b: 118; Peck & Thomas, 1998: 17; Bousquet, 2012: 414
Paraclivina marginipennis (Putzeys, 1846). Bousquet, 2012: 415
Schizogenius Putzeys, 1846
Schizogenius (Schizogenius) arimao Darlington, 1934. Nichols, 1988b: 117; Peck, 2005: 29
Semiardistomis Kult, 1850
Semiardistomis cyaneolimbatus (Chevrolat, 1863). Nichols, 1988b: 118; Peck, 2005: 28
Semiclivina Kult, 1947
Semiclivina addita (Darlington, 1934). Nichols, 1988b: 118
Tribe SCARITINI
Antilliscaris Bänninger, 1949
Antilliscaris darlingtoni (Banninger, 1935: 159). Erwin & Sims, 1984: 426 *H > TL: Mt. Basil, 4700 ft.
Scarites Fabricius, 1801
Scarites (Distichus) orientalis Bonnelli, 1813. Nichols, 1985: 131
Subfamily TRECHINAE
Tribe BEMBIDIINI
Bembidion Latreille, 1802
Bembidion (Notaphus) fastidiosum Jacquelin du Val, 1852. Darlington, 1934: 77
Bembidion (Notaphus) sparsum Bates, 1882. Erwin, 1984: 182
Bembidion rucillum Darlington, 1939: 85. Blackwelder, 1944: 29 *DR > TL: North slope of Loma Rucilla, Cordillera Central,
about 8,000 ft. altitude
Bembidion sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Bembidion spretum Dejean, 1831. Erwin & Sims, 1984: 432
Elaphropus Motschulsky, 1839
Elaphropus tritax (Darlington, 1935: 175). Erwin & Sims, 1984: 429 *H > TL: Camp Perrin
Elaphropus yunax (Darlington, 1939: 87). Erwin & Sims, 1984: 429
Mioptachys Bates, 1882
Mioptachys insularis (Darlington, 1939: 86) *DR > TL: Loma Vieja, south of Constanza, about 6,000 ft. altitude
Mioptachys noctis (Darlington, 1935: 174). Erwin & Sims, 1984: 428 *H > TL: Roche Croix, Massif de la Hotte, 5,000 ft.
Paratachys Casey, 1918
Paratachys abruptus (Darlington, 1934). Peck, 2005: 30
Paratachys carib (Darlington, 1935: 176). Erwin & Sims, 1984: 430
Pericompsus LeConte, 1851
Pericompsus philippi Erwin, 1974: 62. Erwin & Sims, 1984: 430
Tachys Stephens, 1828
Tachys bradycellinus Hayward, 1900. Erwin & Sims, 1984: 430; Bousquet, 2012: 685
Tachys vittiger LeConte, 1852. Bousquet, 2012: 689
Tachyta Kirby, 1837
Tachyta hispaniolae (Darlington, 1934: 77). Erwin, 1975: 31; Erwin & Sims, 1984: 428
Tribe POGONINI
Diplochaetus Chaudoir, 1871
Diplochaetus rutilus (Chevrolat, 1863). Bousquet, 1997: 708; Peck, 2005: 31; Bousquet, 2012: 713
Tribe TRECHINI
Perileptus Schaum, 1860
Perileptus dentifer Darlington, 1935: 177. Erwin & Sims, 1984: 429
Perileptus minutus Darlington, 1935: 178. Erwin & Sims, 1984: 429
Subfamily PSEUDOMORPHINAE

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Tribe PSEUDOMORPHINI
Pseudomorpha Kirby, 1825
Pseudomorpha caribbeana Darlington, 1935: 214. Erwin & Sims, 1984: 425 *H > TL: Port-au-Prince

Family RHYSODIDAE
Clinidium Kirby, 1830
Clinidium (Clinidium) corbis R. Bell, 1970: 322 *H > TL: Tardieu, Morne La Hotte, 3000 ft.
Clinidium (Clinidium) haitiense R. Bell, 1970: 322. Erwin & Sims, 1984: 428 *H > TL: La Visite, La Selle Range, 5000–7000
ft.
Clinidium (Clinidium) trionyx R. Bell & J. Bell, 1985: 124 *DR > TL: Cazabita [Loma Casabito], 1250 m
Clinidium (Clinidium) xenopodium R. Bell, 1970: 316. Erwin & Sims, 1984: 428 *DR > TL: Loma Vieja near Constanza, 6,000
ft.
Clinidium (Clinidium) poinari† R. Bell & J. Bell, 2009: 75 [Fossil in Dominican amber]
Clinidium (Tainoa) grimaldii† R. Bell & J. Bell, 2009: 57 [Fossil in Dominican amber]
Plesioglymmius R. Bell & J. Bell, 1978
Plesioglymmius (Ameroglymmius) zayasi† R. Bell & J. Bell, 2009: 48 [Fossil in Dominican amber]

Suborder POLYPHAGA
Infraorder STAPHYLINIFORMIA
Superfamily HYDROPHILOIDEA
Family HYDROCHIDAE
Hydrochus Leach, 1817
Hydrochus pallipes Chevrolat, 1863. Peck, 2005: 48

Family HYDROPHILIDAE
Subfamily HYDROPHILINAE
Anacaena Thomson, 1859
Anacaena paleodominica† Fikácek & Engel, 2012: 622 [Fossil in Dominican amber]
=Anacaena sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Anacaena suturalis (LeConte, 1866). Komarek, 2005: 280; Perez-Gelabert, 2012: 115
Berosus Leach, 1817
Berosus interstitialis Knisch, 1924. Deler-Hernández et al., 2013: 83
=Berosus stribalus Orchymont, 1946. Peck, 2005: 48
Berosus trilobus Chevrolat, 1863. Deler-Hernández et al., 2013: 95
Derallus Sharp, 1882
Derallus altus (LeConte, 1855). Peck, 2005: 48
Enochrus Thomson, 1859
Enochrus (Methydrus) aequalis (Sharp, 1882). Short, 2004a: 352
Enochrus (Methydrus) bartletti Short, 2004a: 352
Enochrus (Methydrus) debilis (Sharp, 1882). Hansen, 1999: 180; Perez-Gelabert, 2012: 115
Enochrus (Methydrus) ochraceus (Melsheimer, 1844). Hansen, 1999: 185; Short, 2004a: 354
Enochrus (Methydrus) pseudochraceus Gundersen, 1977. Hansen, 1999: 186; Short, 2004a: 355
Enochrus (Methydrus) punctipalpus Short, 2004a: 355
Enochrus (Methydrus) pygmaeus (Fabricius, 1792). Short, 2004a: 356
Hydrobiomorpha Blackburn, 1888
Hydrobiomorpha (Hydrobiomorpha) phallica (Orchymont, 1928: 165). Hansen, 1999: 216; Short, 2004b: 365
Hydrophilus Geoffroy, 1862
Hydrophilus (Hydrophilus) ensifer duvali Hansen, 1999: 229
=Hydrophilus (Hydrophilus) ater Olivier, 1792. Blackwelder, 1944: 171
Hydrophilus (Hydrophilus) insularis Laporte, 1840. Blackwelder, 1944: 171; Hansen, 1999: 230
Laccobius Erichson, 1837
Laccobius (Microlaccobius) antillensis Spangler, 1968. Hansen, 1999: 147; Peck, 2005: 49
Tropisternus Solier, 1824
Tropisternus (Strepitornus) collaris (Fabricius, 1775). Blackwelder, 1944: 170; Hansen, 1999: 225

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Tropisternus (Tropisternus) lateralis (Fabricius, 1775). García Avila & Peña, 1993: 214
Subfamily SPHAERIDIINAE
Cercyon Leach, 1817
Cercyon armatipennis Arriaga-Varela et al., 2017: 51 *DR > TL: 11.3 km NW La Descubierta, 1650 m
Cercyon gimmeli Arriaga-Varela et al., 2017: 47 *DR > TL: Samaná, Salto El Limón
Cercyon insularis Chevrolat, 1863. Arriaga-Varela et al., 2017: 67
Cercyon nigriceps (Marsham, 1802). Fikácek, 2009: 354. Arriaga-Varela et al., 2017: 64
Cercyon praetextatus (Say, 1825). Peck, 2005: 51. Arriaga-Varela et al., 2017: 58
Cercyon spiniventris Arriaga-Varela et al., 2017: 60 *DR > TL: Monseñor Nouel Prov., Parque Nacional La Humeadora, 636 m
Cercyon taino Arriaga-Varela et al., 2017: 52 *DR > TL: Samaná Prov., dam 2.5 km N of Samaná, 58 m
Dactylosternum Wollaston, 1854
Dactylosternum abdominale (Fabricius, 1792). Peck, 2005: 51
Dactylosternum flavicorne (Mulsant, 1844). Blackwelder, 1944: 174; Hansen, 1999: 257
Oosternum Sharp, 1882
Oosternum bacharenge Deler-Hernández et al., 2014: 48 *DR > TL: Hato Mayor Prov., Parque Nacional Los Haitises, 250 m
Oosternum pecki Deler-Hernández et al., 2014: 54 *DR > TL: Barahona, 7 km NW Paraiso, 200 m
Oosternum sharpi Hansen, 1999. Deler-Hernández et al., 2014: 57
Phaenonotum Sharp, 1882
Phaenonotum exstriatum (Say, 1835). Deler-Hernández et al., 2018: 111
Phaenonotum laevicolle Sharp, 1882. Deler-Hernández et al., 2018: 113
Phaenonotum laterale Deler-Hernández et al., 2018: 113 *DR > TL: Barahona, Monumento Natural Miguel Domingo Fuerte
‘Cachote’, 1188 m

Family HISTERIDAE
Subfamily ABRAEINAE
Aeletes Horn, 1873
Aeletes gulliver (Marseul, 1856). Chevrolat, 1864: 598; Blackwelder, 1944: 178; Mazur, 1997: 204
Subfamily DENDROPHILINAE
Carcinops Marseul, 1855
Carcinops troglodytes (Paykull, 1811). Chevrolat, 1864: 594; Blackwelder, 1944: 180 [Introduced]
Carcinops dominicana Marseul, 1855. Blackwelder, 1944: 180; Mazur, 1997: 176
Subfamily HISTERINAE
Atholus Thomson, 1859
Atholus confinis (Erichson, 1834). Peck, 2010: 12
Baconia Lewis, 1885
Baconia riehli (Marseul, 1862). Caterino & Tischechkin, 2013: 60
Hister Linnaeus, 1758
Hister coenosus Erichson, 1834. Chevrolat, 1864: 593; Blackwelder, 1944: 183; Caterino, 1999b: 59
Hister servus Erichson, 1834. Chevrolat, 1864: 593; Blackwelder, 1944: 184
Hololepta Paykull, 1811
Hololepta (Hololepta) cubensis Erichson, 1834. Chevrolat, 1864: 590; Blackwelder, 1944: 176; Mazur, 1997: 55
Hololepta (Hololepta) palisoti Mazur, 1972. Mazur, 1997: 56 *DR > TL: San Domingo
=Hister complanatus Palisot de Beauvois, 1805. Mazur, 1997: 56
Hololepta (Leionota) quadridentata (Olivier, 1789). Wolcott, 1922: 36; Blackwelder, 1944: 176
Omalodes Erichson, 1834
Omalodes (Omalodes) angelo Moura et al., 2016 *DR > TL: Monte Cristi Prov., 5.3 km N Villa Elisa
Omalodes (Omalodes) haitianus Marseul, 1853: 525. Blackwelder, 1944: 182; Mazur, 1997: 87 *H > TL: Haïti (S.-Domingo)
Omalodes (Omalodes) kovariki Moura et al., 2016: 210 *DR > TL: Pr. San Cristóbal; 4 km NW Villa Altagracia, 300 m
Omalodes (Omalodes) laevigatus (Quensel, 1806). Blackwelder, 1944: 182; Perez-Gelabert, 2012: 115
Omalodes soulouquii Marseul, 1861. Blackwelder, 1944: 183; Mazur, 1997: 88
Plaesius Erichson, 1834
Plaesius (Plaesius) javanus Erichson, 1834. Marcano, 1983: 6/83; Mazur, 1997: 84 [Introduced]
Platylister Lewis, 1892
Platylister (Platylister) urvillei (Le Guillou, 1844). Mazur, 1997: 66

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Subfamily TRIBALINAE
Epierus Erichson, 1834
Epierus (Epierus) antillarum Marseul, 1854. Chevrolat, 1864: 594; Blackwelder, 1944: 181; Mazur, 1997: 12
Epierus (Epierus) waterhousei Marseul, 1854. Blackwelder, 1944: 182; Mazur, 1997: 14
Idolia Lewis, 1885
Idolia laevissima (J. L. LeConte, 1852). Chevrolat, 1864: 595; Blackwelder, 1944: 182
Subfamily TRYPANAEINAE
Trypanaeus Eschscholtz, 1829
Trypanaeus hispaniolus† Chatzimanolis et al., 2006: 334 [Fossil in Dominican amber]

Superfamily STAPHYLINOIDEA
Family HYDRAENIDAE
Subfamily OCHTHEBIINAE
Gymnochthebius Orchymont, 1943
Gymnochthebius fossatus (LeConte, 1855). Peck, 2005: 55
Hydraena Kugelann, 1794
Hydraena haitensis Perkins, 1980: 171 *H > TL: Etang Lachaux, SW península, under 1000 ft.
Ochthebius Leach, 1815
Ochthebius (Ochthebius) attritus LeConte, 1878. Perkins, 1980: 516

Family LEIODIDAE
Subfamily LEIODINAE
Tribe AGATHIDIINI
Agathidium Panzer, 1797
Agathidium minutum Peck & Cook, 2014: 13 *DR > TL: Prov. Hato Mayor, Par. Nac. Los Haitises, W. of Sabana de la Mar
Tribe LEIODINI
Isoplastus Horn, 1880
Isoplastus hispaniolensis Peck & Cook, 2014: 14 *DR > TL: La Vega Prov., 10 km NE Jarabacoa, Hotel Montaña, 550 m
Parvocyrtusa Peck & Cook, 2014
Parvocyrtusa hispaniolensis Peck & Cook, 2014: 38 *DR > TL: La Vega Prov., PN A. Bermúdez, Cienaga, 1100 m
Pseudolionothus Peck & Cook, 2014
Pseudolionothus insularis Peck & Cook, 2014: 40 *DR > TL: La Vega Prov., PN A. Bermúdez, Ciénaga, 1010 m
Zeadolopus Broun, 1903
Zeadolopus acinaces Peck & Cook, 2014: 21 *DR > TL: Prov. Pedernales, Las Abejas, 1225 m
Zeadolopus caborojo Peck & Cook, 2014: 24 *DR > TL: Pedernales, 28 km N Cabo Rojo, 760 m
Zeadolopus exiguus Peck & Cook, 2014: 27 *DR > TL: Pedernales, 28 km N Cabo Rojo, 760 m
Zeadolopus hatomayor Peck & Cook, 2014: 28 *DR > TL: Prov. Hato Mayor, Par. Nac. Los Haitises, W. of Sabana de la Mar
Zeadolopus hispaniolensis Peck & Cook, 2014: 29 *DR > TL: La Vega Prov., PN. A. Bermúdez, Cienaga, 1100 m
Zeadolopus ivei Peck & Cook, 2014: 30 *DR > TL: Prov. San Cristóbal, 4 km NW Villa Altagracia, 300 m
Zeadolopus jarabacoa Peck & Cook, 2014: 30 *DR > TL: La Vega Prov., 10 km NE Jarabacoa, Hotel Montaña, 550 m
Zeadolopus lavega Peck & Cook, 2014: 31 *DR > TL: La Vega Prov., 10 km NE Jarabacoa, Racquet Club, 550 m
Zeadolopus longipes Peck & Cook, 2014: 31 *DR > TL: La Vega Prov., PN. A. Bermúdez, Cienaga, 1020 m
Zeadolopus minisculus Peck & Cook, 2014: 32 *DR > TL: Barahona, 7 km NW Paraiso, 200 m
Zeadolopus nanus Peck & Cook, 2014: 33 *DR > TL: San Cristóbal Prov., Cuevas Borbon, Pomier, 200 m
Zeadolopus oviedoensis Peck & Cook, 2014: 34 *DR > TL: Pedernales, 4 km W Oviedo, 10 m
Zeadolopus paulus Peck & Cook, 2014: 36 *DR > TL: Pedernales, 28 km N Cabo Rojo, 760 m
Zeadolopus pedernales Peck & Cook, 2014: 37 *DR > TL: Prov. Barahona, nr. Filipinas, Larimar mine
Tribe SCOTOCRYPTINI
Aglyptinus Cockerell, 1906
Aglyptinus angulatus Peck & Cook, 2014: 46 *DR > TL: San Cristóbal Prov., Cuevas Borbon, Pomier, 200 m
Aglyptinus grandis Peck & Cook, 2014: 50 *DR > TL: La Vega Prov., PN A. Bermúdez, 4 km SW Cienaga, 1200 m
Aglyptinus hispaniolensis Peck & Cook, 2014: 51 *DR > TL: La Vega Prov., PN. A. Bermúdez, Cienaga, 1100 m
Aglyptinus longipalpus Peck & Cook, 2014: 52 *DR > TL: Independencia, 32 km NW La Descubierta, Sabana Real, 1800 m
Aglyptinus poinari† Perkovsky, 2000: S331 [Fossil in Dominican amber]

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Creagrophorus Matthews, 1887
Creagrophorus hispaniolensis Peck & Cook, 2014: 59 *DR > TL: Prov. Pedernales, Las Abejas, 1225 m
Creagrophorus microdenticulus Peck & Cook, 2014: 60 *DR > TL: La Vega Prov., PN. A. Bermúdez, Ciénaga, 1000 m
Subfamily CHOLEVINAE
Tribe EUCATOPINI
Eucatops Portevin, 1903
Eucatops (Eucatops) annulus Peck & Cook, 2014: 5 *DR > TL: Santiago Prov., PN A. Bermúdez, 9 km SW Ciénaga, Redondo,
2000 m
Tribe PTOMAPHAGINI
Proptomaphaginus Szymczakowski, 1969
Proptomaphaginus hispaniolensis Peck, 1983: 255. Peck et al., 1998: 63; Peck, 1999: 610; Peck & Cook, 2014: 7 *DR, *H >
TL: Cave at mouth of Río Chavón, La Romana

Family SILPHIDAE
Subfamily NICROPHORINAE
Nicrophorus Fabricius, 1775
Nicrophorus hispaniola Sikes & Peck, 2000: 391. Perez-Gelabert, 2016: 92 *DR > TL: Pedernales, 8 km NE Los Arroyos,
1,940 m

Family STAPHYLINIDAE
Subfamily ALEOCHARINAE
Aleochara Gravenhorst, 1802
Aleochara bimaculata Gravenhorst, 1802. Blackwelder, 1943: 559
Alisalia Casey, 1911
Alisalia brunnea Cameron, 1922: 126. Blackwelder, 1943: 540
Alisalia picea Cameron, 1922. Blackwelder, 1943: 540 *H > TL: Haiti
Antillusa Pace, 2012
Antillusa dominicana Pace, 2012: 60 *DR > TL: Peravia, 36 km NW San José de Ocoa
Atheta Thomas, 1859
Atheta alternata (Erichson, 1839). Blackwelder, 1943: 550
Atheta anomala Cameron, 1923: 382. Blackwelder, 1943: 550
Atheta cognata Sharp, 1883. Blackwelder, 1943: 551
Atheta plurispinosa Cameron, 1923. Blackwelder, 1943: 554
Atheta suboblita Cameron, 1923: 377. Blackwelder, 1943: 554 *H > TL: Haiti
Atheta tentativa Cameron, 1923. Blackwelder, 1943: 555
Barychara Sharp, 1883
Barychara flavipennis Cameron, 1922: 125. Blackwelder, 1943: 539 *H > TL: Haiti
Coenonica Kraatz, 1857
Coenonica puncticollis Kraatz, 1857. Frank & Thomas, 1984: 416
Dalotia Casey, 1910
Dalotia coriaria (Kraatz, 1856). Pace, 2012: 58
Dalotia pectorina (Casey, 1910). Asenjo et al., 2019: 100
Diestota Mulsant & Rey, 1870
Diestota sperata Sharp, 1876. Blackwelder, 1943: 544
Falagria Samouelle, 1819
Falagria (Leptagria) dominicana Pace, 2012: 55 *DR > TL: Peravia, 15 km E San José de Ocoa
Gyrophaena Mannerheim, 1830
Gyrophaena puncta Cameron, 1922: 647. Blackwelder, 1943: 543 *H > TL: Haiti
Hoplandria Kraatz, 1857
Hoplandria sp.—Blackwelder (1943: 557)
Myllaena Erichson, 1837
Myllaena obscura Cameron, 1922: 651. Blackwelder, 1943: 535 *H > TL: Haiti
Myllaena potawatomi Klimaszewski, 1982: 192
Oligota Mannerheim, 1830

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Oligota haitiana Frank, 1972: 133 *H > TL: Haiti
Oligota maculicornis Cameron, 1922. Blackwelder, 1943: 536; Frank, 1972: 142
Oligota sp.—Blackwelder (1943: 538)
Phanerota Casey, 1906
Phanerota atomaria (Cameron, 1922). Blackwelder, 1943: 540; Peck, 2010: 16
Phloeopora Erichson, 1837
Phloeopora occidentalis Cameron, 1923. Blackwelder, 1943: 557
Phloeopora producta Cameron, 1923: 388. Blackwelder, 1943: 558 *H > TL: Haiti
Pseudothamiaraea Cameron, 1923
Pseudothamiaraea brunnea Cameron, 1923: 364. Blackwelder, 1943: 555
Subfamily OMALIINAE
Omalium Gravenhorst, 1802
Omalium laborium Blackwelder, 1943: 56 *H, *DR > TL: East slope of Morne La Selle
Phloeonomus Heer, 1839
Phloeonomus pedicularius (Erichson, 1840). Blackwelder, 1943: 52
Subfamily OSORIINAE
Antillosorius Irmler, 2010
Antillosorius biarmatus (Darlington, 1937: 287) *H > TL: Northeastern foothills of the Massif de la Hotte
Antillosorius buscki (Notman, 1925: 19). Darlington, 1937: 297; Blackwelder, 1943: 186 *DR, *H > TL: Santo Domingo
Antillosorius cautis (Darlington, 1937: 295). Blackwelder, 1943: 184 *H > TL: Summit of ridge just above Roche Croix, on a
northern spur of Morne La Hotte
Antillosorius daemon (Darlington, 1937: 293). Blackwelder, 1943: 180 *H > TL: Northeastern foothills of the Massif de la
Hotte
Antillosorius fratellus (Darlington, 1937: 291). Blackwelder, 1943: 178 *H > TL: Furcy, La Selle Range
Antillosorius hirtilabris (Darlington, 1937: 292). Blackwelder, 1943: 180 *H > TL: Northeastern foothills of the Massif de la
Hotte
Antillosorius mediocris (Darlington, 1937: 296). Blackwelder, 1943: 185 *H > TL: La Visite and vicinity, La Selle Range
Antillosorius regulus (Darlington, 1937: 288). Blackwelder, 1943: 176 *H > TL: Morne La Hotte
Antillosorius salutator (Darlington, 1937: 290). Blackwelder, 1943: 179 *H > TL: La Visite and vicinity, La Selle Range
Clavilispinus Bernhauer, 1926
Clavilispinus exiguus (Erichson, 1840). Blackwelder, 1943: 158; Irmler, 1991: 86
Espeson Schaufuss, 1882
Espeson moratus Schaufuss, 1882. Scheerpeltz, 1970: 119; Herman, 2001: 1240
Liberiana Blackwelder, 1942
Liberiana sp.†—Irmler (2003c: 5) [Fossil in Dominican amber]
Lispinomimus Irmler, 2003
Lispinomimus atavus† Irmler, 2003c: 11 [Fossil in Dominican amber]
Lispinus Erichson, 1839
Lispinus anguinus Fauvel, 1864. Blackwelder, 1943: 133
Lispinus cordobensis Bernhauer, 1929
Lispinus insularis Fauvel, 1863. Blackwelder, 1943: 136
Mimogonus Fauvel, 1903
Mimogonus fumator (Fauvel, 1889). Blackwelder, 1943: 172; Peck, 2005: 67
=Osorius fauveli Cameron, 1923
Molosoma Say, 1831
Molosoma sp.—Asenjo et al. (2019: 211)
Nacaeus Blackwelder, 1943
Nacaeus dominicanensis† Irmler, 2003c: 4 [Fossil in Dominican amber]
Nacaeus impressicollis (Motschulsky, 1858). [By synonymy of species below]
=Nacaeus irregularis (Blackwelder, 1943: 124)
Nacaeus nigrifrons (Chevrolat & Fauvel, 1863). Blackwelder, 1943: 128
Neosorius Fagel, 1959
Neosorius sp. 1†—Irmler (2003c: 8) [Fossil in Dominican amber]
Neosorius sp. 2†—Irmler (2003c: 9) [Fossil in Dominican amber]

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Osoriellus Fagel, 1959
Osoriellus sp.†—Irmler (2003c: 7) [Fossil in Dominican amber]
Osoriellus eggersi (Bernhauer, 1904). Darlington, 1937: 298; Blackwelder, 1943: 196; Irmler, 2014: 266
=Osorius laeviceps Notman, 1925. Blackwelder, 1943: 195; Irmler, 2014: 266
Osoriellus haitiellus (Darlington, 1937: 299). Blackwelder, 1943: 191; Irmler, 2014: 270
Tannea Blackwelder, 1952
Tannea reversa (Blackwelder, 1943: 122). Irmler, 2003b: 97
Tannea tenella (Erichson, 1840). Blackwelder, 1943: 126
=Nacaeus impar (Cameron, 1913: 122)
Thoracophorus Motschulsky, 1837
Thoracophorus guadalupensis Cameron, 1913. Blackwelder, 1943: 151
Thoracophorus palaeobrevicristatus† Irmler, 2003c: 2 [Fossil in Dominican amber]
Subfamily MICROPEPLINAE
Peplomicrus Berhauer, 1928
Peplomicrus iviei Campbell, 1991: 37 *DR > TL: Las Abejas, 35 km N Cabo Rojo, Pedernales Prov.
Subfamily OXYTELINAE
Anotylus Thomson, 1859
Anotylus glareosus (Wollaston, 1854). Hammond, 1975: 163. Herman, 2001: 1355 [Introduced]
Anotylus insignitus (Gravenhorst, 1806). Blackwelder, 1943: 92
Anotylus scorpio (Fauvel, 1904). Blackwelder, 1943: 99
Bledius Leach, 1819
Bledius caribbeanus Blackwelder, 1943. Blackwelder, 1943: 113; Frank & Ahn, 2011: 31
Bledius ceratus Blackwelder, 1943: 118; Frank & Ahn, 2011: 29
Carpelimus Samouelle, 1819
Carpelimus aridus (Jacquelin du Val, 1857). Blackwelder, 1943: 73
Carpelimus beattyi Blackwelder, 1943: 65
Carpelimus correctus Blackwelder, 1943: 64
Carpelimus flavipes (Erichson, 1840). Blackwelder, 1943: 77
Carpelimus fulvipes (Erichson, 1840). Blackwelder, 1943: 62
Carpelimus rulomus Blackwelder, 1943: 81 *DR > TL: Barahona
Carpelimus testaceipennis (Cameron, 1923). Blackwelder, 1943: 84 *H > TL: Haiti
Carpelimus varicornis (Bernhauer, 1904). Blackwelder, 1943: 73
Dolichoxenus† Engel & Chatzimanolis, 2009
Dolichoxenus newtoni† Engel & Chatzimanolis, 2009: 427 [Fossil in Dominican amber]
Oxytelus Gravenhorst, 1802
Oxytelus incisus Motschulsky, 1857. Blackwelder, 1943: 96 [Introduced]
Oxytelus sculptus Gravenhorst, 1806. Frank & Thomas, 1981: 402
Platystethus Mannerheim, 1830
Platystethus spiculus Erichson, 1840. Blackwelder, 1943: 110
Thinobius Kiesenwetter, 1884
Thinobius exasperatus Blackwelder, 1943: 108
Thinobius opaculus Cameron, 1923: 396. Blackwelder, 1943: 106
Thinobius ornatus Cameron, 1923: 397. Blackwelder, 1943: 105
Thinodromus Kraatz, 1858
Thinodromus croceipes (Fauvel, 1868). Blackwelder, 1943: 61
Thinodromus darlingtoni (Blackwelder, 1943: 65). Makranczy, 2018: 78 *DR > TL: San José de las Matas
Thinodromus hispaniolus (Blackwelder, 1943: 68) *H > TL: Northern foothills of Morne La Hotte
Thinodromus obsolescens (Blackwelder, 1943: 69) *H > TL: Ennery
Thinodromus smithi (Bernhauer, 1908). Blackwelder, 1943: 67
Subfamily PAEDERINAE
Achenomorphus Motschulsky, 1858
Achenomorphus suturalis (Blackwelder, 1943: 254)
Astenus Stephens, 1832
Astenus cinctiventris (Sharp, 1886). Blackwelder, 1943: 366

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 113
Biocrypta Casey, 1905
Biocrypta densa Blackwelder, 1943: 341 *H > TL: La Visite and vicinity in the La Selle Range
Biocrypta dominicana Blackwelder, 1943: 339 *DR > TL: Sánchez
Biocrypta haitia Blackwelder, 1943: 339 *H > TL: Morne La Hotte
Biocrypta hispaniola Blackwelder, 1943: 343 *DR > TL: Loma Rucilla and vicinity
Cylindroxystus Bierig, 1943
Cylindroxystus pluviosum (Blackwelder, 1943: 320) *DR > TL: Vicinity of Valle Nuevo
Echiaster Erichson, 1840
Echiaster microps Blackwelder, 1943. Scheerpeltz, 1969: 100; Perez-Gelabert, 2012: 115
Echiaster waterhousei Cameron, 1913. Blackwelder, 1943: 371; Scheerpeltz, 1969: 99
Homoeotarsus Hochhuth, 1851
Homoeotarsus albipes (Erichson, 1840). Blackwelder, 1943: 328
Homoeotarsus solus Blackwelder, 1943: 327
Homoeotarsus testaceipes (Bierig, 1935). Blackwelder, 1943: 330
Lathrobium Gravenhorst, 1802
Lathrobium dominicanum Blackwelder, 1943: 310 *DR > TL: Loma de la Peña, northwest of Constanza
Lathrobium rubidum (Fauvel, 1863). Blackwelder, 1943: 314
Lithocharis Boisduval & Lacordaire, 1835
Lithocharis dorsalis Erichson, 1840. Blackwelder, 1943: 247
Lithocharis limbata Erichson, 1840. Blackwelder, 1943: 246
Lithocharis secunda Blackwelder, 1943: 244
Lobrathium Mulsant & Rey, 1878
Lobrathium discolor (Erichson, 1840). Blackwelder, 1943: 313
Lobrathium reductum Blackwelder, 1943: 319 *DR > TL: Valle Nuevo, southeast of Constanza
Medon Stephens, 1832
Medon dominicanum Blackwelder, 1943: 272 *DR, *H > TL: Sánchez
Medon matasana Blackwelder, 1943: 272 *DR > TL: San José de las Matas
Oedichirus Erichson, 1839
Oedichirus dominicanensis Herman, 2013: 73 *DR > TL: Prov. Pedernales, Sierra de Bahoruco, N. Cabo Rojo, km 28, 1200 m
Orus Casey, 1884
Orus cameroni Blackwelder, 1943: 278
Paederus Fabricius, 1775
Paederus morio Mannerheim, 1830: 39. Blackwelder, 1943: 324 *H > TL: Insula St. Domingo
Palaminus Erichson, 1840
Palaminus coriaceus Cameron, 1913: 331. Blackwelder, 1943: 389
Pinophilus Gravenhorst, 1802
Pinophilus danforthi Blackwelder, 1943: 378
Pinophilus flavipes Erichson, 1840. Blackwelder, 1943: 383
Pinophilus hispaniolus Blackwelder, 1943: 384 *DR, *H > TL: Sánchez
Pseudolathra Casey, 1905
Pseudolathra nitidum (Erichson, 1840). Peck, 2005: 73
Rugilus Samouelle, 1819
Rugilus chthonus Blackwelder, 1943: 304
Sciocharis Lynch-Arribálzaga, 1884
Sciocharis eidmanni Scheerpeltz, 1970: 241
Scopaeus Erichson, 1840
Scopaeus antennalis Cameron, 1913: 349. Blackwelder, 1943: 284
Scopaeus dominicanus Blackwelder, 1943: 281 *DR > TL: Sánchez
Scopaeus filum Sharp, 1886. Blackwelder, 1943: 288
Stamnoderus Sharp, 1886
Stamnoderus cameroni Blackwelder, 1943: 355 *DR, *H > TL: San José de las Matas
Stamnoderus labeo (Erichson, 1840). Blackwelder, 1943: 352
Stilomedon Sharp, 1886
Stilomedon audanti Blackwelder, 1943: 258

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Stilomedon connexum (Sharp, 1876). Blackwelder, 1943: 257
Stilomedon insularum (Cameron, 1923). Blackwelder, 1943: 256
Stilosaurus Blackwelder, 1943
Stilosaurus rulomus Blackwelder, 1943: 348 *DR > TL: Valle Nuevo
Suniophacis Blackwelder, 1943
Suniophacis thoracica (Cameron, 1923: 398). Blackwelder, 1943: 346
Sunius Stephens, 1829
Sunius darlingtoni Blackwelder, 1943: 264 *DR > TL: Loma Vieja, south of Constanza
Sunius terresi Blackwelder, 1943: 269 *H > TL: Diquini, on coastal plain between Port-au-Prince and Leogane
Thinocharis Kraatz, 1859
Thinocharis chapini Blackwelder, 1943: 234
Thinocharis densicollis (Sharp, 1886). Scheerpeltz, 1970: 241; Perez-Gelabert, 2012: 115
Subfamily PIESTINAE
Hypotelus Erichson, 1840
Hypotelus insulanus Bierig, 1934. Blackwelder, 1943: 42
Piestus Gravenhorst, 1806
Piestus erythropus Erichson, 1840. Scheerpeltz, 1952: 290
Piestus fulvipes Erichson, 1840. Blackwelder, 1943: 45
Piestus penicillatus (Dalman, 1821). Blackwelder, 1943: 46
Piestus pygmaeus Laporte, 1834. Blackwelder, 1943: 49
Subfamily PSELAPHINAE
Apharus Reitter, 1882
Apharus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Arthmius LeConte, 1850
Arthmius sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Balega Reitter, 1881
Balega longiceps Park in Park et al., 1976: 8 *DR > TL: Near Loma Vieja, S. Constanza, at about 6,000 ft.
Barroeuplectoides Park, 1942
Barroeuplectoides sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Brachyglutina Jeannel, 1950
Brachyglutina sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Bythinogaster Schaufuss, 1887
Bythinogaster simplex Schaufuss, 1887: 112. Blackwelder, 1944: 92 *H, *DR > TL: Ins. S. Domingo
Eupsenius LeConte, 1850
Eupsenius dominicanus Schaufuss, 1887: 108. Blackwelder, 1944: 95 *H, *DR > TL: Ins. S. Domingo
Fustiger LeConte, 1866
Fustiger haytiana Mann, 1915: 163. Blackwelder, 1944: 97 *H > TL: Diquini in a nest of Aphaenogaster relicta
Hispanisella Park in Park et al., 1976
Hispanisella haitiana Park in Park et al., 1976: 33 *H > TL: Shada Headquarters near Refuge
Hispanisella hirsuta Park in Park et al., 1976: 34 *H > TL: Tombeau Cheval
Jubus Schaufuss, 1872
Jubus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Lemelba Park, 1953
Lemelba hirsuta Park in Park et al., 1976: 41 *DR > TL: Sánchez
Malleoceps Park, 1954
Malleoceps darlingtoni Park, 1954: 1 *DR > TL: Sánchez
Ocabaraja Schuster & Grigarick, 1980
Ocabaraja romingeri Schuster & Grigarick, 1980: 39 *DR > TL: La Vega Prov., Jarabacoa
Phamisus Aubé, 1844
Phamisus sp.—Asenjo et al. (2019: 31)
Sandersonella Park in Park et al., 1976
Sandersonella transversa Park in Park et al., 1976: 39 *H > TL: Tombeau Cheval
Scalenarthrus LeConte, 1880
Scalenarthrus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 115
Stratus Schaufuss, 1872
Stratus dominicanus Park in Park et al., 1976: 10 *DR > TL: Cloud forest near Valle Nuevo, at about 6,000 ft.
Stratus hirsutus Park in Park et al., 1976: 11 *DR > TL: Constanza, at between 3,000 and 4,000 ft.
Thesium Casey, 1884
Thesium sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Subfamily SCAPHIDIINAE
Cyparium Erichson, 1845
Cyparium anale Reitter, 1880: 42. Blackwelder, 1944: 98 *DR > TL: St. Domingo
Subfamily SCYDMAENINAE
Euconnus Thomson, 1862
Euconnus abejanus Franz, 1991: 28 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
Euconnus abnormeantennatus Franz, 1984: 18 *DR > TL: Cordillera Central bei Capalita [Loma Casabito], 1250 m
Euconnus aceitillari Franz, 1991: 28 *DR > TL: Prov. Pedernales, El Aceitillar, 13 km N Cabo Rojo, 1310 m
Euconnus allosetosipennis Franz, 1991: 34 *DR > TL: Prov. La Vega, 12 km NE Jarabacoa, 550 m
Euconnus allotestaceus Franz, 1991: 31 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
Euconnus almandoensis Franz, 1991: 33 *DR > TL: Prov. La Vega, Parc Natl. Armando Bermúdez, nr. bottom Pico Duarte
trail, 1350 m
Euconnus caborojoanus Franz, 1991: 31 *DR > TL: Prov. Pedernales, 13.5 km N Cabo Rojo, 140 m
Euconnus caborojoi Franz, 1991: 26 *DR > TL: Prov. Pedernales, 13.5 km N Cabo Rojo, 140 m
Euconnus caribicus Franz, 1984: 24 *DR > TL: Boca Chica
Euconnus cazabitae Franz, 1984: 20 *DR > TL: Cordillera Central bei Cazabita, 1250 m
Euconnus constanzanus Franz, 1991: 33 *DR > TL: Prov. La Vega, ca. 10 km E Constanza, 1295 m
Euconnus dominicanus Franz, 1984: 20 *DR > TL: Valle Nuevo, 2400 m
Euconnus lasabejas O’Keefe, 2000: 64 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
=Euconnus lanceolatus Franz, 1991: 29 [Preoccupied name]
Euconnus lasabejanus Franz, 1991: 27 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
Euconnus michaeli Franz, 1991: 35 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
Euconnus nigerrimus Franz, 1984: 21 *DR > TL: Cordillera Central bei Cazabita, 1250 m
Euconnus (Nudatoconnus) banii Franz, 1984: 19 *DR > TL: Baní
Euconnus paenevisus Franz, 1991: 32 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
Euconnus parallelocollis Franz, 1984: 26 *DR > TL: Constanza
Euconnus parasetosipenis Franz, 1984: 25 *DR > TL: Colonia
Euconnus paratestaceus Franz, 1991: 31 *DR > TL: Prov. Pedernales, 24 km N Cabo Rojo, 600 m
Euconnus pedernalesi Franz, 1991: 26 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
Euconnus philipsi Franz, 1991: 34 *DR > TL: Prov. Pedernales, Las Abejas, 35 km N Cabo Rojo, 1250 m
Euconnus pseudotestaceus Franz, 1991: 30 *DR > TL: Prov. Pedernales, Las Abejas, 35.5 km N Cabo Rojo, 1250 m
Euconnus ramfisensis Franz, 1984: 22 *DR > TL: Colonia Ramfis
Euconnus setosipenis Franz, 1984: 24 *DR > TL: Colonia
Euconnus sibylleae Franz, 1984: 27 *DR > TL: Colonia
Euconnus sibyllensis Franz, 1984: 23 *DR > TL: Colonia Ramfis
Euconnus sp.†—Franz (1983: 28) [Fossil in Dominican amber]
Homoconnus Sharp, 1887
Homoconnus conservatus† Franz, 1983: 27 [Fossil in Dominican amber]
Mastigus Latreille, 1802
Mastigus sp.†—Poinar & Poinar (1999: 205) [Misspelled Mastigonius; Possible error, Old World genus] [Fossil in Dominican
amber]
Microscydmus Croissandeau, 1898
Microscydmus cazabitanus Franz, 1984: 30 *DR > TL: Cordillera Central bei Cazabita, 1250 m
Microscydmus (Neoscydmus) dominicanus Franz, 1991: 35 *DR > TL: Prov. Pedernales, Las Abejas, 35 km N Cabo Rojo,
1250 m
Microscydmus ramfis Franz, 1984: 29 *DR > TL: Colonia Ramfis, 1000 m
Microscydmus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Neuraphes Thomson, 1859
Neuraphes fossilis† Franz, 1983: 26 [Fossil in Dominican amber]

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Scydmaenus Latreille, 1802
Scydmaenus dominicanus Franz, 1991: 36. Perez-Gelabert, 2012: 115 *DR > TL: Prov. Pedernales, Las Abejas, 24 km N Cabo
Rojo, 610 m
Subfamily STAPHYLININAE
Acylophorus Nordmann, 1837
Acylophorus aenescens Blackwelder, 1943: 468 *DR > TL: Sánchez
Acylophorus metallicus Blackwelder, 1943: 469 *DR > TL: Sánchez
Atanygnathus Jacobson, 1809
Atanygnathus laticollis (Erichson, 1839). Blackwelder, 1943: 472
Belonuchus Nordmann, 1837
Belonuchus agilis Erichson, 1840. Blackwelder, 1943: 427; Frank, 2004: 92; Chani-Posse et al., 2018: 10
Belonuchus gagates Erichson, 1840. Blackwelder, 1943: 424
Belonuchus hispaniolus Blackwelder, 1943: 422; Chani-Posse et al., 2018: 19 *DR > TL: San Francisco Mountains
Cafius Stephens, 1829
Cafius bistriatus (Erichson, 1840). Frank & Ahn, 2011: 33; Ahn & Frank, 2012: 709
Creophilus Leach in Samouelle, 1819
Creophilus sp.—Asenjo et al. (2019: 301)
Diochus Erichson, 1839
Diochus angustiformis Irmler, 2017: 26
Diochus perplexus Cameron, 1922. Irmler, 2017: 41
Gabronthus Tottenham, 1955
Gabronthus mgogoricus Tottenham, 1955. Frank, 1983: 476; Chani-Posse et al., 2018: 41 [Introduced]
Gabronthus thermarum (Aubé, 1850). Blackwelder, 1943: 403
Heterothops Stephens, 1829
Heterothops cornelli† Chatzimanolis & Engel, 2013: 285 [Fossil in Dominican amber]
Heterothops infernalis† Chatzimanolis & Engel, 2013: 285 [Fossil in Dominican amber]
Holisus Erichson, 1839
Holisus debilis Erichson, 1839. Blackwelder, 1943: 461
Holisus funeratus† Chatzimanolis & Engel, 2013: 287 [Fossil in Dominican amber]
Lithocharodes Sharp, 1876
Lithocharodes strigulata Blackwelder, 1943: 498 *DR, *H > TL: Loma Rucilla and mountains to the north
Neobisnius Ganglbauer, 1895
Neobisnius humilis (Erichson, 1840). Blackwelder, 1943: 441; Chani-Posse et al., 2018: 47
=Neobisnius limbatus Cameron, 1922: 118
Neobisnius nigrocoeruleus (Cameron, 1922: 118). Herman, 2001: 2709; Chani-Posse et al., 2018: 48
Neohypnus Coiffait & Sáiz, 1964
Neohypnus humeralis (Erichson, 1839). Blackwelder, 1943: 479
Neoxantholinus Cameron, 1944
Neoxantholinus apolithomenus† Chatzimanolis & Engel, 2013: 292 [Fossil in Dominican amber]
Neoxantholinus darlingtoni (Blackwelder, 1943: 506) *DR > TL: Loma Rucilla and mountains to the north
Neoxantholinus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Neoxantholinus testaceipennis (Fauvel, 1863). Blackwelder, 1943: 507
Neoxantholinus variabilis (Blackwelder, 1943). Blackwelder, 1943: 508
Philonthus Stephens, 1929
Philonthus alumnus Erichson, 1840. Smetana, 1994: 342; Herman, 2001: 2747; Chani-Posse et al., 2018: 60
Philonthus audanti Blackwelder, 1943: 406; Chani-Posse et al., 2018: 60 *H > TL: 5 miles south of St. Marc
Philonthus discoideus (Gravenhorst, 1802). Blackwelder, 1943: 407 [Introduced]
Philonthus hades† Chatzimanolis & Engel, 2013: 288; Chani-Posse et al., 2018: 67 [Fossil in Dominican amber]
Philonthus havaniensis (Laporte, 1835). Blackwelder, 1943: 418
Philonthus hepaticus Erichson, 1840. Blackwelder, 1943: 401
Philonthus lomatus Erichson, 1840. Frank, 1983: 479; Chani-Posse et al., 2018: 70
Philonthus longicornis Stephens, 1832. Blackwelder, 1943: 410; Chani-Posse et al., 2018: 70
Philonthus rhadamanthus† Chatzimanolis & Engel, 2013: 289; Chani-Posse et al., 2018: 75 [Fossil in Dominican amber]
Philonthus ventralis (Gravenhorst, 1802). Blackwelder, 1943: 404; Frank, 1983: 479 [Introduced]

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Quedius Stephens, 1829
Quedius (Cyrtoquedius) electrodominicanus† Chatzimanolis & Engel, 2013: 289 [Fossil in Dominican amber]
Xantholinus Dejean, 1821
Xantholinus audanti Blackwelder, 1943: 483 *H, *DR > TL: Port-au-Prince
Xantholinus beattyi Blackwelder, 1943: 485
Xantholinus darlingtoni Blackwelder, 1943: 481 *H > TL: La Visite and vicinity, La Selle Range
Xantholinus haitius Blackwelder, 1943: 484 *H > TL: Etang Lachaux
Xantholinus oakleyi Blackwelder, 1943: 476
Xantholinus rulomus Blackwelder, 1943: 482 *DR > TL: Jarabacoa
Subfamily STENINAE
Stenus Latreille, 1797
Stenus gentilis Sharp, 1886
=Stenus hispaniolus Blackwelder, 1943: 224
Stenus haitiensis Blackwelder, 1943: 217 *H > TL: Northeastern foothills of the Massif de La Hotte
Subfamily TACHYPORINAE
Bryoporus Kraatz, 1858
Bryoporus bicolor Blackwelder, 1943: 528. Korge, 1962: 124 *DR > TL: Cloud forest in the vicinity of Valle Nuevo
Bryoporus regulus Blackwelder, 1943: 527. Korge, 1962: 124 *DR, *H > TL: Cloud forest in the vicinity of Valle Nuevo
Coproporus Kraatz, 1857
Coproporus apicalis (Erichson, 1839). Blackwelder, 1943: 514
Coproporus electron† Yamamoto & Takahashi, 2016: 630 [Fossil in Dominican amber]
Coproporus flavicollis Scriba, 1839. Blackwelder, 1943: 518
Coproporus nitidulus (Erichson, 1839). Herman, 2001: 830
Coproporus pulchellus (Erichson, 1839). Blackwelder, 1943: 520
Coproporus rutilus (Erichson, 1839). Blackwelder, 1943: 522
Sepedophilus Gistel, 1856
Sepedophilus interruptus (Erichson, 1839). Blackwelder, 1943: 525
Tachyporus Gravenhorst, 1802
Tachyporus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Subfamily TRICHOPSENIINAE
Prorhinopsenius Pasteels & Kistner, 1971
Prorhinopsenius alzadae† Kistner, 1998: 59 [Fossil in Dominican amber]
Prorhinopsenius poinari† Kistner, 1998: 61 [Fossil in Dominican amber]
Prorhinopsenius seeversi† Kistner, 1998: 61 [Fossil in Dominican amber]

Infraorder SCARABAEIFORMIA
Superfamily SCARABAEOIDEA
Family PASSALIDAE
Key reference: Jiménez-Ferbans et al. (2015).
Passalus Fabricius, 1792
Passalus (Pertinax) affinis Percheron, 1835: 72. Chevrolat, 1865: 409; Blackwelder, 1944: 192; Reyes-Castillo et al., 1995: 30;
Jiménez-Ferbans et al., 2015: 499 *DR, *H > TL: Saint-Domingue
Passalus (Pertinax) doesburgi (Boucher, 2015). Jiménez-Ferbans et al., 2015: 503 *DR > TL: Pedernales, Las Abejas, 38 km
NNW Cabo Rojo, 1160 m
Passalus (Pertinax) dominicanus Van Doesburg, 1953. Reyes-Castillo et al., 1995: 30; Jiménez-Ferbans et al., 2015: 501 *DR
> TL: La Capito, St. Domingo [Possibly La Cavite, as this was one of the collecting localities of R. H. Beck in Haiti]
Paxillus MacLeay, 1819
Paxillus pentaphyllus (Palisot de Beauvois, 1805). Chevrolat, 1865: 410; Reyes-Castillo et al., 1995: 31; Jiménez-Ferbans et
al., 2015: 504 *H, *DR > TL: Saint-Domingue
Spasalus Kaup, 1869
Spasalus crenatus (MacLeay, 1819). Reyes-Castillo et al., 1995: 32

Family TROGIDAE
Omorgus Erichson, 1847

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Omorgus suberosus (Fabricius, 1775). Leng & Mutchler, 1914: 440; Wolcott, 1927: 227; Blackwelder, 1944: 219; Vaurie,
1962: 144
=Trox crenatus Chevrolat, 1865: 416
Trox Fabricius, 1775
Trox monachus Herbst, 1790. Blackwelder, 1944: 219

Family GEOTRUPIDAE
Subfamily ATHYREINAE
Neoathyreus Howden & Martinez, 1963
Neoathyreus biceps (Felsche, 1909: 763). Howden, 1978: 387 *DR, *H > TL: Haiti
Neoathyreus tweedyanus (Westwood, 1848: 387). Howden, 1978: 386 *DR, *H > TL: Insula Hayti
Subfamily GEOTRUPINAE
Geotrupes Latreille, 1796
Geotrupes semiopaca Jekel, 1866. Blackwelder, 1944: 220
Geotrupes splendida (Fabricius, 1775). Blackwelder, 1944: 220
Notes: The names Geotrupes meridionalis Palisot de Beauvois, 1805 and Geotrupes thoracina Palisot de Beauvois, 1805 cited
by Blackwelder, 1944: 220 for Hispaniola, are uncertain species that have not been corroborated with recent records. These are
listed under rejected records and are not considered valid (West Indian Beetle Fauna Project http://wibeetles.org/polyphaga/
scarabaeiformia/scarabaeoidea/geotrupidae.html).

Family CERATOCANTHIDAE
Subfamily CERATOCANTHINAE
Ceratocanthus White, 1842
Ceratocanthus baniensis Howden, 1978: 382 *DR > TL: Baní, 65 m
Germarostes White, 1842
Germarostes emarginatus† (Poinar, 2016: 433). Poinar & Ballerio, 2017: 125 [Fossil in Dominican amber]

Family HYBOSORIDAE
Subfamily ANAIDINAE
Anaides Westwood, 1845
Anaides howdeni† Ocampo, 2006: 40 [Fossil in Dominican amber]
Subfamily HYBOSORINAE
Apalonychus Westwood, 1846
Apalonychus rufulus (Laporte, 1840: 108). Allsop, 1984: 106 *H, *DR > TL: Saint-Domingue
Apalonychus waterhousei Westwood, 1846. Peck, 2005: 84
Hybosorus MacLeay, 1819
Hybosorus illigeri Reiche, 1853. Ocampo, 2002b: 317 [Introduced]
Procoilodes† Ocampo, 2002
Procoilodes adrastus† Ocampo, 2002a: 125 [Fossil in Dominican amber]
Tyrannasorus† Ocampo, 2001
Tyrannasorus rex† Ratcliffe & Ocampo, 2001: 353 [Fossil in Dominican amber]

Family SCARABAEIDAE
Subfamily APHODIINAE
Haroldiellus Gordon & Skelley, 2007
Haroldiellus sallei (Harold, 1863). [By synonymy of species below]
=Aphodius sallei Harold, 1863. Blackwelder, 1944: 213
Labarrus Gordon & Skelley, 2007
Labarrus lividus (Olivier, 1789). [By synonymy of species below. Introduced]
=Aphodius lividus Olivier, 1789. Blackwelder, 1944: 212
Nialaphodius Kolbe, 1908
Nialaphodius nigrita (Fabricius, 1801). [By synonymy of species below. Introduced]
=Aphodius cuniculus Chevrolat, 1864. Blackwelder, 1944: 212
Subfamily SCARABAEINAE

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Rhyparus Westwood, 1845
Rhyparus sp.†—Wu (1997: 187) [Fossil in Dominican amber]
Termitodius Wasmann, 1894
Termitodius sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Tribe EUPARIINI
Ataeniopsis Petrovitz, 1973
Ataeniopsis armasi (Chalumeau, 1982). Stebnicka, 2003a: 109
Ataenius Harold, 1867
Ataenius aciculus Hinton, 1937: 187. Blackwelder, 1944: 213; Stebnicka, 2002: 271 *H, *DR > TL: Diquini
=Ataenius apicecoloratus Balthasar, 1941: 168
Ataenius beattyi Chapin, 1940. Stebnicka, 2006: 107
Ataenius brevicollis (Wollaston, 1854). Stebnicka, 2007: 58
Ataenius carinator Harold, 1874. Peck, 2010: 26
=Ataenius vincentiae Arrow, 1903. Blackwelder, 1944: 216
Ataenius elisaensis Stebnicka, 2002: 270 *DR > TL: Monte Cristi, 8.2 km N Villa Elisa
Ataenius elongatus (Palisot de Beauvois, 1805: 104). Blackwelder, 1944: 213; Stebnicka, 2002: 276
Ataenius gracilis (Melsheimer, 1844). Blackwelder, 1944: 214
Ataenius heinekeni (Wollaston, 1854). Stebnicka, 2004: 216
=Ataenius rhyticephalus (Chevrolat, 1964). Stebnicka, 2004: 216; Peck, 2005: 85
Ataenius hispaniolae Chalumeau, 1982: 325. Stebnicka, 2002: 269 *DR > TL: S. P. Macor., Guayacanes
Ataenius imbricatus (Melsheimer, 1844). Blackwelder, 1944: 214; Stebnicka, 2003b: 225
Ataenius jelineki Chalumeau, 1982: 323 *H > TL: Ennery, 1,000 ft.
Ataenius klapperichi Howden, 1978: 378. Stebnicka, 2002: 279 *DR > TL: La Colonia, 1000 m
Ataenius liogaster Bates, 1887. Stebnicka & Lago, 2005: 60
Ataenius luteomargo Chapin, 1940. Blackwelder, 1944: 215
Ataenius morator Harold, 1869 [By synonymy of species below]
=Ataenius picipes Fleutiaux & Sallé, 1890. Stebnicka, 2003b: 243
=Ataenius tenebrosus Arrow, 1903. Blackwelder, 1944: 216; Stebnicka, 2003b: 243
Ataenius picinus Harold, 1867. Cartwright, 1964: 103; Peck & Thomas, 1998: 59; Stebnicka, 2004: 224
=Ataenius darlingtoni Hinton, 1937. Blackwelder, 1944: 214
=Ataenius edwardsi Chapin, 1940. Blackwelder, 1944: 214
Ataenius raccurti Chalemeau, 1978: 50. Stebnicka, 2002: 276
Ataenius scabrellus Schmidt, 1909. Stebnicka, 2003b: 227
Ataenius scutellaris Harold, 1867. Cartwright, 1978: 14; Stebnicka, 2007: 48
=Ataenius frater Arrow, 1903. Blackwelder, 1944: 214
Ataenius strigicauda Bates, 1887. Blackwelder, 1944: 215. Stebnicka, 2007: 214
Ataenius versicolor Schmidt, 1916. Blackwelder, 1944: 216; Stebnicka, 2002: 273
=Ataenius haitianus Hinton, 1937: 184. Blackwelder, 1944: 216
Note: According to Z. Stebnicka (pers. comm., 2007) Ataenius abditus (Haldeman, 1848), cited by Blackwelder, 1944: 213 and
Ataenius haroldi Steinheil, 1872, cited by Blackwelder, 1944: 214 do not occur in Hispaniola.
Tribe ONTHOPHAGINI
Digitonthophagus Balthasar, 1959
Digitonthophagus gazella (Fabricius, 1787). MacRae & Penn, 2001: 49; Ivie & Philips, 2007: 10 [Introduced]
Onthophagus Latreille, 1802
Onthophagus albicornis Palisot de Beauvois, 1805. Ivie & Philips, 2008: 9
=Onthophagus albicornis albicornis Palisot de Beauvois, 1805. Perez-Gelabert, 2008: 96
Onthophagus antillarum Arrow, 1903. Pulido Herrera & Zunino, 2008: 95
Onthophagus capitatus Laporte, 1840. Ivie & Philips, 2008: 9
=Onthophagus albicornis capitatus Laporte, 1840. Perez-Gelabert, 2008: 96
Tribe SCARABAEINI
Canthidium Erichson, 1847
Canthidium sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Canthochilum Chapin, 1934
Canthochilum alleni† Tarasov & Vaz-de-Mello in Tarasov et al, 2016: 7 [Fossil in Dominican amber]

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Canthochilum brodzinskyorum Philips & Ivie, 2008: 28 *DR > TL: Las Abejas, ca. 35 km N. Cabo Rojo, 1250 m
Canthochilum ciboney Matthews, 1969: 120 *H > TL: Mt. Trou d’Eau
Canthochilum darlingtoni Matthews, 1969: 117 *DR > TL: Sánchez
Canthochilum magnum Philips & Ivie, 2008: 31 *DR > TL: Par. Nac. A. Bermúdez, Los Tablones, 1,290 m
Canthochilum matthewsi Philips & Ivie, 2008: 32 *DR > TL: Par. Nac. A. Bermúdez, Los Tablones, 1,220 m
Canthochilum mimicum Philips & Ivie, 2008: 33 *DR > TL: Prov. Pedernales, 20 km N. Cabo Rojo, 365 m
Canthochilum nebulonemi Philips & Ivie, 2008: 34 *DR > TL: Independencia, 32 km NW La Descubierta, Sabana Real, 1800
m
Canthochilum platycnemis Philips & Ivie, 2008: 35 *DR > TL: Puerto Plata, Pico El Murazo, north slope, near summit, 910 m
Canthochilum philipsivieorum† Tarasov & Vaz-de-Mello in Tarasov et al, 2016: 9 [Fossil in Dominican amber]
Canthochilum xericum Philips & Ivie, 2008: 36 *DR > TL: Pedernales, 4 km W Oviedo, 10 m
Canthon Hoffmannsegg, 1817
Canthon callosus Harold, 1868. Wolcott, 1927: 227; Blackwelder, 1944: 198 *DR, *H > TL: St. Domingo
Canthon signifer Harold, 1868. Wolcott, 1927: 227; Blackwelder, 1944: 201 *DR, *H > TL: St. Domingo
Canthon violaceus (Olivier, 1789). Wolcott, 1927: 227; Blackwelder, 1944: 202 *DR, *H > TL: St.-Domingue
Canthonella Chapin, 1930
Canthonella baorucensis Ivie & Philips, 1990: 430 *DR > TL: Pedernales Prov., Las Abejas, ca. 35 km N Cabo Rojo, 1,250 m
Canthonella constans Matthews, 1966: 81. Matthews, 1969: 124 *DR > TL: 11 km south of Constanza
Canthonella howdeni Ivie & Philips, 1990: 432 *DR > TL: Pedernales Prov., Las Abejas, ca. 35 km N Cabo Rojo, 1,250 m
Canthonella isabellae Matthews, 1966: 79. Ivie & Philips, 1990: 432 *DR > TL: 31 km northwest of Santo Domingo
Canthonella jarmilae Ivie & Philips, 2008: 2 *DR > TL: Independencia, 30 km NW La Descubierta, Sabana Real, 1646 m
Canthonella quesqueya Ivie & Philips, 2008: 3 *DR > TL: PN A. Bermúdez, Ciénaga, 1000 m
Canthonella sikesi Ivie & Philips, 2008: 5 *DR > TL: Prov. Hato Mayor, Par. Nac. Los Haitises
Subfamily MELOLONTHINAE
Key reference: Woodruff and Sanderson (2004).
Phyllophaga Harris, 1826
Phyllophaga aceitillar Woodruff in Woodruff & Sanderson, 2004: 31 *DR > TL: Sierra de Baoruco, Aceitillar, 23.6 km NE
Pedernales, 1560 m
Phyllophaga alcoa Woodruff in Woodruff & Sanderson, 2004: 33 *DR > TL: Pedernales Prov., 17 km N Cabo Rojo, 255 m
Phyllophaga aliada Sanderson, 1951: 279. Woodruff & Sanderson, 2004: 36 *H, *DR > TL: N. Haiti, Mt. Basil, 4700 ft.
Phyllophaga androw Woodruff in Woodruff & Sanderson, 2004: 39 *DR, *H > TL: Prov. San Juan, 9.4 km SSW El Cercado,
Sierra de Neiba, 1973 m
Phyllophaga approxima Woodruff & Sanderson, 2004: 42 *DR > TL: Prov. La Vega, 5–7 mi S. Constanza
Phyllophaga audanti (Wolcott, 1928: 27). Woodruff & Sanderson, 2004: 45 *H, *DR > TL: Port-au-Prince
Phyllophaga baoruco Woodruff in Woodruff & Sanderson, 2004: 45 *DR > TL: Prov. Barahona, Larimar Mine, nr. Filipinas,
3300 ft.
Phyllophaga barrosa Sanderson, 1951: 280. Woodruff & Sanderson, 2004: 50 *H > TL: Fond des Negres
Phyllophaga bonfils Woodruff & Sanderson, 2004: 52 *H > TL: Furcy, Dept. de l’Ouest
Phyllophaga canoa Sanderson, 1951: 279. Woodruff & Sanderson, 2004: 54 *DR > TL: Vicinity of Valle Nuevo, cloudforest,
6000 ft.
Phyllophaga carnegie Woodruff in Woodruff & Sanderson, 2004: 57 *DR > TL: Prov., Elias Piña, 9.3 km WSW Hondo Valle,
Sierra de Neiba, 1901 m
Phyllophaga cartaba Sanderson, 1951: 274. Woodruff & Sanderson, 2004: 60 *DR > TL: S. Dom.
Phyllophaga costura Sanderson, 1951: 264. Woodruff & Sanderson, 2004: 60 *DR > TL: Loma Rucilla and mountains north,
5000–8000 ft.
Phyllophaga davidsoni Woodruff in Woodruff & Sanderson, 2004: 60 *DR > TL: Prov. Pedernales, Las Abejas, 38 km N. Cabo
Rojo, 1250 m
Phyllophaga delphinicauda Androw, 2016: 2 *DR > TL: Eastern Sierra de Bahoruco, Reserva Cachote, 1198 m
Phyllophaga eladio Woodruff in Woodruff & Sanderson, 2004: 65 *DR > TL: Prov. La Vega, 5 km W. Manabao, Finca Eladio
Fernandez, Paso de la Perra, along Río Yaque del Norte, 3050 ft. Elev.
Phyllophaga espina Sanderson, 1951: 264. Woodruff & Sanderson, 2004: 68 *DR > TL: Foothills Cordillera Central south of
Santiago
Phyllophaga esquinada Sanderson, 1951: 268. Woodruff & Sanderson, 2004: 68 *H > TL: La Visite and vicinity, La Selle
Range, 5000–7000 ft.

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Phyllophaga fossoria Sanderson, 1951: 258. Woodruff & Sanderson, 2004: 68 *DR > TL: Constanza to Valle Nuevo, 3000–
7000 ft.
Phyllophaga n. sp. nr. fossoria—Woodruff & Sanderson (2004: 73)
Phyllophaga garrota Sanderson, 1951: 262. Woodruff & Sanderson, 2004: 73 *DR > TL: Loma Rucilla and mountains north,
5000–8000 ft.
Phyllophaga haitiensis Woodruff in Woodruff & Sanderson, 2004: 76 *DR > TL: Departement du Sud, Ville Formon, 31 km
NW Les Cayes, S. Slope Morne Formon, Massif de La Hotte, 1405 m
Phyllophaga hogardi (Blanchard, 1850: 137). Wolcott, 1927: 227; Woodruff & Sanderson, 2004: 79 *H, *DR > TL: Saint-
Domingue
Phyllophaga imprima Sanderson, 1951: 276. Woodruff & Sanderson, 2004: 81 *DR > TL: Mt. Quita Espuela, 2000–3000 ft.
Phyllophaga jaragua Woodruff in Woodruff & Sanderson, 2004: 84 *DR > TL: Prov. Barahona, 6 km S. Cabral, Rd. to Polo
Phyllophaga jimenezi Woodruff & Sanderson, 2004: 87 *DR > TL: La Vega Prov., Jarabacoa
Phyllophaga kenscoffi (Wolcott, 1928: 24). Woodruff & Sanderson, 2004: 90 *H, *DR > TL: Kenscoff (elevation 1,400 m)
Phyllophaga larimar Woodruff in Woodruff & Sanderson, 2004: 90 *DR > TL: Prov. Barahona, nr. Filipinas, Larimar Mine,
3300 ft.
Phyllophaga latiungula (Wolcott, 1928: 28). Woodruff & Sanderson, 2004: 95 *H > TL: Hotel Mon Repos, Carrefour (near
Port-au-Prince)
Phyllophaga leptospica Sanderson, 1951: 262. Woodruff & Sanderson, 2004: 95 *H, *DR > TL: Hayti
Phyllophaga mali (Wolcott, 1928: 25). Woodruff & Sanderson, 2004: 97 *H, *DR > TL: Kenscoff
Phyllophaga marcano Woodruff in Woodruff & Sanderson, 2004: 97 *DR > TL: Prov. Elias Piña, Río Limpio, 2400 ft.
Phyllophaga mella Sanderson, 1951: 278. Woodruff & Sanderson, 2004: 103 *DR > TL: Foothills Cordillera Central, south of
Santiago
Phyllophaga minutissima (Wolcott, 1928: 76). Woodruff & Sanderson, 2004: 106 *H, *DR > TL: Port-au-Prince
Phyllophaga neglecta (Blanchard, 1850: 137). Woodruff & Sanderson, 2004: 108 *H, *DR > TL: Saint-Domingue
Phyllophaga nunezi Woodruff in Woodruff & Sanderson, 2004: 110 *DR > TL: Prov. Monseñor Nouel, Río Blanco, Muro del
Embalse, 600 m
Phyllophaga ortizi Woodruff in Woodruff & Sanderson, 2004: 113 *DR > TL: Prov. Barahona, nr. Filipinas, Larimar Mine
Phyllophaga panicula Sanderson, 1951: 275. Woodruff & Sanderson, 2004: 117 *H, *DR > TL: Ouest Haiti, Kenscoff
Phyllophaga pedernales Woodruff in Woodruff & Sanderson, 2004: 117 *DR > TL: Prov. Pedernales, Cabo Rojo, Alcoa
Phyllophaga permagna (Moser, 1918: 57). Woodruff & Sanderson, 2004: 121 *H, *DR > TL: Port-au-Prince
Phyllophaga probaporra Sanderson, 1951: 263. Woodruff & Sanderson, 2004: 123 *DR > TL: Loma Rucilla and mountains
north, 5000–8000 ft.
Phyllophaga pseudocalcaris (Saylor, 1940: 309). Woodruff & Sanderson, 2004: 123 *DR > TL: Río San Juan
Phyllophaga rawlinsi Woodruff in Woodruff & Sanderson, 2004: 126 *DR > TL: Prov. Azua, 8 km NE Padre las Casas, Río
Las Cuevas, 580 m
Phyllophaga recorta Sanderson, 1951: 268. Woodruff & Sanderson, 2004: 130 *H, *DR > TL: Kenscoff, near Port-au-Prince,
4000–6000 ft.
Phyllophaga rex Woodruff & Sanderson, 2004: 130 *DR > TL: Prov. La Vega, Constanza
Phyllophaga romana (Saylor, 1946: 1). Woodruff & Sanderson, 2004: 137 *DR > TL: Moca, Espaillat Prov.
Phyllophaga rustica Woodruff in Woodruff & Sanderson, 2004: 137 *DR > TL: Prov. La Vega, Convento, 12 km S. Constanza
Phyllophaga santachloe Woodruff in Woodruff & Sanderson, 2004: 141 *DR > TL: Prov. La Altagracia, 5 km W. Laguna
Nisibón, Finca Papagayo
Phyllophaga toni Woodruff in Woodruff & Sanderson, 2004: 143 *DR > TL: Prov. Barahona, nr. Filipinas, Larimar Mine, 3300
ft
Subfamily RUTELINAE
Tribe RUTELINI
Pelidnota MacLeay, 1819
Pelidnota sanctidomini Ohaus, 1905: 317. Leng & Mutchler, 1914: 441; Blackwelder, 1944: 238 *H, *DR > TL: Insel Santo
Domingo
Rutela Latreille, 1802
Rutela dorcyi (Olivier, 1789). Leng & Mutchler, 1914: 441; Blackwelder, 1944: 239
Subfamily DYNASTINAE
Key reference: Ratcliffe & Cave (2015).
Tribe CYCLOCEPHALINI

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Chalepides Casey, 1915
Chalepides barbatus (Fabricius, 1781). Blackwelder, 1944: 253; Ratcliffe & Cave, 2015: 71
Cyclocephala Dejean, 1821
Cyclocephala cerea Burmeister, 1847. Ratcliffe & Cave, 2015: 53
Cyclocephala frontalis Chevrolat, 1844. Ratcliffe & Cave, 2015: 87
Cyclocephala laevis Arrow, 1937: 40. Blackwelder, 1944: 251; Ratcliffe & Cave, 2015: 95
Cyclocephala melanocephala (Fabricius, 1775). Ratcliffe & Cave, 2015: 99
Cyclocephala notata (Illiger, 1806). Wolcott, 1927: 233; Blackwelder, 1944: 252; Endrödi, 1966: 267; Ratcliffe & Cave, 2015:
101
Cyclocephala quisqueya Joly, 1998: 50. Ratcliffe & Cave, 2015: 105 *DR > TL: Convento, 12 km S of Constanza
Cyclocephala rangelana Chapin, 1935. Ratcliffe & Cave, 2015: 107
=Cyclocephala vidua Endrödi, 1966: 331
Cyclocephala striata Endrödi, 1963. Ratcliffe & Cave, 2015: 109
=Cyclocephala striata haitiensis Endrödi, 1963: 328. Endrödi, 1966: 315
Dyscinetus Harold, 1869
Dyscinetus laevicollis Arrow, 1937: 41. Ratcliffe & Cave, 2015: 126
Dyscinetus picipes (Burmeister, 1847). Leng & Mutchler, 1914: 442; Peck, 2005: 73; Ratcliffe & Cave, 2015: 132
Mimeoma Casey, 1915
Mimeoma nigra Endrödi, 1979: 216. Ratcliffe & Cave, 2015: 137 *DR > TL: Dajabón Prov., 13 km S Loma de Cabrera, ca.
400 m
Tribe PENTODONTINI
Endroedianibe Chalumeau, 1981
Endroedianibe bozzolai Chalumeau, 1981: 221. Chalumeau, 1983a: 98; Ratcliffe & Cave, 2011: 4; Ratcliffe & Cave, 2015: 145
*DR, *H > TL: Loma Rucilla
Endroedianibe xerozoyphius Ratcliffe & cave, 2011: 4. Ratcliffe & Cave, 2015: 147 *DR > TL: 1.6 km W Fondo Negro,
Barahona Prov.
Tomarus Erichson, 1847
Tomarus cuniculus (Fabricius, 1801). Cartwright & Chalumeau, 1978: 27; Ratcliffe & Cave, 2015: 156
Tribe ORYCTINI
Hispanioryctes Howden & Endrodi in Howden, 1978
Hispanioryctes paulseni Ratcliffe & Cave, 2011: 6. Ratcliffe & Cave, 2015: 173 *DR > TL: Mano Juan-Catuano, Isla Saona
Hispanioryctes rugulopygus Ratcliffe & Cave, 2011: 8. Ratcliffe & Cave, 2015: 174 *H, *DR > TL: Furcy
Hispanioryctes sculptilis Ratcliffe & Cave, 2011: 10. Ratcliffe & Cave, 2015: 176 *H, *DR > TL: Miragoâne
Hispanioryctes wittmeri Howden & Endrödi in Howden, 1978: 389; Ratcliffe & Cave, 2011: 11; Ratcliffe & Cave, 2015: 178
*H, *DR > TL: Valle Nuevo, 400 m
Strategus Kirby, 1828
Strategus aenobarbus (Fabricius, 1775). Blackwelder, 1944: 257; Ratcliffe & Cave, 2015: 185
=Scarabaeus fossula Palisot de Beauvois, 1819: 210
=Strategus laterispinus Chapin, 1932: 454
Strategus inermis Arrow, 1947: 223. Ratcliffe & Cave, 2015: 201 *H > TL: Haiti
Strategus oblongus (Palisot de Beauvois, 1807). Blackwelder, 1944: 257; Schmutterer, 1990: 125; Ratcliffe, 2014: 1; Ratcliffe
& Cave, 2015: 201
=Scarabaeus monocerus Nicolson, 1776: 360. Ratcliffe, 2014: 1 [Nomen oblitum]
=Strategus quadrifoveatus (Palisot de Beauvois, 1807). Blackwelder, 1944: 257
Tribe PHILEURINI
Hemiphileurus Kolbe, 1910
Hemiphileurus complanatus (Palisot de Beauvois, 1811). Blackwelder, 1944: 258; Ratcliffe & Cave, 2015: 231
Hemiphileurus consimilis Ratcliffe & Cave, 2015: 233 *DR, *H > TL: Pedernales, Las Abejas
Hemiphileurus dispar (Kolbe, 1910). Blackwelder, 1944: 259; Ratcliffe & Cave, 2015: 237
Hemiphileurus phratrius Ratcliffe & Ivie, 1998: 205. Ratcliffe & Cave, 2015: 244 *DR > TL: 26 km N Cabo Rojo, Sierra de
Bahoruco, Pedernales Prov.
Hemiphileurus ryani Ratcliffe & Ivie, 1998: 201. Ratcliffe & Cave, 2015: 247
Hemiphileurus scutellatus Howden & Endrödi in Howden, 1978: 390. Ratcliffe & Cave, 2015: 250 *DR > TL: Baní, 65 m
Homophileurus Kolbe, 1910

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Homophileurus quadrituberculatus (Palisot de Beauvois, 1806). Blackwelder, 1944: 258; Ratcliffe & Cave, 2015: 255
Phileurus Latreille, 1807
Phileurus valgus (Olivier, 1789). Ratcliffe & Cave, 2015: 262
Planophileurus Chapin, 1932
Planophileurus broccus Ratcliffe & Cave, 2015: 267 *DR > TL: Prov. Hato Mayor, El Valle, Hotel Gaurua
Notes: Erroneous records of Dynastinae species in Hispaniola include Cyclocephala discolor (Herbst, 1790)—Endrodi (1966)
listed one record of this species from Haiti with no data. This is a Central and South American species. Also Tomarus
bituberculatus (Palisot de Beauvois, 1811)—Blackwelder (1944) cited Dominican Republic as a locality for this species, but it
occurs from Mexico to Amazonia and not in the West Indies (Ratcliffe & Cave, 2015: 290).
Subfamily CETONINAE
Tribe GYMNETINI
Amithao Thomson, 1878
Amithao incertus (Gory & Percheron, 1833: 73). Blackwelder, 1944: 262; Ratcliffe, 2013: 280 *H, *DR > TL: Saint-Domingue
Gymnetis MacLeay, 1819
Gymnetis chalcipes undata Olivier, 1789. Wolcott, 1927: 236; Blackwelder, 1944: 263
Gymnetis lanius burmeisteri Schürhoff, 1937: 63. Blackwelder, 1944: 263
Gymnetis stellata (Latreille, 1833). Ratcliffe, 2018: 202 [Introduced]
Paleotrichius† Poinar, 2011
Paleotrichius dominicanus† Poinar, 2011: 110 [Fossil in Dominican amber]

Family LUCANIDAE
Subfamily SYNDESINAE
Syndesus MacLeay, 1819
Syndesus ambericus† Woodruff, 2009: 2 [Fossil in Dominican amber]

Infraorder ELATERIFORMIA
Superfamily BUPRESTOIDEA
Family BUPRESTIDAE
Subfamily AGRILINAE
Agrilus Curtis, 1825
Agrilus dominicanus Thomson, 1879. Blackwelder, 1944a: 326
Agrilus hispaniolae Hespenheide, 1998: 197
=Agrilus klapperichi Hespenheide, 1990: 402. Hespenheide, 1998: 197
Aphanisticus Latreille, 1810
Aphanisticus cochinchinae seminulum Obenberger, 1929. Hespenheide, 2007: 164 [Introduced]
Leiopleura Deyrolle, 1864
Leiopleura darlingtoni Fisher, 1940: 162. Blackwelder, 1944a: 340; Hespenheide & Bellamy, 2005: 109 *DR, *H > TL: R.
Froide, Port-au-Prince
Leiopleura gibbipennis (Fisher, 1940: 160). Blackwelder, 1944a: 332; Hespenheide & Bellamy, 2005: 109
Leiopleura iviei Hespenheide & Bellamy, 2004: 107 *DR > TL: La Altagracia Prov., Boca de Yuma, entrance to Parque
Nacional del Este, 12 m
Neotrachys Obenberger, 1923
Neotrachys bilyi Hespenheide, 1990: 403. Hespenheide, 2006: 238 *DR > TL: Cazabita [Loma Casabito], 1250 m
Pachyschelus Solier, 1833
Pachyschelus valentinorum Hespenheide & Bellamy, 2004: 106 *H > TL: Morne Guimby, 22 km SE of Fond Verrettes, 6500 ft.
Sambomorpha Obenberger, 1924
Sambomorpha clarki Hespenheide, 1990: 400 *DR > TL: Baní
Taphrocerus Solier, 1833
Taphrocerus haitiensis Fisher, 1949: 348 *DR, *H > TL: Lac Azuei
Subfamily BUPRESTINAE
Actenodes Lacordaire, 1857
Actenodes auronotatus (Laporte & Gory, 1837). Chevrolat, 1867: 584; Blackwelder, 1944: 319
Actenodes bellulus (Mannerheim, 1837: 79). Blackwelder, 1944a: 319 *DR, *H > TL: Insula St. Domingo
Actenodes embrikstrandi Obenberger, 1936: 138. Blackwelder, 1944a: 319 *H > TL: Port-au-Prince

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Actenodes nobilis (Linnaeus, 1758). Fisher, 1930a: 127; Blackwelder, 1944a: 319
Actenodes oberthuri Bleuzen, 1989: 10
Buprestis Linnaeus, 1758
Buprestis hispaniolae Fisher, 1940: 157. Blackwelder, 1944a: 313 *DR > TL: Between Constanza and Jarabacoa, at an altitude
of 2,000 to 4,000 ft.
Buprestis maculativentris Say, 1824: 272. Blackwelder, 1944a: 313
Buprestis maura Olivier, 1790: 19. Blackwelder, 1944a: 313 *DR > TL: l’Amerique Meridionale
Chrysobothris Solier, 1833
Chrysobothris amberestris† Bellamy, 1999: 321 [Fossil in Dominican amber]
Chrysobothris chlorosticta Thomson, 1879. Blackwelder, 1944a: 316
Chrysobothris dentipes (Germar, 1824). Blackwelder, 1944a: 316
Chrysobothris haitiensis Fisher, 1930b: 7. Blackwelder, 1944a: 317 *DR, *H > TL: Manneville
Chrysobothris hispaniolae Fisher, 1925b: 121. Blackwelder, 1944a: 317; Maier & Ivie, 2013: 85 *H, *DR > TL: Port-au-Prince
Chrysobothris megacephala Laporte & Gory, 1837. Blackwelder, 1944a: 317
Chrysobothris parvofoveata Fisher, 1925b: 116. Blackwelder, 1944a: 318 *H, *DR > TL: Port-au-Prince
Chrysobothris tranquebarica (Gmelin, 1790). Blackwelder, 1944a: 318
=Chrysobothris impressa (Fabricius, 1775). Chevrolat, 1867: 586
Cypriacis Casey, 1909
Cypriacis striata (Fabricius, 1775). Blackwelder, 1944a: 313
Hilarotes Thomas, 1778
Hilarotes mannerheimi (Mannerheim, 1837: 66). Blackwelder, 1944a: 310 *DR > TL: Insula St. Domingo
Hilarotes nitidicollis (Laporte & Gory, 1838): 110. Blackwelder, 1944a: 310 *DR, *H > TL: Santo Domingo
=Psiloptera aureomicans Nonfried, 1891: 272
Melanophila Waterhouse, 1889
Melanophila acuminata (De Geer, 1774). Blackwelder, 1944a: 313 [Introduced]
Mixochlorus Waterhouse, 1887
Mixochlorus elegans Fisher, 1925b: 161. Blackwelder, 1944a: 313 *DR > TL: Sánchez
Peronaemis Waterhouse, 1887
Peronaemis insulicola Fisher, 1940: 159. Blackwelder, 1944a: 313; Bellamy, 2000: 163 *DR > TL: Between Constanza and
Jarabacoa, at an altitude of 2,000 to 4,000 ft.
Spectralia Casey, 1909
Spectralia albonotata (Laporte & Gory, 1838): 149. Blackwelder, 1944a: 312 *DR, *H > TL: Port-au-Prince
Tetragonoschema Thomson, 1857
Tetragonoschema quadratum (Buquet, 1841). Blackwelder, 1944a: 315
Xenorhipis LeConte, 1866
Xenorhipis klapperichorum (Nelson, 1978: 319). Bellamy, 2005: 26 *DR > TL: Baní
Subfamily CHALCOPHORINAE
Chalcophora Solier, 1833
Chalcophora virginiensis (Drury, 1770). Ivie et al., 2014: 712
Dicerca Eschscholtz, 1829
Dicerca divaricata (Say, 1823). Blackwelder, 1944a: 312
Lampetis Spinoa, 1837
Lampetis aurata (Saunders, 1871: 23). Blackwelder, 1944a: 310 *DR, *H > TL: S. Domingo
=Psiloptera (Lampetis) aurata domingoensis Fisher, 1930a: 126. Blackwelder, 1944a: 310
Lampetis aurifera (Olivier, 1790). Chevrolat, 1867: 576; Blackwelder, 1944a: 310
Lampetis torquata (Dalman, 1823). Corona, 2005: 771
Subfamily POLYCESTINAE
Acmaeodera Eschscholtz, 1829
Acmaeodera cruenta (Olivier, 1790: 48). Blackwelder, 1944a: 307 *DR, *H > TL: Saint-Domingue
Acmaeodera gundlachi Fisher, 1925. Westcott, 2006: 157
Micrasta Kerremans, 1893
Micrasta hispaniolae Fisher, 1940: 166. Blackwelder, 1944a: 341 *DR > TL: Foothills of the Cordillera Central, south of
Santiago
Micrasta monticola Fisher, 1940: 165. Blackwelder, 1944a: 341 *DR > TL: Mt. Diego de Ocampo, at an altitude of 3,000 to

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 125
4,000 ft.
Paratyndaris Fisher, 1919
Paratyndaris (Knuliella) antillarum Fisher, 1940: 156. Blackwelder, 1944a: 307; Nelson & Bellamy, 2004: 17
Polycesta Solier, 1833
Polycesta (Jamaiquesia) cyanipes (Fabricius, 1787). MacRae & Nelson, 2003: 66
Polycesta (Jamaiquesia) fisheri Obenberger, 1936: 105. Blackwelder, 1944a: 306 *DR > TL: “Haiti: Hato del Yague”
[probably DR: Cordillera Central]
Polycesta (Polycesta) bilyi Cobos, 1990: 278 *DR > TL: Boca Chica, 10 m
Polycesta (Polycesta) dominicana Cobos, 1990: 281 *DR > TL: Baní
Polycesta (Polycesta) porcata (Fabricius, 1775). Waterhouse, 1904: 254; Wolcott, 1927: 220; Blackwelder, 1944: 306
Polycesta (Polycesta) regularis Waterhouse, 1904: 256. Blackwelder, 1944a: 306 *DR > TL: St. Domingo

Superfamily BYRRHOIDEA
Family ELMIDAE
Subfamily ELMINAE
Anommatelmis Spangler, 1981
Anommatelmis botosaneanui Spangler, 1981: 377 *H > TL: Well of Presbytère Catholique of Limbé
Cylloepus Erichson, 1847
Cylloepus lahottensis (Darlington, 1936: 81). Blackwelder, 1944a: 271; Jäch et al., 2016: 89; Polizei & Barclay, 2019: 94 *H >
TL: Northeastern foothills of the Massif de la Hotte, 2–4,000 ft.
Hexacylloepus Hinton, 1940
Hexacylloepus haitianus (Darlington, 1936: 80). Blackwelder, 1944a: 271; Jäch et al., 2016: 88 *H > TL: Northeastern
foothills of the Massif de la Hotte, 2–4,000 ft.
Lemalelmis Spangler, 1981
Lemalelmis fontana Spangler, 1981: 383 *H > TL: Tessier, karst spring between Beaucalin and Ravine Blanche
Lemalelmis minyops Spangler, 1981: 381 *H > TL: Jérémie (Département de la Grande Anse), Source Débarasse (karst spring)
Neoelmis Musgrave, 1935
Neoelmis minima (Darlington, 1927). Darlington, 1936: 79
Subfamily LARAINAE
Hispaniolara Brown, 1981
Hispaniolara farri Brown, 1981: 88. Spangler & Fragoso, 1992: 59; Manzo, 2013: 206 *DR > TL: Dist. La Vega, mountain
stream w. Jayaco, elev. 1170 m
Phanocerus Sharp, 1882
Phanocerus clavicornis Sharp, 1882. Spangler & Fragoso, 1992: 13
Phanocerus helmoides Darlington, 1936: 74. Blackwelder, 1944a: 273 *H > TL: Northeastern foothills of the Massif de la
Hotte, southwestern peninsula, 2–4,000 ft.
Phanocerus hubbardi Schaeffer, 1911. Darlington, 1936: 74

Family DRYOPIDAE
Pelonomus Erichson, 1847
Pelonomus insularis Grouvelle, 1896: 33

Family LIMNICHIDAE
Eulimnichus Casey, 1889
Eulimnichus plebius Sharp, 1902. Spangler, et al., 2001: 155
Limnichites Casey, 1889
Limnichites rudis Wooldrige, 1977. Peck, 2005: 103
Limnichoderus Casey, 1889
Limnichoderus barberi Wooldridge, 1981a: 174. Spangler, et al., 2001: 156
Limnichoderus insularis Wooldrige, 1981a: 188. Spangler, et al., 2001: 156
Throscinus LeConte, 1876
Throscinus aethiops Darlington, 1936: 75. Blackwelder, 1944: 273; Wooldridge, 1981b: 220; Spangler, et al., 2001: 159

Family LUTROCHIDAE

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Lutrochus Erichson, 1847
Lutrochus geniculatus Chevrolat, 1864. Darlington, 1936: 75

Family HETEROCERIDAE
Subfamily HETEROCERINAE
Heterocerus Fabricius, 1792
Heterocerus meridianus (Pacheco, 1975). Mascagni & Monte, 2010: 160
Neoheterocerus Pacheco, 1964
Neoheterocerus angustatus (Chevrolat, 1864). Peck, 2005: 104
Tropicus Pacheco, 1964
Tropicus lituratus (Kiesenwetter, 1843). Mascagni & Monte, 2010: 165

Family PSEPHENIDAE
Pheneps Darlington, 1936
Pheneps gracilis Darlington, 1936: 67 *H > TL: Tardieu, in the valley just north of Mt. La Hotte (Pic de Macaya), about 3,000
ft.
Psephenops Grouvelle, 1898
Psephenops haitianus Darlington, 1936: 72. Blackwelder, 1944a: 274 *H > TL: From Tardieu, just north of Mt. La Hotte, about
3,000 ft.

Family CHELONARIIDAE
Chelonarium Fabricius, 1801
Chelonarium beauvoisi Latreille, 1807. Méquignon, 1932: 243

Superfamily ELATEROIDEA
Family BRACHYPSECTRIDAE
Brachypsectra LeConte, 1874
Brachypsectra moronei† Branham in Costa et al., 2006: 430 [Fossil in Dominican amber]
Brachypsectra vivafosile Woodruff, 2004: 161. Costa et al., 2006: 429 *DR > TL: Prov. Pedernales, Cabo Rojo, Alcoa
Headquarters

Family EUCNEMIDAE
Subfamily MELASINAE
Microrhagus Dejean, 1833
Microrhagus albofasciatus (Fisher, 1945). Peck, 2005: 106
Subfamily INCERTAE SEDIS
Lissantauga† Poinar, 2013
Lissantauga epicrana† Poinar, 2013: 111 [Fossil in Dominican amber]

Family ELATERIDAE
Subfamily AGRYPNINAE
Tribe HEMIRHIPINI
Alaus Eschscholtz, 1829
Alaus latipennis Casari, 2004: 322 *DR, *H > Prov. Barahona, nr. Filipinas, Larimar Mine
Alaus nobilis Sallé, 1855: 263. Schenkling, 1925: 50; Blackwelder, 1944: 282; Casari, 1996a: 271 *DR > TL: Rancho Abajo
Alaus patricius (Candèze, 1857). Casari, 1996a: 268
Alaus tricolor (Olivier, 1790). Schenkling, 1925: 51; Blackwelder, 1944: 282; Casari, 1996a: 267
Chalcolepidius Eschscholtz, 1829
Chalcolepidius silbermanni Chevrolat, 1835. Wolcott, 1927: 220; Casari, 2002: 329
Pherhimius Fleutiaux, 1942
Pherhimius fascicularis (Fabricius, 1887). Blackwelder, 1944: 282; Casari, 1996b: 383
Tribe OOPHORINI
Aeolus Eschscholtz, 1829
Aeolus cibaensis Candèze, 1859: 329. Schenkling, 1925: 121; Blackwelder, 1944: 289 *H, *DR > TL: Rivières Yaque et

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Santiago
Aeolus depressus Candèze, 1859: 301. Blackwelder, 1944: 289 *H, *DR > TL: San Juan
Aeolus frivolus Candèze, 1859: 302. Schenkling, 1925: 122; Blackwelder, 1944: 289 *H, *DR > TL: San Juan
Aeolus maculatus (De Geer, 1774). Schenkling, 1925: 123; Blackwelder, 1944: 290
=Aeolus angulatus (Fabricius, 1801). Candèze, 1859: 321; Chevrolat, 1867: 607
Aeolus verruculosus Candèze, 1859: 330. Schenkling, 1925: 126; Blackwelder, 1944: 291 *H, *DR > TL: Nisao [Nizao]
Conoderus Eschscholtz, 1829
Conoderus bellus (Say, 1825). Blackwelder, 1944: 287
Conoderus bifoveatus (Palisot de Beauvois, 1805). Chevrolat, 1867: 603; Schenkling, 1925: 112; Blackwelder, 1944: 287;
Becker, 1975: 270
Conoderus curvifrons (Candèze, 1859: 247). Schenkling, 1925: 114; Blackwelder, 1944: 287 *H, *DR > TL: St. Domingue
Conoderus figularis (Candèze, 1881: 50). Schenkling, 1925: 114; Blackwelder, 1944: 287 *H, *DR > TL: St. Domingue
Conoderus lividus (De Geer, 1774). Schenkling, 1925: 115; Blackwelder, 1944: 288
Conoderus posticus (Eschscholtz, 1822). Peck, 2010: 31
=Monocrepidius sticturus Candèze, 1859: 260. Chevrolat, 1867: 605
Conoderus rufidens (Fabricius, 1801). Becker, 1975: 273
Conoderus sericatus (Candèze, 1859: 249). Schenkling, 1925: 118; Blackwelder, 1944: 288
Conoderus sticticus Erichson, 1847. Schenkling, 1925: 119; Blackwelder, 1944: 288
Conoderus substriatus (Candèze, 1859: 254). Schenkling, 1925: 119; Blackwelder, 1944: 288 *H, *DR > TL: St. Domingue
Tribe PLATYCREPIDIINI
Platycrepidius Candèze, 1859
Platycrepidius latus (Fleutiaux, 1902: 114). Schenkling, 1925: 101; Blackwelder, 1944: 291
Platycrepidius wapleri Sallé, 1855: 267. Schenkling, 1925: 101; Blackwelder, 1944: 291 *H, *DR > TL: Environs de la ville de
Santo-Domingo
Tribe PYROPHORINI
Ignelater Costa, 1975
Ignelater dominicanensis Fernández García & Lozada Piña, 2002: 74 *DR > TL: Río Limpio, Prov. Elías Piña
Ignelater glaesum Costa, 1980: 159
Ignelater havaniensis (Laporte, 1840). Costa, 1980: 160
Ignelater luminosus (Illiger, 1807). Wolcott, 1927: 220; Costa, 1980: 162
Ignelater phosphoreus (Linnaeus, 1758). Costa, 1980: 163
Pyrophorus Illiger, 1809
Pyrophorus lychniferus Germar, 1841. Chevrolat, 1867: 612; Schenkling, 1927: 347; Blackwelder, 1944: 285
Pyrophorus lychnus Candèze, 1863. Blackwelder, 1944: 285
Pyrophorus mellifluus Costa, 1972: 211
Pyrophorus noctilucus (Linnaeus, 1758). Blackwelder, 1944: 286
Subfamily CARDIOPHORINAE
Horistonotus Candèze, 1860
Horistonotus asthenicus Candèze, 1860: 266. Chevrolat, 1867: 610; Schenkling, 1925: 256; Blackwelder, 1944: 302
Horistonotus bignoniae Candèze, 1860: 248. Schenkling, 1925: 257; Blackwelder, 1944: 302 *H, *DR > TL: Sur le roble
(Bignonia longissima) à la Boca de Nigua
Subfamily ELATERINAE
Tribe DICREPIDIINI
Anoplischius Candèze, 1859
Anoplischius punctatus Candèze, 1859: 70. Schenkling, 1925: 80; Blackwelder, 1944: 298
Dicrepidius Eschscholtz, 1829
Dicrepidius ramicornis (Palisot de Beauvois, 1805). Chevrolat, 1867: 600; Casari, 2009: 56; Perez-Gelabert, 2012: 110
Dipropus Germar, 1839
Dipropus antennatus (Candèze, 1859: 108). Chevrolat, 1867: 601; Schenkling, 1925: 88; Blackwelder, 1944: 299
Dipropus aper (Candèze, 1859: 108). Schenkling, 1925: 88; Blackwelder, 1944: 299 *H, *DR > TL: Azua
Tribe PHYSORHININI
Anchastomorphus Champion, 1895
Anchastomorphus phedrus (Candèze, 1859). Schenkling, 1925: 189; Blackwelder, 1944: 301
Tribe SYNAPTINI

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Glyphonyx Candèze, 1863
Glyphonyx bivittatus Candèze, 1863: 455. Blackwelder, 1944: 296 *H, *DR > TL: St. Domingue
Glyphonyx fusculus (Erichson, 1842). Chevrolat, 1867: 613; Blackwelder, 1944: 296
Glyphonyx praevius Erichson, 1842: 121. Blackwelder, 1944: 296 *H, *DR > TL: Port-au-Prince
Glyphonyx recticollis (Say, 1811). Chevrolat, 1867: 614
Subfamily LISSOMINAE
Drapetes Redtenbacher, 1849
Drapetes bicolor Laporte, 1833. Chevrolat, 1867: 590
Drapetes tunicatus Bonvouloir, 1859. Blackwelder, 1944: 304

Family LYCIDAE
Subfamily LEPTOLYCINAE
Tribe LEPTOLYCINI
Cessator Kazantsev, 2012
Cessator brodzinskyi† Ferreira & Ivie, 2017: 58 [Fossil in Dominican amber]
Leptolycus Leng & Mutchler, 1922
Leptolycus (Baholycus) barahonensis Bocak, 2001: 206 *DR > TL: Barahona, 4 km N Paraiso, 150 m
Leptolycus (Baholycus) flavoapicalis Bocak, 2001: 207 *DR > TL: San Cristóbal, 500 m, 20 km NW San Cristóbal,
Manomatuey Valley
Leptolycus (Baholycus) masneri Bocak, 2001: 207 *DR > TL: Barahona, Sierra de Bahoruco, Loma Remigio, 800 m
Leptolycus (Leptolycus) nigrostriatus Bocak, 2001: 205 *DR > TL: Barahona, Sierra de Bahoruco, Loma Remigio, 800 m
Subfamily LYCINAE
Tribe LYCINI
Thonalmus Bourgeois, 1883
Thonalmus chevrolati Bourgeois, 1901. Leng & Mutchler, 1922: 422; Wolcott, 1927: 217; Blackwelder, 1945: 343 *H, *DR >
TL: Saint-Domingue
Thonalmus dominicensis (Chevrolat, 1870). Leng & Mutchler, 1922: 421; Wolcott, 1927: 217; Blackwelder, 1945: 343
Subfamily MIMOLIBNETINAE
Tribe DOMINOPTERINI
Dominopteron Kazantsev, 2013
Dominopteron hispaniolum Kazantsev, 2013a: 15 *DR > TL: Hato Mayor, env. Los Haitises, 8 km W Sabana de la Mar, 20 m
Tribe ELECTROPTERINI
Electropteron† Kazantsev, 2012
Electropteron avus† Kazantsev, 2012: 2; Kazantsev, 2013b: 99 [Fossil in Dominican amber]
Electropteron cursarius Kazantsev, 2013a: 18 *DR > TL: Barahona, E. Sierra Bahoruco, NE Paraiso, 1200 m
Electropteron nepos Kazantsev, 2013a: 18 *DR > TL: Hato Mayor, env. Los Haitises, 8 km W Sabana de la Mar, 20 m
Nanolycus Kazantsev, 2013
Nanolycus gnomus Kazantsev, 2013a: 19 *DR > TL: Hato Mayor, env. Los Haitises, 8 km W Sabana de la Mar, 20 m

Family PHENGODIDAE
Subfamily PHENGODINAE
Tribe PENICILLOPHORINI
Acladocera Wittmer, 1981
Acladocera hispaniolae Wittmer, 1981: 105 *DR > TL: Loma Vieja, S. Constanza, ca. 6000 ft.

Family LAMPYRIDAE
Key reference: Kazantsev & Perez-Gelabert (2009).
Subfamily LAMPYRINAE
Tribe CRATOMORPHINI
Aspisoma Laporte, 1833
Aspisoma ignitum Linnaeus, 1767. Blackwelder, 1944: 356; Peck, 2005: 113; Kazantsev & Perez-Gelabert, 2009: 374
Callopisma Motschulsky, 1853
Callopisma altimontana Kazantsev & Perez-Gelabert, 2009: 376 *DR > TL: Caseta 2, Parque Nacional Sierra de Bahoruco,
Pedernales Prov.

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Callopisma dominicana Kazantsev & Perez-Gelabert, 2009: 376 *DR > TL: Casabito, Constanza, La Vega Prov.
Callopisma engombe Kazantsev & Perez-Gelabert, 2009: 378 *DR > TL: Engombe, Santo Domingo Prov.
Callopisma lamellicornis Kazantsev & Perez-Gelabert, 2009: 378 *DR > TL: La Ciénaga de Manabao, Parque Nacional
Armando Bermúdez, La Vega Prov.
Callopisma larimarena Kazantsev & Perez-Gelabert, 2009: 376 *DR > TL: La Travesia, near Larimar mine, Eastern Sierra de
Bahoruco, Barahona Prov.
Callopisma rubicunda Kazantsev & Perez-Gelabert, 2009: 378 *DR > TL: 1 km SE Caseta 1, Parque Nacional Sierra de
Bahoruco, Independencia Prov.
Callopisma rufa (Olivier, 1790). Leng & Mutchler, 1922: 421; Wolcott, 1927: 218; Blackwelder, 1945: 355; McDermot, 1966:
62
Callopisma rufoviolacea Kazantsev, 2006: 378 *DR > TL: Las Abejas, Sierra de Bahoruco
Erythrolychnia Motschulsky, 1853
Erythrolychnia azuensis Kazantsev & Perez-Gelabert, 2009: 380 *DR > TL: La Poza de Agua Nueva, El Curro, Sierra Martín
García, Azua Prov.
Erythrolychnia bipartita (E. Olivier, 1912: 24). Leng & Mutchler, 1922: 447; Blackwelder, 1945: 357; Kazantsev, 2006: 375
*DR > TL: Puerto Plata
=Erythrolychnia albopalpis Leng & Mutchler, 1922: 448. Blackwelder, 1945: 357; Kazantsev, 2006: 375
=Erythrolychnia olivieri Leng & Mutchler, 1922: 448. Blackwelder, 1945: 358
Erythrolychnia caborojensis Kazantsev & Perez-Gelabert, 2009: 380 *DR > TL: La Charca, km 30 Rd. Cabo Rojo—Aceitillar,
Pedernales Prov.
Erythrolychnia clarki Mutchler, 1923b: 11. Blackwelder, 1945: 358; McDermot, 1966: 73 *H
Erythrolychnia cristobalensis Kazantsev & Perez-Gelabert, 2009: 383 *DR > TL: San Cristóbal, Campus Loyola
Erythrolychnia fulgida (Olivier, 1790). Leng & Mutchler, 1922: 447; Blackwelder, 1945: 358; Kazantsev, 2006: 375
Erythrolychnia marcanoi Kazantsev & Perez-Gelabert, 2009: 382 *DR > TL: Isla Beata
Erythrolychnia medranoae Kazantsev & Perez-Gelabert, 2009: 383. Armas & Madruga, 2018: 278 *DR > TL: Caseta 1,
Parque Nacional Sierra de Bahoruco, Pedernales Prov.
=Erythrolychnia medranoi Kazantsev & Perez-Gelabert, 2009: 383
Erythrolychnia nigriventris Kazantsev, 2006: 375 *DR > TL: Constanza, La Vega Prov.
Erythrolychnia pedernalensis Kazantsev & Perez-Gelabert, 2009: 383 *DR > TL: Km 22 Rd. Cabo Rojo—Aceitillar,
Pedernales Prov.
Erythrolychnia quinquenotata (Laporte, 1840: 269). Wolcott, 1927: 218; Blackwelder, 1945: 359 *H, *DR > TL: Saint-
Domingue
Erythrolychnia roseimargo Kazantsev & Perez-Gelabert, 2009: 385 *DR > TL: Casabito, Reserva Ebano Verde, La Vega Prov.
Erythrolychnia unicolor Kazantsev, 2006: 375 *DR > TL: Cabeza de Agua, Pedernales Prov.
Pyractonema Solier, 1849
Pyractonema vitticollis Motschulsky, 1853: 38. Leng & Mutchler, 1922: 454; Wolcott, 1927: 218; Blackwelder, 1945: 357;
Kazantsev, 2006: 372 *DR, *H > TL: Antilles
Pyractonema watsoni Leng & Mutchler, 1922: 453. Blackwelder, 1945: 355 *DR > TL: San Lorenzo
Tribe LAMPROCERINI
Lychnacris Motschulsky, 1853
Lychnacris atrocrocea Kazantsev & Perez-Gelabert, 2009: 368 *DR > TL: Las Abejas, Sierra de Bahoruco, Pedernales Prov.
Lychnacris bahorucoensis Kazantsev & Perez-Gelabert, 2009: 370 *DR > TL: Las Abejas, Sierra de Bahoruco, Pedernales
Prov.
Lychnacris cienagaensis Kazantsev & Perez-Gelabert, 2009: 370 *DR > TL: La Ciénaga de Manabao, La Vega Prov.
Lychnacris hierroi Kazantsev & Perez-Gelabert, 2009: 370 *DR > TL: Rd. Loma del Toro—Puerto Escondido, Independencia
Prov.
Lychnacris konstantinovi Kazantsev, 2006: 372 *DR > TL: 15 km N Cabo Rojo, Pedernales Prov.
Lychnacris mariposa (Leng & Mutchler, 1922: 442). Blackwelder, 1945: 355; McDermot, 1966: 61; Kazantsev, 2006: 390 *DR
> TL: Sánchez
Lychnacris montensis Kazantsev & Perez-Gelabert, 2009: 370 *DR > TL: Monteada Nueva, Barahona Prov.
Lychnacris neslihanae Kazantsev, 2006: 372 *DR > TL: El Guineo, San Cristóbal Prov.
Lychnacris orbis Kazantsev & Perez-Gelabert, 2009: 371 *DR > TL: Boca de Yuma, Parque Nacional del Este, La Altagracia
Prov.
Lychnacris pedernalis Kazantsev, 2006: 372 *DR > TL: 15 km N Cabo Rojo, Pedernales Prov.

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Lychnacris piceonotata Kazantsev & Perez-Gelabert, 2009: 371 *DR > TL: Pirámide 204, Hondo Valle, Elías Piña Prov.
Lychnacris postica (E. Olivier, 1899: 90). Leng & Mutchler, 1922: 444; Blackwelder, 1945: 355; McDermot, 1966: 62;
Kazantsev, 2006: 390
Lychnacris rufocaerulea Kazantsev & Perez-Gelabert, 2009: 372 *DR > TL: La Compartición—Los Tablones, Parque
Nacional Armando Bermúdez, La Vega Prov.
Lychnacris scintilla Kazantsev & Perez-Gelabert, 2009: 373 *DR > TL: ~100 m up from El Sitio del Agua, N of Los Bolos,
Sierra de Neiba, Independencia Prov., 1520 m
Lyhnacris vulpes Kazantsev & Perez-Gelabert, 2013: 267 *DR > TL: Azua Prov., Las Yayitas
Tribe PHOTININI
Heterophotinus E. Olivier, 1894
Heterophotinus alius Kazantsev, 2006: 383 *DR > TL: Los Arroyos, Pedernales Prov.
Heterophotinus constanzae Kazantsev, 2006: 384 *DR > TL: Constanza, La Vega Prov.
Heterophotinus glaucus (Olivier, 1790). Leng & Mutchler, 1922: 460; Blackwelder, 1945: 359; McDermot, 1966: 51
Heterophotinus lengi (Mutchler, 1923a: 2). McDermot, 1966: 51; Blackwelder, 1945: 358
Heterophotinus limpioensis Kazantsev, 2006: 385 *DR > TL: Nalga de Maco, SE Río Limpio, Elías Piña Prov.
Heterophotinus monticola Kazantsev & Perez-Gelabert, 2009: 392 *DR > TL: Cortico, Monteada Nueva, Barahona Prov.
Heterophotinus merielae Kazantsev, 2006: 387 *DR > TL: Nalga de Maco, SE Río Limpio, Elías Piña Prov.
Heterophotinus nigricollis Kazantsev, 2006: 385 *DR > TL: Las Abejas, Pedernales Prov.
Heterophotinus nubilus Kazantsev & Perez-Gelabert, 2009: 392 *DR > TL: Cortico, Monteada Nueva, Barahona Prov.
Heterophotinus quadrimaculatus (Laporte, 1840: 269). Leng & Mutchler, 1922: 473; Blackwelder, 1945: 359; McDermot,
1966: 52; Kasantsev, 2006: 391
=Callopisma interrupta (Motschulsky, 1854). McDermot, 1966: 61
=Photinus divisus Gemminger, 1870. McDermot, 1966: 61
Heterophotinus quadrinotatus (Motschulsky, 1854). Kazantsev, 2006: 387 *DR > TL: Santo Domingo
Heterophotinus striatus Kazantsev & Perez-Gelabert, 2009: 392 *DR > TL: La Toma, San Cristóbal Prov.
Heterophotinus viridicolor Kazantsev, 2006: 387 *DR > TL: Nalga de Maco, SE Río Limpio, Elías Piña Prov.
Heterophotinus vittatus (Olivier, 1790). Leng & Mutchler, 1922: 478; Blackwelder, 1945: 359; McDermot, 1966: 53;
Kasantsev, 2006: 391
Microdiphot Barber, 1941
Microdiphot baorucoensis Kazantsev, 2006: 387 *DR > TL: Los Arroyos, Pedernales Prov.
Robopus Motschulsky, 1853
Robopus acutangulus Kazantsev & Perez-Gelabert, 2009: 387 *DR > TL: Matadero, Honduras, Peravia Prov.
Robopus bastardoae Kazantsev & Perez-Gelabert, 2009: 387. Armas & Madruga, 2018: 278 *DR > TL: Frente a la charca,
Cortico, Barahona Prov.
=Robopus bastardoi Kazantsev & Perez-Gelabert, 2009: 387
Robopus branhami Kazantsev, 2006: 380 *DR > TL: Los Arroyos, Pedernales Prov.
Robopus dissimilis Kazantsev & Perez-Gelabert, 2009: 389 *DR > TL: Cruce de Ocoa, Peravia Prov.
Robopus erythrolytris Kazantsev, 2006: 380 *H > TL: Cap Haitien
Robopus hondovallensis Kazantsev & Perez-Gelabert, 2009: 389 *DR > TL: Pirámide 204, Hondo Valle, Elías Piña Prov.
Robopus kasikus Kazantsev, 2006: 380 *H > TL: Diquini
Robopus nigrifrons Kazantsev & Perez-Gelabert, 2009: 389 *DR > TL: Jardín Botánico Nacional, Santo Domingo Prov.
Robopus oviedoensis Kazantsev & Perez-Gelabert, 2013: 269 *DR > TL: Pedernales Prov., ~2 km way to Alto de la Rosa, 90 m
Robopus peregrinus Kazantsev & Perez-Gelabert, 2009: 390 *DR > TL: La Malanga, Paraiso, Barahona Prov.
Robopus vallinovae Kazantsev & Perez-Gelabert, 2009: 391 *DR > TL: Valle Nuevo, La Vega Prov.
Robopus velozae Kazantsev & Perez-Gelabert, 2013: 268. Armas & Madruga, 2018: 278*DR > TL: Pedernales Prov., 3 km S
Oviedo, km 1 trail Fondo Paradí, 120 m
=Robopus velozi Kazantsev & Perez-Gelabert, 2013: 268
Subfamily PHOTURINAE
Tribe PHOTURINI
Presbyolampis Buck, 1947
Presbyolampis vegaensis Kazantsev, 2006: 389 *DR > TL: La Ciénaga, La Vega Prov.
Presbyolampis mirabilis Kazantsev & Perez-Gelabert, 2009: 399 *DR > TL: Las Abejas, Sierra de Bahoruco, Pedernales Prov.
Subfamily Cheguevariinae
Cheguevaria Kazantsev, 2006

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 131
Cheguevaria angusta Kazantsev, 2006: 372 *DR > TL: 15 km N of Azua

Family CANTHARIDAE
Subfamily SILINAE
Tribe SILINI
Caccodes Sharp, 1885
Caccodes bimaculatus Wittmer, 1991a: 88 *DR > TL: La Vega, 24 km SE Constanza
Caccodes dominicanus Wittmer, 1991a: 90 *DR > TL: Constanza, 3–4000’
Caccodes malthiniformis Wittmer, 1991a: 85 *DR > TL: Colonia, 1000 m
Caccodes rugiceps Wittmer, 1991a: 87 *DR > TL: La Vega, 22 km SE Constanza
Caccodes singularis Wittmer, 1991a: 90 *DR > TL: Constanza, 3–4000’
Caccodes testaceomimus Wittmer, 1980: 464 *DR > TL: Mt. Diego de Ocampo, 3–4000 ft.
Caccodes testaceus Wittmer, 1980: 464 *DR > TL: Constanza, 3–4000 ft.
Polemius LeConte, 1851
Polemius hispaniolae Leng & Mutchler, 1922: 493. Wolcott, 1927: 218; Blackwelder, 1945: 366 *DR > TL: San Francisco
Mts.
Silidiscodon Leng & Mutchler, 1922
Silidiscodon lavagaensis Wittmer, 1998: 314 *DR > TL: La Vega, between Constanza and Salto de Aguas Blancas
Silidiscodon rufiventre Leng & Mutchler, 1922: 493. Blackwelder, 1945: 366 *DR > TL: Romana
Silidiscodon thomasi Wittmer, 1998: 313 *H > TL: Dept. Sud-Oueste: Parc National La Visite, SE slope, 2040–2150 m
Silis Charpentier, 1825
Silis acutispina Wittmer, 1991a: 60 *DR > TL: Constanza, 900–2000 m
Silis argenteosetosa Wittmer, 1991a: 65 *DR > TL: Loma Vieja, S Constanza, ca. 6000’
Silis azurea Wittmer, 1957: 153 *H > TL: Port-au-Prince
Silis debililimbata Wittmer, 1991a: 65 *H > TL: Mt. La Hotte, 5–7800’
Silis dominicana Wittmer, 1955: 270 *DR > TL: Santo Domingo, La Cavite
Silis flavofemorata Wittmer, 1991a: 74 *DR > TL: Valle Nuevo, 2400 m
Silis flavolobata Wittmer, 1991a: 71 *DR > TL: Valle Nuevo, 2400 m
Silis gracilispina Wittmer, 1991a: 63 *H > TL: Morne Guimby, 22 km SE Fond Verrettes, ca. 2000 m
Silis (Hapalocrosilis) flexilobata Wittmer, 1991a: 78 *DR > TL: Colonia, 1000 m
Silis (Hapalocrosilis) haitiana Wittmer 1957: 152
Silis (Hapalocrosilis) parallelilobata Wittmer, 1991a: 81 *DR > TL: Constanza, 1250 m
Silis (Hapalocrosilis) rufithorax Wittmer, 1991a: 83 *H > TL: Morne Guimby, 22 km SE Fond Verrettes, ca. 2000 m
Silis (Hapalocrosilis) violaceotincta Wittmer, 1956: 224 *H > TL: Port-au-Prince
Silis nigrina Wittmer, 1991a: 69 *DR > TL: Valle Nuevo, 2400 m
Silis superba Wittmer, 1998: 314 *DR > TL: Prov. Pedernales: Parque Nac. Baoruco, 1290–1335 m
Silis triangulifera Wittmer, 1991a: 60 *DR > TL: ca. 5 km S Constanza (Prov. La Vega), 1250 m
Silis tricolorata Wittmer, 1991a: 73 *H > TL: Morne Guimby, 22 km SE Fond Verrettes, ca. 2000 m
Tylocerus Dalman, 1823
Tylocerus basitestaceus Wittmer, 1998: 313 *H > TL: Dept. Sud-Oueste: Massif de La Selle, 1850 m, Morne d’Enfer
Tylocerus coriaceus Leng & Mutchler, 1922: 496. Mutchler, 1923b: 9; Blackwelder, 1945: 363 *DR > TL: Sánchez
Tylocerus discoviridis Pic, 1919. Blackwelder, 1945: 363 *H > TL: Haïti
Tylocerus klapperichi Wittmer, 1975: 253 *DR > TL: San Cristóbal, 35 m
Tylocerus purpureus Wittmer, 1975: 254 *DR > TL: La Colonia, 1000 m
Tribe TYTTHONYXINI
Tytthonyx LeConte, 1851
Tytthonyx (Thinalmus) antillarum Wittmer, 1957: 155 *H > TL: Port-au-Prince
Tytthonyx (Thinalmus) haitiensis Wittmer, 1957: 155. Wittmer, 1991b: 118 *H, *DR > TL: Port-au-Prince
Tytthonyx (Thinalmus) hispaniolensis Wittmer, 1991b: 121 *DR > TL: San José de las Matas, 1000–2000 ft.
Tytthonyx (Thinalmus) iviei Wittmer, 1991b: 118 *DR > TL: Prov. Pedernales, 13.5 km N Cabo Rojo, 140 m
Tytthonyx (Thinalmus) neodarlingtoni Wittmer, 1991: 126 *DR > TL: San José de las Matas, 1000–2000 ft.
Tytthonyx (Tytthonyx) armatus Wittmer, 1992: 340 *DR > TL: Constanza to Valle Nuevo, 3–7000’
Tytthonyx (Tytthonyx) baniensis Wittmer, 1992: 375 *DR > TL: Baní, 65 m
Tytthonyx (Tytthonyx) barahonaensis Wittmer, 1998: 316 *DR > TL: Barahona Prov.: near Filipinar, Larimar Mine

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Tytthonyx (Tytthonyx) brunneus Wittmer, 1992: 361 *DR > TL: Baní, 65 m
Tytthonyx (Tytthonyx) coriaceipennis Wittmer, 1998: 315 *H > TL: Dept. Sud-Oueste: Massif de La Selle, Morne d’Enfer, 1850
m
Tytthonyx (Tytthonyx) filamentarius Wittmer, 1992: 363 *DR > TL: Baní, 65 m
Tytthonyx (Tytthonyx) geiseri† Poinar & Fanti, 2016: 5 [Fossil in Dominican amber]
Tytthonyx (Tytthonyx) larimarensis Wittmer, 1998: 316 *DR > TL: Barahona Prov.: near Filipinar, Larimar Mine
Tytthonyx (Tytthonyx) lasellensis Wittmer, 1992: 355 *H > TL: La Visite & vic. (La Selle Range), 5–7000’
Tytthonyx (Tytthonyx) longilobus Wittmer, 1992: 363 *H > TL: Morne Gumby, 22 km SE Fond Verretes, Forêt des Pins, 6500’
Tytthonyx (Tytthonyx) mutabilis Wittmer, 1992: 346 *DR > TL: Santo Domingo, 30 m
Tytthonyx (Tytthonyx) republicanus Wittmer, 1992: 344 *DR > TL: Loma Rucilla and Mts. N, 5–8000 ft.
Tytthonyx (Tytthonyx) spinosus Wittmer, 1992: 352 *DR > TL: Loma Vieja (S Constanza), ca. 6000’
Tytthonyx (Tytthonyx) trifilum Wittmer, 1992: 364 *DR > TL: Mt. Diego de Ocampo, 3–4000’
Tytthonyx (Tytthonyx) valentinei Wittmer, 1992: 342 *H > TL: Morne Gumby, 22 km SE Fond Verretes, Forêt des Pins, 6500’

Infraorder BOSTRICHIFORMIA
Superfamily BOSTRICHOIDEA
Family DERMESTIDAE
Subfamily ATTAGENINAE
Attagenus Latreille, 1802
Attagenus ambericus† Háva & Prokop, 2004: 175 [Fossil in Dominican amber]
Attagenus electron† Poinar & Háva, 2015: 2 [Fossil in Dominican amber]
Limniattagenus Poinar & Háva, 2015
Limniattagenus electron† Poinar & Háva, 2015: 2 [Fossil in Dominican amber]
Subfamily DERMESTINAE
Dermestes Linnaeus, 1758
Dermestes (Dermestes) ater De Geer, 1774. Háva & Kadej, 2006a: 376
Dermestes (Dermestinus) caninus caninus Germar, 1824. Háva & Kadej, 2006a: 376
Dermestes (Dermestinus) maculatus De Geer, 1774. Háva & Kadej, 2006a: 376
Subfamily MEGATOMINAE
Amberoderma† Háva & Prokov, 2004
Amberoderma beali† Háva & Prokop, 2004: 174; Poinar & Háva, 2015: 2 [Fossil in Dominican amber]
Caccoleptus Sharp, 1902
Caccoleptus (Bicaccoleptus) prokopi† Poinar & Háva, 2015: 7 [Fossil in Dominican amber]
Caccoleptus (Caccoleptus) electron† Poinar & Háva, 2015: 7 [Fossil in Dominican amber]
Cryptorhopalum Guérin-Méneville, 1838
Cryptorhopalum acevedoi Kadej & Háva, 2013: 2 *DR > TL: El Cadillar, Reserva Loma Quita Espuela
Cryptorhopalum ambericum† Háva & Prokop, 2004: 176 [Fossil in Dominican amber]
Cryptorhopalum davidsoni Kadej & Háva, 2013: 5 *DR > TL: 2.9 km SW Boca de Yuma
Cryptorhopalum delacruzi Kadej & Háva, 2013: 7 *DR > TL: Aceitillar, 25.2 km ENE Pedernales
Cryptorhopalum dominicanum† Háva & Prokop, 2004: 179 [Fossil in Dominican amber]
Cryptorhopalum jantaricum† Háva & Prokop, 2004: 176; Poinar & Háva, 2015: 2 [Fossil in Dominican amber]
Cryptorhopalum kaliki† Poinar & Háva, 2015: 7 [Fossil in Dominican amber]
Cryptorhopalum macieji† Poinar & Háva, 2015: 2 [Fossil in Dominican amber]
Cryptorhopalum rawlinsoni Kadej & Háva, 2013: 9 *DR > TL: 5.8 km SW Neiba, eastern playa of Lago Enriquillo
Cryptorhopalum thompsonii Kadej & Háva, 2013: 10. Háva & Hermann, 2017: 11 *DR, *H > TL: Cabo Rojo, in swimming
pool
Cryptorhopalum youngi Kadej & Háva, 2013: 11 *DR > TL: Aceitillar, 25.2 km ENE Pedernales
Trogoderma Latreille, 1829
Trogoderma anthrenoides (Sharp, 1902). Háva & Kadej, 2006b: 380 [Introduced]
Trogoderma bicinctum Reitter, 1881. Háva & Kadej, 2006b: 383 [Introduced]
Trogoderma davidsoni Háva & Kadej, 2006b: 382 *DR > TL: Pedernales, Cabo Rojo, sea level
Trogoderma dominicana Háva & Kadej, 2006b: 382 *DR > TL: Elias Piña, Sierra de Neiba, 9 km WSW Hondo Valle, 1843 m
Trogoderma fantastica Háva & Kadej, 2006b: 381 *DR > TL: Pedernales, 9.5 km N. Cabo Rojo, 35 m
Trogoderma granarium Everts, 1898. Zuluaga & Mesa, 2000: 197 [Introduced]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 133
Subfamily TRINODINAE
Apsectus LeConte, 1854
Apsectus sp.†—Poinar & Háva (2015: 9) [Fossil in Dominican amber]

Family BOSTRICHIDAE
Subfamily BOSTRICHINAE
Apate Fabricius, 1775
Apate monachus Fabricius, 1775. Santoro, 1960: 125; Marcano, 1981: 101 [Introduced]
=Apate francisca (Fabricius, 1775). Smith & Audant, 1930: 974
Dendrobiella Casey, 1898
Dendrobiella sericea Mulsant & Wachanru, 1852. Lesne, 1901: 493; Blackwelder, 1945: 400; Borowski & Wegrzynowicz,
2007: 126
Discoclavata† Poinar, 2013
Discoclavata dominicana† Poinar, 2013: 108 [Fossil in Dominican amber]
Tetrapriocera Horn, 1878
Tetrapriocera longicornis (Olivier, 1795). Blackwelder, 1945: 400; Fisher, 1950: 102; Borowski & Wegrzynowicz, 2007: 135
Tetrapriocera tridens Fabricius, 1792. Lesne, 1901: 483
Xylomeira Lesne, 1801
Xylomeira tridens (Fabricius, 1792). [By synonymy of species below]
=Xylomeira torquata (Fabricius, 1801). Lesne, 1901: 503; Blackwelder, 1945: 400; Fisher, 1950: 120 Spilman, 1971: 4
Subfamily DINODERINAE
Dinoderus Stephens, 1830
Dinoderus minutus (Fabricius, 1775). Leng & Mutchler, 1914: 435; Blackwelder, 1945: 398; Spilman, 1971: 3 [Introduced]
Rhizoperta Stephens, 1830
Rhizoperta dominica (Fabricius, 1775). SEA, 1999: 129
Subfamily LYCTINAE
Lyctus Fabricius, 1792
Lyctus brunneus Stephens, 1830. Blackwelder, 1945: 400
Lyctus caribeanus Lesne, 1931. Gerberg, 1957: 19; Spilman, 1971: 6; Borowski & Wegrzynowicz, 2007: 31
Minthea Pascoe, 1866
Minthea rugicollis (Walker, 1856). Blackwelder, 1945: 401; Gerberg, 1957: 33; Borowski & Wegrzynowicz, 2007: 37
Trogoxylon LeConte, 1862
Trogoxylon aequale (Wollaston, 1867). Gerberg, 1957: 37
Trogoxylon rectangulum Lesne, 1921: 229. Blackwelder, 1945: 401; Borowski & Wegrzynowicz, 2007: 43
Subfamily POLYCAONINAE
Melalgus Dejean, 1833
Melalgus gonagrus (Fabricius, 1798). Blackwelder, 1945: 398; Borowski & Wegrzynowicz, 2007: 53

Family ANOBIIDAE
Subfamily ANOBIINAE
Stegobium Motschulsky, 1860
Stegobium paniceum (Linnaeus, 1758). Abud, 1981: 16/81
Subfamily DORCATOMINAE
Protheca LeConte, 1865
Protheca cinerea White, 1979: 9
Subfamily PTININAE
Electrognostus† Philips & Mynhardt, 2011
Electrognostus intermedius† Philips & Mynhardt, 2011: 41 [Fossil in Dominican amber]
Okamninus Mynhardt & Philips, 2013
Okamninus annae† Mynhardt & Philips, 2013: 26 [Fossil in Dominican amber]
Oviedinus Bellés, 2010
Oviedinus hispaniolensis† Bellés, 2010: 81 [Fossil in Dominican amber]
Ptinus Linnaeus, 1766
Ptinus hispaniolaensis Philips & Smiley, 2010: 567 *DR > TL: Pedernales Prov., Sierra Baoruco, 31 km N. Cabo Rojo, 2500’

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Ptinus niveicollis Boieldieu, 1856. Blackwelder, 1945: 402
Subfamily TRICORYNINAE
Lasioderma Stephens, 1835
Lasioderma serricorne (Fabricius, 1792). Wolcott, 1927: 224; Smith & Audant, 1930: 974; Marcano, 1964: 127
Tricorynus Waterhouse, 1849
Tricorynus neltumae (Fisher, 1942). White, 1965: 287
Tricorynus palmarum (Guérin-Méneville, 1851). Blackwelder, 1945: 405
Tricorynus sallei (Guérin-Méneville, 1851). Blackwelder, 1945: 406

Infraorder CUCUJIFORMIA
Superfamily LYMEXYLOIDEA
Family LYMEXYLIDAE
Atractocerus Palisot de Beauvois, 1801
Atractocerus brasiliensis Lepeletier & Audinet-Serville, 1825. Bastardo, 2007: 324
Atractocerus sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]

Superfamily CLEROIDEA
Family TROGOSSITIDAE
Subfamily LOPHOCATERINAE
Antillipeltis Lawrence et al., 2014
Antillipeltis alleni† Lawrence et al., 2014: 445 [Fossil in Dominican amber]
Antillipeltis darlingtoni Lawrence et al., 2014: 439 *H > TL: Kenscoff (near Port-au-Prince)
Antillipeltis iviei† Lawrence et al., 2014: 445 [Fossil in Dominican amber]
Antillipeltis maculata Lawrence et al., 2014: 443 *DR > TL: Foothills of Cordillera Central, S. of Santiago
Antillipeltis minuta Lawrence et al., 2014: 443 *DR > TL: Azua
Antillipeltis pubescens Lawrence et al., 2014: 444 *DR > TL: Loma Vieja, S. Constanza, c. 6000 ft.
Subfamily TROGOSSITINAE
Parallelodera Fairmaire, 1881
Parallelodera parallela (Fairmaire, 1850). Blackwelder, 1945: 394
Temnoscheila Westwood, 1830
Temnoscheila parva (Léveillé, 1888: 417). Leng & Mutchler, 1914: 422; Blackwelder, 1945: 393
Tenebroides Piller & Mitterpacher, 1783
Tenebroides floridanus Schaeffer, 1918. Barron, 1971: 111; Peck, 2005: 123

Family CLERIDAE
Subfamily CLERINAE
Arawakis† Opitz, 2007
Arawakis poinari† Opitz, 2007: 343 [Fossil in Dominican amber]
Subfamily EPIPHLOEINAE
Amboakis Opitz, 2006
Amboakis linitis Opitz, 2006: 132 *DR > TL: Mt. Diego de Ocampo, 3–4000 ft.
Decorosa Opitz, 2008
Decorosa aladecoris Opitz, 2008b: 6 *DR > TL: La Vega, Constanza, 1160 m
Decorosa iviei Opitz, 2008b: 8 *DR > TL: Prov. San Juan, Pico Duarte, weather sta., 2885 m
Decorosa limatula Opitz, 2008b: 8 *DR > TL: Loma Vieja, S. Constanza
Decorosa neiba Opitz, 2008b: 11 *DR > TL: Elías Piña, Sierra de Neiba, 9.1 km WSW Hondo Valle, 1856 m
Madoniella Pic, 1935
Madoniella adona Opitz, 2011: 156 *DR > TL: Mt. Diego de Ocampo, 4–4000 ft.
Madoniella basilaris Opitz, 2011: 161 *DR > TL: Prov. La Vega, 1 km NW Manabao
Madoniella bullalis Opitz, 2011: 164 *DR > TL: Monte Cristi Prov., 8.6 km N Villa Elisa
Madoniella cavina Opitz, 2011: 204 *DR > TL: Pedernales, 23.5 km N Cabo Rojo, 540 m
Madoniella cerviculina Opitz, 2011: 206 *DR > TL: Pedernales, 5.3 km NE Los Arroyos, 1540 m
Madoniella cymatilis Opitz, 2011: 207 *DR > TL: Cap Haitien, Dept. du Nord
Madoniella ebena Opitz, 2011: 208 *DR > TL: Hato Mayor, Parque Los Haitises, 3 km W Cueva de Arena, 20 m

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 135
Madoniella infula Opitz, 2011: 222 *DR > TL: La Vega, Cordillera Central, 4.1 km SW El Convento, 1733 m
Madoniella lurida Opitz, 2011: 209 *DR > TL: Pedernales, La Abeja, 38 km NNW Cabo Rojo, 1250 m
Pericales Opitz, 2008
Pericales albogilvus Opitz, 2008a: 33 *H, *DR > TL: Massif de la Selle, Morne d’Enfer
Subfamily KORYNETINAE
Corinthiscus Fairmaire & Germain, 1861
Corinthiscus velutinus Klug, 1842: 358. Blackwelder, 1945: 391
Necrobia Olivier, 1795
Necrobia rufipes (De Geer, 1775). SEA, 1999: 128 [Introduced]
Pelonium Spinola, 1844
Pelonium caborojo Opitz, 2018: 471 *DR > TL: Pedernales, Cabo Rojo, 10 m
Pelonium condylum Opitz, 2018: 450 *DR > TL: Pedernales, 26 km N Cabo Rojo, 730 m
Pelonium denotatum Opitz, 2018: 452 *DR > TL: Pedernales Prov., PN Sierra de Bahoruco, Las Abejas, 1150 m
Pelonium inagua Opitz, 2018: 419
Pelonium murinum (Klug, 1842: 358). Blackwelder, 1945: 391 *H > TL: Port-au-Prince
Pelonium rallum Opitz, 2018: 457 *DR > TL: Baoruco, 5.8 km SW Neiba, eastern playa of Lago Enriquillo, 5 m
Pelonium woodruffi Opitz, 2018: 460 *DR > TL: Prov. Dajabón, 5 km N Dajabón, and 3 km E of Cañongo, 200 ft.
Subfamily TILLINAE
Onychotillus Chapin, 1945
Onychotillus cubana De Zayas, 1988. Burke et al., 2015: 23; Burke & Zolnerowich, 2017: 138
Onychotillus vittatus Chapin, 1945. Burke et al., 2015: 23; Burke & Zolnerowich, 2017: 146

Family MELYRIDAE
Subfamily MALACHIINAE
Ablechroides Wittmer, 1976
Ablechroides flabellatus Wittmer, 1976: 286 *DR > TL: Baní
Ablechroides haitianus Wittmer, 1984: 233 *H > TL: Port-au-Prince
Ablechroides sarasini Wittmer, 1976: 287. Wittmer, 1999: 228 *DR > TL: La Colonia, 1000 m
Ablechrus Waterhouse, 1877
Ablechrus cameroni (Wittmer, 1957: 157). Wittmer, 1961: 36 *H > TL: Chester Vale [Incertae sedis]
Ablechrus dentatus Wittmer, 1976: 284
Ablechrus grandis Wittmer, 1999: 227 *H > TL: Parc National La Visite, vicinity Park headquarters, Dept. Sud-Ouest
Ablechrus klapperichi Wittmer, 1976: 285 *DR > TL: Baní, 65 m
Ablechrus opacicollis Wittmer, 1976: 284 *H > TL: Kenscoff Pass, 800–2000 m
Ablechrus? testaceus Wittmer, 1976: 286 *DR > TL: La Colonia, 1000 m
Ablechrus testaceovarius Wittmer, 1999: 224 *H > TL: Morne La Visite, Dept. de l’Ouest
Ablechrus woodruffi Wittmer, 1999: 227 *DR > TL: Larimar mine, near Filipinas, Barahona Prov.
Attalomimus Wittmer, 1976
Attalomimus cephalotes Wittmer, 1976: 289 *DR > TL: La Colonia, 1000 m
Collops Erichson, 1840
Collops ludicrus Erichson, 1840: 60. Champion, 1914: 39; Blackwelder, 1945: 376 *H, *DR > TL: Port-au-Prince
Lemphus Erichson, 1840
Lemphus viridis Wittmer, 1957: 156 *H > TL: Port-au-Prince
Opsablechrus Wittmer, 1967
Opsablechrus dominicanus Wittmer, 1976: 288 *DR > TL: Baní
Opsablechrus politus Wittmer, 1967: 431 *H > TL: Port-au-Prince

Superfamily CUCUJOIDEA
Family SPHINDIDAE
Carinisphindus McHugh, 1990
Carinisphindus leptosphinctos McHugh, 1990: 317 *DR > TL: Samaná

Family NITIDULIDAE
Subfamily CARPOPHILINAE

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Carpophilus Stephens, 1829
Carpophilus dimidiatus (Fabricius, 1792). SEA, 1999: 105
Carpophilus hemipterus Linnaeus, 1758. Blackwelder, 1945: 412; SEA, 1999: 128
Carpophilus mutilatus Erichson, 1843. Peck, 2010: 36
Subfamily CILLAEINAE
Cillaeus Laporte, 1835
Cillaeus insularis Grouvelle, 1898: 350. Blackwelder, 1945: 410 *H, *DR > TL: St-Domingue
Conotelus Erichson, 1843
Conotelus zayasi Jelínek & Evans, 1982: 231
Subfamily NITIDULINAE
Camptodes Erichson, 1843
Camptodes communis Cline, 2003: 101
Pallodes Erichson, 1843
Pallodes ruficollis Reitter, 1873. Peck, 2005: 128
Pocadius Erichson, 1843
Pocadius dominicus Cline, 2008: 54 *DR > TL: Prov. Hato Mayor, Par. Nac. Los Haitises
Stelidota Erichson, 1843
Stelidota ruderata Erichson, 1843. Chevrolat, 1864: 604; Blackwelder, 1945: 413
Subfamily PROMETOPINAE
Palaeometopia† Kirejtshuk in Kirejtshuk & Poinar, 2007
Palaeometopia colorata† Kirejtshuk & Poinar, 2007: 639 [Fossil in Dominican amber]
Palaeometopia dominicana† Kirejtshuk & Poinar, 2007: 637 [Fossil in Dominican amber]

Family CYBOCEPHALIDAE
Cybocephalus Erichson, 1844
Cybocephalus sp.—Smith & Cave (2007: 164); Perez-Gelabert, 2019: 525

Family SMICRIPIDAE
Smicrips LeConte, 1878
Smicrips exilis (Murray, 1864). Blackwelder, 1945: 424; Price, 2002: 317
=Tisiphone nitiduloides Reitter, 1876: 299

Family MONOTOMIDAE
Subfamily RHIZOPHAGINAE
Europs Wollaston, 1854
Europs maculata Grouvelle, 1896: 210. Blackwelder, 1945: 424

Family SILVANIDAE
Subfamily BRONTINAE
Brontes Fabricius, 1801
Brontes sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Platamus Erichson, 1846
Platamus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]
Telephanus Erichson, 1846
Telephanus consimilis Nevermann, 1937: 13. Blackwelder, 1945: 422 *H > TL: Miragoâne
Telephanus declivis Nevermann, 1937: 12. Blackwelder, 1945: 422 *H > TL: North-east foothills La Hotte, 2–4000 ft.
Telephanus dentatus Nevermann, 1937: 8. Blackwelder, 1945: 422 *H > TL: Port-au-Prince, Swamps north of Dessalines
Telephanus haitianus Nevermann, 1932: 32. Blackwelder, 1945: 422 *H > TL: Port-au-Prince
Telephanus nigrolateralis Nevermann, 1937: 14. Blackwelder, 1945: 422 *H, *DR > TL: Haiti, Santo Domingo
Telephanus sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Telephanus tabaciphilus Nevermann, 1937: 5. Blackwelder, 1945: 423 *H, *DR > TL: Haiti, Santo Domingo
Subfamily SILVANINAE
Ahasverus Gozis, 1881
Ahasverus advena (Waltl, 1832). Wolcott, 1927: 221 [Introduced]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 137
Catharthus Reiche, 1854
Catharthus quadricollis Guérin-Méneville, 1844. Thomas, 1993: 35
Catharthosilvanus Grouvelle, 1912
Catharthosilvanus opaculus (LeConte, 1854). Peck, 2005: 130
Monanus Sharp, 1879
Monanus concinnulus (Walker, 1858). Thomas, 1993: 31 [Introduced]
Nausibius Lentz, 1857
Nausibius clavicornis (Kugelann, 1794). Wolcott, 1927: 221; Blackwelder, 1945: 421; Thomas, 1993: 16
=Nausibius intermedius Smith, 1851. Blackwelder, 1945: 421
Orizaephilus Ganglbauer, 1899
Orizaephilus surinamensis (Linnaeus, 1758). Wolcott, 1927: 221 [Introduced]
Silvanoprus Reitter, 1911
Silvanoprus scuticollis (Walker, 1859). Thomas, 1993: 29 [Introduced]

Family PASSANDRIDAE
Catogenus Westwood, 1830
Catogenus cayman Slipinsky, 1989. Thomas, 1995: 476
Catogenus klapperichorum Burckhardt & Slipinski, 1995: 79 *DR > TL: Baní
=Catogenus slipinskii Thomas, 1995: 476

Family LAEMOPHLOEIDAE
Charaphloeus Casey, 1916
Charaphloeus bituberculatus (Reitter, 1876). Thomas, 1993: 64; Peck, 2005: 132
Cryptolestes Ganglbauer, 1899
Cryptolestes ferrugineus (Stephens, 1831). SEA, 1999: 128
Cryptolestes pusillus (Schönherr, 1817). Peck, 2010: 39
=Laemophloeus minutus (Olivier, 1791). Marcano, 1964: 147; SEA, 1999: 128
Cryptolestes uncicornis (Reitter, 1876). Peck, 2010: 39
Laemophloeus Dejean, 1835
Laemophloeus lecontei Grouvelle, 1876. Thomas, 2014: 10
Lathropus Erichson, 1846
Lathropus parvulus Grouvelle, 1878. Thomas, 2010: 3
Lathropus pictus Schwarz, 1878. Thomas, 2010: 3
Lathropus rhabdophloeoides Thomas, 2010: 18
Placonotus MacLeay, 1871
Placonotus majus Lefkovitch, 1962. Thomas, 1984: 5
Placonotus politissimus (Wollaston, 1867). Thomas, 1984: 6

Family PHALACRIDAE
Acylomus Sharp, 1888
Acylomus latisternus (Guillebeau, 1894: 303). Leng & Mutchler, 1914: 409; Blackwelder, 1945: 430 *H, *DR > TL: Saint-
Domingue
Stilbus Seidlitz, 1872
Stilbus sublineatus (Guillebeau, 1894). Leng & Mutchler, 1914: 409; Blackwelder, 1945: 430 *H, *DR > TL: Saint-Domingue
Xanthocomus Guillebeau, 1894
Xanthocomus grouveillei Guillebeau, 1894: 306. Blackwelder, 1945: 430 *H, *DR > TL: Saint-Domingue

Family CRYPTOPHAGIDAE
Subfamily CRYPTOPHAGINAE
Tribe CRYPTOPHAGINI
Cryptophagus Herbst, 1892
Cryptophagus sinuatus Grouvelle, 1919: 201. Blackwelder, 1945: 429 *H > TL: Haïti
Note: In the opinion of expert coleopterists, this genus probably belongs to a different family. Apparently the situation has not
been reviewed through examination of the type, which would presumably be located at the Paris Museum.

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Family EROTYLIDAE
Subfamily EROTYLINAE
Tribe DACNINI
Dacne Latreille, 1796
Dacne (Ameridacne) brodzinskyi† Skelley, 1997: 51; Skelley, 2009: 13 [Fossil in Dominican amber]
Tribe EROTYLINI
Aegithus Fabricius, 1801
Aegithus clavicornis (Linnaeus, 1758). Medrano-Cabral, Bastardo & Skelley, 2009: 1; Skelley, 2009: 22 [Introduced]
Tribe MEGALODACNINI
Antillengis Skelley, 2009
Antillengis epochthidius Skelley, 2009: 18 *DR > TL: Prov. La Vega, La Ciénaga, Parque Nac. HQ, 1100 m
Tribe TRITOMINI
Altisessor Skelley, 2009
Altisessor ater Skelley, 2009: 37 *DR, *H > TL: Prov. Barahona, nr. Filipinas, Larimar Mine
Altisessor centralis Skelley, 2009: 38 *DR > TL: Prov. La Vega, ca. 10 km E. Constanza, 1295 m
Altisessor hottensis Skelley, 2009: 41 *H > TL: Desbarriere, Mt. La Hotte, nr. 4000 ft.
Notaepytus Skelley, 2009
Notaepytus baorucoensis Skelley, 2009: 58 *DR > TL: Pedernales, La Abeja, 38 km NNW Cabo Rojo, 1160 m
Notaepytus cyclosignatus Skelley, 2009: 61 *DR > TL: Duarte, Reserva Loma Quita Espuela, El Cadillar
Notaepytus decoregens Skelley, 2009: 64 *DR > TL: Pr. La Altagracia, Nisibón, Finca Papagayo
Notaepytus elongatus Skelley, 2009: 66 *DR > TL: Pedernales, 37 km N. Cabo Rojo, 1500 m
Notaepytus flavitarsis (Lacordaire, 1842). Curran, 1944: 3; Alvarenga, 1994: 18; Skelley, 2009: 68
Notaepytus fulvitarsis (Lacordaire, 1842). Curran, 1944: 3; Alvarenga, 1994: 18; Skelley, 2009: 69
Notaepytus haitensis (Curran, 1944: 2). Alvarenga, 1994: 18; Skelley, 2009: 72 *H > Northeast Foothills, La Hotte, 2000 to
4000 feet
Notaepytus ignotensis Skelley, 2009: 72 *H > TL: La Cavite
Notaepytus inversus Skelley, 2009: 74 *DR > TL: Pedernales, 24 km N. Cabo Rojo, 610 m
Notaepytus lavegaensis Skelley, 2009: 74 *DR > TL: Valle Nuevo, SE. Constanza
Notaepytus modestus (Olivier, 1807: 483). Curran, 1944: 3; Alvarenga, 1994: 18; Skelley, 2009: 75 *H, *DR > TL: Saint-
Domingue
Notaepytus neibaensis Skelley, 2009: 77 *DR > TL: Sierra de Neiba, Sabana del Silencio, 10 km SSW El Cercado
Notaepytus quisqueya† Keller & Skelley, 2019: 192 [Fossil in Dominian amber]
Notaepytus tarsalis (Lacordaire, 1842). Curran, 1944: 4; Alvarenga, 1994: 18; Skelley, 2009: 78 *H > TL: Hayti
Subfamily TORAMINAE
Loberus LeConte, 1861
Loberus testaceus Reitter, 1875. Blackwelder, 1945: 428; Peck, 2010: 39

Superfamily COCCINELLOIDEA
Family BOTHRIDERIDAE
Subfamily BOTHRIDERINAE
Tribe BOTHRIDERINI
Lithophorus Sharp, 1884
Lithophorus gemmatus (Reitter, 1877). Blackwelder, 1945: 473; Slipinski et al., 1989: 144
Lithophorus succineus (Pascoe, 1860). Blackwelder, 1945: 473

Family CERYLONIDAE
Subfamily CERYLONINAE
Cerylcautomus Sen Gupta & Crowson, 1973
Cerylcautomus ocellatus Slipinski, 1982b: 61 *DR > TL: Loma Rucilla and mts N, 5–8000 ft.
Lapethinus Slipinski, 1984
Lapethinus dominicanus Slipinski, 1984: 93 *DR > TL: San José de las Matas, 1000–2000 ft.
Lapethus Casey, 1890
Lapethus sharpi Champion, 1913. Slipinski, 1984: 74

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Lapethus sulcimargo (Champion, 1913). Slipinski, 1984: 86
Metacerylon Grouvelle, 1906
Metacerylon amaroides (Chevrolat, 1863). Slipinski, 1991: 80
Metacerylon dufai Grouvelle, 1912. Slipinski, 1991: 80
Philothermus Aubé, 1843
Philothermus dentipes Slipinski, 1982a: 40
Philothermus puberulus Schwarz, 1878. Lawrence & Stephan, 1975: 153
Subfamily EUXESTINAE
Euxestus Wollaston, 1858
Euxestus erithacus (Chevrolat, 1864). Leng & Mutchler, 1914: 412; Blackwelder, 1945: 474; Peck, 2005: 135

Family COCCINELLIDAE
Subfamily CHILOCORINAE
Chilocorus Leach, 1815
Chilocorus cacti (Linnaeus, 1767). Schmutterer, 1990: 123
Subfamily COCCIDULINAE
Tribe COCCIDULINI
Psorolyma Sicard, 1922
Psorolyma baorucensis Gordon, 1994: 228 *DR > TL: Prov. Pedernales, 25 km N Cabo Rojo, 915 m
Psorolyma cyanella Gordon, 1974: 230 *DR, *H > TL: Puerto Libertad
Psorolyma doyeni Gordon, 1994: 228 *DR > TL: La Vega Prov., 24 km S Jarabacoa, 4,000 feet
Tribe NOVIINI
Rodolia Mulsant, 1850
Rodolia cardinalis (Mulsant, 1850). Wolcott, 1927: 206; Gómez-Menor, 1941: 127; Zuluaga & Mesa, 2000: 11 [Purposely
introduced]
Subfamily COCCINELLINAE
Tribe COCCINELLINI
Coelophora Mulsant, 1850
Coelophora inaequalis (Fabricius, 1775). Marcano, 1989: 5/89
Coleomegilla Timberlake, 1920
Coleomegilla cubensis (Casey, 1908). Marcano, 1989: 5/89; Alvarez et al., 1993: 36
Cycloneda Crotch, 1871
Cycloneda sanguinea limbifer Casey, 1899. Blackwelder, 1945: 453; Knight, 1944: 100; Alvarez et al., 1993: 36
Hippodamia Mulsant, 1846
Hippodamia convergens (Guérin-Méneville, 1842). Alvarez et al., 1993: 36
Mulsantina Weise, 1906
Mulsantina labyrinthica (Sicard, 1929: 517). Blackwelder, 1945: 453; Chapin, 1985: 364 *H > TL: Haïti
Procula Mulsant, 1850
Procula ferruginea (Olivier, 1808). Blackwelder, 1945: 453
Tribe HALYZIINI
Psyllobora Chevrolat, 1836
Psyllobora nana Mulsant, 1850. Wolcott, 1927: 196; Blackwelder, 1945: 455; Chapin, 1957: 89
Psyllobora schwarzi Chapin, 1957: 89
Subfamily HYPERASPIDINAE
Tribe HYPERASPIDINI
Hyperaspis Redtenbacher, 1844
Hyperaspis connectens (Thunberg, 1808). Marcano, 1990: 3/90
Hyperaspis festiva Mulsant, 1850. Blackwelder, 1945: 447
Hyperaspis luteola Mulsant, 1850. Blackwelder, 1945: 447
Subfamily SCYMNINAE
Tribe DIOMINI
Decadiomus Chapin, 1933
Decadiomus bahamicus (Casey, 1899). Chapin, 1933: 97
Diomus Mulsant, 1850

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Diomus roseicollis (Mulsant, 1853). Perez-Gelabert, 2008: 109
Tribe SCYMNILLINI
Zilus Mulsant, 1850
Zilus atrox (Chapin, 1930: 493). Blackwelder, 1945: 445 *H > TL: Camp Perrin, La Prise
Tribe SCYMNINI
Cryptolaemus Mulsant, 1853
Cryptolaemus montrouzieri Mulsant, 1853. Gómez-Menor, 1941: 127; Marcano, 1989: 4/89 [Purposely introduced]
Nephaspis Casey, 1899
Nephaspis lynx Gordon, 1997: 47 *DR > TL: Prov. Pedernales, 24 km N. Cabo Rojo, 610 m
Scymnus Kugelann, 1794
Scymnus panamensis Jacoby, 1884. Gómez-Menor, 1941: 127
Scymnus sp.—Gómez-Menor (1941: 127)
Tribe STETHORINI
Stethorus Weise, 1885
Stethorus (Stethorus) caribus Gordon & Chapin, 1983: 245 *DR > TL: Baní
Subfamily STICHOLOTIDINAE
Tribe SERANGIINI
Delphastus Casey, 1899
Delphastus pallidus (LeConte, 1878). Gordon, 1970: 360; Marcano, 1990: 3/90; Alvarez et al., 1993: 36
Delphastus quinculus Gordon, 1994: 120
Tribe STICHOLOTIDINI
Bura Mulsant, 1850
Bura cuprea Mulsant, 1850. Blackwelder, 1945: 443; Vandenberg & Perez-Gelabert, 2007: 39 *H, *DR > TL: Haïti
Nelasa Gordon, 1991
Nelasa dominicensis Gordon, 1991: 304 *DR > TL: La Matica, Boca Chica
Nelasa haitiensis Gordon, 1991: 304 *H > TL: Kenscoff
Parinesa Gordon, 1991
Parinesa whiteheadi Gordon, 1991: 315 *DR > TL: S. Francisco

Family CORYLOPHIDAE
Subfamily CORYLOPHINAE
Hoplicnema Matthews, 1899
Hoplicnema darlingtoni Pakaluk, 1987: 89 *H > TL: Sud-Ouest: Desbarriere, Mt. La Hotte, near 4000 ft.
Hoplicnema thomasi Pakaluk, 1987: 108 *H > TL: Sud-Ouest: Parc National La Visite, Morne La Visite, 2100 m

Family DISCOLOMATIDAE
Subfamily APHANOCEPHALINAE
Fallia Sharp, 1902
Fallia iviei Cline & Shockley, 2012: 94 *DR > TL: 35 km N Cabo Rojo, Las Abejas, 1250 m
Fallia palaeodominicana† Szawaryn & Kupryjanowicz, 2019: 2 [Fossil in Dominican amber]

Family ENDOMYCHIDAE
Subfamily ANAMORPHINAE
Discolomopsis† Shockley, 2006
Discolomopsis dominicana† Shockley, 2006: 212 [Fossil in Dominican amber]
Micropsephodes Champion, 1913
Micropsephodes sp.—Leschen & Carlton (2000: 237)
Subfamily STENOTARSINAE
Stenotarsus Perty, 1832
Stenotarsus nigrivestis Shockley, 2007: 480 *DR > TL: La Agüita, N of Pedernales, Pedernales Prov.

Family LATRIDIIDAE
Subfamily CORTICARIINAE
Displotera Reitter, 1887

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Displotera maderae (Wollaston, 1854). Mann, 1915: 164; Blackwelder, 1945: 435; Dajoz, 1970: 652
Holoparamecus Curtis, 1833
Holoparamecus kunzei Aubé, 1843. Blackwelder, 1945: 435; Dajoz, 1971: 653
Melanophthalma Motschulsky, 1866
Melanophthalma lohsei Rucker, 1987: 4 *DR > TL: Azua
Melanophthalma scitula Rucker, 1987: 4 *DR > TL: Cordillere Central, Colonia Ramfis
Melanophthalma venusta Rucker, 1987: 7 *DR > TL: Cazabita [Loma Casabito], 1250 m
Metophthalmus Wollaston, 1854
Metophthalmus columbusi Andrews, 1994: 191 *DR > TL: Cabo Rojo, Pedernales Prov.
Metophthalmus schusteri Andrews, 1994: 194 *DR > TL: Carbonara, Monte Cristi Prov.
Metophthalmus trilineatus Andrews, 1994: 194 *DR > TL: El Aceitillar, ~35 km NNW Cabo Rojo, Pedernales Prov.

Superfamily TENEBRIONOIDEA
Family CIIDAE
Subfamily CIINAE
Tribe CIINI
Ceracis Mellié, 1848
Ceracis curtus (Mellié, 1848). Lawrence, 1967: 110
Ceracis minutus Drury, 1917. Lawrence, 1967: 114
Cis Latreille, 1796
Cis hirsutus Casey, 1898. Peck, 2005: 141
Tribe XYLOGRAPHELLINI
Scolytocis Blair, 1928
Scolytocis kiskeyensis Lopes-Andrade, 2008: 24

Family MORDELLIDAE
Subfamily MORDELLINAE
Tribe MORDELLINI
Conalia Mulsant & Rey, 1858
Conalia ebenina Champion, 1891. Blackwelder, 1945: 477
Glipa LeConte, 1857
Glipa hilaris (Say, 1835). Blackwelder, 1945: 475
Mordella Linnaeus, 1758
Mordella divergens Ray, 1939: 279. Blackwelder, 1945: 476 *H > TL: Mount Pilboreau
Mordella haitiensis Ray, 1939: 282. Blackwelder, 1945: 476 *H > TL: Mariani
Mordella summermanae Ray, 1939: 283. Blackwelder, 1945: 477 *DR > TL: Constanza
Mordellistena Costa, 1854
Mordellistena epidendrana Ray, 1937: 239. Blackwelder, 1945: 478 *DR > TL: Dominican Republic [Secured in Epidendrum
sp. at San Francisco, California, the port of entry]

Family SCRAPTIIDAE
Scraptia Latreille, 1807
Scraptia sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]

Family RIPIPHORIDAE
Subfamily RHIPIPHORINAE
Macrosiagon Hentz, 1830
Macrosiagon mutilata (Gerstaecker, 1855). Falin, 2004: 8
Subfamily RIPIDIINAE
Undetermined larvae†—Poinar (1992: 151); Wu (1997: 174); Kaupp et al. (2001: 188) [Fossil in Dominican amber]
Neorhipidius Viana, 1958
Neorhipidius seicherti† Batelka et al., 2011: 282 [Fossil in Dominican amber]
Quasipirhidius Zaragoza Caballero, 1991
Quasipirhidius luzziae† Batelka et al., 2011: 276 [Fossil in Dominican amber]

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Family ZOPHERIDAE
Subfamily COLYDIINAE
Bitoma Herbst, 1793
Bitoma carinata (LeConte, 1863). Blackwelder, 1945: 470
Bitoma exarata (Pascoe, 1863). Grouvelle, 1896: 185; Blackwelder, 1945: 470
Colydium Fabricius, 1792
Colydium sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Endeitoma Sharp, 1894
Endeitoma granulata (Say, 1827). Blackwelder, 1945: 470
Subfamily ZOPHERINAE
Aspathines Champion, 1888
Aspathines aeneus aeneus (Thomson, 1860). Freude, 1955: 694; Freude, 1993: 216
Aspathines aeneus ovatus Champion, 1888. Freude, 1993: 216
Hyporhagus Thomson, 1860
Hyporhagus antillicus Freude, 1955: 725. Freude, 1993: 222 *DR > TL: San Domingo
Hyporhagus cicatricosus Freude, 1955: 722. Freude, 1993: 221 *DR > TL: Jarabacoa
Hyporhagus hispaniolae Freude, 1977: 30. Freude, 1993: 219 *H, *DR > TL: Palmar de Ocoa
Hyporhagus marginatus fabricii Freude, 1955: 724. Spilman, 1971: 7
Pycnomerus Erichson, 1842
Pycnomerus biimpresus Reitter, 1877. Ivie & Slipinsky, 1989: 69
Pycnomerus hottae Ivie & Slipinski, 1989: 73 *H > TL: Mt. La Hotte
Pycnomerus valentinei Ivie & Slipinski, 1989: 76

Family MYCETOPHAGIDAE
Litargus Erichson, 1846
Litargus sp.†—Poinar & Poinar (1999: 205) [Fossil in Dominican amber]

Family TENEBRIONIDAE
Classification follows catalogue of Bousquet et al. (2017).
Subfamily LAGRIINAE
Tribe BELOPINI
Rhypasma Pascoe, 1862
Rhypasma haitianum Marcuzzi, 1953. Marcuzzi, 1984: 75
Tribe LAGRIINI
Subtribe STATIRINA
Statira Serville, 1825
Statira dermoidea† Doyen & Poinar, 1994: 30 [Fossil in Dominican amber]
Statira haitensis Champion, 1917: 255. Blackwelder, 1945: 499 *H > TL: Port-au-Prince
Tribe LUPROPINI
Lorelus Sharp, 1876
Lorelus angulatus† Doyen & Poinar, 1994: 28 [Fossil in Dominican amber]
Lorelus foraminosus† Doyen & Poinar, 1994: 28 [Fossil in Dominican amber]
Lorelus minutulis† Doyen & Poinar, 1994: 30 [Fossil in Dominican amber]
Lorelus wolcotti† Doyen & Poinar, 1994: 30 [Fossil in Dominican amber, specimen presumably conspecific with extant
species]
Subfamily NILIONINAE
Nilio Latreille, 1802
Nilio dominicana† Poinar & Brown, 2010: 233 [Fossil in Dominican amber]
Subfamily PIMELIINAE
Tribe EDROTINI
Trientoma Solier, 1835
Trientoma laevis Allard, 1883: 14
Trientoma nascens† Doyen & Poinar, 1994: 31 [Fossil in Dominican amber]
Trientoma rugifrons Champion, 1884. Leng & Mutchler, 1914: 460

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Trientoma ryticephala Allard, 1883: 14
Trientoma sallei Kraatz, 1865. Allard, 1883: 14; Marcuzzi, 1984: 74
Tribe EPITRAGINI
Epitragus Latreille, 1802
Epitragus aurulentus aurulentus Kirsch, 1866. Marcano, 1981: 36; Schmutterer, 1990: 210; Bousquet et al., 2017: 127
=Epitragus jamaicensis Champion, 1896. Marcuzzi, 1998a: 152; Bousquet et al., 2017: 127
Tribe STENOSINI
Caribanosis Nabozhenko, Kirejtshuk, Merkl, Varela, Aalbu & Smith, 2016
Caribanosis quisqueyanus (Garrido & Varela, 2011: 32). Nabozhenko et al., 2016: 569 *DR > TL: Cueva del Guano,
Pedernales Prov.
Subfamily TENEBRIONINAE
Tribe ALPHITOBIINI
Alphitobius Stephens, 1829
Alphitobius diaperinus (Panzer, 1797). SEA, 1999: 128
Tribe AMARYGMINI
Cymatothes Dejean, 1834
Cymatothes dominicus† Doyen & Poinar, 1994: 36 [Fossil in Dominican amber]
Cymatothes nebulosa (Fabricius, 1781). Marcuzzi, 1984: 105
Cymatothes undata (Fabricius, 1792). Blackwelder, 1945: 544
Tribe BOLITOPHAGINI
Rhipidandrus LeConte, 1866
Rhipidandrus cornutus (Arrow, 1904). Blackwelder, 1945: 527; Marcuzzi, 1984: 86
Rhipidandrus quadripapillatus† Doyen & Poinar, 1994: 33 [Fossil in Dominican amber]
Rhipidandrus sulcatus Gorham, 1898. Blackwelder, 1945: 527; Marcuzzi, 1984: 86
Tribe HELOPINI
Helops Fabricius, 1775
Helops sp.—Garrido (2004: 44)
Tribe OPATRINI
Subtribe OPATRINA
Blapstinus Dejean, 1821
Blapstinus dominicus Marcuzzi, 1962. Marcuzzi, 1977: 27; Ivie & Hart, 2016: 463
Blapstinus domingoensis (Marcuzzi, 1998b: 222). Ivie & Hart, 2016: 463
Blapstinus genaroi (Garrido, 2004: 41). Ivie & Hart, 2016: 463 *DR > TL: 3.5 km al norte de Guayabal, Postrer Río, Provincia
Independencia, 350 msnm
Blapstinus haitensis Marcuzzi, 1962: 34. Ivie & Hart, 2016: 463 *H, *DR > TL: “Haiti, Santiago”
Blapstinus hispaniolensis (Marcuzzi, 1998b: 220). Ivie & Hart, 2016: 463 *DR > TL: Palmar de Ocoa, southern part of the
island
Blapstinus klapperichi (Marcuzzi, 1998b: 219). Ivie & Hart, 2016: 465 *DR > TL: Boca Chica, southern coast of the island
Cenophorus Mulsant & Rey, 1859
Cenophorus viduus Mulsant & Rey, 1859: 113. Blackwelder, 1945: 524; Ivie & Hart, 2016: 468 *H > TL: Saint-Domingue
Diastolinus Mulsant & Rey, 1859
Diastolinus azuensis Hart & Ivie, 2016: 506 *DR > TL: Azua Prov, 18 km E. Azua
Diastolinus coarctatus (Mulsant & Rey, 1859: 106). Blackwelder, 1945: 525; Marcuzzi, 1984: 82; Hart & Ivie, 2016: 507
=Diastolinus estebani Garrido, 2004: 42. Hart & Ivie, 2016: 507
Diastolinus espoloni Garrido, 2007: 46. Hart & Ivie, 2016: 512 *DR > TL: Carretera de Duvergé a Jimaní, Prov. Pedernales
Diastolinus gladiator (Garrido, 2004: 120). Hart & Ivie, 2016: 514
Diastolinus vaderi Hart & Ivie, 2016: 518 *H > TL: Dept. Sud-Oueste, Parc National La Visite, Morne La Visite, 2100 m
Xerolinus Ivie & Hart, 2016
Xerolinus sallei (Mulsant & Rey, 1859: 80). Blackwelder, 1945: 524; Marcuzzi, 1977: 22; Ivie & Hart, 2016: 474
=Diastolinus costipennis Mulsant & Rey, 1859: 85. Blackwelder, 1945: 524; Marcuzzi, 1984: 77; Ivie & Hart, 2016: 474
=Diastolinus puncticollis Mulsant & Rey, 1859: 83. Blackwelder, 1945: 524; Marcuzzi, 1977: 22; Ivie & Hart, 2016: 474
=Diastolinus assoi Garrido, 2004: 40. Ivie & Hart, 2016: 474
Tribe PEDININI
Subtribe PLATYNOTINA

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Alaetrinus Iwan, 1995
Alaetrinus pullus (Sahlberg, 1823). Iwan, 1995: 38; Iwan, 2002: 225
Tribe TENEBRIONINI
Zophobas Dejean, 1834
Zophobas atratus Fabricius, 1775. Ferrer, 2011: 297; Bousquet et al., 2017: 216
=Zophobas morio (Fabricius, 1775). Wolcott, 1927: 223
Zophobas costatus Pic, 1921: 10. Blackwelder, 1945: 534; Bousquet et al., 2017: 218 *DR, *H > TL: St. Domingue
Tribe TOXICINI
Subtribe DYSANTINA
Wattius Kaszab, 1982
Wattius emmabaconae Smith & Sanchez, 2015: 121 *DR > TL: Barahona, nr. Filipinas, Mt. Tutu
Wattius reflexus† Doyen & Poinar, 1994: 33 [Fossil in Dominican amber]
Tribe TRIBOLIINI
Hypogena Dejean, 1834
Hypogena biimpresa (Latreille, 1813). De Armas & Garrido, 2012: 107
Hypogena marginalis† Doyen & Poinar, 1994: 35 [Fossil in Dominican amber]
Latheticus Watherhouse, 1880
Latheticus oryzae Watherhouse, 1880. SEA, 1999: 128
Tribolium MacLeay, 1825
Tribolium castaneum (Herbst, 1797). Blackwelder, 1945: 531 [Introduced]
Tribolium confusum Jacquelin du Val, 1868. Smith & Audant, 1930: 975; Blackwelder, 1945: 531 [Introduced]
Tribe ULOMINI
Uloma Dejean, 1821
Uloma sp.—Garrido (2004: 45)
Subfamily ALLECULINAE
Tribe ALLECULINI
Subtribe ALLECULINA
Allecula Fabricius, 1801
Allecula ramosi Campbell, 1971. Campbell, 1978: 207
Allecula sp.†—Doyen & Poinar (1994: 36) [Fossil in Dominican amber]
Hymenorus Mulsant, 1852
Hymenorus haitiellus Campbell, 1971: 94 *H > TL: Diquini
Hymenorus haitius Campbell, 1971: 93 *H, *DR > TL: Manneville, about 60 ft. alt.
Hymenorus hispaniolensis Campbell, 1971: 78 *H, *DR > TL: Barahona
Hymenorus oculatus† Doyen & Poinar, 1994: 37 [Fossil in Dominican amber]
Hymenorus striatus (Pic, 1930: 26). Campbell, 1971: 95 *H, *DR > TL: Haiti
Lobopoda Solier, 1835
Lobopoda annosa† Doyen & Poinar, 1994: 37 [Fossil in Dominican amber]
Lobopoda (Gabrilopoda) darlingtoni Campbell, 1971: 40 *DR > TL: Loma Vieja, S Constanza, 6,000 ft.
Lobopoda (Lobopoda) colona Campbell, 1971: 56 *H > TL: La Visite 7 vic., La Selle Range, 5–7,000 ft.
Lobopoda (Lobopoda) haitensis Campbell, 1966: 158 *H, *DR > TL: Shada Refuge Headquarters
Lobopoda (Lobopoda) hispaniolensis Campbell, 1971: 46 *DR > TL: Loma Vieja, S. Constanza, 6,000 ft.
Lobopoda (Lobopoda) micans Campbell, 1971: 50 *DR > TL: Loma Rucilla & mts. N, 5–8000 ft.
Lobopoda (Lobopoda) notapuncta Campbell, 1971: 54 *DR > TL: Desbarriere, Mt. La Hotte, nr. 4000 ft.
Lobopoda (Lobopoda) picipennis Campbell, 1971: 61 *DR > TL: Villa Altagracia
Lobopoda (Lobopoda) sandersoni Campbell, 1971: 60 *DR > TL: Jarabacoa, La Vega Prov.
Lobopoda (Lobopoda) wittmeri Campbell, 1978: 204 *DR > TL: La Colonia, San Cristóbal Prov.
Lobopoda sp.†—Doyen & Poinar (1994: 39) [Fossil in Dominican amber]
Lobopoda (Lobopoda) substriatus Campbell, 1966: 159
Parahymenorus Campbell, 1971
Parahymenorus sp.†—Doyen & Poinar (1994: 39) [Fossil in Dominican amber]
Subfamily DIAPERINAE
Tribe CRYPTICINI
Gondwanocrypticus Español, 1955

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Gondwanocrypticus filicornis (Chevrolat, 1878). Marcuzzi, 1998a: 158
Tribe DIAPERINI
Subtribe ADELININA
Adelina LeConte, 1851
Adelina bidens (Schaeffer, 1915). Ardoin, 1977: 10
Adelina dominicana (Ardoin, 1977: 18) *DR > TL: Baní
Adelina klapperichi (Ardoin, 1977: 12) *DR > TL: Baní
Adelina plana (Olivier, 1795). Ardoin, 1977: 3
Loxostethus Triplehorn, 1962
Loxostethus erythroscelis Triplehorn & Merkl, 1997: 739 *DR > TL: Prov. La Vega, Parque Nac. HQ
Loxostethus gowdeyi (Pic, 1930). Triplehorn & Merkl, 1997: 739
Loxostethus oblongus Triplehorn & Merkl, 1997: 737 *DR > TL: Prov. La Vega, Par. Nac. Armando Bermúdez
Loxostethus unicolor Triplehorn, 1997. Triplehorn & Merkl, 1997: 737
Subtribe DIAPERINA
Diaperis Geoffroy, 1762
Diaperis maculata Olivier, 1791. Triplehorn, 1965: 370; Marcuzzi, 1984: 87
Lelegeis Champion, 1886
Lelegeis hispaniolae Triplehorn, 1962: 503 *H, *DR > TL: Ennery, near 1000 ft.
Liodema Horn, 1870
Liodema phalacroides† Doyen & Poinar, 1994: 43 [Fossil in Dominican amber]
Neomida Latreille, 1829
Neomida cioides (Champion, 1886). Triplehorn, 2006: 314
Neomida ferruginea (LeConte, 1866). Marcuzzi, 1984: 88; Triplehorn, 2006: 322
Neomida lecontei (Bates, 1873). Triplehorn, 2006: 326
Neomida senicula† Doyen & Poinar, 1994: 41 [Fossil in Dominican amber]
Platydema Laporte & Brullé, 1831
Platydema antennatum Laporte & Brullé, 1831. Blackwelder, 1945: 527; Marcuzzi, 1984: 88
Platydema excavatum (Say, 1823). Blackwelder, 1945: 528; Triplehorn, 1965: 395; Marcuzzi, 1984: 88
Platydema ferruginea (LeConte, 1866). Triplehorn, 1965: 384
Platydema micans Horn, 1870. Triplehorn, 1965: 430; Marcuzzi, 1984: 89
Platydema picicorne (Fabricius, 1792). Blackwelder, 1945: 528; Marcuzzi, 1984: 89
Platydema sp. 1†—Doyen & Poinar (1994: 43) [Fossil in Dominican amber]
Platydema sp. 2†—Doyen & Poinar (1994: 43) [Fossil in Dominican amber]
Platydema sp. 3†—Doyen & Poinar (1994: 43) [Fossil in Dominican amber]
Ulomoides Blackburn, 1888
Ulomoides ocularis (Casey, 1891). Perez-Gelabert, 2008: 113 [Introduced]
Tribe GNATHIDIINI
Subtribe ANOPIDIINA
Caecophloeus Dajoz, 1972
Caecophloeus darlingtoni Dajoz, 1975: 115 *H > TL: La Visite & vic., La Selle Range, 5000–7000 pieds
Sphaerognathium Dajoz, 1975
Sphaerognathium globosum Dajoz, 1975: 113 *H > TL: Mont La Hotte, 5000 à 7000 pieds
Tyrtaeus Champion, 1913
Tyrtaeus azureus† Doyen & Poinar, 1994: 39 [Fossil in Dominican amber]
Tyrtaeus cupreorutilans† Vitali, 2008: 12 [Fossil in Dominican amber]
Tyrtaeus elongatus† Doyen & Poinar, 1994: 40 [Fossil in Dominican amber]
Tyrtaeus flavoantennatus† Doyen & Poinar, 1994: 40 [Fossil in Dominican amber]
Tyrtaeus thoracicus† Doyen & Poinar, 1994: 41 [Fossil in Dominican amber]
Tribe HYPOPHLOEINI
Corticeus Piller & Mitterpacher, 1783
Corticeus tensicollis Triplehorn, 1979: 46
Corticeus tertiarius† Vitali, 2007: 2 [Fossil in Dominican amber]
Tribe PHALERIINI
Phaleria Latreille, 1802

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Phaleria fulva Fleautiaux & Sallé, 1889. Watrous & Triplehorn, 1982: 17
Phaleria picipes Say, 1824. Watrous & Triplehorn, 1982: 19
Phaleria testacea Say, 1824. Watrous & Triplehorn, 1982: 19
=Phaleria angustata Chevrolat, 1878. Blackwelder, 1945: 526; Watrous & Triplehorn, 1982: 19
Phaleria thinophila Watrous & Triplehorn, 1982: 15. Marcuzzi, 1984: 84
Subfamily STENOCHIINAE
Tribe CNODALONINI
Cnodalon Latreille, 1796
Cnodalon viride Latreille, 1804: 521. Blackwelder, 1945: 538; Marcuzzi, 1984: 102 *H > TL: Saint-Domingue
Nesocyrtosoma Marcuzzi, 1976
Nesocyrtosoma altagracia Hopp & Ivie, 2009: 40 *DR > TL: Prov. La Altagracia; Boca de Yuma, 12 m
Nesocyrtosoma antiquum† (Kaszab & Schawaller, 1984: 3). Doyen & Poinar, 1994: 45; Hopp & Ivie, 2009: 78 [Fossil in
Dominican amber]
Nesocyrtosoma basilense Hopp & Ivie, 2009: 51 *H > TL: Mt. Basil, N. Haiti, 4,700 ft.
Nesocyrtosoma bestiola Hopp & Ivie, 2009: 20 *DR > TL: Prov. Barahona, 7 km NE Enriquillo
Nesocyrtosoma celadonum† Doyen & Poinar, 1994: 46. Hopp & Ivie, 2009: 78 [Fossil in Dominican amber]
Nesocyrtosoma crenulatum Hopp & Ivie, 2009: 44. Hopp, 2011: 242 *DR, *H > Independencia Prov., ESE Jimaní, La Florida,
20 m
=Nesocyrtosoma bromelicolus Garrido & Varela, 2010: 33. Hopp, 2011: 242 *DR > TL: Las Mercedes, Prov. Pedernales
Nesocyrtosoma darlingtoni Hopp & Ivie, 2009: 47 *H > NE foothills; La Hotte, 2–4,000 ft
Nesocyrtosoma dolosum Hopp & Ivie, 2009: 42 *H > TL: La Hotte, 2–4,000 ft.
Nesocyrtosoma guerreroi Hopp & Ivie, 2009: 69 *DR > Prov. Pedernales, 26 km N. Cabo Rojo, 915 m
Nesocyrtosoma hadratum† Doyen & Poinar, 1994: 47. Hopp & Ivie, 2009: 79 [Fossil in Dominican amber]
Nesocyrtosoma hispaniolae (Marcuzzi, 1999: 83). Hopp & Ivie, 2009: 28 *DR > TL: La Altag.; 9 km SW Campo Nuevo
Nesocyrtosoma impensum† Doyen & Poinar, 1994: 47. Hopp & Ivie, 2009: 79 [Fossil in Dominican amber]
Nesocyrtosoma larseni Hopp & Ivie, 2009: 50
Nesocyrtosoma merkli (Marcuzzi, 1999: 82). Hopp & Ivie, 2009: 33 *DR > La Vega, 53 km SE Constanza
Nesocyrtosoma minisculum† Hopp & Ivie, 2009: 80 [Fossil in Dominican amber]
Nesocyrtosoma mutabile Hopp & Ivie, 2009: 48 *DR > Pedernales Prov., PN Sierra de Bahoruco, Las Abejas, 1,150 m
Nesocyrtosoma nearnsi Hopp & Ivie, 2009: 65 *DR, *H > TL: La Vega Prov. PN Armando Bermúdez, km 1–3 along trail W of
La Ciénaga, 900—1,100 m
Nesocyrtosoma neibaense Hopp & Ivie, 2009: 29 *DR > TL: Sierra de Neiba, Sabana del Silencio, 2,009 m
Nesocyrtosoma otus Hopp & Ivie, 2009: 45 *DR > TL: Mt. Diego de Ocampo, 4,000 ft.
Nesocyrtosoma phthanatum† Doyen & Poinar, 1994: 48. Hopp & Ivie, 2009: 80 [Fossil in Dominican amber]
Nesocyrtosoma productum Hopp & Ivie, 2009: 67 *DR > TL: La Ciènaga, Parque Nac. HQ, 1100 m
Nesocyrtosoma purpureum Hopp & Ivie, 2009: 43 *DR > TL: Prov. Barahona, nr. Filipinas, Larimar Mine
Nesocyrtosoma scabrosum Hopp & Ivie, 2009: 35 *H > TL: Tardieu, Mt. La Hotte, 3,000 ft
Nesocyrtosoma serratum Hopp & Ivie, 2009: 62 *DR > TL: Pedernales Prov., ~10 km on trail to Carlitos, 172 m
Nesocyrtosoma simplex Hopp & Ivie, 2009: 37 *H, *DR > TL: La Visite and vic., La Selle Range, 5–7,000 ft.
Nesocyrtosoma skelleyi Hopp & Ivie, 2009: 63 *DR > TL: Prov. Barahona, nr. Filipinas, Larimar Mine
Tribe STENOCHIINI
Strongylium Kirby, 1819
Strongylium elongatum Garrido, 2012: 65 *DR > TL: 25 km al norte de Cabo Rojo, Prov. Pedernales
Strongylium haetianum Garrido, 2012: 64 *H > TL: Viarte, entre el Parque Nacional headquarters y Morne de Ember, 1750
msnm
Strongylium quisqueyanum Garrido, 2012: 66 *DR > TL: Cumbre de Puerto Plata (914 msnm) [evidentemente Pico Isabel de
Torres]
Strongylium woodruffi Garrido, 2012: 67 *DR > TL: Finca de Eladio Fernández, 5 km al oeste de Manabao
Tribe TALANINI
Talanus Mäklin, 1878
Talanus sp.—Garrido & Gutiérrez (2004: 63)
Notes: The species “Amarygmus” desfontainesi (Pic, 1952: 2) cited for Hispaniola by. Bremer, 2001a: 57 is considered to have
been incorrectly recorded (Bousquets et al., 2017: 385) and therefore is removed from the Hispaniolan faunal list.

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 147
Family OEDEMERIDAE
Subfamily OEDOMERINAE
Tribe HYPASCLERINI
Hypasclera Kirsch, 1866
Hypasclera costata (Champion, 1896). Arnett, 1957: 4; Arnett, 1984: 2; Peck & Thomas, 1998: 111
Hypasclera nesiotes (Arnett, 1951). Arnett, 1984: 3; Peck & Thomas, 1998: 111
Hypasclera pleuralis (LeConte, 1866). Arnett, 1984: 3; Peck & Thomas, 1998: 111
Hypasclera spinosus (Arnett, 1957). Arnett, 1983: 119.6
Tribe OEDOMERINI
Ananca Fairmaire & Germain, 1863
Ananca despaignei (Pic, 1934: 22). Blackwelder, 1945: 490 *H, *DR > TL: Saint-Domingue
Tribe OXACIINI
Oxacis LeConte, 1866
Oxacis laeta (Waterhouse, 1878: 306). Blackwelder, 1945: 490
Paroxacis Arnett, 1951
Paroxacis albomarginata (Pic, 1934: 21). Arnett, 1957: 2
Paroxacis geniculata (Chevrolat, 1877). Arnett, 1983: 119.5
Tribe OXYCOPIINI
Oxycopis Arnett, 1951
Oxycopis elegans (Waterhouse, 1878: 307). Blackwelder, 1945: 490 *H, *DR > TL: St. Domingo
Oxycopis falli (Blatchley, 1928). Arnett, 1984: 2
Oxycopis lineata (Waterhouse, 1878). Arnett, 1957: 7
Oxycopis metallica (Pic, 1930). Blackwelder, 1945: 490
Oxycopis multiguttata (Pic, 1930). Blackwelder, 1945: 490
Oxycopis tenella (Waterhouse, 1878). Blackwelder, 1945: 490; Arnett, 1957: 6
Oxycopis thoracica (Fabricius, 1801). Arnett, 1957: 7
Oxycopis vittata (Fabricius, 1775). Arnett, 1957: 6
=Copidita lateralis Waterhouse, 1878: 307. Blackwelder, 1945: 490

Family MELOIDAE
Subfamily MELOINAE
Tribe MELOINI
Meloe Linnaeus, 1758
Meloe (Meloe) dominicanus† Poinar, 2009: 146 [Fossil in Dominican amber]
Meloe (Treiodous) laevis Leach, 1813. Blackwelder, 1945: 488; Pinto & Selander, 1970: 111
Subfamily NEMOGNATHINAE
Tribe HORIINI
Cissites Latreille, 1804
Cissites maculata (Swederus, 1787). Blackwelder, 1945: 482; Enns, 1959: 62; Genaro, 1996: 384
Tribe NEMOGNATHINI
Pseudozonitis Dillon, 1952
Pseudozonitis marginata (Fabricius, 1781). Genaro, 1996: 384
Tribe TETRAONYCINI
Tetraonyx Latreille, 1805
Tetraonyx quadrimaculata (Fabricius, 1792). Blackwelder, 1945: 487; Genaro, 1996: 384

Family MYCTERIDAE
Subfamily HEMIPEPLINAE
Hemipeplus Berthold, 1827
Hemipeplus dominicensis Pollock, 1999: 65 *DR > TL: Baní
Hemipeplus insularis Grouvelle, 1896: 196. Blackwelder, 1945: 423; Pollock, 1999: 66 *H, *DR > TL: Saint-Domingue
Subfamily LACCONOTINAE
Physcius Champion, 1889
Physcius maculatus Pic, 1906. Blackwelder, 1945: 494 *H, *DR > TL: Saint-Domingue

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Family SALPINGIDAE
Subfamily DACODERINAE
Myrmecoderus Aalbu et al., 2005
Myrmecoderus dominicensis (Horn, 1876: 219). Blackwelder, 1945: 515; Aalbu et al., 2005: 30 *DR > TL: Santo Domingo
Subfamily INOPEPLINAE
Inopeplus Smith, 1851
Inopeplus wolcotti Blackwelder, 1943: 139 *H > TL: Aquin

Family ANTHICIDAE
Tribe ANTHICINI
Acanthinus Laferté-Sénectère, 1849
Acanthinus concinnus (Laferté-Sénectère, 1849). Werner, 1983: 231
Acanthinus quinquemaculatus (Laferté-Sénectère, 1848). Werner, 1983: 231
Acanthinus scitulus (LeConte, 1852). Werner, 1983: 232
Anthicus Paykull, 1798
Anthicus antilleorum Werner, 1983: 218. Pollock & Ivie, 1996: 232
Anthicus blackwelderi Werner, 1983: 219
Anthicus crinitus Laferté-Sénectère, 1848. Werner, 1983: 232; Pollock & Ivie, 1996: 233
Anthicus darlingtoni Werner, 1983: 224 *H > TL: Etang Lachaux (under 1000’)
Anthicus hispaniolae Werner, 1983: 224 *DR > TL: Constanza to Jarabacoa (2–4000’)
Anthicus pallidus Say, 1826. Werner, 1964: 230; Werner, 1983: 233
Anthicus russoi Krekich-Strassoldo in Menozzi, 1929: 93. Werner, 1983: 220 *DR > TL: Moca
Anthicus subtilis Laferté-Sénectère, 1849. Werner, 1983: 222
Cyclodinus Mulsant & Rey, 1866
Cyclodinus californicus (Laferté-Sénectère, 1849). Chandler, 2005: 7
Cyclodinus mundulus (Sharp, 1885). Chandler, 2005: 13 [Introduced]
Omonadus Mulsant & Rey, 1866
Omonadus floralis (Linnaeus, 1758). Werner, 1983: 232; Pollock & Ivie, 1996: 235 [Introduced]
Sapintus Casey, 1895
Sapintus teapensis (Champion, 1890). Werner, 1983: 233; Pollock & Ivie, 1996: 236
Sapintus pallidus (Say, 1826). Peck, 2006: 191
Squamanotoxus Chandler, 2001
Squamanotoxus hispaniolae (Werner, 1983: 227). Chandler, 2001: 487 *DR > TL: Las Salinas, Baní
Sticticomus Pic, 1894
Sticticomus tobias (Marseul, 1879). Werner, 1983: 233 [Introduced]
Thicanus Casey, 1895
Thicanus texanus (Laferté-Sénectère, 1848). Werner, 1983: 234
Vacusus Casey, 1895
Vacusus vicinus (Laferté-Sénectère, 1849). Werner, 1983: 234; Pollock & Ivie, 1996: 235 [Introduced]

Superfamily CHRYSOMELOIDEA
Family CERAMBYCIDAE
Key references: Monné and Hovore (2003), Monné (2005a, 2005b).
Subfamily ASEMINAE
Tribe ASEMINI
Arhopalus Audinet-Serville, 1834
Arhopalus hispaniolae (Fisher, 1942: 4) *DR > TL: Valle Nuevo, southeast of Constanza
Subfamily CERAMBYCINAE
Tribe ACHRYSONINI
Achryson Serville, 1833
Achryson surinamum (Linnaeus, 1767). Blackwelder, 1946: 560
Enosmaeus Thomson, 1878
Enosmaeus cubanus Thomson, 1878. Monné, 2005a: 15

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 =Pseudoeme poolei Fisher, 1932: 5. Monné, 2005a: 15
Tribe ANAGLYPTINI
Tilloclytus Bates, 1885
Tilloclytus baoruco Lingafelter, 2011a: 65 *DR > TL: Las Abejas, Pedernales Prov.
Tylloclytus neiba Lingafelter, 2011a: 69 *DR > TL: Trail to Sabana del Silencio, San Juan Prov.
Tribe CALLICHROMATINI
Plinthocoelium Schmidt, 1824
Plinthocoelium domingoensis (Fisher, 1922: 52). Russo, 1930: 142; Blackwelder, 1946: 578
Plinthocoelium virens (Drury, 1770). Blackwelder, 1946: 578
Mionochroma Schmidt, 1924
Mionochroma vittatum (Fabricius, 1775). Blackwelder, 1946: 579
Tribe CALLIDIINI
Callidium Fabricius, 1775
Callidium biguttatum Sallé, 1856: 688. Blackwelder, 1946: 580 *DR > TL: Santo-Domingo
Tribe CALLIDIOPINI
Caribbomerus Vitali, 2003
Caribbomerus asperatus (Fisher, 1932: 52). Blackwelder, 1946: 571; Lingafelter, 2011b: 34
Caribbomerus decoratus (Zayas, 1975). Lingafelter, 2011b: 34
Caribbomerus elongatus (Fisher, 1932). Lingafelter, 2011b: 29
Caribbomerus sp.†—Poinar (1992: 138) [As Merostenus] [Fossil in Dominican amber]
Ceresium Newman, 1842
Ceresium compressipenne (Fisher, 1932). Blackwelder, 1946: 565; Lingafelter, 2008: 357; Perez-Gelabert, 2008: 118
Ceresium miserum (Thomson, 1878: 8). Monné, 2005a: 49 *H, *DR > TL: Sancto-Domingo
Tribe CLYTINI
Euryscelis Dejean, 1835
Euryscelis suturalis (Olivier, 1795). Russo, 1930: 143; Blackwelder, 1946: 581
Isotomus Mulsant, 1863
Isotomus comptus (Mannerheim, 1825). Blackwelder, 1946: 582 [Introduced]
Neoclytus Thomson, 1860
Neoclytus araneiformis (Olivier, 1795). Russo, 1930: 143; Blackwelder, 1946: 581
Neoclytus pallidicornis Fisher, 1932: 58. Blackwelder, 1946: 582 *H > TL: La Vanneau
Neoclytus podagricus (White, 1855). Blackwelder, 1946: 582 *H, *DR > TL: St. Domingo
Placosternus Hopping, 1937
Placosternus difficilis (Chevrolat, 1862). Marcano, 1981: 132
Tribe CURIINI
Curius Newman, 1840
Curius punctatus (Fisher, 1932). Nearns & Tavakilian, 2012: 22
Tribe PLECTROMERINI
Plectromerus Haldeman, 1847
Plectromerus bidentatus Fisher, 1942: 16. Blackwelder, 1946: 572 *DR > TL: Loma de la Peña, northwest of Constanza
Plectromerus distinctus (Cameron, 1910: 136). Blackwelder, 1946: 572; C. Micheli & Nearns, 2005: 25 *H, *DR > TL: Near
Port-au-Prince
=Plectromerus crenulatus Cazier, 1952: 1. Nearns & Branham, 2005: 29
Plectromerus dominicanus (Micheli, 1983). Nearns & Branham, 2005: 31 *DR > TL: 20 km SE Constanza, La Vega Prov.
=Curiosa dominicana Micheli, 1983: 262. Nearns & Branham, 2008: 31
Plectromerus exis Zayas, 1975. Nearns & Turnbow, 2005: 158; Nearns & Branham, 2008: 32
Plectromerus grimaldii† Nearns & Branham, 2005: 19. Vitali, 2006: 8 [Fossil in Dominican amber]
Plectromerus josephi Nearns & Branham, 2005: 40 *DR > TL: La Cumbre de Puerto Plata
Plectromerus lingafelteri C. Micheli & Nearns, 2005: 25 *DR > TL: Pico Duarte Trail
Plectromerus serratus (Cameron, 1910: 135). Blackwelder, 1946: 572 *H > TL: Near Port-au-Prince
Plectromerus tertiarius† Vitali, 2004: 453. Nearns & Branham, 2005: 23; Vitali, 2006: 5; Vitali, 2009b: 354 [Fossil in
Dominican amber]
Plectromerus thomasi Nearns & Branham, 2005: 51 *H > TL: Morne Guimby, 22 km SE Fond Verrettes
Plectromerus turnbowi Nearns & Branham, 2005: 52 *DR > TL: PN Sierra de Bahoruco, Las Abejas

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Pterolophosoma† Vitali, 2006
Pterolophosoma otiliae† Vitali, 2006: 3; Vitali, 2007: 36 [Fossil in Dominican amber]
Tribe EBURIINI
Eburia Lepeletier & Audinet-Serville, 1830
Eburia didyma Olivier, 1795. Peck, 2005: 167
Eburia fisheri Russo, 1930: 144. Blackwelder, 1946: 563
Eburia pseudostigma Lingafelter & Nearns, 2007: 182 *DR > TL: Punta Cana near Ecological Reserve, La Altagracia Prov.
Eburia sericea Sallé, 1855. Blackwelder, 1946: 563
Eburia sp.†—Poinar (1992: 138) [Fossil in Dominican amber]
Eburia stigma (Olivier, 1795). Blackwelder, 1946: 563
Eburiola Thomson, 1864
Eburiola geminata (Fabricius, 1787). Monné & Bezark, 2008: 87
Tribe ELAPHIDIINI
Anelaphus Linsley, 1936
Anelaphus cinereus (Olivier, 1795). Ivie, 1985b: 307
Anelaphus crispulum (Fisher, 1947). Lingafelter, 2008: 354
Anelaphus hispaniolae (Fisher, 1932: 30). Blackwelder, 1946: 565 *H, *DR > TL: Port-au-Prince
Anelaphus mutatum (Gahan, 1890). Lingafelter, 2008: 357
=Elaphidion pilosum Fisher, 1932: 36. Lingafelter, 2008: 357
Anelaphus praeclarus Lingafelter, 2008: 359 *DR > TL: Guaraguao, PN del Este, La Altagracia Prov.
Anelaphus velteni† Vitali, 2009b: 352 [Fossil in Dominican amber]
Anopliomorpha Linsley, 1936
Anopliomorpha antillarum (Fisher, 1932: 42). Blackwelder, 1946: 564 *H > TL: Source Matelas
Curtomerus Stephens, 1839
Curtomerus flavus (Fabricius, 1775). Ivie, 1985b: 309
Curtomerus puncticollis (Fisher, 1932: 50) *H > TL: Port-au-Prince
Elaphidion Audinet-Serville, 1834
Elaphidion androsensis Fisher, 1942. Lingafelter, 2008: 358
Elaphidion auricoma Lingafelter, 2008: 366 *DR > TL: Guaraguao, PN del Este, La Altagracia Prov.
Elaphidion compressipenne Fisher, 1932: 35. Blackwelder, 1946: 565 *H > TL: Mariani
Elaphidion confusum Fisher, 1932: 32. Blackwelder, 1946: 565 *H > TL: Hinche
Elaphidion conspersum Newman, 1841. Russo, 1930: 142; Blackwelder, 1946: 565
Elaphidion costipenne Fisher, 1932: 39. Blackwelder, 1946: 565 *H > TL: Fond Parisien, south shore of Lake Assuei
Elaphidion fullonium Newman, 1841: 110. Blackwelder, 1946: 565; Lingafelter, 2008: 357 *H > TL: Island of Hayti
=Elaphidion confusum Fisher, 1932. Lingafelter, 2008: 357
Elaphidion inclusum† Vitali, 2007: 30 [Fossil in Dominican amber]
Elaphidion irroratum irroratum (Linnaeus, 1767). Russo, 1930: 142; Blackwelder, 1946: 565
Elaphidion iviei Lingafelter, 2008: 367
Elaphidion lucidum (Olivier, 1795: 59). Blackwelder, 1946: 565 *H, *DR > TL: Saint-Domingue
Elaphidion nearnsi Lingafelter, 2008: 370 *DR > TL: Punta Cana near Ecological reserve, La Altagracia Prov.
Elaphidion rotundipenne Fisher, 1932: 23. Blackwelder, 1946: 565 *DR > TL: Haina
Elaphidion sp.†—Poinar (1992: 138) [Fossil in Dominican amber]
Elaphidion spinicorne (Drury, 1773). Blackwelder, 1946: 565
Elaphidion splendidum Fisher, 1932: 19. Blackwelder, 1946: 565
Elaphidion tocanum† Vitali, 2009a: 224 [Fossil in Dominican amber]
Elaphidion tomentosum Chevrolat, 1862. Blackwelder, 1946: 565
Elaphidion wappesi Lingafelter, 2008: 371 *DR > TL: 14 km W Puerto Plata
Nesanoplium Chemsak, 1966
Nesanoplium puberulum (Fleutiaux & Sallé, 1889). Blackwelder, 1946: 571
Nesiosphaerion Martins & Napp, 1982
Nesiosphaerion charynae Lingafelter, 2008: 361 *DR > TL: 150 m N bridge on road Cabo Rojo—Aceitillar, Pedernales Prov.
Nesiosphaerion testaceum (Fisher, 1932: 45). Blackwelder, 1946: 566; Martins & Napp, 1982: 62 *H > TL: Manneville
Psyrassa Pascoe, 1866
Psyrassa woodleyi Lingafelter, 2008: 363 *DR > TL: Road 47 between Los Pinos and Angel Félix, Independencia Prov.

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Stizocera Serville, 1834
Stizocera dozieri (Fisher, 1947: 33). Monné, 2005a: 253 *H > TL: Kenscoff
Stizocera evanescens† Vitali, 2010: 340 [Fossil in Dominican amber]
Trichophoroides Linsley, 1935
Trichophoroides dozieri (Fisher, 1932: 38). Blackwelder, 1946: 565 *H > TL: Port-au-Prince
Trichophoroides signaticolle (Chevrolat, 1862). Monné, 2005a: 259
Tribe HESPEROPHANINI
Parandraceps Giesbert, 1998
Parandraceps turnbowi Giesbert, 1998: 75 *DR > TL: 20.5 km N Cabo Rojo, Pedernales Prov.
Tribe HETEROPSINI
Heterops Blanchard, 1842
Heterops hispaniolae Fisher, 1932: 72. Blackwelder, 1946: 586 *DR, *H > TL: Santo Domingo
Neopoeciloderma Monné & Martins, 1981
Neopoeciloderma lepturoides (Jacquelin du Val, 1857). Blackwelder, 1946: 586
Plectrocerum Dejean, 1835
Plectrocerum cribratum Sallé, 1856: 689. Blackwelder, 1946: 586 *DR > TL: Santo Domingo
Plectrocerum spinicorne (Olivier, 1795). Blackwelder, 1946: 586
Trichrous Chevrolat, 1858
Trichrous bicolor (Sallé, 1856: 687). Blackwelder, 1946: 586 *DR > TL: Santiago
Trichrous dimidiatipennis (Chevrolat, 1838). Blackwelder, 1946: 586
Trichrous fisheri Monné & Giesbert, 1992: 251 *H > TL: Port-au-Prince
=Trichrous bicolor Fisher, 1932: 70 [Preoccupied name]
Trichrous irroratus (Olivier, 1795). Blackwelder, 1946: 586
Trichrous jaegeri Chevrolat, 1858: 211. Blackwelder, 1946: 586 *DR > TL: Santo Domingo
Trichrous nigripes Fisher, 1942: 21. Blackwelder, 1946: 586 *DR > TL: Foothills of the Cordillera Central, south of Santiago
Trichrous violaceipennis Fisher, 1942: 22. Blackwelder, 1946: 586
Tribe IBIDIONINI
Heterachthes Newman, 1840
Heterachthes? zappii† Vitali, 2015: 8 [Fossil in Dominican amber]
Hormathus Gahan, 1890
Hormathus cinctellus Gahan, 1890: 33. Blackwelder, 1946: 569 *DR > TL: St. Domingo
Hormathus giesberti Lingafelter & Nearns, 2007: 180 *DR > TL: Punta Cana near Ecological Reserve, La Altagracia Prov.
Tribe METHIINI
Methia Newman, 1842
=Tessaropa Haldeman, 1847
Methia dolichoptera Lingafelter, 2010: 266 *DR > TL: Loma Hundida, Parque Juan B. Pérez Rancier, La Vega Prov.
Methia hispaniolae (Lingafelter, 2010: 265) *DR > TL: 25.5 km N Cabo Rojo, Pedernales Prov.
Methia necydalea (Fabricius, 1798). Blackwelder, 1946: 559
=Methia pallida Fisher, 1932: 6. Philips & Ivie, 1998: 72
Methia sp.†—Poinar (1992: 138) [Fossil in Dominican amber]
Tribe OEMINI
Malacopterus Audinet-Serville, 1834
Malacopterus tenellus (Fabricius, 1801). Lingafelter, 2010: 268
Tribe SMODICINI
Metaphrenon Martins, 1975
Metaphrenon impressicolle (Lacordaire, 1869). Russo, 1930: 142
Smodicum Haldeman, 1847
Smodicum brunneum Thomson, 1878. Russo, 1930: 142
Tribe TILLOMORPHINI
Calliclytus Fisher, 1932
Calliclytus macoris Lingafelter, 2011a: 63 *DR > TL: 12 km W San Pedro de Macorís, San Pedro de Macorís Prov.
Licracantha Lingafelter, 2011
Licracantha formicaria Lingafelter, 2011a: 60 *DR > TL: 4.7 km SE Jarabacoa, La Vega Prov.
Pentanodes Schaeffer, 1904

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Pentanodes sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Tribe TORNEUTINI
Chlorida Audinet-Serville, 1834
Chlorida festiva (Linnaeus, 1758). Wolcott, 1927: 238; Russo, 1930: 142; Blackwelder, 1946: 562
Tribe TRACHYDERINI
Trachyderes Dalman, 1817
Trachyderes (Trachyderes) succinctus succinctus (Linnaeus, 1758). Marcano, 1990: 5/90
Phoenicus Lacordaire, 1869
Phoenicus sanguinipennis Lacordaire, 1869. Lingafelter & Nearns, 2006: 201 *DR > TL: Originally described from an
unknown location. More recently collected at Punta Cana
Subfamily DISTENIINAE
Tribe DISTENIINI
Novantinoe Santos-Silva & Hovore, 2007
Novantinoe darlingtoni (Fisher, 1942: 1). Blackwelder, 1946: 557
Subfamily LAMIINAE
Tribe ACANTHOCININI
Alcidion Sturm, 1843
Alcidion unicolor (Fisher, 1932: 87). Blackwelder, 1946: 617 *H > TL: Furcy
Eugamandus Fisher, 1926
Eugamandus darlingtoni Fisher, 1942: 38. Blackwelder, 1946: 619 *DR > TL: Foothills of the Cordillera Central, south of
Santiago
Eugamandus? sp.†—Poinar (1992: 138) [Fossil in Dominican amber]
Kallyntrosternidius† Vitali, 2009
Kallyntrosternidius bucarensis† Vitali, 2009a: 228 [Fossil in Dominican amber]
Lagocheirus Dejean, 1835
Lagocheirus araneiformis araneiformis (Linnaeus, 1767). Wolcott, 1927: 238; Russo, 1930: 143; Dillon, 1957: 149
Leptostylopsis Dillon, 1956
Leptostylopsis annulipes (Fisher, 1942: 28). Blackwelder, 1946: 613; Lingafelter & Micheli, 2009: 7 *DR > TL: Sánchez
=Leptostylopsis hispaniolae (Fisher, 1942: 30). Gilmour, 1963: 59; Lingafelter & Micheli, 2009: 7
Leptostylopsis argentatus (Jacquelin du Val, 1857). Lingafelter & Micheli, 2009: 13
Leptostylopsis basifulvus Lingafelter & Micheli, 2009: 31 *DR > TL: Punta Cana, near Ecological Reserve
Leptostylopsis caliginosus Lingafelter & Micheli, 2009: 33 *DR, *H > TL: Pico Duarte Trail, La Vega Prov.
Leptostylopsis chlorescens Lingafelter & Micheli, 2009: 37 *DR > TL: Las Abejas, Parque Nacional Sierra de Bahoruco
Leptostylopsis cristatus (Fisher, 1925a: 7). Gilmour, 1963: 58; Lingafelter & Micheli, 2009: 16 *DR > TL: San Lorenzo
Leptostylopsis humerofulvus Lingafelter & Micheli, 2009: 39 *DR > TL: Las Abejas, Parque Nacional Sierra de Bahoruco
Leptostylopsis milleri (Fisher, 1932: 83). Gilmour, 1963: 59; Lingafelter & Micheli, 2009: 19 *DR > TL: Río San Juan
Leptostylopsis perfasciatus Lingafelter & Micheli, 2009: 43 *DR > TL: Filipinas, Barahona Prov.
Leptostylopsis poeyi (Fisher, 1925a: 4). Blackwelder, 1946: 614; Lingafelter & Micheli, 2009: 23 *DR, *H > TL: San
Francisco Mountains [San Cristóbal]
Leptostylopsis puntacanaensis Lingafelter & Micheli, 2009: 47 *DR > TL: Punta Cana, near Ecological Reserve
Leptostylopsis thomasi Lingafelter & Micheli, 2009: 50 *DR, *H > TL: Las Abejas, Parque Nacional Sierra de Bahoruco
Leptostylopsis viridicomus (Fisher, 1942: 29). Blackwelder, 1946: 615; Lingafelter & Micheli, 2009: 27 *DR, *H > TL: Valle
Nuevo
Styloleptus Dillon, 1956
Styloleptus atrovittatus (Fisher, 1925a). Gilmour, 1963: 66
Styloleptus biustus (LeConte, 1852). Russo, 1930: 142; Blackwelder, 1946: 614
Styloleptus dozieri (Fisher, 1932: 86). Gilmour, 1963: 66 *H > TL: Port-au-Prince
Styloleptus infuscatus (Fisher, 1932: 84). Gilmour, 1963: 66 *DR, *H > TL: Samaná
Styloleptus laticollis (Fisher, 1925a: 12). Russo, 1930: 142; Gilmour, 1963: 66
Styloleptus nigrofasciatus Gilmour, 1963: 67 *DR > TL: Jarabacoa
Styloleptus planicollis (Fisher, 1935). Monné, 2005b: 132
=Styloleptus divisus Gilmour, 1963: 70
=Styloleptus maraicola (Fisher, 1942: 35). Gilmour, 1963: 67
Trypanidius Blanchard, 1846

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Trypanidius insularis Fisher, 1925. Devesa & Barro, 2018: 3
Urgleptes Dillon, 1956
Urgleptes charinae Ravin & Lingafelter, 2015: 68 *DR > TL: Pico Duarte trail below La Compartición
Urgleptes conjunctus Ravin & Lingafelter, 2015: 71 *DR > TL: 36 km NW San José de Ocoa
Urgleptes curtipennis Ravin & Lingafelter, 2015: 74 *DR > TL: Sierra de Bahoruco, Rabo de Gato
Urgleptes marionae Ravin & Lingafelter, 2015: 76 *DR > TL: Monseñor Nouel Prov., Cabo Vito
Urgleptes obliteratus Ravin & Lingafelter, 2015: 79 *DR > TL: Pedernales Prov., 25 km N of Cabo Rojo
Urgleptes puertoricensis Gilmour, 1963. Ravin & Lingafelter, 2015: 57
Urgleptes sandersoni Gilmour, 1963. Ravin & Lingafelter, 2015: 60
=Urgleptes haitiensis Gilmour, 1963: 82
Tribe ACANTHODERINI
Oreodera Serville, 1835
Oreodera glauca glauca (Linnaeus, 1758). Blackwelder, 1946: 608
Steirastoma Lacordaire, 1830
Steirastoma acutipenne Sallé, 1856: 691. Blackwelder, 1946: 609 *H, *DR > TL: Dicayagua
Tribe AGAPANTHIINI
Spalacopsis Newman, 1842
Spalacopsis (Euthuorus) filum filum (Klug, 1829). Blackwelder, 1946: 606; Tyson, 1973: 121
Spalacopsis (Euthuorus) filum brevialatum Tyson, 1973: 125 *DR > TL: “Bosco del Infierno”, Samana Bay
Tribe APOMECYNINI
Acrepidopterum Fisher, 1926
Acrepidopterum pilosum Fisher, 1932: 75. Blackwelder, 1946: 599 *DR > TL: San Cristóbal
Dorcasta Pascoe, 1858
Dorcasta gracilis Fisher, 1932: 82. Blackwelder, 1946: 606 *H > TL: Fourgu
Parmenonta Thomson, 1868
Parmenonta dominicana Galileo & Martins, 2004: 57 *DR > TL: Las Abejas, 38 km NNW Cabo Rojo, Pedernales Prov.
Tribe CYRTININI
Cyrtinus Leconte, 1852
Cyrtinus fauveli (Cameron, 1909: 78). Blackwelder, 1946: 620 *H > TL: Carrefour, near Port-au-Prince
Tribe CALLIINI
Drycothaea Thomson, 1868
Drycothaea indistincta Lingafelter & Nearns, 2007: 188 *DR > TL: Punta Cana near Ecological Reserve, La Altagracia Prov.
Tribe DESMIPHORINI
Atimiola Bates, 1880
Atimiola rickstanleyi Lingafelter & Nearns, 2007: 185 *DR > TL: Las Abejas, PN Sierra de Bahoruco, Pedernales Prov.
Estola Fairmaire & Germain, 1859
Estola annulicornis Fisher, 1942: 26. Blackwelder, 1946: 600 *DR > TL: Foothills of the Cordillera Central, south of Santiago
Eupogonius LeConte, 1852
Eupogonius haitiensis Fisher, 1935: 197. Blackwelder, 1946: 600 *H > TL: Plaisance
Tribe HEMILOPHINI
Adesmus Lepeletier & Audinet-Serville, 1825
Adesmus fortunei Lingafelter, 2013: 58 *DR > TL: Pedernales Prov., N. of Cabo Rojo, Parque Nacional Sierra de Bahoruco,
km 26 on Carr. Alcoa, 691 m
Calocosmus Chevrolat, 1862
Calocosmus contortus Lingafelter, 2013: 63 *DR > TL: San Cristóbal Prov., Colonia Ramfis
Calocosmus chevrolati Fisher, 1925a. Lingafelter, 2013: 73
Calocosmus hispaniolae Fisher, 1925a: 15. Blackwelder, 1946: 625; Lingafelter, 2013: 75 *DR > TL: San Lorenzo
Calocosmus janus Bates, 1881. Lingafelter, 2013: 76
=Calocosmus holosericeus Gahan, 1889: 395. Blackwelder, 1946: 625
Calocosmus magnificus Fisher, 1932: 89. Blackwelder, 1946: 625; Lingafelter, 2013: 76 *H, *DR > TL: Valley Rivière Froide
Calocosmus melanurus Gahan, 1889: 395. Blackwelder, 1946: 625; Lingafelter, 2013: 78 *H, *DR > TL: San Domingo
Calocosmus nigripennis Chevrolat, 1862. Lingafelter, 2013: 79
Calocosmus nigritarsis Fisher, 1942: 41. Blackwelder, 1946: 625 *DR > TL: San José de las Matas
Calocosmus punctatus Lingafelter, 2013: 65 *DR > TL: Peravia Prov., 12.4 km E. Río Ocoa

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Calocosmus rawlinsi Lingafelter, 2013: 67 *DR > TL: Elias Piña Prov., Sierra de Neiba, 9.1 km WSW Hondo Valle, 1856 m
Calocosmus robustus Lingafelter, 2013: 69 *DR > TL: La Vega Prov., Parque Nacional Armando Bermúdez, km 1–3 along trail
W of La Ciénaga, 900–1100 m
Calocosmus semimarginatus Bates, 1881. Lingafelter, 2013: 81
Calocosmus thonalmus Lingafelter, 2013: 70 *DR > TL: La Altagracia Prov., El Verón, road to Hoyo Azul, 25–40 m
Oedudes Thomson, 1868
Oedudes anulatus Lingafelter, 2013: 61 *DR > TL: Peravia Prov., 5 km W of road to El Río, S. of Pedregal, 52 m
Paleohemilophus† Martins & Galileo, 1999
Paleohemilophus dominicanus† Martins & Galileo, 1999: 309; Vitali, 2007: 34; Lingafelter, 2013: 82 [Fossil in Dominican
amber]
Tribe POGONOCHERINI
Ecyrus LeConte, 1852
Ecyrus hirtipes Gahan, 1895. Ivie & Chemsak, 1983: 199
=Ecyrus hoffmanni Fisher, 1932: 78. Blackwelder, 1946: 600; Ivie & Chemsak, 1983: 199
Tribe PTEROPLIINI
Ataxia Haldeman, 1847
Ataxia haitiensis Fisher, 1932: 73. Blackwelder, 1946: 597 *H > TL: Hinche
Ataxia hovorei Lingafelter & Nearns, 2007: 184. Nearns & Tavakilian, 2012: 22 *DR > TL: Punta Cana near Ecological
Reserve, La Altagracia Prov.
Subfamily PARANDRINAE
Tribe PARANDRINI
Birandra Santos-Silva, 2002
Birandra (Yvesandra) laevis Santos-Silva & Shute, 2009: 32
=Parandra (Parandra) laevis Schoenherr, 1817. Blackwelder, 1946: 551
Birandra (Yvesandra) latreillei Santos-Silva & Shute, 2009: 24. Deler-Hernández & Santos-Silva, 2011: 224
Subfamily PRIONINAE
Tribe CALLIPOGONINI
Callipogon Audinet-Serville, 1832
Callipogon (Callomegas) sericeum (Olivier, 1795). Russo, 1930: 141; Blackwelder, 1946: 553; Linfafelter, 2015: 355
Orthomegas Audinet-Serville, 1832
Orthomegas cinnamomeus (Linnaeus, 1758). Lingafelter, 2015: 354
Tribe MACROTOMINI
Hovorodon Santos-Silva, Swift & Nearns, 2010
Hovorodon bituberculatum (Palisot de Beauvois, 1805: 216). Monné, 2005c: 54; Santos-Silva, Swift & Nearns, 2010: 27;
Lingafelter, 2015: 357
Stenodontes Audinet-Serville, 1832
Stenodontes exsertus (Olivier, 1795). Russo, 1930: 141; Blackwelder, 1946: 552; Lingafelter, 2015: 357
=Prionus laevigatus Palisot de Beauvois, 1805. Gahan, 1890: 24
Strongylaspis White, 1853
Strongylaspis corticarium (Erichson, 1848). Lingafelter, 2015: 355
Xixuthrus Thomson, 1864
Xixuthrus domingoensis Fisher, 1932: 1. Blackwelder, 1946: 552; Lingafelter & Woodley, 2007: 173; Lingafelter, 2015: 355
*DR > TL: Santiago
Tribe MEROSCELISINI
Monodesmus Audinet-Serville, 1832
Monodesmus atratus Fisher, 1932. Lingafelter, 2015: 358
Sarifer Kirsch, 1870
Sarifer seabrai Fragoso & Monné, 1982: 519. Monné, 2005c: 76; Lingafelter & Woodley, 2007: 172; Lingafelter, 2015: 358
*DR > TL: La Vega, Constanza, 1,200 m
Tribe PRIONINI
Trichoprionus Fragoso & Monné, 1982
Trichoprionus aureopilosus (Fragoso & Monné, 1982: 522). Monné, 2005c: 88; Lingafelter, 2015: 358; Santos-Silva et al.,
2016: 8 *DR > TL: La Vega, Constanza, Yaquecillo
Tribe SOLENOPTERINI

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Derancistrachroma Lingafelter, 2015
Derancistrachroma melanoleuca Lingafelter, 2015: 375 *DR > TL: 10 Km N of Oviedo
Derancistrus Audinet-Serville, 1832
Derancistrus anthracinus (Gahan, 1890). Russo, 1930: 141; Blackwelder, 1946: 554; Galileo & Martins, 1993: 201;
Lingafelter, 2015: 358
Derancistrus elegans (Palisot de Beauvois, 1805: 217). Blackwelder, 1946: 554; Galileo & Martins, 1993: 200; Lingafelter,
2015: 360 *H, *DR > TL: Saint-Domingue
Derancistrus hovorei Lingafelter & Woodley, 2007: 167; Lingafelter, 2015: 360 *DR > TL: 25 km N Cabo Rojo, Pedernales
Prov.
Elateropsis Chevrolat, 1862
Elateropsis antennatus Galileo & Martins, 1994: 126. Galileo & Martins, 1994: 126; Lingafelter, 2015: 360 *DR, *H > TL:
Sánchez
Elateropsis dichroma Lingafelter, 2015: 364 *DR > TL: 23 Km North of Cabo Rojo, Pedernales Prov.
Elateropsis femoratus (Sallé, 1856: 270). Blackwelder, 1946: 554; Galileo & Martins, 1994: 124; Lingafelter & Woodley,
2007: 172; Lingafelter, 2015: 361 *H, *DR > TL: Environs de la ville de Santo-Domingo
Elateropsis lineatus (Linnaeus, 1758). Galileo & Martins, 1994: 112; Lingafelter, 2015: 361
Elateropsis quinquenotatus Chevrolat, 1862. Lingafelter, 2015: 361
Elateropsis sericeiventris Chevrolat, 1862. Lingafelter, 2015: 363
Elateropsis trimarginatus (Cazier & Lacey, 1952). Lingafelter, 2015: 363
Elateropsis woodleyi Lingafelter, 2015: 365 *DR > TL: Punta Cana near Ecological Reserve
Solenoptera Audinet-Serville, 1832
Solenoptera dominicensis (Gahan, 1890: 26). Blackwelder, 1946: 554; Galileo & Martins, 1993c: 444; Lingafelter & Woodley,
2007: 170; Lingafelter, 2015: 366 *H, *DR > TL: St. Domingo
Solenoptera furfurosa (Galileo & Martins, 1993b: 205); Lingafelter, 2015: 366 *DR > TL: Prov. La Vega
Solenoptera helbi Lingafelter, 2015: 370 *DR > TL: Parque Nacional Armando Bermúdez, Río Bao
Solenoptera rugosa Lingafelter, 2015: 372 *DR > TL: 3–4 km S Constanza
Solenoptera scutellata (Gahan, 1890: 25). Blackwelder, 1946: 554; Monné, 2005c: 100; Lingafelter, 2015: 369
Solenoptera tomentosa Lingafelter, 2015: 373 *DR > TL: 3 km South of Los Tres Charcos
Solenoptera vittata (Olivier, 1795). Galileo & Martins, 1993a: 99; Lingafelter, 2015: 369
=Prosternodes oberthuri Gahan, 1895: 85

Family CHRYSOMELIDAE
Key reference: Takizawa (2003).
Subfamily GALERUCINAE
Tribe ALTICINI
Acallepitrix Bechyné, 1959
Acallepitrix sp.—Santiago-Blay & Craig (1999: 20)
Acrocyum Jacoby, 1885
Acrocyum haitiensis (Blake, 1938: 48). Takizawa, 2003: 55 *H > TL: Cape Haitien
Aedmon Clark, 1860
Aedmon albicincta (Blake, 1945a: 89). Takizawa, 2003: 55 *H > TL: Morne La Hotte, elev., 5,000–7,800 ft.
Aedmon calva (Blake, 1943: 426). Takizawa, 2003: 56 *DR > TL: Mt. Diego de Ocampo, 3–4,000 ft.
Aedmon comosa (Blake, 1943: 419). Takizawa, 2003: 56 *DR > TL: Loma Rucilla and mountains north, at 5,000 ft.
Aedmon constanzae (Blake, 1943: 432). Takizawa, 2003: 56 *DR > TL: Constanza, 3–4000 ft.
Aedmon corrugata (Blake, 1943: 425). Takizawa, 2003: 56 *DR > TL: Loma Rucilla and mountains north, 5–8000 ft.
Aedmon crispula (Blake, 1943: 426). Takizawa, 2003: 56 *DR > TL: Foothills of Cordillera Central, south of Santiago
Aedmon darlingtoni Blake, 1943: 420. Takizawa, 2003: 56 *DR > TL: Loma Rucilla and mountains north, 5–8,000 ft.
Aedmon elachia (Blake, 1943: 431). Takizawa, 2003: 56 *DR > TL: Puerto Plata
Aedmon flavicoma (Blake, 1943: 436). Takizawa, 2003: 57 *DR > TL: Foothills of Cordillera Central, south of Santiago
Aedmon glabroguttata Blake, 1943: 421. Takizawa, 2003: 57 *DR > TL: Loma Rucilla, alt. 8–10,000 ft.
Aedmon gracilenta (Blake, 1943: 429). Takizawa, 2003: 57 *H > TL: La Visite and vicinity, La Selle Range, 5–7000 ft. alt.
Aedmon guttata (Blake, 1943: 419). Takizawa, 2003: 57 *DR > TL: Loma Vieja, south of Constanza, at about 6000 ft.
Aedmon hoffmani (Blake, 1943: 435). Takizawa, 2003: 57 *H > TL: Río Froide, 1300 ft. alt.
Aedmon hugonis (Blake, 1943: 439). Takizawa, 2003: 57 *DR > TL: Cloudforest, vicinity of Valle Nuevo, circa 6000 ft.

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Aedmon minuta (Blake, 1943: 430). Takizawa, 2003: 57 *DR > TL: Villa Altagracia
Aedmon pallida (Blake, 1943: 435). Takizawa, 2003: 58 *DR > TL: Loma Rucilla and mountains north, 5–8000 ft. alt.
Aedmon robusta (Blake, 1943: 431). Takizawa, 2003: 58 *DR > TL: Constanza to Jarabacoa, 2–4000 ft. alt.
Aedmon verrucosa (Blake, 1943: 428). Takizawa, 2003: 59 *DR > TL: Vicinity of Valle Nuevo, cloudforest, circa 6000 ft.
Alagoasa Bechyné, 1955
Alagoasa bicolor (Linnaeus, 1767: 593). Takizawa, 2003: 59
=Oedionychis lantanae Blake, 1928: 95
=Oedinychis purpurella Blake, 1940: 172
Alagoasa cinctus (Linnaeus, 1767). Takizawa, 2005: 200
Alagoasa dimidiatus (Olivier, 1808: 676). Takizawa, 2003: 59 *DR > TL: St. Domingo
Alagoasa fasciatus (Fabricius, 1798: 96). Takizawa, 2003: 60
Alagoasa megalopia (Blake, 1939: 233) *DR > TL: San José de las Matas, 1–2000 ft.
Alagoasa viridipennis (Blake, 1971: 278). Takizawa, 2003: 60 *DR > TL: Aceitillar, Pedernales
Aphthona Chevrolat, 1842
Aphthona compressa (Suffrian, 1868). Takizawa, 2003: 61
Aphthona elachia Blake, 1948a: 276. Takizawa, 2003: 61 *H > TL: Morne Trou d’Eau
Aphthona fraterna Blake, 1948b: 147. Takizawa, 2003: 61 *H > TL: Desbarriere, Mt. La Hotte, near 4000 ft.
Apraea Baly, 1877
Apraea aenea (Blake, 1948c: 124). Takizawa, 2003: 62 *DR > TL: Loma Vieja, south of Constanza, about 6,000 ft. elevation
Apraea fulgens Blake, 1963: 20. Takizawa, 2003: 63 *H > TL: Port-au-Prince
Apraea metallica Medvedev, 2004: 124 *DR > TL: Baní, 65 m
Apraea oculata Medvedev, 2004: 124 *DR > TL: Baní, 65 m
Argopistes Motschulsky, 1860
Argopistes woodleyi Blanco & Konstantinov, 2013: 262 *DR > TL: La Altagracia Prov., El Verón, road to Hoyo Azul
Bonfilsus Scherer, 1967
Bonfilsus sofiae Konstantinov & Micheli in Micheli & Konstantinov, 2019: 3 *DR > TL: Pedernales Prov., 3 km (st3) N of
Cabo Rojo, 1125 m
Centralaphthona Bechyné & Bechyné, 1960
Centralaphthona dominicana Medvedev, 2004: 123 *DR > TL: Boca Chica, 10 m
Chaetocnema Stephens, 1831
Chaetocnema apricaria (Suffrian, 1878). Takizawa, 2003: 65
Chaetocnema brunnescens brunnescens Horn, 1889. Takizawa, 2003: 66
Chaetocnema obesula LeConte, 1878. Takizawa, 2003: 66
Chaetocnema platicephala Blake, 1969. Takizawa, 2003: 67
Chaetocnema plicipennis Blake, 1941: 176. Takizawa, 2003: 67 *DR > TL: Valle Nuevo, southeast of Constanza
Cyrsylus Jacoby, 1891
Cyrsylus hispaniolae Blake, 1949: 371. Takizawa, 2003: 67 *H > TL: Froide River Valley
Disonycha Chevrolat, 1844
Disonycha comma White, 1990: 362. Takizawa, 2003: 68 *DR > TL: Taken at West Palm Beach, Fla., with Cucumis melo fruit
[exported from Dominican Republic]
Disonycha eximia Harold, 1876. Takizawa, 2003: 68; Takizawa, 2005: 204
Disonycha spilotrachela Blake, 1928: 96. Takizawa, 2003: 69; Takizawa, 2005: 201
Distigmoptera Blake, 1943
Distigmoptera antennata Medvedev, 2004: 122 *DR > TL: Loma Casabito, 1250 m
Epitrix Foudras, 1860
Epitrix cucumeris (Harris, 1851). Bienkowski & Orlova-Bienkowskaja, 2016: 414 [Introduced]
Epitrix tuberis Gentner, 1944. Bienkowski & Orlova-Bienkowskaja, 2016: 421 [Introduced]
Heikertingerella Csiki, 1940
Heikertingerella darlingtoni Blake, 1963: 19. Takizawa, 2003: 72 *DR > TL: Mt. Diego de Ocampo, 3–4000 ft.
Heikertingerella minima (Suffrian, 1868). Blake, 1960: 101; Takizawa, 2003: 72
Hemilactica Blake, 1937
Hemilactica graphica Blake, 1939: 233. Takizawa, 2003: 73 *DR > TL: Mt. Diego de Ocampo, 3–4000 ft.
Hirtasphaera Medvedev, 2004
Hirtasphaera hirsuta Medvedev, 2004: 121 *DR > TL: Constanza, 1250 m

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Homoschema Blake, 1950
Homoschema biscutatum Blanco & Duckett, 2001: 7 *H > TL: 32 km SW Mirebalais
Homoschema buscki Blake, 1950b: 20. Blanco & Duckett, 2001: 8 *DR > TL: San Francisco Mts.
Homoschema furthi Blanco & Duckett, 2001: 9 *DR > TL: Río Cumayasa, San Pedro de Macorís-La Romana, km 14 (bajo
Puente)
Homoschema hoffmani Blake, 1950b: 15. Blanco & Duckett, 2001: 10 *H > TL: Bizoton
Homoschema leucurum Blake, 1950b: 15. Blanco & Duckett, 2001: 11 *H > TL: La Vanneau
Homoschema lingulatum Blanco & Duckett, 2001: 12 *DR > TL: Peravia, 15 km E. San José de Ocoa
Homoschema opimum Blake, 1950b: 23. Blanco & Duckett, 2001: 14 *DR > TL: San José de las Matas, 1000–2000 ft. alt.
Kiskeya Konstantinov & Chamorro-Lacayo, 2006
Kiskeya baorucae Konstantinov & Chamorro-Lacayo, 2006: 284 *DR > TL: Pedernales Prov., PN Sierra de Bahoruco, Las
Abejas, 1270 m
Kiskeya neibae Konstantinov & Chamorro-Lacayo, 2006: 286 *DR > TL: Elías Piña Prov., Sierra de Neiba, 1.5 km E of
military post 204, 1597 m
Leptophysa Baly, 1877
Leptophysa hispaniolae (Blake, 1941: 174). Takizawa, 2003: 78 *DR > TL: Cloud forest, vicinity of Valle Nuevo, about 6000
ft.
Leptophysa hottensis (Blake, 1948b: 145). Takizawa, 2003: 78 *H > TL: Desbarriere, Mt. La Hotte, about 4000 ft. elevation
Leptophysa ornata Medvedev, 2004: 124 *DR > TL: Loma Casabito, 1250 m
Leptophysa punctatissima (Blake, 1948b: 146). Takizawa, 2003: 78 *H > TL: Desbarriere, Mt. La Hotte, about 4000 ft.
elevation
Longitarsus Latreille, 1829
Longitarsus hoffmani Blake, 1965b: 107. Takizawa, 2003: 79 *H > TL: Mt. Cabrite
Longitarsus photinus Blake, 1965b: 107. Takizawa, 2003: 80 *DR > TL: Loma Rucilla, 8,000–10,000 ft. alt.
Lupraea Jacoby, 1885
Lupraea hoffmani (Blake, 1950c: 183). Takizawa, 2003: 81 *H > TL: La Vanneau
Lysathia Bechyné, 1959
Lysathia occidentalis (Suffrian, 1868). Takizawa, 2003: 81; Takizawa, 2005: 197
Macrohaltica Bechyné, 1959
Macrohaltica amethystina (Olivier, 1808). Takizawa, 2003: 81
Macrohaltica jamaicensis (Fabricius, 1792). Takizawa, 2003: 81; Takizawa, 2005: 198
Megistops Boheman, 1859
Megistops dissita Blake, 1931: 81. Takizawa, 2003: 82 *H > TL: Camp Perrin, La Prise
Megistops liturata (Olivier, 1808). Blake, 1928: 97; Takizawa, 2003: 82
Monomacra Chevrolat, 1837
Monomacra porphyrea (Blake, 1948b: 143). Takizawa, 2003: 85 *DR > TL: Mt. Diego de Ocampo, circa 3–4000 ft. elevation
Normaltica Konstantinov, 2002
Normaltica iviei Konstantinov, 2002: 15 *DR > TL: La Vega, 53 km SE Constanza
Omophoita Chevrolat, 1836
Omophoita albicollis (Fabricius, 1787). Takizawa, 2003: 65
Omophoita cyanipennis (Fabricius, 1798). Takizawa, 2003: 88
Pseudodisonycha Blake, 1954
Pseudodisonycha hispaniolae Blake, 1954: 249. Takizawa, 2003: 89 *DR > TL: Mount Diego de Ocampo, 3,000–4,000 ft.
altitude
Strabala Chevrolat, 1837
Strabala ferruginea (Olivier, 1808: 697). Blake, 1953: 132; Takizawa, 2003: 90
Syphraea Baly, 1876
Syphraea constanzae (Blake, 1965b: 108). Takizawa, 2003: 91 *DR > TL: Constanza, 3,000–4,000 ft. alt.
Syphraea darlingtoni (Blake, 1965b: 108). Takizawa, 2003: 91 *H > TL: Desbarriere, Mt. La Hotte, near 4,000 ft. alt.
Syphraea punctata (Blake, 1965b: 109). Takizawa, 2003: 92 *DR > TL: San José de las Matas, 1,000–2,000 ft. alt.
Systena Chevrolat, 1837
Systena basalis (Jacquelin du Val, 1856). Wolcott, 1927: 239; Takizawa, 2003: 92
Wanderbiltiana Bechyné, 1955
Wanderbiltiana wawasita† Santiago-Blay et al., 2004: 275 [Fossil in Dominican amber]

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 =Walterianella sp.†—Santiago-Blay et al. (1996: 421) [Fossil in Dominican amber]
Tribe GALERUCINI
Acalymma Barber, 1947
Acalymma bivittatum (Fabricius, 1801). Takizawa, 2003: 45
Cerotoma Chevrolat, 1837
Cerotoma ruficornis (Olivier, 1791). Schmutterer, 1990: 371; Takizawa, 2003: 45
Dicoelotrachelus Blake, 1941
Dicoelotrachelus brevicollis Blake, 1941: 173. Takizawa, 2003: 45 *DR > TL: Foothills of Cordillera Central, south of
Santiago
Dicoelotrachelus darlingtoni Blake, 1941: 172. Takizawa, 2003: 47 *DR > TL: Loma Rucilla and mountains north, alt. 5–
8,000 ft.
Dicoelotrachelus depilatus Blake, 1941: 173. Takizawa, 2003: 47 *H > TL: La Visite, La Selle Range, 5–7000 ft. alt.
Ectmesopus Blake, 1940
Ectmesopus angusticollis Blake, 1940a: 101. Takizawa, 2003: 47 *DR > TL: Duarte, Santo Domingo city
Ectmesopus crassicornis Blake, 1940a: 97. Takizawa, 2003: 47 *DR > TL: Puerto Plata
Ectmesopus darlingtoni Blake, 1940a: 103. Takizawa, 2003: 47 *H > TL: Port-au-Prince and vicinity
Ectmesopus leonardorum Blake, 1958: 97. Takizawa, 2003: 47 *H > TL: Bassin Bleu, Tortue Island
Ectmesopus longicornis Blake 1940a: 103. Takizawa, 2003: 48 *H > TL: Camp Perrin
Ectmesopus pallidus Blake, 1940a: 98. Takizawa, 2003: 48 *DR > TL: Foothills Cordillera Central, south of Santiago
Gonaives Clark, 1987
Gonaives buenae Clark, 1987: 168. Takizawa, 2003: 49 *H > TL: 10 mi. SE Gonaives
Isotes Weise, 1922
Isotes albicornis (Fabricius, 1798: 96). Takizawa, 2003: 49
Isotes darlingtoni (Blake, 1937: 69). Takizawa, 2003: 49 *H > TL: Kenscoff (near Por-au-Prince), 4–6000 ft.
Leptonesiotes Blake, 1958
Leptonesiotes sp.†—Santiago-Blay et al. (1996: 25) [Fossil in Dominican amber]
Leptonesiotes virkkii† Santiago-Blay et al., 1996: 419 [Fossil in Dominican amber]
Miraces Jacoby, 1888
Miraces glaber (Blake, 1946: 113). Takizawa, 2003: 51 *H > TL: Trou Caiman
Nyctiplanctus Blake, 1963
Nyctiplanctus ferruginea (Blake, 1963: 14). Takizawa, 2003: 51 *H > TL: Port-au-Price and vicinity
Nyctiplanctus hispaniolae (Blake, 1948c: 122). Takizawa, 2003: 51 *DR > TL: Constanza to Jarabacoa, elevation 2–4,000 ft.
Oedionychus Berthold, 1827
Oedionychus fasciatus (Fabricius, 1798). Blake, 1940b: 170
Paranapiacaba Bechyné, 1958
Paranapiacaba hispaniolae (Blake, 1963: 18). Takizawa, 2003: 52 *DR, *H > TL: Valle del Río, Prov. La Vega
Paranapiacaba marcanoi (Blake, 1971: 277). Takizawa, 2003: 52 *DR > TL: Los Tablones, Jarabacoa
Yingaresca Bechyné, 1956
Yingaresca bowditchi (Blackwelder, 1946: 688). Takizawa, 2003: 53
=Galerucella conjuncta Blake, 1937: 68
Yingaresca chrysura (Blake, 1939: 232). Takizawa, 2003: 53 *DR > TL: Constanza to Jarabacoa, 2–4000 ft.
Yingaresca constanzae Blake, 1939: 231. Takizawa, 2003: 53 *DR > TL: Constanza, 3–4000 ft.
Yingaresca decemvittata (Blake, 1971: 274). Takizawa, 2003: 53 *DR > TL: Loma de Sicajagua, Jánico
Subfamily CASSIDINAE
Agathispa Weise, 1905
Agathispa dimidiata (Olivier, 1808: 776). Blake, 1940b: 171; Takizawa, 2003: 93 *H, *DR > TL: Saint-Domingue
Agroiconota Spaeth, 1913
Agroiconota propinqua (Boheman, 1855). Takizawa, 2003: 97; Borowiec, 2009b: 621
Anisostena Weise, 1910
Anisostena cyanoptera (Suffrian, 1868: 227). Staines, 1994: 130; Takizawa, 2003: 93
Asteriza Chevrolat, 1836
Asteriza blakeae Shin et al., 2012: 48 *DR > TL: Prov. Barahona: Eastern Sierra de Bahoruco, Cachote, 12.8 km NE Paraiso,
1230 m
Asteriza darlingtoni Blake, 1939: 238. Takizawa, 2003: 97; Shin et al., 2012: 44 *DR > TL: Constanza to Jarabacoa and

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 159
Constanza, 2–4000 ft.
Asteriza flavicornis (Olivier, 1790: 393). Boheman, 1854: 496; Wolcott, 1927: 242; Takizawa, 2003: 97; Shin et al., 2012: 43
Asteriza tainosa Shin et al., 2012: 45 *DR > TL: Prov. Independencia: Sierra de Bahoruco, north slope, 13.5 km S. E. Puerto
Escondido, alt. 1807 m
Cassida Linnaeus, 1758
Cassida (Deloyala) fuliginosa (Olivier, 1808). Takizawa, 2003: 100; Borowiec, 2009b: 664
Cassida (Deloyala) guttata (Olivier, 1790). Blackwelder, 1946: 748
Chalepus Thunberg, 1805
Chalepus sanguinicollis (Linnaeus, 1771). Olivier, 1808: 765; Takizawa, 2003: 94
Charidotella Weise, 1896
Charidotella dominicanensis Borowiec, 2011: 496 *DR > TL: La Vega, Constanza, 5 km SE of Jarabacoa, 600 m
Charidotella latevittata (Boheman, 1855: 151). Takizawa, 2003: 98; Borowiec, 2009b: 636
Charidotella praeusta (Boheman, 1855: 152). Takizawa, 2003: 98; Borowiec, 2009b: 637
Charidotella quadrisignata (Boheman, 1855: 150). Takizawa, 2003: 98
Charidotella subsignata (Boheman, 1862: 403). Takizawa, 2003: 99
Chelymorpha Boheman, 1854
Chelymorpha cribraria (Fabricius, 1775). Thomas, 1994: 1; Takizawa, 2003: 99; Borowiec, 2009b: 649
Chelymorpha testaceomarginata Boheman, 1854. Borowiec, 2009b: 653
Convexocoleus Shin, 2013
Convexocoleus rileyi Shin, 2013: 526 *H > TL: Dept. Grand’Anse, Jérémie, 0.5 km West of summit of Pic Macaya, 2,300 m
Delocrania Guérin-Méneville, 1844
Delocrania sp.†—Farrell et al. (1992: 37); Santiago-Blay et al. (1996: 422) [Fossil in Dominican amber]
Demotispa Baly, 1858
Demotispa sallei Baly, 1858: 167. Takizawa, 2003: 94 *H, *DR > TL: St. Domingo
Elytrogona Chevrolat, 1837
Elytrogona bacca Boheman, 1850: 441. Chaboo, 2000: 386; Takizawa, 2003: 101 *DR, *H > TL: Insula St. Domingo
Elytrogona bulla Boheman, 1862: 193. Borowiec, 1999: 114; Sekerka, 2016: 328
=Elytrogona gemmata Blake, 1930: 222. Chaboo, 2000: 388; Takizawa, 2003: 101; Sekerka, 2016: 328
Elytrogona nigrodorsata Blake, 1937: 78. Chaboo, 2000: 389; Takizawa, 2003: 101 *DR, *H > TL: Mt. Basil, northern Haiti,
4700 ft.
Elytrogona quatuordecimmaculata (Latreille, 1802). Wolcott, 1927: 243; Chaboo, 2000: 386; Takizawa, 2003: 102 *H, *DR >
TL: Saint-Domingue
=Cassida ampulla Olivier, 1808. Boheman, 1850: 440
Eurypedus Gistel, 1834
Eurypedus nigrosignatus (Boheman, 1854). Simões & Peterson, 2018: 11 [Introduced]
Hemisphaerota Chevrolat, 1837
Hemisphaerota palmarum (Boheman, 1862: 40). Takizawa, 2003: 103; Chaboo & Nguyen, 2004: 171; Borowiec, 2009b: 673
*DR, *H > TL: St. Domingo
Octhispa Chapuis, 1877
Octhispa gibba (Olivier, 1792: 98). Takizawa, 2003: 95
Odontota Dejean, 1835
Odontota dorsalis (Thunberg, 1805). Takizawa, 2003: 96
Omoteina Chevrolat, 1837
=Paratrikona Spaeth, 1923
Omoteina aculeata (Boheman, 1854: 170). Takizawa, 2003: 101; Simões & Sekerka, 2015: 232; Simões, 2018: 10 *H, *DR >
TL: Insula St. Domingo
=Dorynota pubescens (Blake, 1939: 234). Takizawa, 2003: 101; Simões & Sekerka, 2015: 232 *DR > TL: Constanza, 3–4000
ft.
Omoteina humeralis (Olivier, 1808: 964). Boheman, 1862: 242; Wolcott, 1927: 243; Takizawa, 2003: 104; Simões, 2018: 10
Omoteina lerouxii (Boheman, 1854). Simões, 2017: 418; Simões, 2018: 10
Omoteina ovata (Blake, 1938: 51). Takizawa, 2003: 104; Simões, 2017: 420; Simões, 2018: 10 *H > TL: Grande Rivière
Omoteina rubescens (Blake, 1939: 238). Blake, 1971: 282; Takizawa, 2003: 104; Sekerka, 2016: 300; Simões, 2017: 421;
Simões, 2018: 10 *DR > TL: Jarabacoa, 1500–4000 ft.
=Paratrikona albomaculata Borowiec, 2009a: 567; Sekerka, 2016: 300; Simões, 2017: 421

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Omoteina turrifera (Boheman, 1854: 171). Takizawa, 2003: 105; Simões, 2017: 424; Simões, 2018: 10 *DR > TL: St.
Domingo
Omoteina variegata (Blake, 1939: 236). Takizawa, 2003: 105; Simões, 2017: 424; Simões, 2018: 10 *DR > TL: Constanza, 3–
4000 ft.
Sceloenopla Chevrolat, 1835
Sceloenopla ambarensis† Santiago-Blay et al., 1996: 421 [Fossil in Dominican amber]
Sceloenopla dominicana† Samuelson & Staines, 2000: 61 [Fossil in Dominican amber]
Sceloenopla stainesorum† Santiago-Blay & Craig, 1999: 22 [Fossil in Dominican amber]
Xenochalepus Weise, 1910
Xenochalepus cyanura Blake, 1971: 279. Takizawa, 2003: 96 *H > TL: Corail
Subfamily CHRYSOMELINAE
Leucocera Chevrolat, 1837
Leucocera aeneomicans (Stål, 1857). Takizawa, 2003: 41
Leucocera cubana Weise, 1916. Takizawa, 2003: 43
Leucocera cyanea Blake, 1971: 273. Takizawa, 2003: 41 *DR > TL: Esperanza
Leucocera hoffmani Blake, 1928: 93. Takizawa, 2003: 42 *H > TL: Hinche
Leucocera insulana (Jacquelin du Val, 1856: 302). Takizawa, 2003: 42
Leucocera octopustulata Stål, 1858. Stål, 1862: 202; Takizawa, 2003: 42
Leucocera quadriguttata Chevrolat, 1882: lxxx. Blackwelder, 1946: 677 *H, *DR > TL: Santo-Domingo ?
Leucocera quinquepunctata (Linnaeus, 1767: 586). Takizawa, 2003: 43 *H, *DR > TL: America
=Plagiodera semicaerulea Achard, 1925. Blackwelder, 1946: 678
Leucocera spilota Blake, 1971: 273. Takizawa, 2003: 43 *DR > TL: San Cristóbal
Stenaspidiotus† Poinar, 2013
Stenaspidiotus microptilus† Poinar, 2013: 102 [Fossil in Dominican amber]
Subfamily CRIOCERINAE
Lema Fabricius, 1798
Lema extersignata Pic, 1947: 10 *H > TL: Haiti
Lema insignata Pic, 1947: 9 *H > TL: Haiti
Lema mannerheimi Lacordaire, 1845. Takizawa, 2003: 7
Neolema Monrós, 1951
Neolema dorsalis (Olivier, 1791). Takizawa, 2003: 6
Neolema gundlachiana (Suffrian, 1874). Takizawa, 2003: 6
Subfamily CRYPTOCEPHALINAE
Chlamisus Rafinesque, 1815
Chlamisus haitiensis (Bowditch, 1913: 3). Takizawa, 2003: 21 *H, *DR > TL: Haiti
=Chlamisus insularis Jacoby, 1901: 157
Cryptocephalus Geoffroy, 1762
Cryptocephalus balteatus Suffrian, 1851: 264. Takizawa, 2003: 9 *H, *DR > TL: St. Domingo
Cryptocephalus cameroni Bryant, 1957: 828. Takizawa, 2003: 9 *H > TL: Carrefour
Cryptocephalus chevrolati Bryant, 1957: 830. Takizawa, 2003: 10 *H, *DR > TL: St. Domingo
Cryptocephalus chloroticus (Olivier, 1808). Takizawa, 2003: 10
Cryptocephalus crabroniformis (Suffrian, 1852: 94). Takizawa, 2003: 11 *H, *DR > TL: St. Domingo
Cryptocephalus dissectus Suffrian, 1851: 262. Takizawa, 2003: 11 *H, *DR > TL: St. Domingo
Cryptocephalus exaratus Suffrian, 1851: 271. Takizawa, 2003: 12 *H > TL: Port-au-Prince
Cryptocephalus fuliginosus Suffrian, 1851: 257. Takizawa, 2003: 12 *H, *DR > TL: St. Domingo
Cryptocephalus kheelorum† Chamorro & Bukejs in Bukejs & Chamorro, 2015: 121 [Fossil in Dominican amber]
Cryptocephalus haitiensis Jacoby, 1908: 843. Takizawa, 2003: 13 *H > TL: Port-au-Prince
Cryptocephalus hilaris Suffrian, 1851: 283. Takizawa, 2003: 13 *H, *DR > TL: St. Domingo
Cryptocephalus multiguttatus Suffrian, 1851. Takizawa, 2003: 14
Cryptocephalus octodecimguttatus Suffrian, 1851: 253. Takizawa, 2003: 15 *H, *DR > TL: St. Domingo
Cryptocephalus quatordecimsignatus Suffrian, 1851: 260. Takizawa, 2003: 15 *H, *DR > TL: St. Domingo
Cryptocephalus rubricus Suffrian, 1851: 316. Takizawa, 2003: 16 *H > TL: Port-au-Prince
Cryptocephalus sp. or Pachybrachis sp.†—Santiago-Blay et al. (1996: 415) [Fossil in Dominican amber]
Cryptocephalus strigicollis Suffrian, 1851. Takizawa, 2003: 17 *H, *DR > TL: St. Domingo

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 161
Diachus LeConte, 1880
Diachus sp.†—Santiago-Blay et al. (1996: 416) [Fossil in Dominican amber]
Pachybrachis Chevrolat, 1837
Pachybrachis carmelita Suffrian, 1852: 183. Takizawa, 2003: 19 *H > TL: Port-au-Prince
Pachybrachis sparsuta Suffrian, 1852: 145. Takizawa, 2003: 20 *H > TL: Port-au-Prince
Subfamily EUMOLPINAE
Alethaxius Lefevre, 1885
Alethaxius darlingtoni Blake, 1945b: 324. Takizawa, 2003: 24 *DR > TL: Loma Rucilla and mountains north, 5,000–8,000 ft.
Alethaxius hispaniolae Blake, 1945b: 323. Takizawa, 2003: 24 *DR > TL: Loma Vieja, ca. 6,000 ft., south of Constanza
Alethaxius integer Blake, 1945b: 324. Takizawa, 2003: 24 *DR > TL: Mount Diego de Ocampo, 3,000–4,000 ft. altitude
Alethaxius sp.†—Santiago-Blay et al. (1996: 417) [Fossil in Dominican amber]
Alethaxius tortuensis Blake, 1947a: 314. Takizawa, 2003: 24 *H > TL: Bassin Bleu, Isle Tortue, north of Haiti
Chalcosicya Blake, 1930
Chalcosicya acuminata Blake, 1951a: 291. Takizawa, 2003: 25 *DR > TL: Loma Rucilla and mountains north, 5,000–8,000 ft.
Chalcosicya aptera Blake, 1951a: 300. Takizawa, 2003: 25 *DR > TL: Azua
Chalcosicya constanzae Blake, 1951a: 303. Takizawa, 2003: 25 *DR > TL: Constanza
Chalcosicya convexa Blake, 1951a: 304. Takizawa, 2003: 25 *DR > TL: San José de las Matas, 1,000–2,000 ft.
Chalcosicya darlingtoni Blake, 1951a: 303. Takizawa, 2003: 26 *DR > TL: Constanza, 3,000–4,000 ft.
Chalcosicya fraterna Blake, 1951a: 309. Takizawa, 2003: 26 *DR > TL: Loma Rucilla and mountains north, 5,000–8,000 ft.
Chalcosicya gemina Blake, 1951a: 308. Takizawa, 2003: 26 *DR > TL: Loma Rucilla and mountains north, 5,000–8,000 ft.
Chalcosicya glabra Blake, 1951a: 307. Takizawa, 2003: 26 *DR > TL: Foothills of Cordillera Central, south of Santiago
Chalcosicya humeralis Blake, 1971: 270. Takizawa, 2003: 27 *DR > TL: Los Ramones proximo Palo Alto, Santiago en
Cordillera Septentrional, 1000 meters alt.
Chalcosicya parvula Blake, 1951a: 311. Takizawa, 2003: 27 *DR > TL: Loma Rucilla and mountains north, 5,000–8,000 ft.
Chalcosicya senilis Blake, 1951a: 306. Takizawa, 2003: 27 *DR > TL: Loma Vieja, south of Constanza, about 6,000 ft.
Chalcosicya setosella Blake, 1971: 269. Takizawa, 2003: 28 *DR > TL: Pico Duarte
Chalcosicya sp.†—Santiago-Blay et al. (1996: 18) [Fossil in Dominican amber]
Chalcosicya teres Blake, 1951a: 308. Takizawa, 2003: 28 *DR > TL: Loma Rucilla and mountains north, 5,000–8,000 ft.
Chalcosicya truncata Blake, 1951a: 310. Takizawa, 2003: 28 *DR > TL: Loma Rucilla and mountains north, 5,000–8,000 ft.
Chalcosicya villosa Blake, 1951a: 305. Takizawa, 2003: 28 *DR > TL: Valle Nuevo, S. E. of Constanza, about 7,000 ft.
Colaspis Fabricius, 1801
Colaspis amplicollis Blake, 1971: 272. Takizawa, 2003: 28 *DR > TL: Majagual, Sánchez
Colaspis chevrolati Lefèvre, 1891: cclvii. Takizawa, 2003: 28 *H, *DR > TL: St. Domingue
Colaspis darlingtoni Blake, 1967: 234. Takizawa, 2003: 29 *DR > TL: Foothills of the Cordillera Central, south of Santiago
Colaspis purpurea Blake, 1971: 271. Takizawa, 2003: 30 *DR > TL: El Cachón, Jimaní
Colaspis smaragdula Olivier, 1808: 883. Takizawa, 2003: 30 [Haiti, *Endemic]
Colaspoides Laporte, 1833
Colaspoides? sp.†—Santiago-Blay et al. (1996: 417) [Fossil in Dominican amber]
Glyptoscelis LeConte, 1859
Glyptoscelis brevicornis (Olivier, 1808: 914). Takizawa, 2003: 31 *H, *DR > TL: Saint-Domingue
Glyptoscelis sp.†—Santiago-Blay et al. (1996: 417) [Fossil in Dominican amber]
Metachroma Chevrolat, 1837
Metachroma barahonense Blake, 1946: 23. Takizawa, 2003: 32 *DR > TL: Barahona
Metachroma brevistriatum Blake, 1946: 23. Takizawa, 2003: 32 *DR > TL: San José de las Matas, 1,000–2,000 ft.
Metachroma cinctipes Blake, 1946: 26. Takizawa, 2003: 32 *DR > TL: Mt. Diego de Ocampo, altitude 3,000–4,000 ft.
Metachroma darlingtoni Blake, 1946: 24. Takizawa, 2003: 33 *DR > TL: Constanza to Jarabacoa, 2,000–4,000 ft. altitude
Metachroma dentatum Blake, 1965a: 5. Takizawa, 2003: 33 *DR > TL: Santo Domingo
Metachroma florissantensis† Santiago-Blay, 1994: 19 [Fossil in Dominican amber]
Metachroma hottense Blake, 1948a: 279. Takizawa, 2003: 35 *H > TL: Desbarriere, Morne La Hotte, near 4,000 ft.
Metachroma laevius Blake, 1946: 22. Takizawa, 2003: 35 *DR > TL: Mt. Diego de Ocampo, 3,000–4,000 ft. altitude
Metachroma leonardi Blake, 1970: 61. Takizawa, 2003: 35 *H > TL: St. Louis, Tortue Island
Metachroma leucurum Blake, 1970: 66. Takizawa, 2003: 35 *H > TL: Port-au-Prince
Metachroma longipunctatum Blake, 1966: 217. Takizawa, 2003: 36 *H > TL: La Visite and vicinity, La Selle Range, 5,000–
7,000 ft. alt.

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Metachroma melanochrotum Blake, 1970: 66. Takizawa, 2003: 36 *H > TL: Carrefour
Metachroma nigromaculatum Blake, 1962: 175. Takizawa, 2003: 37 *H > TL: Kenscoff
Metachroma nigroviride Blake, 1948a: 278. Takizawa, 2003: 37 *DR > TL: Puerto Plata
Metachroma obesum Blake, 1946: 23. Takizawa, 2003: 37 *DR > TL: Constanza, 3,000–4,000 ft. altitude
Metachroma sp.†—Santiago-Blay et al. (1996: 417) [Fossil in Dominican amber]
Metachroma wolcotti Bryant, 1926: 403. Takizawa, 2003: 39 *H > TL: Port-au-Prince
Myochrous Erichson, 1874
Myochrous hispaniolae Blake, 1947b: 24. Takizawa, 2003: 40 *H, *DR > TL: Swamps north of Dessalines
Subfamily LAMPROSOMATINAE
Oomorphus Curtis, 1831
Oomorphus darlingtoni Monrós, 1956: 48. Takizawa, 2003: 23 *H > TL: Mts. near Port-au-Prince, up to 2000 ft.
Subfamily BRUCHINAE
Acanthoscelides Schilsky, 1905
Acanthoscelides apicalis (Sharp, 1885). Johnson, 1990: 327
Acanthoscelides argillaceus (Sharp, 1885). Udayagiri & Wadhi, 1989: 37; Johnson, 1990: 330; Romero & Johnson, 2004: 618
Acanthoscelides dominicana Johnson, 1990: 371
Acanthoscelides flavescens (Fahraeus, 1839). Maes & Kingsolver, 1991: 218; Peck, 2005: 183
Acanthoscelides indigoferestes Johnson, 1983. Johnson, 1990: 399
Acanthoscelides johnique Johnson, 1986: 265
Acanthoscelides megacornis Kingsolver, 1980. Johnson, 1990: 427; Romero & Johnson, 2004: 620
Acanthoscelides obtectus (Say, 1831). Wolcott, 1927: 244; Johnson, 1983: 144; Johnson, 1990: 438 [Introduced]
=Bruchus tetricus Gyllenhal, 1839: 22
=Acanthoscelides tetricus (Gyllenhal, 1839). Blackwelder, 1946: 761
Acanthoscelides obvelatus Bridwell, 1942. Kingsolver, 1995: 170
Acanthoscelides schrankiae (Horn, 1873). Romero & Johnson, 2004: 621
Acanthoscelides suramerica Johnson, 1990. Kingsolver, 1992: 78
Amblycerus Thunberg, 1815
Amblycerus eustrophoides (Schaeffer, 1904). Kingsolver, 1992: 78
Amblycerus martorelli Bridwell, 1944. Blackwelder, 1946: 762; Kingsolver, 1970: 479; Udayagiri & Wadhi, 1989: 301
Amblycerus sallei (Jekel, 1855). Blackwelder, 1946: 763; Johnson & Kingsolver, 1981: 410; Kingsolver, 1992: 78
Amblycerus schwarzi Kingsolver, 1970. Kingsolver, 1992: 78
Bruchus Linnaeus, 1767
Bruchus pisorum (Linnaeus, 1758). Wolcott, 1927: 244 [Introduced]
Callosobruchus Pic, 1902
Callosobruchus chinensis (Linnaeus, 1758). Kingsolver, 1992: 78 [Introduced]
Caryedon Schönherr, 1826
Caryedon serratus (Olivier, 1790). Nilsson & Johnson, 1992: 62; Maes & Kingsolver, 1991: 32 [Introduced]
Caryobruchus Schönherr, 1823
Caryobruchus dominicanus† Poinar, 1999: 221 [Fossil in Dominican amber]
Caryobruchus gleditsiae (Johansson & Linnaeus, 1767). Nilsson & Johnson, 1993: 23
Ctenocolum Kingsolver & Whitehead, 1974
Ctenocolum podagricus (Fabricius, 1801). Manfio et al., 2019: 30
Megacerus Fahraeus, 1839
Megacerus deceptor Terán & Kingsolver, 1977. Terán & Johnson, 2002: 54
Megacerus incisithorax (Pic, 1932). Terán & Kingsolver, 1977: 95; Johnson & Kingsolver, 1981: 411; Kingsolver, 1992: 78
Megacerus tricolor (Suffrian, 1870). Terán & Kingsolver, 1977: 137; Kingsolver, 1992: 78
Merobruchus Bridwell, 1946
Merobruchus lysilomae Kingsolver, 1988: 25. Kingsolver, 1992: 78
Merobruchus paquetae Kingsolver, 1980. Kingsolver, 1992: 78
Mimosestes Bridwell, 1946
Mimosestes insularis Kingsolver & Johnson, 1978. Johnson & Kingsolver, 1981: 418; Kingsolver, 1992: 78
Mimosestes mimosae (Fabricius, 1781). Kingsolver & Johnson, 1978: 42; Udayagiri & Wadhi, 1989: 92; Kingsolver, 1992: 78;
Maes & Kingsolver, 1991: 28
=Acanthoscelides dominicanus (Jekel, 1855). Blackwelder, 1946: 759; Kingsolver & Johnson, 1978: 42

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 163
Pygiopachymerus Pic, 1911
Pygiopachymerus lineola (Chevrolat, 1871). Da Silva & Ribeiro-Costa, 2008: 803
Sennius Bridwell, 1946
Sennius rufomaculatus (Motschulsky, 1874). Maes & Kingsolver, 1991: 29; Kingsolver, 1992: 78
=Sennius instabilis (Sharp, 1885). Johnson & Kingsolver, 1973: 68
Stator Bridwell, 1946
Stator chalcodermus Kingsolver, 1972: 223. Johnson & Kingsolver, 1981: 420; Kingsolver, 1992: 78
Zabrotes Horn, 1885
Zabrotes subfasciatus (Boheman, 1833). Kingsolver, 1992: 78 [Introduced]
Notes: The records of Amblycerus hoffmanseggi (Gyllenhal, 1833) for Hispaniola cited by Blackwelder, 1946: 762; Kingsolver,
1970: 485; Johnson & Kingsolver, 1981: 410 have been deemed to be erroneous (Viana & Ribeiro, 2014: 67), therefore this
species is hereby deleted from the fauna of Hispaniola.

Superfamily CURCULIONOIDEA
Classification follows Alonso-Zarazaga & Lyal (1997).
Family ANTHRIBIDAE
Key reference: Valentine (2003).
Subfamily CHORAGINAE
Tribe ARAECERINI
Araecerus Schönherr, 1823
Araecerus fasciculatus (De Geer, 1775). Abud Antún, 1985: 4/85; Valentine, 2003: 52 [Introduced]
New genus A, nr. Cyptoxenus Valentine, 1982
New genus A, nr. Cyptoxenus n. sp. 1—Valentine (2003: 53)
Habroxenus Valentine, 1998
Habroxenus sarmenticola Valentine, 1999: 257. Valentine, 2003: 53 *H > TL: Morne Guimby, 22 km SE. Fond Verrettes,
6000’, Forêt des Pins
Tribe CHORAGINI
Choragus Kirby, 1819
Choragus exsertus† Poinar & Legalov, 2016: 26 [Fossil in Dominican amber]
Holostilpna Jordan, 1906
Holostilpna n. sp. 2—Valentine (2003: 54)
Holostilpna n. sp. 3—Valentine (2003: 54)
Holostilpna n. sp. 4—Valentine (2003: 54)
Tribe VALENFRIESIINI
Cyptoxenus Valentine, 1982
Cyptoxenus bruchelus† Poinar & Legalov, 2016: 30 [Fossil in Dominican amber]
Cyptoxenus ovatus† Poinar & Legalov, 2016: 32 [Fossil in Dominican amber]
Neoxenus Valentine, 1999
Neoxenus globosus† Poinar & Legalov, 2016: 28 [Fossil in Dominican amber]
Subfamily ANTHRIBINAE
Tribe DECATAPHANINI
Ptychoderes Schönherr, 1823
Ptychoderes n. sp. 1—Valentine (2003: 55)
Tribe PIESOCORYNINI
Piesocorynus Dejean, 1834
Piesocorynus n. sp. 1—Valentine (2003: 58)
Piesocorynus n. sp. 2—Valentine (2003: 58)
Piesocorynus bibullus† Poinar & Legalov, 2016: 10 [Fossil in Dominican amber]
Piesocorynus hamus† Poinar & Legalov, 2016: 14 [Fossil in Dominican amber]
Piesocorynus tribullus† Poinar & Legalov, 2016: 12 [Fossil in Dominican amber]
Piezobarra Jordan, 1906
Piezobarra brevisensoriata† Poinar & Legalov, 2016: 23 [Fossil in Dominican amber]
Brachycorynus Valentine, 1999
Brachycorynus n. sp. 1—Valentine (2003: 58)

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Brachycorynus n. sp. 2—Valentine (2003: 58)
Brachycorynus n. sp. 3—Valentine (2003: 58)
Brachycorynus n. sp. 4—Valentine (2003: 58)
Phaenotheriopsis Wolfrum, 1931
Phaenotheriopsis tuberculatus Valentine, 1991: 334. Valentine, 2003: 58 *DR > TL: 53 km SE Constanza, La Vega Prov.
Phaenotheriopsis umbronatus Valentine, 1991: 338. Valentine, 2003: 59 *DR > TL: 53 km SE Constanza, La Vega Prov.
Phaenotheriopsis verrucosus Valentine, 1991: 336. Valentine, 2003: 59 *DR > TL: 18 km SE Constanza, La Vega Prov.
Tribe EUGONINI
Eugonus Lacordaire, 1866
Eugonus adustus† Poinar & Legalov, 2016: 5 [Fossil in Dominican amber]
Eugonus angustus† Poinar & Legalov, 2016: 7 [Fossil in Dominican amber]
Tribe PLATYRHININI
New genus F, nr. Goniocloeus Jordan, 1904
New genus F, nr. Goniocloeus tuberculatus (Olivier, 1795). Valentine, 2003: 59
New genus F, nr. Goniocloeus sp. 1—Valentine (2003: 59)
New genus F, nr. Goniocloeus sp. 2—Valentine (2003: 59)
Tribe PLATYSTOMINI
Toxonotus Lacordaire, 1866
Toxonotus bidens (Olivier, 1795: 13). Valentine, 1955: 102; Valentine, 2003: 60 *H, *DR > TL: Saint Domingue
Toxonotus comatus† Poinar & Legalov, 2016: 4 [Fossil in Dominican amber]
Toxonotus n. sp. 3—Valentine (2003: 60)
Toxonotus n. sp. 4—Valentine (2003: 60)
Tribe RHINOTROPIDINI
Homocloeus Jordan, 1904
Homocloeus conspersus Wolfrum, 1930. Valentine, 2003: 56
Homocloeus n. sp. 6—Valentine (2003: 57)
Homocloeus n. sp. 7—Valentine (2003: 57)
Homocloeus n. sp. 8—Valentine (2003: 57)
Homocloeus n. sp. 9—Valentine (2003: 57)
Homocloeus n. sp. 10—Valentine (2003: 57)
Homocloeus n. sp. 11—Valentine (2003: 57)
Homocloeus n. sp. 12—Valentine (2003: 57)
Homocloeus n. sp. 13—Valentine (2003: 57)
Homocloeus n. sp. 14—Valentine (2003: 57)
Homocloeus n. sp. 15—Valentine (2003: 57)
Homocloeus n. sp. 16—Valentine (2003: 57)
Homocloeus n. sp. 17—Valentine (2003: 57)
Homocloeus n. sp. 18—Valentine (2003: 57)
Monocloeus Jordan, 1904
Monocloeus n. sp. 1—Valentine (2003: 58)
Tribe ZYGAENODINI
Eugonops Jordan, 1904
Eugonops n. sp. 1—Valentine (2003: 61)
Ormiscus Waterhouse, 1845
Ormiscus angulatus (Suffrian, 1870). Valentine, 2003: 61
Ormiscus n. sp. 9—Valentine (2003: 62)
Ormiscus n. sp. 10—Valentine (2003: 62)
Ormiscus n. sp. 11—Valentine (2003: 62)
Ormiscus n. sp. 12—Valentine (2003: 62)
Ormiscus n. sp. 13—Valentine (2003: 62)
Ormiscus n. sp. 14—Valentine (2003: 62)
Ormiscus n. sp. 15—Valentine (2003: 62)
Ormiscus n. sp. 16—Valentine (2003: 62)
Ormiscus n. sp. 17—Valentine (2003: 62)

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Ormiscus n. sp. 18—Valentine (2003: 62)
Ormiscus n. sp. 19—Valentine (2003: 62)
Ormiscus n. sp. 20—Valentine (2003: 62)
Ormiscus n. sp. 21—Valentine (2003: 62)
Ormiscus n. sp. 22—Valentine (2003: 62)
Ormiscus n. sp. 23—Valentine (2003: 62)
Ormiscus n. sp. 24—Valentine (2003: 62)
Ormiscus n. sp. 25—Valentine (2003: 62)
Ormiscus n. sp. 26—Valentine (2003: 62)
Ormiscus n. sp. 27—Valentine (2003: 62)
Ormiscus n. sp. 28—Valentine (2003: 62)
Ormiscus n. sp. 29—Valentine (2003: 62)
Ormiscus n. sp. 30—Valentine (2003: 62)
Ormiscus n. sp. 31—Valentine (2003: 63)
Ormiscus n. sp. 32—Valentine (2003: 63)
Ormiscus n. sp. 33—Valentine (2003: 63)
Ormiscus n. sp. 34—Valentine (2003: 63)
Ormiscus n. sp. 35—Valentine (2003: 63)
Ormiscus n. sp. 36—Valentine (2003: 63)
Ormiscus n. sp. 37—Valentine (2003: 63)
Ormiscus n. sp. 38—Valentine (2003: 63)
Ormiscus n. sp. 39—Valentine (2003: 63)
Ormiscus n. sp. 40—Valentine (2003: 63)
Ormiscus n. sp. 41—Valentine (2003: 63)
Ormiscus n. sp. 51—Valentine (2003: 63)

Family ATTELABIDAE
Subfamily ATTELABINAE
Attelabus Linnaeus, 1758
Attelabus canaliculatus Olivier, 1807: 7. Blackwelder, 1947: 832; O’Brien & Wibmer, 1982: 11 *H, *DR > TL: Saint-
Domingue
Euscelus Germar in Schönherr, 1833
Euscelus armatus (Gyllenhal, 1833). Voss, 1925: 43; Wolcott, 1927: 245; Blackwelder, 1947: 831; O’Brien & Wibmer, 1982:
11; Hamilton, 2007: 5
=Euscelus callosipennis Voss, 1925: 44. Blackwelder, 1947: 831; O’Brien & Wibmer, 1982: 12
Euscelus aureolus (Gyllenhal, 1833). Hamilton, 2007: 20
Euscelus cribrarius (Olivier, 1807: 8). Voss, 1925: 50; Blackwelder, 1947: 831; O’Brien & Wibmer, 1982: 12; Hamilton, 2007:
15
Euscelus dentipes (Fabricius, 1798). O’Brien & Wibmer, 1982: 12; Hamilton, 2007: 9
=Euscelus coccolobae Wolcott, 1924. Blackwelder, 1947: 831
Euscelus dominicanus Hamilton, 2007: 22 *DR, *H > TL: La Vega, 1.4–2.6 km E Manabao
Euscelus fornicatus (Olivier, 1807: 9). Voss, 1925: 68; Blackwelder, 1947: 831; O’Brien & Wibmer, 1982: 12
Euscelus haitensis Hamilton, 2007: 21 *H, *DR > TL: Camp Perrin
Euscelus postoculidens Hamilton, 2007: 8 *DR > TL: Elias Piña, Sierra de Neiba, 9.3 km WSW Hondo Valle, 190 m
Euscelus scrobiculatus Voss, 1925: 41. Blackwelder, 1947: 831; O’Brien & Wibmer, 1982: 12; Hamilton, 2007: 12 *DR > TL:
S. Domingo
Euscelus scutellatus (Klug, 1829). O’Brien & Wibmer, 1982: 12; Hamilton, 2007: 7
Euscelus spinifex (Olivier, 1807: 9). O’Brien & Wibmer, 1982: 12 *H, *DR > TL: Saint-Domingue
Euscelus uviferae Marshall, 1926b: 53. Blackwelder, 1947: 832; O’Brien & Wibmer, 1982: 12; Hamilton, 2007: 13 *H, *DR >
TL: St. Louis on Coccoloba uviferae
Hamiltoniauletes Legalov, 2001
Hamiltoniauletes trifasciatus (Suffrian, 1870). Legalov, 2010: 18
Pseudopilolabus Legalov, 2003
Pseudopilolabus othnius† Poinar et al., 2016: 12 [Fossil in Dominican amber]

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Family APIONIDAE
Subfamily APIONINAE
Tribe APIONINI
Apionion Kissinger, 1998
Apionion homochronon† Poinar & Legalov, 2015b: 150 [Fossil in Dominican amber]
Apionion kallimosum† Poinar & Legalov, 2015b: 151 [Fossil in Dominican amber]
Apionion formoculus† Poinar & Legalov, 2015b: 153 [Fossil in Dominican amber]
Neapion Alonzo-Zarazaga, 1990
Neapion vestitum (Gyllenhal, 1833: 267). Blackwelder, 1947: 831; O’Brien & Wibmer, 1982: 28 *H, *DR > TL: Insula St.
Domingo
Stenapion Wagner, 1912
Stenapion levigatum† Poinar & Legalov, 2015b: 154 [Fossil in Dominican amber]
Toxorhynchus Scudder, 1894
Toxorhynchus acolum (Kissinger, 1974: 22). O’Brien & Wibmer, 1982: 20; Poinar & Legalov, 2015: 2 *H > TL: 10 kms north
Ennery
Toxorhynchus amphioculus† Poinar & Legalov, 2015b: 137 [Fossil in Dominican amber]
Toxorhynchus atriantenus† Poinar & Legalov, 2015b: 139 [Fossil in Dominican amber]
Toxorhynchus atriartus† Poinar & Legalov, 2015b: 138 [Fossil in Dominican amber]
Toxorhynchus convexoculus† Poinar & Legalov, 2015b: 148 [Fossil in Dominican amber]
Toxorhynchus convexus† Poinar & Legalov, 2015b: 145 [Fossil in Dominican amber]
Toxorhynchus dominicanus† Poinar & Legalov, 2015b: 144 [Fossil in Dominican amber]
Toxorhynchus fuscocorpus† Poinar & Legalov, 2015b: 146 [Fossil in Dominican amber]
Toxorhynchus hispaniolicus† Poinar & Legalov, 2015b: 140 [Fossil in Dominican amber]
Toxorhynchus hispidulus† Poinar & Legalov, 2015b: 147 [Fossil in Dominican amber]
Toxorhynchus leptorhinus† Poinar & Legalov, 2015b: 135 [Fossil in Dominican amber]
Toxorhynchus microsomus† Poinar & Legalov, 2015b: 142 [Fossil in Dominican amber]
Toxorhynchus picipes (Gerstaecker, 1854). O’Brien & Wibmer, 1982: 26
=Apion stabile Sharp, 1890. Kissinger, 1974: 27
Toxorhynchus robustus† Poinar & Legalov, 2015b: 136 [Fossil in Dominican amber]
Toxorhynchus sandersoni (Kissinger, 1974: 26). O’Brien & Wibmer, 1982: 27 *DR, *H > TL: Valle Nuevo, Prov. La Vega,
1,500 m
Toxorhynchus stenelytrus† Poinar & Legalov, 2015b: 143 [Fossil in Dominican amber]
Toxorhynchus valentinei (Kissinger, 1974: 23). O’Brien & Wibmer, 1982: 28 *DR, *H > TL: Pico Isabel de Torres, near Puerto
Plata, Puerto Plata Prov.
Trichapion Wagner, 1912
Trichapion arcanum (Kissinger, 1974: 15). O’Brien & Wibmer, 1982: 20 *H > TL: 10 kilometers southeast Gonaives
Trichapion enigmaticum (Kissinger, 1974: 9). O’Brien & Wibmer, 1982: 22 *H > TL: Port-au-Prince, up to 2000 ft.
Trichapion nesium (Kissinger, 1974: 11). O’Brien & Wibmer, 1982: 25 *DR, *H > TL: Pico Isabel de Torres, near Puerto Plata,
Puerto Plata Prov.
Trichapion salarium (Kissinger, 1974). O’Brien & Wibmer, 1982: 27
Trichapion silendum (Kissinger, 1974: 12). O’Brien & Wibmer, 1982: 27 *H > TL: Mountains near Port-au-Prince, up to 2,000
ft.
INCERTAE SEDIS
Apion portorium Kissinger, 1974: 29. O’Brien & Wibmer, 1982: 26

Family BRENTIDAE
Key reference: Sforzi and Bartolozzi (2004).
Subfamily BRENTINAE
Tribe BRENTINI
Brentus Fabricius, 1887
Brentus forcipitigerus (Gyllenhal, 1839). Damoiseau, 1989: 62; Sforzi & Bartolozzi, 2004: 152
Brentus linearis Olivier, 1790. Sforzi & Bartolozzi, 2004: 154; Sforzi & Bartolozzi, 2004: 154
=Brentus volvulus (Fabricius, 1792). Wolcott, 1927: 245; Blackwelder, 1947: 777; Perez-Gelabert, 2008: 131

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Brentus turbatus Boheman, 1840. Sforzi & Bartolozzi, 2004: 157
INCERTAE SEDIS
Brentus monilis (Olivier, 1807). Blackwelder, 1947: 774; Perez-Gelabert, 2008: 131; Sforzi & Bartolozzi, 2004: 717
Tribe ARRHENODINI
Belopherus Schönherr, 1833
Belopherus maculatus (Olivier, 1807). Blackwelder, 1947: 774; Sforzi & Bartolozzi, 2004: 180
Belopherus militaris (Olivier, 1807). Sforzi & Bartolozzi, 2004: 180
=Belopherus spinosus Gyllenhal in Schönherr, 1833. Sforzi & Bartolozzi, 2004: 131; Perez-Gelabert, 2008: 131
Belopherus rostralatus Damoiseau, 1989: 53. Sforzi & Bartolozzi, 2004: 181 *DR > TL: Boca Chica, 10 m
Belopherus schoenherrii Mannerheinn, 1840. Blackwelder, 1947: 774; Sforzi & Bartolozzi, 2004: 181
Estenorhinus Gemminger & Harold, 1872
Estenorhinus turbatus (Gyllenhal, 1833). Blackwelder, 1947: 773; Sforzi & Bartolozzi, 2004: 211
Raphirhynchus Dejean, 1834
Raphirhynchus cylindricornis (Fabricius, 1787). Sforzi & Bartolozzi, 2004: 262; Perez-Gelabert, 2012: 115
Stratiorrhynchus Damoiseau, 1962
Stratiorrhynchus strigicollis (Lacordaire, 1866). Blackwelder, 1947: 774; Sforzi & Bartolozzi, 2004: 278
Subfamily CYLADINAE
Cylas Latreille, 1802
Cylas formicarius (Fabricius, 1798). Wolcott, 1927: 246; Santoro, 1960: 72; O’Brien & Wibmer, 1982: 19; Schmutterer, 1990:
501 [Introduced]
Subfamily CYPHAGOGINAE
Tribe DOMINIBRENTINI†
Dominibrentus† Poinar, 2009
Dominibrentus leptus† Poinar, 2009: 52 [Fossil in Dominican amber]
Tribe STEREODERMINI
Stereodermus Lacordaire, 1866
Stereodermus exilis Suffrian, 1870. Mantilleri & Sforzi, 2006: 27
Subfamily TRACHELIZINAE
Tribe TRACHELIZINI
Hephebocerus Schönherr, 1840
Hephebocerus mexicanus Sharp, 1895. Sforzi & Bartolozzi, 2004: 494; Perez-Gelabert, 2012: 115
Nemocephalus Güerin Méneville, 1827
Nemocephalus ayitiensis Mantilleri, 2014: 121 *RD > TL: S Francesco Mts, St. Domingo
Nemocephalus uncimanus (Boheman, 1840). Mantilleri, 2014: 122
=Nemocephalus moderatus (Kleine, 1922). Blackwelder, 1947: 775; Sforzi & Bartolozzi, 2004: 580
Note: The species Nemocephalus avarus (Kleine, 1927) cited by Blackwelder, 1947: 775 and Sforzi & Bartolozzi, 2004: 579
and Nemocephalus monilis (Fabricius, 1787) cited for Hispaniola by Sforzi & Bartolozzi, 2004: 580 have been restricted to
other islands and therefore taken out of the Hispaniolan faunal list.
Tribe ITHYSTENINI
Belorhynchus Berthold, 1827
Belorhynchus nasutus (Fabricius, 1781). Blackwelder, 1947: 774; Sforzi & Bartolozzi, 2004: 593; Perez-Gelabert, 2008: 131
=Belopherus nasutus (Fabricius, 1781)
Tribe TYCHAEINI
Nemorhinus Schönherr, 1840
Nemorhinus myrmecophaga (Herbst, 1797). Sforzi & Bartolozzi, 2004: 705; Perez-Gelabert, 2012: 115

Family BELIDAE
Subfamily OXYCORYNINAE
Pleurambus† Poinar & Legalov, 2014
Pleurambus strongylus† Poinar & Legalov, 2014c: 671 [Fossil in Dominican amber]

Family CURCULIONIDAE
Subfamily ANTHONOMINAE
Tribe ANTHONOMINI

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Anthonomus Germar, 1817
Anthonomus (Anthomorphus) browni† Poinar & Legalov, 2015c: 7 [Fossil in Dominican amber]
Anthonomus (Anthomorphus) cruraluma† Poinar & Legalov, 2015c: 3 [Fossil in Dominican amber]
Anthonomus (Anthonomocyllus) argentatus Gyllenhal, 1836. Clark, 1990b: 568
Anthonomus (Anthonomocyllus) guanita Clark, 1990b: 574
Anthonomus (Anthonomus) agerochus Clark, 1991c: 45
Anthonomus (Anthonomus) alboannulatus Boheman, 1843. Clark, 1991b: 210
Anthonomus (Anthonomus) albocapitis Clark, 1991a: 267
Anthonomus (Anthonomus) brenthus Clark, 1991c: 49 *DR > TL: Altagracia, Guaraguao
Anthonomus (Anthonomus) convexifrons Hustache, 1929. Clark, 1991c: 50
Anthonomus (Anthonomus) curvirostris Champion, 1903. Clark, 1991c: 44
Anthonomus (Anthonomus) eugenii Cano in Cano & Alcacio, 1894. Serra et al., 2011: 147 [Introduced]
Anthonomus (Anthonomus) grandis Boheman, 1843. Santoro, 1960: 24; O’Brien & Wibmer, 1982: 107; Schmutterer, 1990:
308 [Introduced]
Anthonomus (Anthonomus) macromalus Gyllenhal, 1836. Clark, 1992: 286
=Anthonomus malpighiae Clark & Burke, 1985: 121
Anthonomus (Anthonomus) nanus Gyllenhal, 1836: 351. Blackwelder, 1947: 839; O’Brien & Wibmer, 1982: 108; Clark, 1992:
287 *H, *DR > TL: Insula St. Domingo
=Anthonomus incanus Champion, 1903. Clark, 1988: 335
Anthonomus (Anthonomus) oraapis Clark, 1988: 338 *DR > TL: Barahona, 11 km SW Paraiso
Anthonomus pulicarius Boheman, 1843. Wolcott, 1927: 252; Blackwelder, 1947: 839; Schmutterer, 1990: 537
Anthonomus (Anthonomus) quisqueyensis Clark & Burke, 1985: 129 *DR > TL: Peravia, 6 km W. Baní
Anthonomus (Anthonomus) sisyphus Clark, 1987: 75
Anthonomus sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Anthonomus (Anthonomus) trisicifer Clark, 1991b: 222
Anthonomus (Anthonomus) tricuspis Clark, 1991b: 224 *DR > TL: Guaraguao, La Altagracia Prov.
Anthonomus (Anthonomus) varipes Jacquelin du Val, 1857. Clark, 1996: 63
Anthonomus (Anthonomus) ylem Clark, 1991d: 159
Anthonomus (Anthonomus) zagluli Clark & Burke, 1985: 124 *DR > TL: Peravia, 6 km W. Baní
Atractomerus Duponchel & Chevrolat, 1849
Atractomerus knabi Clark, 1989: 372
Atractomerus punctipennis (Gyllenhal, 1836). Clark, 1989: 320
Huaca Clark, 1993
Huaca apian Clark, 1993a: 17
Huaca payan Clark, 1993a: 12 *DR > TL: Santiago: Loma Rucilla & mts. N, 5–8,000 ft.
Huaca puquiu Clark, 1993a: 24 *DR > TL: Peravia, 6 km W. Baní
Neomastix Dietz, 1891
Neomastix numerus Clark, 1993b: 14
Pseudoanthonomus Dietz, 1891
Pseudoanthonomus photon Clark, 1990a: 678
Pseudoanthonomus testaceus (Boheman, 1843). Clark, 1990a: 677
Subfamily BARIDINAE
Tribe BARIDINI
Baris Germar, 1817
Baris grossacavis† Poinar & Legalov, 2015a: 32 [Fossil in Dominican amber]
Baris penicillata (Boheman, 1836). O’Brien & Wibmer, 1982: 184
Eurhinus Illiger, 1807
Eurhinus festivus (Fabricius, 1792). Wolcott, 1927: 252; Blackwelder, 1947: 891; O’Brien & Wibmer, 1982: 181
Rhaptinus Casey, 1920
Rhaptinus tabaci (Sallé, 1855: 269). Blackwelder, 1947: 895; O’Brien & Wibmer, 1982: 185
Rhaptinus torquatus (Olivier, 1807). Leng & Mutchler, 1914: 436; Marcano, 1964: 144
Tribe MADOPTERINI
Nicentrus Casey, 1892
Nicentrus saccharinus Marshall, 1951. Woodruff, 1972: 1

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Tribe PERIDINETINI
Peridinetus Schönherr, 1837
Peridinetus concentricus (Olivier, 1807). O’Brien & Wibmer, 1982: 128; Chevrolat, 1882: 81; Blackwelder, 1947: 886; Prena,
2009: 55
=Peridinetus signatus Rosenschöld, 1837. Marcano & Abud, 1995: 10/95; SEA, 1999: 88; Prena, 2009: 56
Peridinetus decipiens Prena, 2009: 57 *DR > TL: La Vega Prov., 10 km W Jima
Peridinetus fuscosignum Prena, 2009: 58 *DR, *H > TL: Pedernales, 25 km N Cabo Rojo, 700 m
Subfamily CONODERINAE
Tribe ZYGOPINI
Eulechriops Faust, 1896
Eulechriops argyrosoma† Poinar & Legalov, 2014b: 556 [Fossil in Dominican amber]
Geratozygops† Davis & Engel, 2006
Geratozygops arsinotus† Poinar & Legalov, 2014b: 558 [Fossil in Dominican amber]
Geratozygops atropos† Davis & Engel, 2006: 255 [Fossil in Dominican amber]
Geratozygops platysoma† Poinar & Legalov, 2014b: 558 [Fossil in Dominican amber]
Geratozygops stenosoma† Poinar & Legalov, 2014b: 559 [Fossil in Dominican amber]
Subfamily Cossoninae
Acamptus LeConte, 1876
Acamptus exilipes† Poinar & Legalov, 2015e: 492 [Fossil in Dominican amber]
Ogygius Poinar & Legalov, 2015
Ogygius obrieni† Poinar & Legalov, 2015e: 498 [Fossil in Dominican amber]
Tribe COSSONINI
Catolethrus Dejean, 1836
Catolethrus sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Cossonus Clairville, 1798
Cossonus hinojosai† Davis & Engel, 2007: 812 [Fossil in Dominican amber]
Cossonus pyrirostris Boheman, 1838: 1014. O’Brien & Wibmer, 1982: 224 *H, *DR > TL: St.-Domingo
Cossonus spathula Boheman, 1838. O’Brien & Wibmer, 1982: 224
Himatium Wollaston, 1873
Himatium sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Micromimus Wollaston, 1873
Micromimus orcus† Davis & Engel, 2007: 804 [Fossil in Dominican amber]
Paralicus O’Brien, 1984
Paralicus abnormis† Davis & Engel, 2007: 809 [Fossil in Dominican amber]
Proeces Schönherr, 1838
Proeces longirostrum† Davis & Engel, 2007: 809 [Fossil in Dominican amber]
Rhinanisus Broun, 1883
Rhinanisus sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Stenotrupis Wollaston, 1873
Stenotrupis acicula Wollaston, 1873. O’Brien & Wibmer, 1982: 226
Stenotrupis breviscapus† Davis & Engel, 2007: 811 [Fossil in Dominican amber]
Stenotrupis pumilis† Poinar & Legalov, 2015e: 500 [Fossil in Dominican amber]
Tribe DRYOTRIBINI
Caulophilus Wollaston, 1854
Caulophilus ashei† Davis & Engel, 2006: 102 [Fossil in Dominican amber]
Caulophilus bennetti† Davis & Engel, 2007: 807 [Fossil in Dominican amber]
Caulophilus camptus† Poinar & Legalov, 2015: 494 [Fossil in Dominican amber]
Caulophilus elongatus† Poinar & Legalov, 2015: 496 [Fossil in Dominican amber]
Caulophilus falini† Davis & Engel, 2007: 805 [Fossil in Dominican amber]
Caulophilus orizae (Gyllenhall, 1838) [By synonymy of species below. Introduced]
=Caulophilus latinasus (Say, 1831). Abud, 1993: 5/93
Caulophilus ruidipunctus† Poinar & Legalov, 2015: 495 [Fossil in Dominican amber]
Caulophilus swensoni† Davis & Engel, 2007: 806 [Fossil in Dominican amber]
Dryotribus Horn, 1873

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Dryotribus amplioculus† Davis & Engel, 2007: 808 [Fossil in Dominican amber]
Dryotribus mimeticus Horn, 1873. O’Brien & Wibmer, 1982: 227
Dryotribus pedanus† Poinar & Legalov, 2015: 492 [Fossil in Dominican amber]
Subfamily CRYPTORHYNCHINAE
Anlemmus† Poinar & Legalov, 2014
Anlemmus leptorhinus† Poinar & Legalov, 2014a: 532 [Fossil in Dominican amber]
Apharosoma† Poinar & Legalov, 2014
Apharosoma euryrhina† Poinar & Legalov, 2014a: 530 [Fossil in Dominican amber]
Cophes Champion, 1905
Cophes armipes (Boheman, 1837). O’Brien & Wibmer, 1982: 153; Peck, 2010: 62
Cryptorhynchus Illiger, 1807
Cryptorhynchus otiosus Boheman, 1837: 81. Blackwelder, 1947: 872 *DR > TL: St. Domingo
Episcirrus† Poinar & Legalov, 2014
Episcirrus isolepus† Poinar & Legalov, 2014a: 502 [Fossil in Dominican amber]
Euscepes Schönherr, 1833
Euscepes frontalis Chevrolat, 1879b: cl. O’Brien & Wibmer, 1982: 142 *DR > TL: Sancto Domingo
Euscepes porcellus Boheman, 1844. O’Brien & Wibmer, 1982: 142
Euscepes postfaciatus (Fairmaire, 1849) [By synonymy of species below. Introduced]
=Euscepes batatae (Waterhouse, 1849). SEA, 1999: 32
Gerstaeckeria Champion, 1905
Gerstaeckeria (Philopuntia) insulana Fisher, 1928: 6. O’Brien & Wibmer, 1982: 136 *H > TL: Port-au-Prince
Lemmasomus† Poinar & Legalov, 2013
Lemmasomus anodontotus† Poinar & Legalov, 2014a: 529 [Fossil in Dominican amber]
Lembodes Schönherr, 1844
Lembodes solitarius Boheman, 1844. O’Brien & Wibmer, 1982: 136
Lembodes ulula Chevrolat, 1879b: cxlix. O’Brien & Wibmer, 1982: 136 *DR, *H > TL: Sancto Domingo, Haïti
Neoulosomus O’Brien & Wibmer, 1982
Neoulosomus laticaudis (Suffrian, 1872). O’Brien & Wibmer, 1982: 143
Neoulosomus (Neoulosomus) conthorhinus† Poinar & Legalov, 2014a: 518 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) leptosomus† Poinar & Legalov, 2014a: 520 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) megaholcus† Poinar & Legalov, 2014a: 528 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) megus† Poinar & Legalov, 2014a: 525 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) microholcus† Poinar & Legalov, 2014a: 528 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) pedinus† Poinar & Legalov, 2014a: 526 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) pediosomus† Poinar & Legalov, 2014a: 527 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) platystegus† Poinar & Legalov, 2014a: 526 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) scambosomus† Poinar & Legalov, 2014a: 527 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) scambus† Poinar & Legalov, 2014a: 524 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) stenocalypus† Poinar & Legalov, 2014a: 527 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) stylolepus† Poinar & Legalov, 2014a: 525 [Fossil in Dominican amber]
Neoulosomus (Neoulosomus) tanyrhinus† Poinar & Legalov, 2014a: 516 [Fossil in Dominican amber]
Odontamera† Poinar & Legalov, 2014
Odontamera dolichosoma† Poinar & Legalov, 2014a: 533 [Fossil in Dominican amber]
Palaeopus Faust, 1896
Palaeopus costicollis Marshall, 1918. Anonymous, 1925: 654
Paracamptopsis† Poinar & Legalov, 2014
Paracamptopsis stenis† Poinar & Legalov, 2014a: 506 [Fossil in Dominican amber]
Paraulosomus Hustache, 1930
Paraulosomus adenolepus† Poinar & Legalov, 2014a: 506 [Fossil in Dominican amber]
Pseudomoides† Poinar & Legalov, 2014
Pseudomoides clisaulis† Poinar & Legalov, 2014a: 504 [Fossil in Dominican amber]
Pseudomopsis Champion, 1905
Pseudomopsis inflata (LeConte, 1884). O’Brien & Wibmer, 1982: 148
Pseudomus Schönherr, 1844

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Pseudomus nitidicutis Chevrolat, 1880c: 279. O’Brien & Wibmer, 1982: 137 *DR, *H > TL: Haiti
Pseudomus proximus Chevrolat, 1880c: 279. O’Brien & Wibmer, 1982: 137 *H > TL: Haiti
Pseudomus punctatissimus Chevrolat, 1880c: 279. O’Brien & Wibmer, 1982: 137
Pseudomus semicribratus Boheman, 1844. O’Brien & Wibmer, 1982: 137
Semnorhynchus Faust, 1896
Semnorhynchus brachyrhinus† Poinar & Legalov, 2014a: 512 [Fossil in Dominican amber]
Semnorhynchus campostegus† Poinar & Legalov, 2014a: 513 [Fossil in Dominican amber]
Semnorhynchus contorhinus† Poinar & Legalov, 2014a: 510 [Fossil in Dominican amber]
Semnorhynchus euryaspus† Poinar & Legalov, 2014a: 514 [Fossil in Dominican amber]
Semnorhynchus eurystegus† Poinar & Legalov, 2014a: 507 [Fossil in Dominican amber]
Semnorhynchus leptostegus† Poinar & Legalov, 2014a: 511 [Fossil in Dominican amber]
Semnorhynchus megasomus† Poinar & Legalov, 2014a: 509 [Fossil in Dominican amber]
Semnorhynchus stenostegus† Poinar & Legalov, 2014a: 508 [Fossil in Dominican amber]
Semnorhynchus tanyrhinus† Poinar & Legalov, 2014a: 511 [Fossil in Dominican amber]
Tyloderma Say, 1831
Tyloderma schoenherri Wibmer, 1989: 74 *DR > TL: Prov. La Altagracia, 31 kms north of Higuey
Tyloderma sp.†—Poinar & Poinar (1999: 204) [Fossil in Dominican amber]
Subfamily CURCULIONINAE
Tribe OTIDOCEPHALINI
Micromyrmex Sleeper, 1953
Micromyrmex asclepia Ivie & Sikes, 1995: 241 *DR > TL: ~5 km S Constanza, La Vega Prov.
Micrormex formicarius (Olivier, 1807: 236). Blackwelder, 1947: 835; O’Brien & Wibmer, 1982: 98; Ivie & Sikes, 1995: 236
*H, *DR > TL: Saint-Domingue
Micromyrmex spatulata Sikes & Ivie, 1998: 156 *DR > TL: Peravia, 36 km NW San José de Ocoa
Tribe PYROPINI
Pyropus Schönherr, 1838
Pyropus cyaneus (Herbst, 1795). O’Brien & Wibmer, 1982: 125
Pyropus sapphirinus Gyllenhal, 1836. O’Brien & Wibmer, 1982: 125
Tribe TYCHIINI
Derelomus Schoenherr, 1825
Derelomus thalioculus† Poinar & Legalov, 2015c: 2 [Fossil in Dominican amber]
Neosibinia Poinar & Legalov, 2015
Neosibinia lepidosoma† Poinar & Legalov, 2015c: 10 [Fossil in Dominican amber]
Plocetes LeConte, 1876
Plocetes simulacrifer Clark, 1982: 136 *DR > TL: Cloudforest vic. Valle Nuevo, 6,000 ft.
Sibinia Germar, 1817
Sibinia (Microtychius) hispaniolae Clark, 1978: 274. O’Brien & Wibmer, 1982: 119 *H > TL: Port-au-Prince
Subfamily CYCLOMINAE
Listronotus Jekel, 1865
Listronotus cryptops (Dietz, 1889). O’Brien, 1981: 94
Subfamily DRYOPHTHORINAE
Tribe DIOCALANDRINI
Bicalcasura† Poinar & Legalov, 2014
Bicalcasura maculata† Poinar & Legalov, 2014: 450 [Fossil in Dominican amber]
Tribe DRYOPHTHORINI
Dryophthorus Germar, 1824
Dryophthorus acarophilus† Davis & Engel, 2006: 195 [Fossil in Dominican amber]
Dryophthorus microtremus† Poinar & Legalov, 2015d: 508 [Fossil in Dominican amber]
Stenommatus Wollaston, 1873
Stenommatus leptorhinus† Poinar & Legalov, 2015d: 511 [Fossil in Dominican amber]
Stenommatus tanyrhinus† Poinar & Legalov, 2015d: 510 [Fossil in Dominican amber]
Stenommatus pulvereus† Davis & Engel, 2006: 193 [Fossil in Dominican amber]
Tribe SIPALINI
Rhinostomus Rafinesque, 1815

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Rhinostomus oblitus (Jacquelin du Val, 1857). Vaurie, 1970: 28; O’Brien & Wibmer, 1982: 220; Morrone & Cuevas, 2002: 382
Rhinostomus scrutator (Olivier, 1807). Blackwelder, 1947: 917; Vaurie, 1970: 32; Morrone & Cuevas, 2002: 382
Subfamily ENTIMINAE
Tribe EUDIAGOGINI
Promecops Sahlberg, 1823
Promecops (Promecodes) divarichela† Poinar & Legalov, 2017: 10 [Fossil in Dominican amber]
Promecops tumidirostris† Poinar & Brown, 2010: 231 [Fossil in Dominican amber]
Tribe EUSTYLINI
Brachyomus Lacordaire, 1863
Brachyomus octotuberculatus (Fabricius, 1787). Perez-Gelabert, 2008: 134
Diaprepes Schönherr, 1823
Diaprepes abbreviatus (Linnaeus, 1758). O’Brien & Wibmer, 1982: 55; Morrone, 1999: 126
Diaprepes anticus† Franz & Zhang, 2017: 17 [Fossil in Dominican amber]
Diaprepes doublierii Guérin-Méneville, 1847: 9. O’Brien & Wibmer, 1982: 55 *H > TL: L’île d’Haïti
Diaprepes squamula† Poinar & Legalov, 2017: 2 [Fossil in Dominican amber]
Exophthalmus Schönherr, 1823
Exophthalmus albosquamosus (Sallé, 1855: 268). Gemminger & Harold, 1871: 2233; O’Brien & Wibmer, 1982: 56
Exophthalmus chrysopus Chevrolat, 1880b: 165. O’Brien & Wibmer, 1982: 56 *DR, *H > TL: S. Domingo, Haiti
Exophthalmus cinerascens (Fabricius, 1792). Gemminger & Harold, 1871: 2233; O’Brien & Wibmer, 1982: 56 *DR, *H > TL:
Original says “Cajennae”, but endemic to Hispaniola
Exophthalmus costatus (Gyllenhal, 1834: 18). Gemminger & Harold, 1871: 2233; O’Brien & Wibmer, 1982: 56 *H, *DR >
TL: Insula St. Domingo
Exophthalmus depressicollis (Gyllenhal, 1834: 22). O’Brien & Wibmer, 1982: 56 *H, *DR > TL: Insula St. Domingo
Exophthalmus frenatus (Marshall, 1934: 624). O’Brien & Wibmer, 1982: 57 *H, *DR > TL: Port-au-Prince
Exophthalmus haitiensis Bovie, 1907: 68. O’Brien & Wibmer, 1982: 57 *H > TL: Haïti
Exophthalmus hieroglyphicus Chevrolat, 1879a: xcvii. O’Brien & Wibmer, 1982: 57 *Hispaniola > TL: Sancto-Domingo
Exophthalmus inaequalis (Gyllenhal, 1834: 23). Gemminger & Harold, 1871: 2233; O’Brien & Wibmer, 1982: 57 *H, *DR >
TL: Insula St. Domingo
Exophthalmus laetus (Olivier, 1807: 326). O’Brien & Wibmer, 1982: 57 *H, *DR > TL: Saint-Domingue
Exophthalmus leucographus (Gyllenhal, 1834: 19). Gemminger & Harold, 1871: 2233; O’Brien & Wibmer, 1982: 57 *H, *DR
> TL: Insula St. Domingo
Exophthalmus mannerheimi Boheman, 1840: 338. Gemminger & Harold, 1871: 2233; O’Brien & Wibmer, 1982: 57 *DR, *H >
TL: S. Domingo
Exophthalmus obsoletus (Olivier, 1807: 325). Gemminger & Harold, 1871: 2234; O’Brien & Wibmer, 1982: 57 *H, *DR > TL:
Saint-Domingue
Exophthalmus pugnax (Olivier, 1807: 349). O’Brien & Wibmer, 1982: 57 *H > TL: l’Amerique Meridionale
Exophthalmus quadrivittatus (Olivier, 1807: 325). Boheman in Schönherr, 1840: 337; Wolcott, 1927: 248; O’Brien & Wibmer,
1982: 57; Schmutterer, 1990: 93 *H, *DR > TL: Saint-Domingue
Exophthalmus roseipes (Chevrolat, 1876). SEA, 1999: 112
Exophthalmus sphacelatus (Olivier, 1807). Gemminger & Harold, 1871: 2234; Wolcott, 1927: 248; O’Brien & Wibmer, 1982:
58
Exophthalmus squamipennis Gemminger, 1871: 119. Gemminger & Harold, 1871: 2234. O’Brien & Wibmer, 1982: 58 *H,
*DR > TL: St. Domingo
Hormotrophus Schönherr, 1843
Hormotrophus aureomixtus Boheman, 1843: 44. O’Brien & Wibmer, 1982: 55 *H, *DR > TL: St.-Domingo
Phaops Sahlberg, 1823
Phaops albugo Sahlberg, 1823. O’Brien & Wibmer, 1982: 56
Tribe GEONEMINI
Apotomoderes Schönherr, 1840
Apotomoderes anodontos Franz, 2010: 45 *DR > TL: Pedernales, Along Rd. Cabo Rojo to Aceitillar, 12 km uphill, 395 m
Apotomoderes chariedris Franz, 2010: 58 *DR > TL: ~2 km N Bayahibe, La Altagracia Prov.
Apotomoderes hadroprion Franz, 2010: 62 *DR > TL: El Verón, rd. to Hoyo Azul, 40 m
Apotomoderes lateralis (Gyllenhal, 1834: 45). O’Brien & Wibmer, 1982: 41; Franz, 2010: 56 *DR > TL: Santo Domingo
Apotomoderes menocrater Franz, 2010: 49 *DR > TL: Las Cuevas, Bahía de Las Aguilas station, 40 m

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Artipus Sahlberg, 1823
Artipus calceatus Marshall, 1926a: 530. Blackwelder, 1947: 792; O’Brien & Wibmer, 1982: 31 *H > TL: Port-au-Prince
Artipus psittacinus Gyllenhal, 1834: 95. Wolcott, 1927: 251; Blackwelder, 1947: 792; O’Brien & Wibmer, 1982: 31 *H, *DR >
TL: Insula St. Domingo
Eugeonemus Buchanan, 1947
Eugeonemus sp.—Buchanan (1947: 44); O’Brien & Wibmer (1982: 39)
Ischionoplus Chevrolat, 1878
Ischionoplus viridiguttatus Chevrolat, 1878: 11. O’Brien & Wibmer, 1982: 41 *DR > TL: Sancto Domingo
Lachnopus Schönherr, 1840
Lachnopus aereus (Gyllenhal, 1834: 40). Morrone, 1999: 140; Girón et al., 2018: 44 *H, *DR > TL: Insula St. Domingo
Lachnopus albomaculatus (Gyllenhal, 1834: 37). Morrone, 1999: 141; Girón et al., 2018: 44 *H, *DR > TL: Insula St.
Domingo
Lachnopus atramentarius (Gyllenhal, 1834: 33). Morrone, 1999: 141; Girón et al., 2018: 45 *H, *DR > TL: Insula St.
Domingo
Lachnopus aulicus (Gyllenhal, 1834: 35). Morrone, 1999: 141; Girón et al., 2018: 45 *H, *DR > TL: Insula St. Domingo
Lachnopus bellus Marshall, 1926b: 54. Morrone, 1999: 141; Girón et al., 2018: 46 *H, *DR > TL: St. Louis on Coccoloba
uviferae
Lachnopus canescens (Gyllenhal in Schönherr, 1840: 388). Morrone, 1999: 141; Girón et al., 2018: 48 *H, *DR > TL: Insula
St. Domingo
Lachnopus chirographus (Olivier, 1807: 334). Morrone, 1999: 141; Girón et al., 2018: 48 *H, *DR > TL: Saint-Domingue
Lachnopus chlorophanus (Gyllenhal, 1834: 39). Morrone, 1999: 141; Girón et al., 2018: 48 *H, *DR > TL: Insula St. Domingo
Lachnopus consentaneus Perroud, 1853: 487. Morrone, 1999: 141; Girón et al., 2018: 49 *H, *DR > TL: St-Domingue
Lachnopus curvipes (Fabricius, 1787). Morrone, 1999: 141; Girón et al., 2018: 52
Lachnopus dentipes Perroud, 1853: 490. Morrone, 1999: 141; Girón et al., 2018: 58 *H, *DR > TL: St-Domingue
Lachnopus granicollis (Gyllenhal Schönherr, 1840: 390). Morrone, 1999: 141; Girón et al., 2018: 60 *H, *DR > TL: Insula St.
Domingo
Lachnopus hirtus Perroud, 1853: 484. Morrone, 1999: 141; Girón et al., 2018: 61 *H, *DR > TL: St-Domingue
Lachnopus histrio Marshall, 1926a: 534. Morrone, 1999: 141; Girón et al., 2018: 62 *H > TL: Port-au-Prince
Lachnopus inconditus Rosenschold, 1840: 383. Morrone, 1999: 141; Girón et al., 2018: 62
Lachnopus interruptus Perroud, 1853: 475. Morrone, 1999: 141; Girón et al., 2018: 63 *H, *DR > TL: St-Domingue
Lachnopus luxurians (Olivier, 1807: 334). Morrone, 1999: 141; Girón et al., 2018: 66 *H, *DR > TL: Saint-Domingue
Lachnopus mercator (Olivier, 1807: 335). Wolcott, 1927: 251; Morrone, 1999: 141; Girón et al., 2018: 66 *H, *DR > TL: St.
Domingo
Lachnopus mundus (Gyllenhal, 1834: 31). Morrone, 1999: 141; Girón et al., 2018: 67 *H, *DR > TL: Insula St. Domingo
Lachnopus planifrons (Gyllenhal in Schönherr, 1840: 385). Morrone, 1999: 141; Girón et al., 2018: 68 *H, *DR > TL: St.
Domingo
Lachnopus plebejus (Gyllenhal, 1834: 23). Morrone, 1999: 141; Girón et al., 2018: 68 *H, *DR > TL: Insula St. Domingo
Lachnopus plumipes Perroud, 1853: 471. Morrone, 1999: 141; Girón et al., 2018: 68 *H, *DR > TL: St-Domingue
Lachnopus proteus (Olivier, 1807: 336). Wolcott, 1927: 250; Morrone, 1999: 141; Girón et al., 2018: 71 *H, *DR > TL: Saint-
Domingue
Lachnopus pruinosus (Gyllenhal, 1834: 33). Morrone, 1999: 141; Girón et al., 2018: 72 *H, *DR > TL: Insula St. Domingo
Lachnopus serraticrus† Poinar & Legalov, 2017: 3. Girón et al., 2018: 79 [Fossil in Dominican amber]
Lachnopus spretus (Gyllenhal, 1834). Morrone, 1999: 142; Girón et al., 2018: 75 *H, *DR > TL: Insula St. Domingo
Melathra Franz, 2011
Melathra huyenae Franz, 2011: 359 *DR > TL: Las Cuevas, Bahia de las Aguilas Station, Pedernales Prov.
Scelianoma Franz & Girón, 2009
Scelianoma compacta† Franz & Zhang, 2017: 8 [Fossil in Dominican amber]
Tetrabothynus Labram & Imhoff, 1852
Tetrabothynus regalis (Linnaeus, 1767). O’Brien & Wibmer, 1982: 41
Tropirhinus Schönherr, 1823
Tropirhinus novemdecimpunctatus (Fabricius, 1781). O’Brien & Wibmer, 1982: 41
Tropirhinus palpebratus† Franz & Zhang, 2017: 12 [Fossil in Dominican amber]
Tribe NAUPACTINI
Brachycamacus† Poinar, Legalov & Brown, 2013

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Brachycamacus gyrommatus† Poinar, Legalov & Brown, 2013: 3 [Fossil in Dominican amber]
Naupactus Dejean, 1821
Naupactus cervinus Boheman, 1840. Woodruff & Bullock, 1979: 1
=Pantomorus godmani Crotch, 1867. Valentine, 1957: 30
Tribe POLYDRUSINI
Apodrosus Marshall, 1922
Apodrosus andersoni Girón & Franz, 2010: 356 *DR > TL: 60 km N. W. Cabo Rojo, 1200, Las Abejas, cloud forest
Apodrosus argentatus Wolcott, 1924. Girón & Franz, 2010: 372
Apodrosus artus Girón & Franz, 2010: 353 *DR > TL: Jarabacoa-Constanza Rd., km 16, 1150 m
Apodrosus canalinasus† Poinar & Legalov, 2017: 8 [Fossil in Dominican amber]
Apodrosus earinusparsus Girón & Franz, 2010: 359 *H > TL: Dept. Sud-Oueste, Parc National La Visite, Morne La Visite,
2100 m
Apodrosus eximius Girón & Franz, 2010: 369 *DR > TL: La Altagracia Province, Guaraguao, 5 m
Apodrosus quisqueyanus Girón & Franz, 2010: 388 *DR > TL: Pedernales, Las Cuevas, Bahía de Las Aguilas station, 40 m
Apodrosus stenoculus Girón & Franz, 2010: 385 *DR > TL: Independencia, Lake Enriquillo National Park, La Azufrada, 4 km
east of La Descubierta, -20 m
Apodrosus tinctocorpus† Poinar & Legalov, 2017: 6 [Fossil in Dominican amber]
Apodrosus viridium Girón & Franz, 2010: 381 *DR > TL: Santiago, Cordillera Septentrional, 4.0 km N of Jacagua on rd. to
Pico Diego de Ocampo, 460 m
Tribe TANYMECINI
Pachnaeus Schönherr, 1826
Pachnaeus litus (Germar, 1824). Perez-Gelabert, 2008: 136
Pachnaeus psittacus (Olivier, 1807). Blackwelder, 1947: 799 *H > TL: Original says ‘Puerto Rico’. But species is endemic to
Hispaniola
Pandeleteius Schönher, 1834
Pandeleteius metallicus Anderson & Ivie, 2015: 322 *DR > TL: Puerto Plata: Pico El Murazo
Paululusus Howden, 1970
Paululusus calypso Howden, 1970: 34. O’Brien & Wibmer, 1982: 47 *H > TL: Port-au-Prince
Paululusus constanzae Howden, 1970: 38. O’Brien & Wibmer, 1982: 47 *DR > TL: Constanza, 3–4000 ft.
Paululusus hispaniolae Howden, 1970: 36. O’Brien & Wibmer, 1982: 47 *H > TL: Port-au-Prince
Scalaventer Howden, 1970
Scalaventer gelinasus Howden, 1970: 27. O’Brien & Wibmer, 1982: 47 *DR > TL: Loma Rucilla and mountains North, 5–
8000 ft.
Notes: The following records for Hispaniola resulted from erroneous reading of the literature when species recorded for the
island of Dominica in the Lesser Antilles were confused and listed for the Dominican Republic: Polydacris scansorius (Klug,
1829) listed by Morrone, 1999: 137 and Diaprepes famelicus (Olivier, 1790) listed by Morrone, 1999: 126. These are hereby
deleted from the fauna of Hispaniola.
Subfamily ERIRRHININAE
Tribe STENOPELMINI
Helodytes Kuschel, 1952
Helodytes foveolatus (Jacquelin du Val, 1945). Kuschel, 1951: 59; O’Brien & Wibmer, 1982: 94
Lissorhoptrus LeConte, 1876
Lissorhoptrus isthmicus Kuschel, 1952. O’Brien & Wibmer, 1982: 93; O’Brien, 1996: 123
Lissorhoptrus oryzophilus Kuschel, 1952. Schmutterer, 1990: 43
Subfamily HYPERINAE
Tribe CEPURINI
Cephalalges Schönherr, 1840
Cephalalges murinus Boheman in Schönherr, 1840: 468. Copiomont, 1867: 443; O’Brien & Wibmer, 1982: 71 *DR, *H > TL:
Saint-Domingue
Subfamily MOLYTINAE
Tribe ANCHONINI
Anchonus Schönherr, 1825
Anchonus suillus (Fabricius, 1792). O’Brien & Wibmer, 1982: 84
Velatis† Poinar & Voisin, 2002

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Velatis dominicana† Poinar & Voisin, 2002: 375 [Fossil in Dominican amber]
Tribe ERODISCINI
Sicoderus Vanin, 1986
Sicoderus aeneus Anderson, 2018: 306 *H > TL: Department du Sud, Formond, 1650 m
Sicoderus alternatus Anderson, 2018: 307 *H > TL: Pedernales, 3.3 km N. E. Los Arroyos, 1450 m
Sicoderus bautistai Anderson, 2018: 308 *H > TL: Prov. La Vega, Constanza, 1160 m
Sicoderus championi Vanin, 1986: 593; Anderson, 2018: 312 *DR > TL: Samaná
Sicoderus detonnancouri Anderson, 2018: 316 *DR > TL: Province María Trinidad Sánchez, Cabrera
Sicoderus guanyangi Anderson, 2018: 317 *DR > TL: Province Monseñor Nouel, 25 km E El Rio (La Vega), 945 m
Sicoderus humeralis Anderson, 2018: 320 *DR > TL: Province Santiago Rodríguez, Sabaneta, Gurabo
Sicoderus medranae Anderson, 2018: 323 *H > TL: Province Monseñor Nouel, La Presa de Blanco
Sicoderus perezi Anderson, 2018: 325 *DR > TL: Province Barahona, 4.5 km S. Barahona
Sicoderus pseudostriatolateralis Anderson, 2018: 328 *DR, *H > TL: Province Pedernales, Parque Nacional Sierra de
Bahoruco, Las Abejas, 1240 m
Sicoderus ramosi (Sleeper, 1954: 593). O’Brien & Wibmer, 1982: 101; Vanin, 1986: 593; Anderson, 2018: 328 *DR > TL:
Colonia Ramfis
Sicoderus schoenherri Vanin, 1986: 596; Anderson, 2018: 330 *DR > TL: La Estrelleta Prov., 4 km SE Río Limpio, ca. 760 m
Sicoderus striatolateralis Anderson, 2018: 332 *DR > TL: Province Barahona, near Filipinas, Mt. Tutu
Sicoderus thomasi Anderson, 2018: 333 *H > TL: Department Sud-Oueste, Parc National La Visite, Morne La Visite, 2100 m
Sicoderus truncatipennis Vanin, 1986: 594; Anderson, 2018: 336 *DR, *H > TL: Port-au-Prince
Sicoderus turnbowi Anderson, 2018: 337 *DR > TL: Province Barahona, Upper Paraiso Valley, 600–800 m
Tribe HYLOBIINI
Hylobius Germar, 1817
Hylobius sp.—Valentine & Valentine (1957: 29); O’Brien & Wibmer (1982: 79)
Marshallius Kuschel, 1955
Marshallius guttatus (Boheman, 1843). O’Brien & Wibmer, 1982: 78
Tribe ITHYPORINI
Cleogonus Schönherr, 1825
Cleogonus distinctus Chevrolat, 1880. Prena & Whitehead, 2012: 74
Cleogonus fratellus Fiedler, 1951. O’Brien & Wibmer, 1982: 135
Cleogonus proximus Chevrolat, 1880a: lxvii. O’Brien & Wibmer, 1982: 135
Cleogonus rubetra (Fabricius, 1787). Prena & Whitehead, 2012: 68
Cleogonus trochilus (Olivier, 1807). O’Brien & Wibmer, 1982: 135
Conotrachelus Dejean, 1835
Conotrachelus cariniscutum Marshall, 1934: 630. Blackwelder, 1947: 848
Conotrachelus sapotae Barber, 1923. Domínguez, 1992: 10/92; Abud, 1995: 7/95; Serra, 2005: 121 [Introduced]
Conotrachelus serpentinus (Klug, 1829). O’Brien, 1982: 132
Nettarhinus Schönherr, 1826
Nettarhinus bilobus (Olivier, 1807: 178). Fiedler, 1954: 188; O’Brien & Wibmer, 1982: 135 *H, *DR > TL: Saint-Domingue
Nettarhinus domingonus Fiedler, 1954: 190. O’Brien & Wibmer, 1982: 135 *DR > TL: S. Domingo
Rhyssomatus Schönherr, 1837
Rhyssomatus aciculaticollis Boheman, 1837: 375. O’Brien & Wibmer, 1982: 125 *DR > TL: St. Domingo
Tribe LYMANTINI
Kuschelaxius Howden, 1992
Kuschelaxius discifer Howden, 1992: 46 *DR > TL: Parque Nacional Armando Bermúdez, Pico Duarte trail, 1335 m
Tribe PETALOCHILINI
Nanus Schönherr, 1844
Nanus uniformis Boheman, 1844. Chevrolat, 1880b: 295; O’Brien & Wibmer, 1982: 97
Tribe PISSODINI
Pissodes Germar, 1817
Pissodes sp.—Valentine & Valentine (1957); O’Brien & Wibmer (1982: 87)
Tribe STERNECHINI
Chalcodermus Dejean, 1835
Chalcodermus ebeninus Boheman in Schoenherr, 1837. Valdez, Matos & Álvarez, 2016: 89

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Subfamily RHYNCHOPHORINAE
Tribe SPHENOPHORINI
Cosmopolites Chevrolat, 1885
Cosmopolites sordidus (Germar, 1824). Wolcott, 1927: 253; O’Brien & Wibmer, 1982: 218 [Introduced]
Metamasius Horn, 1873
Metamasius hemipterus sericeus (Olivier, 1807). Wolcott, 1922: 36; Wolcott, 1927: 252; O’Brien & Wibmer, 1982: 218
Metamasius mosieri Barber, 1920. O’Brien & Wibmer, 1982: 218
Scyphophorus Schönherr, 1838
Scyphophorus acupunctatus Gyllenhal, 1838. O’Brien & Wibmer, 1982: 219 [Introduced]
Sitophilus Schönherr, 1838
Sitophilus granarius (Linnaeus, 1758). SEA, 1999: 129 [Introduced]
Sitophilus linearis (Herbst, 1795). Wolcott, 1927: 256 [Introduced]
Sitophilus oryzae (Linnaeus, 1763). Wolcott, 1927: 256; Blackwelder, 1947: 916 [Introduced]
Sitophilus zeamais Motschulsky, 1855. Schmutterer, 1990: 19 [Introduced]
Sphenophorus Schönherr, 1838
Sphenophorus tenuivittatus (Buchanan, 1936: 149). Blackwelder, 1947: 915; O’Brien & Wibmer, 1982: 213 *DR > TL:
Sánchez
Sphenophorus venatus vestitus Chittenden, 1904. O’Brien & Wibmer, 1982: 214
Subfamily SCOLYTINAE
Tribal classification of Bright (2019) has been kept, all under subfamily Scolytinae.
Tribe SCOLYTINI
Cnemonyx Eichhoff, 1868
Cnemonyx priscus† Bright & Poinar, 1994: 180 [Fossil in Dominican amber]
Loganius Chapuis, 1869
Loganius vagabundus Wood, 1961. Bright, 1981: 153; Bright, 1985: 171; Bright, 2019: 10
Scolytus Geoffroy, 1762
Scolytus poinari† Bright & Poinar, 1994: 181 [Fossil in Dominican amber]
Tribe HYLASTINI
Hylastes Erichson, 1836
Hylastes tenuis Eichhoff, 1868. Bright & Skidmore, 2002: 514; Bright, 2019: 19
Tribe BOTHROSTERNINI
Cnesinus LeConte, 1868
Cnesinus amplus Bright, 2019: 23 *DR > TL: La Vega, vic. La Ciénaga
Cnesinus guadeloupensis Eggers, 1940. Bright, 2019: 25
Cnesinus strigicollis LeConte, 1868. Bright, 2019: 28
Pagiocerus Eichhoff, 1868
Pagiocerus frontalis (Fabricius, 1801). Bright, 2019: 29
Tribe PHLOEOTRIBINI
Phloeotribus Latreille, 1804
Phloeotribus antiguus† Bright & Poinar, 1994: 176 [Fossil in Dominican amber]
Phloeotribus atlanticus Schedl, 1951. Bright, 2019: 33
Tribe PHLOEOSININI
Chramesus LeConte, 1868
Chramesus atlanticus Bright & Torres, 2006. Bright, 2019: 37
Cladoctonus Strohmeyer, 1911
Cladoctonus angustostriatus† Bright & Poinar, 1994: 176 [Fossil in Dominican amber]
Cladoctonus brevisetosus Bright, 1972. Bright, 2019: 45
Cladoctonus cubensis (Wood, 1961). Bright, 2019: 46
Cladoctonus ruber† Bright & Poinar, 1994: 178 [Fossil in Dominican amber]
Dendrosinus Chapuis, 1869
Dendrosinus bourreriae Schwarz, 1920. Bright, 2019: 52
Tribe HYPOBORINI
Chaetophloeus LeConte, 1876
Chaetophloeus cubensis Bright, 1981. Bright, 2019: 57

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Chaetophloeus montanus Bright, 2019: 62 *DR > TL: Sierra de Bahoruco, 4–5000’
Chaetophloeus woodruffi Bright, 2019: 64 *DR > TL: Pedernales, km 19, N. Cabo Rojo, 1000 ft.
Liparthrum Wollaston, 1854
Liparthrum hispaniolum Bright, 1981: 161. Bright, 1985: 171; Bright, 2019: 66 *DR > TL: San Cristóbal, 35 m
Tribe MICRACIDINI
Hylocurus Eichhoff, 1872
Hylocurus elegans Eichhoff, 1872: 134. Bright, 1985: 172; Bright, 2019: 75
Hylocurus quadrispinosus Blackman, 1928. Bright, 2019: 76
Micracis LeConte, 1868
Micracis cubensis Blackman, 1928. Bright, 2019: 80
Parathysanoes Bright, 2019
Parathysanoes absonus Bright, 2019: 82
Pseudothysanoes Blackman, 1920
Pseudothysanoes amoenus Bright, 2019: 87 *DR > TL: La Altagracia, Parque del Este, 2.9 km SW Boca de Yuma, 11 m
Pseudothysanoes caribbeanensis Bright, 2019: 87
Pseudothysanoes dislocatus (Blackman, 1920). Bright & Skidmore, 2002: 514
Pseudothysanoes incertissimus Bright, 2019: 90 *DR > TL: Dominican R., via Canada, ON, Thunder Bay, wood with bark
from Mamajuana
Pseudothysanoes marginatus Bright, 2019: 94
Pseudothysanoes masneri Bright, 2019: 95 *DR > TL: Pedernales, 4 km W Oviedo, 10 m
Pseudothysanoes muricatus Bright, 2019: 97 *DR > TL: Pr. El Seibo, Loma de Chivo, 7 mi. N. of Pedro Sánchez, 5000’
Pseudothysanoes perexiguus Bright, 2019: 97 *DR > TL: San Cristóbal Province, Borbón, Cuevas Pomier, 200 m
Pseudothysanoes securigerus (Blackman, 1943). Bright, 1985: 172; Bright & Torres, 2006: 400; Bright, 2019: 98
Pseudothysanoes trunculus Bright, 2019: 100 *DR > TL: Pedernales, Cabo Rojo
Tribe CRYPHALINI
Allothenemus Bright & Torres, 2006
Allothenemus exquisitus Bright, 2019: 105 *DR > TL: Province La Altagracia, Nisibón, “Papagallo”
Atomothenemus Bright, 2019
Atomothenemus unicus Bright, 2019: 107 *DR > TL: La Altagracia, Parque del Este, caseta Guaraguao, 4.4 km SE Bayahibe, 3
m
Cryptocarenus Eggers, 1937
Cryptocarenus heveae (Hagedorn, 1912). Bright & Skidmore, 2002: 512; Bright, 2019: 108
Cryptocarenus seriatus Eggers, 1933. Bright, 1985: 174; Bright & Torres, 2006: 409; Bright, 2019: 110
Hypocryphalus Hopkins, 19115
Hypocryphalus mangiferae (Stebbing, 1914). Bright, 2019: 112
Hypothenemus Westwood, 1834
Hypothenemus africanus (Hopkins, 1915). Bright & Torres, 2006: 404; Bright, 2019: 123
Hypothenemus atomus Hopkins, 1915. Bright, 2019: 126
Hypothenemus avitus† Bright & Poinar, 1994: 186 [Fossil in Dominican amber]
Hypothenemus columbi Hopkins, 1915. Bright, 2019: 129
Hypothenemus crudiae (Panzer, 1791). Bright & Skidmore, 2002: 514; Bright, 2019: 131 [Introduced]
Hypothenemus discordis Bright, 2019: 133
Hypothenemus erectus LeConte, 1876. Bright, 2019: 134
Hypothenemus eruditus (Westwood, 1836). Bright & Skidmore, 2002: 514; Bright, 2019: 136
=Hypothenemus plumeriae (Nördlinger, 1856). Blackwelder, 1947: 778; Wood & Bright, 1992: 932
Hypothenemus exceptus Bright, 2019: 139
Hypothenemus hampei (Ferrari, 1867). Zuluaga & Mesa, 2000: 197; Vega et al., 2002: 374; Contreras & Camilo, 2007: 44
[Introduced]
Hypothenemus hirsutus (Wood, 1954). Bright, 2019: 146
Hypothenemus interstitialis (Hopkins, 1915). Bright, 2019: 148
Hypothenemus javanus (Eggers, 1908). Wood & Bright, 1992: 932; Peck, 2005: 238; Bright, 2019: 150 [Introduced]
=Stephanoderes prosper Schedl, 1951. Schedl, 1960: 76
Hypothenemus nesiotus Bright, 2019: 154
Hypothenemus obscurifrons Bright, 2019: 155

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Hypothenemus obscurus (Fabricius, 1801). Bright, 1985: 175; Bright & Torres, 2006: 407; Bright, 2019: 156
=Hypothenemus seriatus (Eichhoff, 1872). SEA, 1999: 105; Bright & Torres, 2006: 408
Hypothenemus opacus Eichhoff, 1872). Bright, 2019: 159
Hypothenemus pubescens Hopkins, 1915. Bright, 2019: 165
Hypothenemus rotundicollis (Eichhoff, 1878). Bright, 2019: 167
Hypothenemus setosus (Eichhoff, 1868). Bright & Torres, 2006: 408; Bright, 2019: 170 [Introduced]
Hypothenemus striatus (Atkinson, 1993). Bright, 2019: 172
Scolytogenes Eichhoff, 1878
Scolytogenes jalappae (Letzner, 1848). Bright, 2019: 180 [Introduced]
=Scolytogenes knabi (Hopkins, 1915). Bright, 1985: 175; Bright & Torres, 2006: 402
Tribe HEXACOLINI
Microborus Blandford, 1897
Microborus lautus Wood, 1961. Bright, 2019: 189
Microborus rawlinsi Bright, 2019: 189 *DR > TL: Pedernales, 26 km N Cabo Rojo, 730 m
Pycnarthrum Eichhoff, 1878
Pycnarthrum hispidum (Ferrari, 1867). Bright & Skidmore, 2002: 514; Bright, 2019: 191
Pycnarthrum senectum† Bright & Poinar, 1994: 183 [Fossil in Dominican amber]
Scolytodes Ferrari, 1867
Scolytodes aridus Bright, 2019: 199 *DR > TL: Monte Cristi, 5 km NNE Botoncillo, 50 m
Scolytodes atlanticus Bright & Torres, 2006. Bright, 2019: 200
Scolytodes bellus Bright, 2019: 201 *DR > TL: Province Barahona, nr. Filipinas, Larimar Mine
Scolytodes cubensis (Schedl, 1972). Bright, 2019: 202
Scolytodes electrosimus† Bright & Poinar, 1994: 183 [Fossil in Dominican amber]
Scolytodes glaber (Eichhoff, 1868). Bright, 2019: 203
Scolytodes neoschwarzi† Bright & Poinar, 1994: 181 [Fossil in Dominican amber]
Scolytodes striatulus Wood, 1940. Bright, 2019: 216
Tribe IPINI
Acanthotomicus Blandford, 1894
Acanthotomicus mimicus (Schedl, 1961). Bright, 1981: 152; Bright, 1985: 173; Bright, 2019: 222
Ips De Geer, 1775
Ips cribricollis (Eichhoff, 1869). Bright, 2019: 225
Ips grandicollis (Eichhoff, 1868). Bright, 1985: 173; Bright, 2019: 226
Ips interstitialis (Eichhoff, 1869). Bright, 1985: 173; Haack, 1999: 18; Lanier et al., 1991: 1105; Bright, 2019: 224
Tribe DRYOCOETINI
Coccotrypes Eichhoff, 1878
Coccotrypes advena Blandford, 1894. Bright & Skidmore, 2002: 511; Bright, 2019: 230 [Introduced]
Coccotrypes carpophagus (Hornung, 1842). Bright & Skidmore, 2002: 511; Bright & Torres, 2006: 411; Bright, 2019: 231
[Introduced]
Coccotrypes cyperi (Beeson, 1929). Bright & Skidmore, 2002: 511; Bright, 2019: 232
Coccotrypes dactyliperda (Fabricius, 1801). Bright, 2019: 234
Coccotrypes distinctus (Motschulsky, 1866). Bright & Skidmore, 2002: 511; Bright, 2019: 235
Coccotrypes robustus Eichhoff, 1878. Blackwelder, 1947: 778; Bright, 2019: 239
Dendrocranulus Schedl, 1937
Dendrocranulus ambiguus Bright, 2019: 241
Dendrocranulus carbonarius (Ferrari, 1867). Bright, 2019: 243
Dendrocranulus hispaniolus Bright, 2019: 246 *H > TL: Fermathe, Baptist Haiti Mission
Tribe PREMNOBINI
Premnobius Eichhoff, 1878
Premnobius cavipennis Eichhoff, 1878. Bright, 2019: 249
Tribe XYLEBORINI
Ambrosiodmus Hopkins, 1915
Ambrosiodmus devexulus (Wood, 1977). Bright, 1985: 173; Bright & Torres, 2006: 413; Bright, 2019: 252
Ambrosiodmus hagedorni (Iglesias, 1914). Bright, 2019: 252
=Ambrosiodmus lecontei Hopkins, 1915. Bright, 1985: 173; Bright & Torres, 2006: 413

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 179
Ambrosiodmus infidelis Bright, 2019: 254 *DR > TL: La Vega, Estación Casabito, Reserva Científica Ebano Verde
Ambrosiodmus obliquus (LeConte, 1878). Bright, 1981: 157;
Bright, 1985: 173; Bright & Torres, 2006: 413; Bright, 2019: 255
=Ambrosiodmus klapperichi Bright, 1985: 178
Dryocoetoides Hopkins, 1915
Dryocoetoides capucinus (Eichhoff, 1869). Bright, 1981: 154; Bright, 1985: 173; Bright, 2019: 259
Dryocoetoides cristatus (Fabricius, 1801). Bright & Skidmore, 2002: 513; Bright & Torres, 2006: 413; Bright, 2019: 260
Dryocoetoides pseudosolitarius (Eggers, 1933). Wood & Bright, 1992: 658; Bright, 2019: 262
Euwallacea Hopkins, 2019
Euwallacea elevatus (Eggers, 1931). Bright & Skidmore, 2002: 521; Bright, 2019: 265
Euwallacea simulatus (Bright, 1972). Bright & Skidmore, 2002: 521; Bright, 2019: 270
Theoborus Hopkins, 1915
Theoborus atlanticus (Bright & Torres, 2006). Bright, 2019: 273
Theoborus puertoricensis Bright & Torres, 2006. Bright, 2019: 275
Theoborus ricini (Eggers, 1932). Peck, 2005: 236; Bright, 2019: 275
=Theoborus solitariceps (Schedl, 1954). Bright, 1982: 165; Bright, 1985: 173
Theoborus theobromae Hopkins, 1915. Bright, 1985: 173; Bright, 2019: 276
Xyleborinus Reitter, 1913
Xyleborinus gracilis (Eichhoff, 1868). Bright, 2019: 281
Xyleborinus intersetosus (Blandford, 1898). Bright, 2019: 283
Xyleborus Eichhoff, 1864
Xyleborus affinis Eichhoff, 1868. Bright & Torres, 2006: 417; Bright, 2019: 288
Xyleborus bispinatus Eichhoff, 1868. Bright, 2019: 291
Xyleborus ferrugineus (Fabricius, 1801). Bright & Torres, 2006: 419; Bright, 2019: 294
Xyleborus pubescens Zimmermann, 1868. Bright, 2019: 298
=Xyleborus intrusus Blandford, 1898. Bright, 1985: 174
Xyleborus spinulosus Blanford, 1898. Bright, 1982: 165; Bright, 1985: 174; Bright & Torres, 2006: 419; Bright, 2019: 301
Xyleborus volvulus (Fabricius, 1775). Blackwelder, 1947: 780; Bright & Torres, 2006: 419; Bright, 302
Xylosandrus Reitter, 1913
Xylosandrus compactus (Eichhoff, 1876). Valdez, Matos & Álvarez, 2016: 28; Bright, 2019: 306
Xylosandrus curtulus (Eichhoff, 1869). Bright, 2019: 309
Tribe PITYOPHTHORINI
Araptus Eichhoff, 1878
Araptus beckeri Bright, 2019: 318
Araptus ciseruditus Bright, 2019: 320 *DR > TL: M. Nouel, Bonao
Araptus fuscus Bright, 2019: 324 *DR > TL: Independencia, Sierra de Neiba just south of crest, 5 km NW Angel Feliz, 1780 m
Araptus hymenaeae (Eggers, 1933). Wood & Bright, 1992: 957; Bright & Torres, 2006: 421; Bright, 2019: 329
=Neodryocoetes guianae Blackman, 1942: 186. Blackwelder, 1947: 781
Araptus ineditus Bright, 2019: 331
Araptus niger Bright, 1972. Bright, 1981: 152; Wood & Bright, 1992: 959; Bright, 2019: 335
Araptus pallidus (Blackman, 1942). Bright, 1981: 152; Bright, 1985: 176; Wood & Bright, 1992: 960; Bright, 2019: 336
Araptus politus (Blanford, 1904). Bright, 1985: 176: Bright, 2019: 337
Araptus turnbowi Bright, 2019: 338 *DR > TL: Barahona, 2 km E Payaso
Conophthorus Hopkins, 1915
Conophthorus insulatus Bright, 2019: 340 *DR > TL: Province Santiago, Par. Nac. Armando Bermúdez, 2575 m, Agüita Fría
Gnatholeptus Blackman, 1943
Gnatholeptus concinnus Bright, 2019: 343 *DR > TL: Duarte, Reserva Loma Quita Espuela, Canela, 13.2 km NNE San
Francisco de Macorís, 523 m
Gnatholeptus hispanicus Bright, 2019: 344 *DR > TL: Province Pedernales, N. of Pedernales, border rd. rio, Banano, S. of
Arroyos
Pityophthorus Eichhoff, 1864
Pityophthorus antillicus Bright, 1981: 162. Bright, 1985: 176; Bright, 2019: 357
Pityophthorus antiquarius† Bright & Poinar, 1994: 190 [Fossil in Dominican amber]
Pityophthorus aphelofacies† Bright & Poinar, 1994: 190 [Fossil in Dominican amber]

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Pityophthorus auspicatus Bright, 2019: 358 *DR > TL: La Vega, 2.5 km SW Pinar Bonito, 1430 m
Pityophthorus confusus sequestus Bright, 2019: 364 *DR > TL: Province Pedernales, ca. 35 km NNW Cabo Rojo, 1370 m, El
Aceitillar
Pityophthorus congruus Bright, 2019: 364 *DR > TL: Independencia, Sierra de Neiba, south slope near summit, 4 km N Angel
Feliz, 1825 m
Pityophthorus eggersianus Schedl, 1958. Bright, 2019: 368
Pityophthorus favorabilis Bright, 2019: 370 *DR > TL: Province San Juan, 2,545 m, Agüita Fría, Par. Nac. A. Bermúdez
Pityophthorus gratus Bright, 2019: 372 *DR > TL: La Altagracia, Parque del Este, caseta Guaraguao, 4.4 km SE Bayahibe, 3
m
Pityophthorus illuminus Bright, 2019: 373 *DR > TL: La Vega, 10 km NE Jarabacoa
Pityophthorus laevis (Schedl, 1938). Bright, 2019: 375
=Pityophthorus pudens (Blackman, 1942). Bright, 1981: 156; Bright, 1985: 176
Pityophthorus liquidambarus Blackman, 1921. Bright, 2019: 376
Pityophthorus masneri Bright, 2019: 378 *DR > TL: Pedernales, 3.3 km NE Los Arroyos, 1450 m
Pityophthorus pinavorus Bright, 1985: 182. Bright, 2019: 381
Pityophthorus senticosus Bright, 2019: 386 *DR > TL: La Altagracia, Parque del Este, 2.9 km SW Boca de Yuma, 11 m
Pityophthorus subconcentralis Schedl, 1928. Bright, 2019: 387
=Pityophthorus hispaniolus Bright, 1985: 181. Bright, 1985: 176; Bright & Torres, 2006: 423
Pityophthorus subtilis Bright, 2019: 388 *DR > TL: La Altagracia, Parque del Este, caseta Guaraguao, 4.4 km SE Bayahibe, 3
m
Pityophthorus temporarius† Bright & Poinar, 1994: 190 [Fossil in Dominican amber]
Pityophthorus youngi Bright, 2019: 391 *DR > TL: Pedernales, 26 km N Cabo Rojo, 730 m
Sphenoceros Schedl, 1939
Sphenoceros antillicus Bright, 2019: 394
Tribe CORTHYLINI
Amphicranus Erichson, 1836
Amphicranus hispaniolus Bright, 2019: 396. *DR > TL: Province Pedernales, Sierra de Bahoruco, 1240 m, Las Abejas
Amphicranus taino Bright, 2019: 396. *DR > TL: Province La Vega, La Cienaga, 1100 m
Corthylus Erichson, 1836
Corthylus alpestris Bright, 2019: 400 *DR > TL: Independencia, Sierra de Neiba just south of crest, 5 km NNW Angel Feliz,
1780 m
Corthylus monticellus Bright, 2019: 402 *DR > TL: Barahona, Eastern Sierra Bahoruco, Reserva, 12.8 km NE Paraiso, 1230 m
Corthylus papulans Eichhoff, 1868. Bright & Skidmore, 2002: 512; Bright, 2019: 403
Corthylus tuberculatus Eggers, 1940. Bright, 2019: 406
Corthylus versicolor Bright, 2019: 407 *DR > TL: La Vega, Res. Cient. Ebano Verde, 1000 m
Gnathotrichus Eichhoff, 1869
Gnathotrichus hispaniolus Bright, 2019: 408 *DR >TL: Pr. La Vega, La Ciénaga de Manabao, park hdqt., 3000 ft.
Gnathotrupes Schedl, 1951
Gnathotrupes megapunctatus Bright, 2019: 410
Microcorthylus Ferrari, 1867
Microcorthylus antiquarius† Bright & Poinar, 1994: 188 [Fossil in Dominican amber]
Microcorthylus minutissimus Schedl, 1952. Bright, 2019: 413
Monarthrum Kirsch, 1866
Monarthrum bifoveatum Wood, 1974. Bright, 1985: 176
Monarthrum bullatum Bright, 2019: 418 *DR > TL: Puerto Plata, Pico El Murazo, north slope near summit, 910 m
Monarthrum collinum Bright, 2019: 419 *DR > TL: Province Barahona, Larimar Mine, Filipinas, 3300 ft.
Monarthrum mali (Fitch, 1856). Bright, 2019: 420
Tricolus Blandford, 1905
Tricolus animatus Bright, 2019: 423 *DR > TL: La Colonia, 1000 m
Tricolus gracilis Eggers, 1937. Bright, 2019: 424
=Tricolus perdiligens Schedl, 1950. Bright, 1981: 156; Bright, 1985: 176
Tricolus incomptus Bright, 2019: 426 *DR > TL: Province Barahona, nr. Filipinas, Larimar Mine
Tribe HYLESININI
Electroborus† Cognato, 2013

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Electroborus brighti† Cognato, 2013: 503 [Fossil in Dominican amber]
Tribe INCERTAE SEDIS
Corthylites† Bright & Poinar, 1994
Corthylites bicolor† Bright & Poinar, 1994: 188 [Fossil in Dominican amber]
Dryomites† Bright & Poinar, 1994
Dryomites incognitus† Bright & Poinar, 1994: 188 [Fossil in Dominican amber]
Micracites† Bright & Poinar, 1994
Micracites squamifera† Bright & Poinar, 1994: 186 [Fossil in Dominican amber]
Paleophthorus† Bright & Poinar, 1994
Paleophthorus bispinus† Bright & Poinar, 1994: 190 [Fossil in Dominican amber]
Paleosinus† Bright & Poinar, 1994
Paleosinus fossulatus† Bright & Poinar, 1994: 178 [Fossil in Dominican amber]
Protosinus† Bright & Poinar, 1994
Protosinus hispaniolensis† Bright & Poinar, 1994: 180 [Fossil in Dominican amber]
Subfamily PLATYPODINAE
Cenocephalus Chapuis, 1861
Cenocephalus antillicus† Bright & Poinar, 1994: 171 [Fossil in Dominican amber]
Cenocephalus antiguus† Bright & Poinar, 1994: 172 [Fossil in Dominican amber]
Cenocephalus biconicus† Bright & Poinar, 1994: 172 [Fossil in Dominican amber]
Cenocephalus exquisitus† Bright & Poinar, 1994: 172. Peris et al., 2015: 8 [Fossil in Dominican amber]
Cenocephalus quasiexquisitus† Davis & Engel, 2007: 121. Peris et al., 2015: 9 [Fossil in Dominican amber]
Cenocephalus rhinoceroide† (Schawaller, 1994: 10). Davis & Engel, 2007: 124; Peris et al., 2015: 9 [Fossil in Dominican
amber]
Cenocephalus senectus† Bright & Poinar, 1994: 172 [Fossil in Dominican amber]
Cenocephalus spinatus† Bright & Poinar, 1994: 174. Peris et al., 2015: 9 [Fossil in Dominican amber]
Euplatypus Wood, 1993
Euplatypus pulicaris (Chapuis, 1865). Bright & Skidmore, 2002: 522
Gnathotrichus Eichhoff, 1868
Gnathotrichus fossor† Bright & Poinar, 1994: 186 [Fossil in Dominican amber]
Gnathotrichus materiarius (Fitch, 1858). Bright, 1985: 176
Platypus Herbst, 1793
Platypus parallelus (Fabricius, 1801). Peck, 2005: 239
Platypus poeyi Guérin-Méneville, 1844. Blackwelder, 1947: 790
Platypus subcostatus Jacquelin du Val, 1856. Blackwelder, 1947: 790
Teloplatypus Wood, 1993
Teloplatypus excisus (Chapuis, 1865). Bright & Skidmore, 2002: 522
Teloplatypus ustulatus (Chapuis, 1865). Haack, 1999: 19
Tesserocerus Saunders, 1837
Tesserocerus primus† Bright & Poinar, 1994: 176 [Fossil in Dominican amber]

Family RHYNCHITIDAE
Subfamily RHYNCHITINAE
Tribe AULETINI
Rhynchitobius Sharp, 1889
Rhynchitobius tanyrhinus† Poinar & Legalov, 2015: 71 [Fossil in Dominican amber]
Rhynchitobius xuthocolus† Poinar & Legalov, 2015: 72 [Fossil in Dominican amber]

Order DERMAPTERA
Key reference: Steinmann (1989).

Family CARCINOPHORIDAE
Subfamily ANTISOLABINAE
Antisolabis Burr, 1911
Antisolabis allardi (Brindle, 1971: 31). Steinmann, 1989: 330 *H, *DR > TL: Loma Constanza, 5200 ft. in altitude

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Subfamily CARCINOPHORINAE
Anisolabis Fieber, 1853
Anisolabis maritima (Bonelli, 1832). Burr, 1911: 448; Steinmann, 1989: 222
Carcinophora Scudder, 1876
Carcinophora americana (Palisot de Beauvois, 1817). Steinmann, 1989: 243
Euborellia Burr, 1910
Euborellia ambigua (Borelli, 1905). Steinmann, 1989: 261
Euborellia caraibea Hebard, 1921. Steinmann, 1981: 159

Family FORFICULIDAE
Doru Burr, 1907
Doru albipes (Fabricius, 1787). Steinmann, 1989: 799
Doru taeniatum (Dohrn, 1862). Steinmann, 1981: 166; Steinmann, 1989: 805

Family LABIDURIDAE
Labidura Leach, 1815
Labidura riparia (Pallas, 1773). Steinmann, 1989: 377

Family LABIIDAE
Subfamily LABIINAE
Paralabella Steinmann, 1989
Paralabella curvicauda (Motchulsky, 1793). Rehn & Hebard, 1917: 640; Steinmann, 1989: 578
Paralabella dorsalis (Burmeister, 1838). Brindle, 1971d: 175; Steinmann, 1989: 580
Subfamily SPARATTINAE
Sparatta Audinet-Serville, 1838
Sparatta nigrina Stål, 1855. Steinmann, 1989: 451
Subfamily SPONGIPHORINAE
Formicilabia Rehn & Hebard, 1917
Formicilabia caribea Rehn & Hebard, 1917: 642. Steinmann, 1989: 485 *DR > TL: San Francisco Mountains, San Domingo
Marava Burr, 1911
Marava rogersi (De Bormans, 1893). Steinmann, 1989: 509
Marava unidentata (Palisot de Beauvois, 1805). Brindle, 1971a: 558; Steinmann, 1989: 513
Spongovostox Burr, 1911
Spongovostox ghiliani (Dohrn, 1864). Steinmann, 1989: 538
Subfamily STRONGYLOPSALINAE
Strongylopsalis Burr, 1900
Strongylopsalis haitica Steinmann, 1986: 371. Steinmann, 1989: 417 *H > TL: La Visite vic., La Selle Range, 5–7000 ft.

Family PYGIDICRANIDAE
Pyragropsis Borelli, 1908
Pyragropsis buscki (Caudell, 1907). Steinmann, 1989: 111
Pyragropsis tristani Borelli, 1908. Steinmann, 1989: 113
Note: The citation of Sparatta dominicana for Hispaniola by Steinmann (1989: 448) which was subsequently followed by
Perez-Gelabert (2008: 142) is an error. This species was described from Dominica in the Lesser Antilles.

Family SPONGIPHORIDAE
Nesogerax† Engel, 2017
Nesogerax polytrichum† Engel, 2018: 35 [Fossil in Dominican amber]

Superorder DICTYOPTERA

Taxonomic summary:
Order Blattodea
Superfamily Blattoidea

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 183
 Family Blattidae
Superfamily Blaberoidea
Family Blaberidae
Family Ectobiidae
Family Corydiidae
Family Polyphagidae
Epifamily Termitoidae
Family Kalotermitidae
Family Mastotermitidae
Family Rhinotermitidae
Family Termitidae
Order Mantodea
Family Epaphroditidae
Family Mantidae

Order BLATTODEA
Key reference: Gutiérrez and Perez-Gelabert (2000). Classification follows the Cockroach Species File online. Beccaloni, G.
W. 2014. Cockroach Species File Online. Version 5.0/5.0. World Wide Web electronic publication. http://
cockroach.speciesfile.org/HomePage/Cockroach/HomePage.aspx

Superfamily BLATTOIDEA
Family BLATTIDAE
Subfamily BLATTINAE
Blatta Linnaeus, 1758
Blatta orientalis Linnaeus, 1758. Marcano, 1995: 3/95; Barnes & Brenner, 1996: 70; Gutiérrez & Perez-Gelabert, 2000: 426
[Introduced]
Pelmatosilpha Dohrn, 1887
Pelmatosilpha sp.—Gutiérrez & Perez-Gelabert (2000: 427)
Periplaneta Burmeister, 1838
Periplaneta americana (Linnaeus, 1758). Rehn & Hebard, 1927: 188; Gutiérrez & Perez-Gelabert, 2000: 426
Periplaneta australasiae (Fabricius, 1775). Rehn & Hebard, 1927: 190; Gutiérrez & Perez-Gelabert, 2000: 426
Periplaneta brunnea Burmeister, 1838. Rehn & Hebard, 1927: 189; Gutiérrez & Perez-Gelabert, 2000: 426
Subfamily POLYZOSTERIINAE
Eurycotis Stål, 1874
Eurycotis adamesi Gutiérrez, 2014: 6 *DR > TL: Parque Nacional Armando Bermúdez, Ciénaga de Manabao, Prov. La Vega,
camino a La Virgen
=Eurycotis sp. 2—Gutiérrez & Perez-Gelabert (2000: 427)
Eurycotis barahonensis Gutiérrez, 2013a: 20 *DR > TL: Entronque El Cruce de Monteada Nueva, carretera de Cabral a Polo,
Prov. Barahona
Eurycotis hebardi Gutiérrez, 2013a: 26 *DR > TL: Loma Casabito, Reserva Científica Ebano Verde, Prov. La Vega, ~1000
msnm
=Eurycotis sp. 4—Gutiérrez & Perez-Gelabert (2000: 427)
Eurycotis hierroi Gutiérrez, 2014: 12 *DR > TL: ~100 m up from El Sitio del Agua, bosque nublado N Los Bolos, Sierra de
Neiba, Prov. Independencia, 1520 m
Eurycotis histrio Rehn, 1937: 226. Gutiérrez & Perez-Gelabert, 2000: 426; Gutiérrez, 2013a: 18 *H > TL: Northeastern
foothills of Massif de La Hotte
Eurycotis isabeltorres Gutiérrez, 2014: 15 *DR > TL: Loma Isabel de Torres, Puerto Plata
Eurycotis jaragua Gutiérrez, 2013a: 22 *DR > TL: Parque Nacional Jaragua, Isla Beata
Eurycotis mercedes Gutiérrez, 2014: 18 *DR > TL: Las Mercedes, Prov. Pedernales
Eurycotis neiba Gutiérrez, 2014: 10 *DR > TL: Los Bolos (1100 m), Sierra de Neiba, Postrer Río, Prov. Independencia
Eurycotis quitaespuela Gutiérrez, 2014: 14 *DR > TL: Loma Quita Espuela, San Francisco de Macorís, Prov. Duarte
Eurycotis rehni Gutiérrez, 2013a: 29 *DR > TL: La Sal, Reserva Científica Ebano Verde, Prov. La Vega, 1043 msnm
=Eurycotis sp. 3—Gutiérrez & Perez-Gelabert (2000: 427)
Eurycotis rothi Gutiérrez, 2001: 72 *DR > TL: Parque Nacional Nalga de Maco, cerca de Río Limpio, camino a Lenó, Prov.

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Elias Piña
Eurycotis ruthae Gutiérrez, 2013a: 25 *DR > TL: Chaquito Prieto, El Curro, Sierra Martín García, 731 msnm
Eurycotis samana Gutiérrez, 2014: 2 *DR > TL: La Laguna, N El Valle, Prov. Samaná
Eurycotis tibialis Hebard, 1916: 264. Rehn & Hebard, 1927: 178; Gutiérrez & Perez-Gelabert, 2000: 427; Gutiérrez, 2013a: 18;
Gutiérrez, 2014: 2 *DR > TL: San Francisco Mountains, San Domingo

Superfamily BLABEROIDEA
Family BLABERIDAE
Subfamily BLABERINAE
Aspiduchus Rehn & Hebard, 1927
Aspiduchus rothi Gutiérrez & Perez-Gelabert, 2001: 73 *DR > TL: Pedernales prov., cueva Robinson
Blaberus Serville, 1831
Blaberus craniifer Burmeister, 1838. Gutiérrez & Perez-Gelabert, 2000: 428
Blaberus discoidalis (Serville, 1839). Rehn & Hebard, 1927: 260; Gutiérrez & Perez-Gelabert, 2000: 428
Blaberus giganteus Linnaeus, 1758. Gutiérrez & Perez-Gelabert, 2000: 428
Hemiblabera Saussure, 1893
Hemiblabera tenebricosa Rehn & Hebard, 1927: 271. Gutiérrez & Perez-Gelabert, 2000: 428
Phoetalia Stål, 1874
Phoetalia circumvagans (Burmeister, 1838). Rehn & Hebard, 1927: 203; Gutiérrez & Perez-Gelabert, 2000: 428
Phoetalia pallida (Brunner, 1865). Gutiérrez & Perez-Gelabert, 2000: 429
Subfamily EPILAMPRINAE
Tribe EPILAMPRINI
Epilampra Burmeister, 1838
Epilampra haitensis Rehn & Hebard, 1927: 228. Gutiérrez & Perez-Gelabert, 2000: 430 *H, *DR > TL: Port-au-Prince,
Pétionville (elev. ~2,250 ft.)
Epilampra quisqueiana Rehn & Hebard, 1927: 233. Gutiérrez & Perez-Gelabert, 2000: 430 *DR > TL: San Lorenzo, Prov.
Samaná
Epilampra sabulosa Walker, 1868: 70. Rehn & Hebard, 1927: 231; Gutiérrez & Perez-Gelabert, 2000: 430 *H, *DR > TL: St.
Domingo
Epilampra sp. 1—Gutiérrez & Perez-Gelabert (2000: 430)
Epilampra sp. 2—Gutiérrez & Perez-Gelabert (2000: 430)
Epilampra sp. 3—Gutiérrez & Perez-Gelabert (2000: 430)
Tribe COLAPTEROBLATTINI
Colapteroblatta Hebard, 1919
Colapteroblatta ariasae Gutiérrez, 2012b: 45 *DR > TL: Honduras, Matadero, Prov. Peravia
Colapteroblatta darlingtoni Roth & Gutiérrez, 1998: 195. Gutiérrez & Perez-Gelabert, 2000: 429; Gutiérrez, 2012: 40 *DR >
TL: Valle Nuevo, SE Constanza, c. 7000 ft.
Colapteroblatta rehni Roth & Gutiérrez, 1998: 198. Gutiérrez & Perez-Gelabert, 2000: 429; Gutiérrez, 2012: 45 *DR > TL:
Mt. Diego de Ocampo, 3–4000 ft.
Colapteroblatta sp. 1—Gutiérrez & Perez-Gelabert (2000: 429)
Colapteroblatta sp. 2—Gutiérrez & Perez-Gelabert (2000: 429)
Tribe INCERTAE SEDIS
Calolampra Saussure, 1893
Calolampra signatura (Walker, 1871). Gutiérrez & Perez-Gelabert, 2000: 429
Subfamily OXYHALOINAE
Tribe NAUPHOETINI
Nauphoeta Burmeister, 1838
Nauphoeta cinerea (Olivier, 1789). Rehn & Hebard, 1927: 254; Gutiérrez & Perez-Gelabert, 2000: 430
Rhyparobia Krauss, 1892
Rhyparobia maderae (Fabricius, 1781). Rehn & Hebard, 1927: 242; Gutiérrez & Perez-Gelabert, 2000: 431
Subfamily PANCHLORINAE
Panchlora Burmeister, 1838
Panchlora nivea (Linnaeus, 1758). Gutiérrez & Perez-Gelabert, 2000: 431
Panchlora viridis (Fabricius, 1775). Gutiérrez & Perez-Gelabert, 2000: 431

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=Panchlora antillarum Saussure, 1862. Rehn & Hebard, 1927: 249
Subfamily PYCNOSCELINAE
Pycnoscelus Scudder, 1862
Pycnoscelus surinamensis (Linnaeus, 1758). Rehn & Hebard, 1927: 243; Wolcott, 1927: 140; Gutiérrez & Perez-Gelabert,
2000: 431

Family ECTOBIIDAE
Subfamily ANAPLECTINAE
Anaplecta Burmeister, 1838
Anaplecta sp.†—Gutiérrez & Perez-Gelabert (2000: 441) [Fossil in Dominican amber]
Anaplecta sp.—Gutiérrez (2002: 14)
Subfamily BLATTELLINAE
Blattella Caudell, 1903
Blattella germanica (Linnaeus, 1767). Rehn & Hebard, 1927: 98; Wolcott, 1927: 143; Gutiérrez & Perez-Gelabert, 2000: 432
[Introduced]
Ischnoptera Burmeister, 1838
Ischnoptera aglandis Roth, 2001: 538. Roth, 2002: 128 *H > TL: Port-au-Prince
Ischnoptera caborojoensis Roth, 2001. Roth, 2002: 123 *DR > TL: Pedernales, 37 km N Cabo Rojo, 1500 m
Ischnoptera lestrelleta Roth, 2001: 528. Roth, 2002: 115 *DR > TL: La Estrelleta Prov., 4 km SE of Río Limpio, ca. 760 m
Ischnoptera ligula Rehn & Hebard, 1927: 116. Gutiérrez & Perez-Gelabert, 2000: 432
Ischnoptera pedernalesensis Roth, 2001: 525. Roth, 2002: 115 *DR > TL: Pedernales, La Abeja, 38 km NNW Cabo Rojo, 1160
m
Ischnoptera rufa rufa (De Geer, 1773). Rehn & Hebard, 1927: 112; Gutiérrez & Perez-Gelabert, 2000: 432
Litoblatta Hebard, 1921
Litoblatta sp.—Gutiérrez & Perez-Gelabert (2000: 433)
Nesomylacris Rehn & Hebard, 1927
Nesomylacris haitiensis Gutiérrez, 2013c: 18 *H > TL: Nr. P[ort]-au P[rince], 4–6000 ft.
=Nesomylacris sp. 2—Gutiérrez & Perez-Gelabert (2000: 433)
Nesomylacris sp. 1—Gutiérrez & Perez-Gelabert (2000: 433)
Symploce Hebard, 1916
Symploce bicolor (Palisot de Beauvois, 1805). Rehn & Hebard, 1927: 137; Gutiérrez & Perez-Gelabert, 2000: 433
Symploce cristata Rehn & Hebard, 1927: 129. Gutiérrez & Perez-Gelabert, 2000: 434
Symploce flagellata Hebard, 1916. Rehn & Hebard, 1927: 136; Gutiérrez & Perez-Gelabert, 2000: 434
Symploce jamaicana (Rehn, 1903). Gutiérrez & Perez-Gelabert, 2000: 434
Symploce morsei Hebard, 1916. Rehn & Hebard, 1927: 131; Gutiérrez & Perez-Gelabert, 2000: 434
Symploce sp. 1—Gutiérrez & Perez-Gelabert (2000: 434)
Symploce sp. 2—Gutiérrez & Perez-Gelabert (2000: 434)
Subfamily NYCTIBORINAE
Nyctibora Burmeister, 1838
Nyctibora laevigata (Palisot de Beauvois, 1805). Gutiérrez & Perez-Gelabert, 2000: 435
Nyctibora sp. 1—Gutiérrez & Perez-Gelabert (2000: 435)
Nyctibora stygia Walker, 1868. Rehn & Hebard, 1927: 195; Gutiérrez & Perez-Gelabert, 2000: 435
Subfamily PSEUDOPHYLODROMINAE
Cariblatta Hebard, 1916
Cariblatta advena Rehn, 1932: 125. Gutiérrez & Perez-Gelabert, 2000: 435 *H > TL: Fond des Negres, Massif de la Hotte
Cariblatta delicatula (Guérin-Méneville, 1857: 436). Rehn & Hebard, 1927: 25
Cariblatta islacolonis Rehn & Hebard, 1927: 35. Gutiérrez & Perez-Gelabert, 2000: 436 *H, *DR > TL: San Lorenzo, Prov.
Samaná
Cariblatta leucops Rehn & Hebard, 1927: 24. Gutiérrez & Perez-Gelabert, 2000: 436 *H > TL: Port-au-Prince
Cariblatta sp. 1—Gutiérrez & Perez-Gelabert (2000: 436)
Cariblatta sp. 2—Gutiérrez & Perez-Gelabert (2000: 436)
Cariblatta sp. 1†—Gutiérrez & Perez-Gelabert (2000: 441) [Fossil in Dominican amber]
Cariblatta sp. 2†—Gutiérrez & Perez-Gelabert (2000: 441) [Fossil in Dominican amber]
Cariblatta unguiculata Rehn & Hebard, 1927: 22. Gutiérrez & Perez-Gelabert, 2000: 436 *DR > TL: San Francisco Mountains

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Euthlastoblatta Hebard, 1917
Euthlastoblatta diaphana (Fabricius, 1793). Rehn & Hebard, 1927: 8; Gutiérrez & Perez-Gelabert, 2000: 436
Euthlastoblatta sp.†—Gutiérrez & Perez-Gelabert (2000: 441) [Fossil in Dominican amber]
Euthlastoblatta sp. 1—Gutiérrez & Perez-Gelabert (2000: 436)
Euthlastoblatta sp. 2—Gutiérrez & Perez-Gelabert (2000: 437)
Lophoblatta Hebard, 1929
Lophoblatta sp.—Gutiérrez & Perez-Gelabert (2000: 437)
Nahublattella Bruijning, 1959
Nahublattella laodamia (Rehn & Hebard, 1927). Lopes & Oliveira, 2004: 263 [Record needs confirmation]
Neoblattella Shelford, 1911
Neoblattella carcinus Rehn & Hebard, 1927: 61. Gutiérrez & Perez-Gelabert, 2000: 437 *DR > TL: St. Domingo
Neoblattella detersa (Walker, 1868). Rehn & Hebard, 1927: 74; Gutiérrez & Perez-Gelabert, 2000: 437
Neoblattella infausta Rehn & Hebard, 1927: 87. Gutiérrez & Perez-Gelabert, 2000: 437 *H > TL: Haiti
Neoblattella proserpina Rehn & Hebard, 1927. Gutiérrez & Perez-Gelabert, 2000: 437
Neoblattella sp. 1—Gutiérrez & Perez-Gelabert (2000: 438)
Neoblattella sp. 2—Gutiérrez & Perez-Gelabert (2000: 438)
Neoblattella sp. 3—Gutiérrez & Perez-Gelabert (2000: 438)
Neoblattella sp. 4—Gutiérrez & Perez-Gelabert (2000: 438)
Neoblattella sp. 5—Gutiérrez & Perez-Gelabert (2000: 438)
Paranocticola Bonfils, 1977
Paranocticola sp. 1—Gutiérrez & Perez-Gelabert (2000: 432)
Paranocticola sp. 2—Gutiérrez & Perez-Gelabert (2000: 432)
Paranocticola sp. 3—Gutiérrez & Perez-Gelabert (2000: 432)
Plectoptera Saussure, 1864
Plectoptera pygmaea (Palisot de Beauvois, 1805). Rehn & Hebard, 1927: 303; Gutiérrez & Perez-Gelabert, 2000: 439
Plectoptera electrina† Gorochov & Aniyutkin, 2007: 162 [Fossil in Dominican amber]
Pseudosymploce Rehn & Hebard, 1927
Pseudosymploce elongata (Palisot de Beauvois, 1805: 183). Rehn & Hebard, 1927: 104; Gutiérrez & Perez-Gelabert, 2000:
439 *H, *DR > TL: Saint-Domingue
Pseudosymploce sp.†—Gutiérrez & Perez-Gelabert (2000: 441) [Fossil in Dominican amber]
Supella Shelford, 1911
Supella longipalpa (Fabricius, 1798). Gutiérrez & Perez-Gelabert, 2000: 439 [Introduced]
Subfamily INCERTAE SEDIS
Nymphodromia Rehn & Hebard, 1927
Nymphodromia miranda Rehn & Hebard, 1927: 100. Gutiérrez & Perez-Gelabert, 2000: 438 *DR > TL: San Francisco
Mountains, 13 km north of San Cristóbal

Family CORYDIIDAE
Compsodes Hebard, 1917
Compsodes perezgelaberti Gutiérrez, 2012a: 57 *DR > TL: La Altagracia, 2 km N Bayahibe, 10 m
=Compsodes sp.—Gutiérrez & Perez-Gelabert (2000: 439)
Erucoblatta† Gorochov & Aniyutkin, 2007
Erucoblatta semicaeca† Gorochov & Aniyutkin, 2007: 158 [Fossil in Dominican amber]

Family POLYPHAGIDAE
Subfamily HOLOCOMPSINAE
Holocompsa Burmeister, 1838
Holocompsa abbreviata† Gorochov & Aniyutkin, 2007: 159 [Fossil in Dominican amber]
Holocompsa cyanea Burmeister, 1838. Gutiérrez & Perez-Gelabert, 2000: 440
Holocompsa nitidula (Fabricius, 1781). Gutiérrez & Perez-Gelabert, 2000: 440
Holocompsa nigra† Gorochov & Aniyutkin, 2007: 159 [Fossil in Dominican amber]

Epifamily TERMITOIDAE (Isoptera)

Key references: Scheffrahn et al. (1994), Constantino (1998).

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 187
Family KALOTERMITIDAE
Cryptotermes Banks, 1906
Cryptotermes brevis (Walker, 1853). Scheffrahn et al., 1994: 219
Cryptotermes cavifrons Banks, 1906. Scheffrahn et al., 1994: 219
Cryptotermes chasei Scheffrahn, 1993: 501. Scheffrahn et al., 1994: 219; Scheffrahn & Krecek, 1999: 130 *DR > TL: Juanillo,
La Altagracia Prov.
Cryptotermes juliani Scheffrahn & Krecek, 1999: 150 *H, *DR > TL: Peravia Prov., Las Avispas, ca. 1000 m
Cryptotermes longicollis (Banks, 1918). Scheffrahn et al., 1994: 219
Cryptotermes mangoldi Scheffrahn & Krecek, 1999: 152 *H, *DR > TL: La Altagracia Prov., 0.5 km W Club Mediterráneo
Cryptotermes rotundiceps Scheffrahn & Krecek, 1999: 163
Cryptotermes spathifrons Scheffrahn & Krecek, 1999: 165
Cryptotermes yamini† Krishna & Bachus, 1987: 2 [Fossil in Dominican amber]
Glyptotermes Froggatt, 1897
Glyptotermes glaesarius† Engel & Krishna, 2007: 269 [Fossil in Dominican amber]
Glyptotermes grimaldii† Engel & Krishna, 2007: 267 [Fossil in Dominican amber]
Glyptotermes liberatus (Snyder, 1929). Scheffrahn et al., 1994: 219
Glyptotermes paleoliberatus† Engel & Krishna, 2007: 266 [Fossil in Dominican amber]
Incisitermes Krishna, 1961
Incisitermes bequaerti (Snyder, 1929). Scheffrahn et al., 1994: 219
Incisitermes milleri (Emerson, 1943). Scheffrahn et al., 1994: 219
Incisitermes peritus† Engel & Krishna, 2007: 264 [Fossil in Dominican amber]
Neotermes Holmgren, 1911
Neotermes castaneus (Burmeister, 1839). Scheffrahn et al., 1994: 219
Neotermes mona (Banks, 1919). Krececk et al., 2000: 269
Neotermes platyfrons Krececk & Scheffrahn, 2001: 70 *DR > TL: 2.6 km SW of Punta Cana, La Altagracia Prov.
Procryptotermes Holmgren, 1911
Procryptotermes corniceps (Snyder, 1923). Scheffrahn et al., 1994: 219

Family MASTOTERMITIDAE
Mastotermes Froggatt, 1897
Mastotermes electrodominicus† Krishna & Grimaldi, 1991: 3. De Salle et al., 1992: 1933; Wu, 1997: 148; Wier et al., 2002:
1410 [Fossil in Dominican amber]

Family RHINOTERMITIDAE
Subfamily COPTOTERMITINAE
Coptotermes Wasmann, 1896
Coptotermes hirsutus† Krishna & Grimaldi, 2009: 4 [Fossil in Dominican amber]
Coptotermes paleodominicanus† Krishna & Grimaldi, 2009: 5 [Fossil in Dominican amber]
Coptotermes priscus† Emerson, 1971: 268; Krishna & Grimaldi, 2009: 6 [Fossil in Dominican amber]
Subfamily HETEROTERMITINAE
Heterotermes Froggatt, 1897
Heterotermes convexinotatus (Snyder, 1924). Szalanski et al., 2004: 558
Heterotermes sp.—Scheffrahn et al. (1994: 219)
Subfamily RHINOTERMITINAE
Dolichorhinotermes Snyder & Emerson, 1949
Dolichorhinotermes dominicus† Schlemmermeyer & Cancello, 2000: 305 [Fossil in Dominican amber]
Rhinotermes Hagen, 1858
Rhinotermes marginalis (Linnaeus, 1758). Scheffrahn et al., 1994: 220

Family TERMITIDAE
Subfamily APICOTERMITINAE
Anoplotermes Müller, 1873
Anoplotermes bohio† Krishna & Grimaldi, 2009: 11 [Fossil in Dominican amber]

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Anoplotermes cacique† Krishna & Grimaldi, 2009: 11 [Fossil in Dominican amber]
Anoplotermes carib† Krishna & Grimaldi, 2009: 11 [Fossil in Dominican amber]
Anoplotermes maboya† Krishna & Grimaldi, 2009: 12 [Fossil in Dominican amber]
Anoplotermes meridianus Emerson, 1925. Scheffrahn et al., 1994: 220
Anoplotermes naboria† Krishna & Grimaldi, 2009: 13 [Fossil in Dominican amber]
Anoplotermes nitaino† Krishna & Grimaldi, 2009: 15 [Fossil in Dominican amber]
Anoplotermes quisqueya† Krishna & Grimaldi, 2009: 15 [Fossil in Dominican amber]
Anoplotermes taino† Krishna & Grimaldi, 2009: 16 [Fossil in Dominican amber]
Anoplotermes sp. A—Scheffrahn et al. (1994: 219)
Anoplotermes sp. B—Scheffrahn et al. (1994: 219)
Subfamily NASUTITERMITINAE
Antillitermes Roisin, Scheffrahn & Krecek, 1996
Antillitermes subtilis (Scheffrahn & Krecek, 1993). Roisin et al., 1996: 786
Atlantitermes Fontes, 1979
Atlantitermes antillea† Krishna & Grimaldi, 2009: 24 [Fossil in Dominican amber]
Atlantitermes magnoculus† Krishna & Grimaldi, 2009: 26 [Fossil in Dominican amber]
Caribitermes Roisin, Scheffrahn & Krecek, 1996
Caribitermes discolor (Banks, 1919). Roisin et al., 1996: 786
Caribitermes hispaniola† Krishna & Grimaldi, 2009: 39 [Fossil in Dominican amber]
Constrictotermes Holmgren, 1910
Constrictotermes electroconstrictus† Krishna, 1996: 3 [Fossil in Dominican amber]
Nasutitermes Dudley, 1890
Nasutitermes amplioculatus† Krishna & Grimaldi, 2009: 29 [Fossil in Dominican amber]
Nasutitermes costalis (Holmgren, 1910). Scheffrahn et al., 1994: 220; Scheffrahn, 2002: 273
Nasutitermes corniger (Motschulsky, 1855). Scheffrahn et al., 2005: 274
Nasutitermes electronasutus† Krishna, 1996: 8 [Fossil in Dominican amber]
Nasutitermes hubbardi Banks, 1919. Scheffrahn et al., 1994: 220
Nasutitermes incisus† Krishna & Grimaldi, 2009: 29 [Fossil in Dominican amber]
Nasutitermes lividus (Burmeister, 1839). Scheffrahn et al., 1994: 220
Nasutitermes magnocellus† Krishna & Grimaldi, 2009: 32 [Fossil in Dominican amber]
Nasutitermes medioculatus† Krishna & Grimaldi, 2009: 33 [Fossil in Dominican amber]
Nasutitermes pilosus† Krishna & Grimaldi, 2009: 33 [Fossil in Dominican amber]
Nasutitermes rotundicephalus† Krishna & Grimaldi, 2009: 38 [Fossil in Dominican amber]
Nasutitermes seminudus† Krishna & Grimaldi, 2009: 34 [Fossil in Dominican amber]
Parvitermes Emerson, 1949
Parvitermes antillarum (Holmgren, 1910: 310). Roisin et al., 1996: 786 *H, *DR > TL: San Domingo
Parvitermes collinsae Scheffrahn & Roisin, 1995: 587. Roisin et al., 1996: 786 *DR > TL: 2 km W Fondo Negro, Barahona
Prov.
Parvitermes dominicanae Scheffrahn et al., 1998: 180 *DR > TL: Bomba de Guanuma, Distrito Nacional
Parvitermes flaveolus (Banks, 1919: 488). Roisin et al., 1996: 786 *H, *DR > TL: Pétionville
Parvitermes longinasus† Krishna & Grimaldi, 2009: 39 [Fossil in Dominican amber]
Parvitermes n. sp.—Scheffrahn et al. (1994: 220); Roisin et al. (1996: 786)
Parvitermes pallidiceps (Banks, 1919: 487). Scheffrahn & Roisin, 1995: 592; Roisin et al., 1996: 786 *H, *DR > TL: Port-au-
Prince
Parvitermes subtilis Scheffrahn & Krecek, 1993: 604
Parvitermes toussainti (Banks, 1919: 488). Spaeth, 1967: 852; Roisin et al., 1996: 786 *H, *DR > TL: Milot
Subulitermes Holmgren, 1910
Subulitermes hispaniola† Krishna & Grimaldi, 2009: 38 [Fossil in Dominican amber]
Subulitermes insularis† Krishna & Grimaldi, 2009: 37 [Fossil in Dominican amber]
Velocitermes Holmgren, 1912
Velocitermes bulbus† Krishna & Grimaldi, 2009: 40 [Fossil in Dominican amber]
Subfamily TERMITINAE
Amitermes Silvestri, 1901
Amitermes lucidus† Krishna & Grimaldi, 2009: 18 [Fossil in Dominican amber]

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Microcerotermes Silvestri, 1901
Microcerotermes arboreus Emerson, 1925. Scheffrahn et al., 1994: 220
Microcerotermes insulanus† Krishna & Grimaldi, 2009: 20 [Fossil in Dominican amber]
Microcerotermes setosus† Krishna & Grimaldi, 2009: 21 [Fossil in Dominican amber]
Termes Linnaeus, 1758
Termes hispaniolae (Banks, 1918). Scheffrahn et al., 1994: 220
Termes primitivus† Krishna & Grimaldi, 2009: 21 [Fossil in Dominican amber]

Order MANTODEA
Family EPAPHRODITIDAE
Epaphrodita Serville, 1831
Epaphrodita lobivertex Lombardo & Perez-Gelabert, 2004: 46; Otte & Spearman, 2005: 84 *DR > TL: Santo Domingo Prov.,
Sierra Prieta
Epaphrodita musarum Palisot de Beauvois, 1805. Wolcott, 1927: 144; Lombardo & Perez-Gelabert, 2004: 44; Otte &
Spearman, 2005: 84 *H, *DR > TL: Saint-Domingue
=Epaphrodita dentifrons Saussure, 1872. Lombardo & Perez-Gelabert, 2004: 46

Family MANTIDAE
Callimantis Stål, 1877
Callimantis antillarum Saussure, 1859. Wolcott, 1927: 144; Lombardo & Perez-Gelabert, 2004: 40
Gonatista Saussure, 1869
Gonatista jaiba Lombardo & Perez-Gelabert, 2004: 38; Otte & Spearman, 2005: 128 *DR > TL: Pedernales Prov., Oviedo
Gonatista major Caudell, 1912: 161. Lombardo & Perez-Gelabert, 2004: 38; Otte & Spearman, 2005: 128 *H, *DR > TL: San
Domingo
Gonatista phryganoides Serville, 1839: 198. Caudell, 1912: 162; Lombardo & Perez-Gelabert, 2004: 36; Otte & Spearman,
2005: 128 *H, *DR > TL: “Amerique septentrionale”
Gonatista reticulata (Thunberg, 1815). Lombardo & Perez-Gelabert, 2004: 36; Otte & Spearman, 2005: 128
Paramusonia Rehn, 1904
Paramusonia cubensis (Saussure, 1869). Lombardo & Perez-Gelabert, 2004: 40; Perez-Gelabert, 2007: 220
Phyllovates Kirby, 1904
Phyllovates cingulata (Drury, 1773). Lombardo & Perez-Gelabert, 2004: 35; Otte & Spearman, 2005: 312
Stagmomantis Saussure, 1869
Stagmomantis domingensis (Palisot de Beauvois, 1805: 61). Lombardo & Perez-Gelabert, 2004: 42; Otte & Spearman, 2005:
209; Maxwell, 2014: 515
Stagmomantis marginata (Palisot de Beauvois, 1805). Otte & Spearman, 2005: 210; Maxwell, 2014: 517 [This is an uncertain
species]

Order DIPTERA

Taxonomic summary:
Suborder Nematocera
Infraorder Tipulomorpha
Family Tipulidae
Suborder Psychodomorpha
Family Psychodidae
Family Scatopsidae
Family Valeseguyidae
Infraorder Culicomorpha
Family Ceratopogonidae
Family Chironomidae
Family Corethrellidae
Family Culicidae
Family Dixidae
Family Simuliidae

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Infraorder Bibionomorpha
Family Anisopodidae
Family Bibionidae
Family Cecidomyiidae
Family Keroplatidae
Family Lygistorrhinidae
Family Sciaridae
Suborder Brachycera
Infraorder Stratiomyomorpha
Family Stratiomyidae
Infraorder Tabanomorpha
Family Acroceridae
Family Rhagionidae
Family Tabanidae
Superfamily Asiloidea
Family Asilidae
Family Bombyliidae
Family Mydidae
Family Mythicomyiidae
Family Scenopinidae
Family Therevidae
Superfamily Empidoidea
Family Dolichopodidae
Family Empididae
Family Hybotidae
Infraorder Muscomorpha
Family Phoridae
Family Pipunculidae
Family Platypezidae
Family Syrphidae
Calyptratae
Family Anthomyiidae
Family Calliphoridae
Family Hippoboscidae
Family Streblidae
Family Fanniidae
Family Muscidae
Family Oestridae
Family Mesembrinellidae
Family Sarcophagidae
Family Scatophagidae
Family Tachinidae
Acalyptratae
Superfamily Nerioidea
Family Cypselosomatidae
Family Micropezidae
Family Neriidae
Superfamily Conopoidea
Family Conopidae
Superfamily Diopsoidea
Family Syringogastridae
Family Tanypezidae
Superfamily Tephritoidea
Family Lonchaeidae

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 Family Richardiidae
Family Tephritidae
Family Ulidiidae
Superfamily Lauxanioidea
Family Lauxaniidae
Superfamily Sciomyzoidea
Family Sciomyzidae
Family Sepsidae
Superfamily Opomyzoidea
Family Agromyzidae
Family Anthomyzidae
Family Asteiidae
Family Aulacigastridae
Family Clusiidae
Family Odiniidae
Family Periscelididae
Superfamily Carnoidea
Family Canacidae
Family Carnidae
Family Chloropidae
Family Milichiidae
Superfamily Sphaeroceroidea
Family Sphaeroceridae
Superfamily Ephydroidea
Family Curtonotidae
Family Drosophilidae
Family Ephydridae

Suborder NEMATOCERA
Infraorder TIPULOMORPHA
Family TIPULIDAE
Subfamily LIMONIINAE
Tribe ERIOPTERINI
Erioptera Meigen, 1803
Eriptera (Mesocyphona) caliptera caliptera Say, 1823. Gavryushin, 2018: 63
Erioptera (Mesocyphona) immaculata immaculata Alexander, 1913. Gavryushin, 2018: 65
Erioptera sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Gnophomyia Osten Sacken, 1859
Gnophomyia asymmetrica† Podenas & Poinar, 1999: 602 [Fossil in Dominican amber]
Gnophomyia ieva† Podenas & Poinar, 1999: 600 [Fossil in Dominican amber]
Gnophomyia martyni† Podenas & Poinar, 1999: 602 [Fossil in Dominican amber]
Gnophomyia sp.†—Krzeminski (1992: 108) [Fossil in Dominican amber]
Gonomyia Meigen, 1818
Gonomyia (Gonomyia) sp.—Gavryushin (2018: 66)
Gonomyia (Leiponeura) puer Alexander, 1913: 506. Alexander & Alexander, 1970: 4.137
Gonomyia (Leiponeura) orthomera Alexander, 1937. Gavryushin, 2018: 65
Gonomyia (Leiponeura) producta Alexander, 1919. Gavryushin, 2018: 65
Gonomyia (Paralipophleps) peracuta peracuta Alexander, 1928. Gavryushin, 2018: 65
Gonomyia (Paralipophleps) pleuralis (Williston, 1896). Alexander, 1933: 374; Gavryushin, 2018: 66
Molophilus Curtis, 1831
Molophilus (Miomolophilus) theischingeri† Krzeminski, 1992: 108 [Fossil in Dominican amber]
Rhabdomastix Skuse, 1890
Rhabdomastix (Rhabdomastix) parvula Alexander, 1947. Alexander & Alexander, 1970: 4.155
Styringomyia Loew, 1845

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Styringomyia dominicana† Podenas & Poinar, 1999: 598 [Fossil in Dominican amber]
Styringomyia optiva† Podenas & Poinar, 2001: 872 [Fossil in Dominican amber]
Toxorhina Loew, 1845
Toxorhina (Ceratocheilus) sp.†—Krzeminski (1992: 108) [Fossil in Dominican amber]
Toxorhina (Toxorhina) domingensis Alexander, 1937: 186. Alexander & Alexander, 1970: 4.167; Gavryushin, 2018: 66 *DR >
TL: Santo Domingo
Trentepohila Bigot, 1854
Trentepohila (Paramongoma) agri† Podenas & Poinar, 1999: 604 [Fossil in Dominican amber]
Tribe HEXATOMINI
Australolimnophila Alexander, 1920
Australolimnophila sp.†—Krzeminski (1992: 107) [Fossil in Dominican amber]
Elephantomyia Osten Sacken, 1859
Elephantomyia (Elephantomyia) grata† Podenas & Poinar, 2001: 867 [Fossil in Dominican amber]
Elephantomyia sp.†—Krzeminski (1992: 107) [Fossil in Dominican amber]
Elephantomyia (Elephantomyia) westwoodi antillarum Alexander, 1933: 386.
Epiphragma Osten Sacken, 1859
Epiphragma aurora† Podenas & Poinar, 2001: 869 [Fossil in Dominican amber]
Epiphragma (Epiphragma) auricosta Alexander, 1939: 116. Alexander & Alexander, 1970: 4.95 *DR > TL: Valle Nuevo,
southeast of Constanza, altitude about 7,000 ft.
Epiphragma (Epiphragma) buscki Alexander, 1913: 540. Alexander & Alexander, 1970: 4.96 *DR > TL: San Francisco
Mountains [San Cristóbal], Santo Domingo
Hexatoma Latreille, 1809
Hexatoma (Eriocera) aetherea (Alexander, 1916: 346). Alexander & Alexander, 1970: 4.100 *DR > TL: San Lorenzo
Hexatoma (Eriocera) domingensis (Alexander, 1916: 346). Alexander & Alexander, 1970: 4.101 *DR > TL: Sánchez
Hexatoma (Eriocera) multiguttula Alexander, 1939: 124. Alexander & Alexander, 1970: 4.103 *DR > TL: Valle Nuevo,
southeast of Constanza, altitude about 7,000 ft.
Polymera Wiedemann, 1821
Polymera sp.†—Podenas & Poinar (1999: 598) [Fossil in Dominican amber]
Polymera (Polymera) specula† Podenas & Poinar, 1999: 596 [Fossil in Dominican amber]
Polymera (Polymera) virgo† Podenas & Poinar, 1999: 596 [Fossil in Dominican amber]
Shannonomyia Alexander, 1928
Shannonomyia (Shannonomyia) batesi Alexander, 1939: 119. Alexander & Alexander, 1970: 4.114; Gavryushin, 2018: 65 *H >
TL: La Visite and vicinity, La Selle Range, altitude 5,000–7,000 ft.
Shannonomyia (Shannonomyia) haitensis Alexander, 1939: 120. Alexander & Alexander, 1970: 4.115 *H > TL: La Visite and
vicinity, La Selle Range, altitude 5,000–7,000 ft.
Shannonomyia (Shannonomyia) septempunctata Alexander, 1939: 121. Alexander & Alexander, 1970: 4.116 *DR > TL: Valle
Nuevo, southeast of Constanza, altitude about 7,000 ft.
Tribe LIMONIINI
Dicranomyia Stephens, 1829
Dicranomyia (Dicranomyia) brevivena torrida (Alexander, 1933). Gavryushin, 2018: 65
Dicranomyia (Dicranomyia) calliergon calliergon Alexander, 1939: 109. Alexander & Alexander, 1970: 4.51 *H > TL: La
Visite and vicinity, La Selle Range, altitude 5,000–7,000 ft.
Dicranomyia (Dicranomyia) calliergon polygrapha Alexander, 1939: 111. Alexander & Alexander, 1970: 4.51 *DR > TL:
Valle Nuevo, southeast of Constanza, altitude about 7,000 ft.
Dicranomyia (Dicranomyia) distans Osten Sacken, 1860. Gavryushin, 2018: 65
Dicranomyia (Dicranomyia) divisa Alexander, 1929. Alexander & Alexander, 1970: 4.52; Gavryushin, 2018: 65
Dicranomyia (Dicranomyia) fera† Podenas & Poinar, 1999: 606 [Fossil in Dominican amber]
Dicranomyia (Dicranomyia) indefensa Alexander, 1939: 112. Alexander & Alexander, 1970: 4.53 *H > TL: La Visite, La Selle
Range, altitude 5,000–7,000 ft.
Dicranomyia (Dicranomyia) lema† Podenas & Poinar, 1999: 607 [Fossil in Dominican amber]
Geranomyia Haliday, 1833
Geranomyia banksiana (Alexander, 1939). Gavryushin, 2018: 66
Geranomyia cinereinota (Alexander, 1913). Alexander & Alexander, 1970: 4.61
=Geranomyia domingensis Alexander, 1916. Alexander & Alexander, 1970: 4.61

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Geranomyia constanza Gavryushin, 2018: 62 *DR > TL: Constanza
Geranomyia euchara† Podenas & Poinar, 2001: 864 [Fossil in Dominican amber]
Geranomyia sp.†—Krzeminski (1992: 108) [Fossil in Dominican amber]
Geranomyia tibialis (Loew, 1851). Alexander & Alexander, 1970: 4.66; Gavryushin, 2018: 66
Helius Lepeletier & Serville, 1828
Helius (Helius) phasmatis Alexander, 1945: 56. Alexander & Alexander, 1970: 4.46 *DR > TL: Santo Domingo
Limonia Meigen, 1803
Limonia sp.†—Krzeminski (1992: 108) [Fossil in Dominican amber]
Neolimonia Alexander, 1964
Neolimonia domballah (Alexander, 1939: 108). Alexander & Alexander, 1970: 4.71 *DR > TL: Loma Rucilla and mountains to
north, altitude 5,000–8,000 ft.
Orimarga Osten-Sacken, 1869
Orimarga (Orimarga) perpallens Alexander, 1964. Gavryushin, 2018: 66
Rhipidia Meigen, 1818
Rhipidia (Rhipidia) domestica domestica Osten Sacken, 1860. Alexander, 1933: 360; Gavryushin, 2018: 66
Rhipidia (Rhipidia) mira† Podenas & Poinar, 1999: 607 [Fossil in Dominican amber]
Rhipidia (Rhipidia) pratti (Alexander, 1950). Gavryushin, 2018: 66
Rhipidia (Rhipidia) schwarzi (Alexander, 1912). Alexander & Alexander, 1970: 4.78; Gavryushin, 2018: 66
Subfamily TIPULINAE
Brachypremna Osten Sacken, 1886
Brachypremna brodzinskii† Krzeminski, 1996: 276 [Fossil in Dominican amber]
Brachypremna unicolor Osten Sacken, 1887. Alexander & Alexander, 1970: 4.7
Dolichopeza Curtis, 1825
Dolichopeza (Megistomastix) domingensis Alexander, 1939: 106. Alexander & Alexander, 1970: 4.8 *DR > TL: Loma Rucilla,
altitude 5,000–10,000 ft.
Megistocera Wiedemann, 1828
Megistocera longipennis (Macquart, 1838). Alexander, 1933: 354
Nephrotoma Meigen, 1803
Nephrotoma glabricristata Alexander, 1939: 102. Alexander & Alexander, 1970: 4.17
Tipula Linnaeus, 1758
Tipula (Eumicrotipula) darlingtoniana Alexander, 1939: 101. Alexander & Alexander, 1970: 4.24 *DR > TL: Valle Nuevo,
southeast of Constanza, altitude about 7,000 ft.

Suborder PSYCHODOMORPHA
Family PSYCHODIDAE
Subfamily BRUCHOMYIINAE
Boreofairchildia Wagner & Stuckenberg, 2016
Boreofairchildia antillarum (Fairchild, 1952: 270). Wagner & Stuckenberg, 2016: 163; Wagner, 2017: 112 *DR > TL: Sabana
de la Mar, 100 ft. elev., 4 km south of town
Boreofairchildia dominicana† Wagner, 2017: 110 [Fossil in Dominican amber]
Boreofairchildia hennigianus† (Schlüter, 1978: 243); Wagner, 2017: 110 [Fossil in Dominican amber]
Boreofairchildia scheveni† (Wagner, 2006: 83). Wagner, 2017: 110 [Fossil in Dominican amber]
Boreofairchildia youngi (Wagner, 2000: 234). Wagner & Stuckenberg, 2016: 164 *DR > TL: Pedernales, 26 km N Cabo Rojo,
730 m
Subfamily PHLEBOTOMINAE
Micropygomyia Barreto, 1962
Micropygomyia brandaoi† Andrade et al., 2008: 344. Andrade Filho, Galati & Brazil, 2009: 977; Shimabukuro et al., 2017: 76
[Fossil in Dominican amber]
Micropygomyia (Micropygomyia) cayennensis hispaniolae (Fairchild & Trapido, 1950: 415). Young & Duncan, 1994: 645;
Shimabakuro et al., 2017: 73
Micropygomyia (Sauromyia) dorafeliciangeliae† Andrade Filho, Galati & Brazil, 2009: 970; Shimabukuro et al., 2017: 75
[Fossil in Dominican amber]
Pintomyia Costa Lima, 1932
Pintomyia adiketis† (Poinar, 2008: 1); Shimabukuro et al., 2017: 86 [Fossil in Dominican amber]

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Pintomyia brazilorum† Andrade Filho et al., 2006b: 141. Andrade Filho, Galati & Brazil, 2009: 977; Shimabukuro et al., 2017:
86 [Fossil in Dominican amber]
Pintomyia dissimilis† Andrade Filho, Serra e Meira, Castillo Sanguinette & Brazil, 2009: 2; Shimabukuro et al., 2017: 86
[Fossil in Dominican amber]
Pintomyia dominicana† Andrade Filho, Galati & Brazil, 2009: 973; Shimabukuro et al., 2017: 86 [Fossil in Dominican amber]
Pintomyia falcaorum† (Brazil & Andrade Filho, 2002: 501). Peñalver & Grimaldi, 2006: 180; Andrade Filho, Galati & Brazil,
2009: 975; Shimabukuro et al., 2017: 86 [Fossil in Dominican amber]
Pintomyia filipalpis† (Peñalver & Grimaldi, 2006: 180). Andrade Filho, Galati & Brazil, 2009: 975; Shimabukuro et al., 2017:
86 [Fossil in Dominican amber]
Pintomyia killickorum† (Andrade Filho & Falcão in Andrade Filho et al., 2004: 71). Andrade Filho, Galati & Brazil, 2009: 975;
Shimabukuro et al., 2017: 86 [Fossil in Dominican amber]
Pintomyia miocena† (Peñalver & Grimaldi, 2006: 181). Andrade Filho, Galati & Brazil, 2009: 976; Shimabukuro et al., 2017:
86 [Fossil in Dominican amber]
Pintomyia paleopestis† (Peñalver & Grimaldi, 2006: 182). Andrade Filho, Galati & Brazil, 2009: 976; Shimabukuro et al.,
2017: 86 [Fossil in Dominican amber]
Pintomyia succini† (Peñalver & Grimaldi, 2006: 181). Andrade Filho, Galati & Brazil, 2009: 975; Shimabukuro et al., 2017: 86
[Fossil in Dominican amber]
Pintomyia (Pifanomyia) christophei (Fairchild & Trapido, 1950: 409). Young & Duncan, 1994: 176; Shimabukuro et al., 2017:
83 *DR, *H > TL: 16 km south of Hato Mayor, Prov. Seibo
Pintomyia (Pifanomyia) paleotownsendi† Andrade Filho et al., 2006a: 57. Andrade Filho, Galati & Brazil, 2009: 977;
Shimabukuro et al., 2017: 84 [Fossil in Dominican amber]
Pintomyia (Pifanomyia) paleotrichia† Andrade Filho et al., 2007: 901. Andrade Filho, Galati & Brazil, 2009: 977;
Shimabukuro et al., 2017: 84 [Fossil in Dominican amber]
Psathyromyia Barreto, 1962
Psathyromyia (Forattiniella) schleei† (Peñalver & Grimaldi, 2006: 183). Andrade Filho, Galati & Brazil, 2009: 976;
Shimabukuro et al., 2017: 93 [Fossil in Dominican amber]
Subfamily PSYCHODINAE
Atrichobrunettia Satchell, 1953
Atrichobrunettia dominicana Wagner, 2000: 238 *DR > TL: Pedernales, 23.5 km N Cabo Rojo, 540 m
Psychoda Latreille, 1796
Psychoda montana Wagner, 2000: 240 *DR > TL: Independencia, 32 km NW La Descubierta, Sierra de Neiba, 1850 m
Psychoda sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Quatiella Botosaneanu & Vaillant, 1970
Quatiella interdicta (Dyar, 1928). Wagner, 2000: 243
Setomima Enderlein, 1937
Setomima (Setomima) triangularis Wagner, 2000: 242
Succinarisemus Wagner, 2002
Succinarisemus scheveni† Wagner, 2002: 30 [Fossil in Dominican amber]
Telmatoscopus Eaton, 1904
Telmatoscopus (Clogmia) albipunctata (Williston, 1893). Smith & Audant, 1930: 977; Duckhouse, 1973: 6A.14
Telmatoscopus sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Telmatoscopus thompsoni Wagner, 2000: 240 *DR > TL: Pedernales, 9.5 km N Cabo Rojo, 35 m
Tinearia Schellenberg, 1803
Tinearia alternata (Say, 1824). Wagner, 2000: 244
Subfamily TRICHOMYIINAE
Trichomyia Haliday, 1839
Trichomyia masneri Wagner, 2000: 236 *DR > TL: Pedernales, 23–26 km N Cabo Rojo, 500–700 m
Trichomyia rawlinsi Wagner, 2000: 238 *DR > TL: Pedernales, 23.5 km N Cabo Rojo, 540 m

Family SCATOPSIDAE
Ectaetia Enderlein, 1912
Ectaetia sp.†—Amorim (1998: 5) [Fossil in Dominican amber]
Procolobostema Cook, 1971
Procolobostema roseni† Amorim, 1998: 5 [Fossil in Dominican amber]

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Rhegmoclemina Enderlein, 1936
Rhegmoclemina (Neorhegmoclemina?) sp.†—Cook (1990: 389); Amorim (1998: 5) [Fossil in Dominican amber]
Scatopse Geoffroy, 1762
Scatopse sp.†—Peñalver (1995: 61) [Fossil in Dominican amber]

Family VALESEGUYIDAE
Valeseguya Colless, 1990
Valeseguya disjuncta† Grimaldi, 1991: 5. Amorim & Grimaldi, 2006: 512 [Fossil in Dominican amber]

Infraorder CULICOMORPHA
Family CERATOPOGONIDAE
Subfamily CERATOPOGONINAE
Tribe CULICOIDINI
Culicoides Latreille, 1809
Culicoides borinqueni Fox & Hoffman, 1944. Delecolle et al., 1986: 112
Culicoides (Drymodesmyia) haitiensis Delecolle et al., 1986: 108 *H > TL: Thomassin, entre Pétionville et Kenscoff, 1000 m
Culicoides (Hoffmania) foxi Ortiz, 1950. Spinelli et al., 1993: 42
Culicoides (Hoffmania) insignis Lutz, 1913. Wirth, 1974: 57
Culicoides (Hoffmania) trinidadensis Hoffman, 1925. Wirth & Blanton, 1956: 324; Wirth, 1974: 82
=Culicoides (Hoffmania) oliveri Fox & Hoffman, 1944: 108. Fox, 1946: 255
Culicoides (Oecacta) ambericus† Szadziewski & Grogan, 1998a: 41 [Fossil in Dominican amber]
Culicoides (Oecacta) antilleanus† Szadziewski & Grogan, 1998a: 40 [Fossil in Dominican amber]
Culicoides (Oecacta) barbosai Wirth & Blanton, 1956. Lowrie & Raccurt, 1984: 1275
Culicoides (Oecacta) brodzinskyi† Szadziewski & Grogan, 1998a: 40 [Fossil in Dominican amber]
Culicoides (Oecacta) furens (Poey, 1851). Wolcott, 1927: 312; Wirth & Blanton, 1956: 159; Wirth, 1974: 43
Culicoides (Oecacta) hispaniolicus† Szadziewski & Grogan, 1998a: 41 [Fossil in Dominican amber]
Culicoides mammalicolus† Szadziewski & Grogan, 1998a: 40 [Fossil in Dominican amber]
Tribe CERATOPOGONINI
Allaudomyia Kieffer, 1913
Allaudomyia caribbeana Spinelli & Wirth, 1984: 678
Baeodasymyia Clastrier & Raccurt, 1979
Baeodasymyia dominicana† Szadziewski & Grogan, 1994: 220 [Fossil in Dominican amber]
Baeodasymyia modesta Clastrier & Raccurt, 1979: 100. Borkent & Craig, 1999: 20 *H > TL: Bois d’Eau, entre Port-Margot et
Bayeaux
Brachypogon Kieffer, 1899
Brachypogon (Brachypogon) americanus† Szadziewski & Grogan, 1998a: 42 [Fossil in Dominican amber]
Brachypogon (Brachypogon) woodruffi Spinelli & Grogan, 1998: 72 *DR > TL: Barahona Prov., nr. Filipinas, Larimar Mine
Brachypogon (Isohelea) dominicanus† Szadziewski & Grogan, 1998a: 42 [Fossil in Dominican amber]
Brachypogon (Isohelea) prominuloides† Szadziewski & Grogan, 1998a: 42 [Fossil in Dominican amber]
Ceratopogon Meigen, 1803
Ceratopogon eriophorus Williston, 1896. Curran, 1928: 11
Nannohelea Grogan & Wirth, 1980
Nannohelea sp. A†—Szadziewski & Grogan (1998a: 44) [Fossil in Dominican amber]
Parabezzia Malloch, 1915
Parabezzia arenosa Clastrier & Raccurt, 1979: 172 *H > TL: L’Eau Spinique, 480 m
Parabezzia caribbeana Clastrier & Raccurt, 1979: 173 *H > TL: Ennery, 320 m
Parabezzia haitiensis Clastrier & Raccurt, 1979: 170 *H > TL: Plaisance, 380 m
Parabezzia wirthi Clastrier & Raccurt, 1979: 169 *H > TL: Grande Rivière du Nord, 70 m
Stilobezzia Kieffer, 1911
Stilobezzia (Acanthohelea) dominicana† Szadziewski & Grogan, 1998a: 45 [Fossil in Dominican amber]
Stilobezzia (Acanthohelea) sp. A†—Szadziewski & Grogan (1998a: 45) [Fossil in Dominican amber]
Stilobezzia (Stilobezzia) antilleana† Szadziewski & Grogan, 1998a: 46 [Fossil in Dominican amber]
Tribe HETEROMYIINI
Heteromyia Say, 1825

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Heteromyia dominicana† Szadziewski & Grogan, 1996: 254 [Fossil in Dominican amber]
Heteromyia sp.†—Schlee (1980: 51); Szadziewski & Grogan (1994: 221) [Fossil in Dominican amber]
Tribe PALPOMYIINI
Bezzia Kieffer, 1899
Bezzia jamaicensis Spinelli & Wirth, 1989. Spinelli & Wirth, 1989: 115
Bezzia punctipennis (Williston, 1896). Spinelli & Wirth, 1989: 118
Palpomyia Meigen, 1818
Palpomyia wirthorum† Szadziewski & Grogan, 1996: 257 [Fossil in Dominican amber]
Phaenobezzia Haeselbarth, 1965
Phaenobezzia wirthi† Szadziewski & Grogan, 1996: 259 [Fossil in Dominican amber]
Tribe STENOXENINI
Stenoxenus Coquillett, 1899
Stenoxenus setiger Macfie, 1939. Wirth & Ratanaworabhan, 1972: 1387
Subfamily DASYHELEINAE
Dasyhelea Kieffer, 1911
Dasyhelea (Sebessia) antilleana† Szadziewski & Grogan, 1998: 257 [Fossil in Dominican amber]
Dasyhelea (Sebessia) dominicana† Szadziewski & Grogan, 1998: 258 [Fossil in Dominican amber]
Dasyhelea (Sebessia) hispaniolae† Szadziewski & Grogan, 1998: 260 [Fossil in Dominican amber]
Dasyhelea (Sebessia) minuticola† Szadziewski & Grogan, 1998: 262 [Fossil in Dominican amber]
Subfamily FORCIPOMIINAE
Atrichopogon Kieffer, 1906
Atrichopogon (Atrichopogon) dominicanus† Szadziewski & Grogan, 1998: 264 [Fossil in Dominican amber]
Atrichopogon sp.†—Szadziewski & Grogan (1994: 221) [Fossil in Dominican amber]
Atrichopogon (Psilokempia) woodruffi Spinelli, Marino & Huerta, 2015: 58 *DR > TL: Nr. Filipinas, Larimar mine
Forcipomyia Meigen, 1818
Forcipomyia (Forcipomyia) fusiparamera† Szadziewski & Grogan, 1998: 266 [Fossil in Dominican amber]
Forcipomyia (Forcipomyia) grimaldii† Szadziewski & Grogan, 1998: 268 [Fossil in Dominican amber]
Forcipomyia (Lasiohelea) americana† Szadziewski & Grogan, 1998: 270 [Fossil in Dominican amber]
Forcipomyia (Lasiohelea) woodruffi† Szadziewski & Grogan, 1998: 272 [Fossil in Dominican amber]
Forcipomyia (Lepidohelea) antilleana† Szadziewski & Grogan, 1998: 274 [Fossil in Dominican amber]
Forcipomyia (Lepidohelea) chrysosuccinea† Szadziewski & Grogan, 1998: 274 [Fossil in Dominican amber]
Forcipomyia (Lepidohelea) domibicolor† Szadziewski & Grogan, 1998: 276 [Fossil in Dominican amber]
Forcipomyia (Lepidohela) lepidosuccinea† Szadziewski & Grogan, 1998: 278 [Fossil in Dominican amber]
Forcipomyia (Microhelea) tettigonaris Wirth & Castner, 1990. Perez-Gelabert & Grogan, 1999: 312
Forcipomyia (Synthyridomyia) tertiaricola† Szadziewski & Grogan, 1998: 281 [Fossil in Dominican amber]
Forcipomyia (Thyridomyia) frutetosuccinea† Szadziewski & Grogan, 1998: 280 [Fossil in Dominican amber]
Forcipomyia (Trichohelea) sp.†—Szadziewski & Grogan (1998: 284) [Fossil in Dominican amber]
Subfamily LECTOCONOPINAE
Leptoconops Skuse, 1889
Leptoconops (Holoconops) bequaerti (Kieffer, 1925). Raccurt, 1984: 13
Note: Bezzia flinti Spinelli & Wirth (1989) was erroneously included as endemic to Hispaniola in the list of Perez-Gelabert
(2008: 149). This is a species from Dominica, Lesser Antilles.

Family CHIRONOMIDAE
Subfamily CHIRONOMINAE
Tribe CHIRONOMINI
Chironomus Meigen, 1803
Chironomus redeuns Walker, 1856. Curran, 1928: 13
Chironomus sp.—Silva et al. (2015: 14)
Dicrotendipes Kieffer, 1913
Dicrotendipes sp.†—Grund (2006a: 412) [Fossil in Dominican amber]
Einfeldia Kieffer, 1924
Einfeldia sp.—Silva et al. (2015: 14)
Goeldichironomus Fittkau, 1965

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Goeldichironomus sp. 1—Silva et al. (2015: 14)
Goeldichironomus sp. 2—Silva et al. (2015: 14)
Microchironomus Kieffer, 1926
Microchironomus sp.—Silva et al. (2015: 14)
Parachironomus Lenz, 1921
Parachironomus yanomami Spies, Fittkau & Reiss, 1994. Silva et al., 2015: 15
Paralauterborniella Lenz, 1941
Paralauterborniella nigrohalteralis (Malloch, 1915). Silva et al., 2015: 15
Polypedilum Kieffer, 1913
Polypedilum (Asheum) beckae (Sublette, 1964). Silva et al., 2015: 15
Polypedilum sp.†—Grund (2006a: 412) [Fossil in Dominican amber]
Polypedilum sp. 1—Silva et al. (2015: 15)
Polypedilum sp. 2—Silva et al. (2015: 15)
Polypedilum sp. 3—Silva et al. (2015: 15)
Polypedilum sp. 4—Silva et al. (2015: 15)
Polypedilum sp. 5—Silva et al. (2015: 15)
Stenochironomus Kieffer, 1919
Stenochironomus sp.†—Grund (2006a: 412) [Fossil in Dominican amber]
Stenochironomus sp. 1—Silva et al. (2015: 15)
Stenochironomus sp. 2—Silva et al. (2015: 15)
Xestochironomus Sublette & Wirth, 1972
Xestochironomus spp.†—Grund (2006a: 412) [Fossil in Dominican amber]
Xestochironomus sp.—Silva et al. (2015: 15)
Tribe PSEUDOCHIRONOMINI
Manoa Fittkau, 1963
Manoa pahayokeensis Jacobsen in Jacobsen & Perry, 2002. Silva et al., 2015: 15
Pseudochironomus Malloch, 1915
Pseudochironomus sp.—Silva et al. (2015: 15)
Tribe TANYTARSINI
Tanytarsus Van der Wulp, 1874
Tanytarsus excavatus group—Silva et al. (2015: 15)
Tanytarsus sp.—Silva et al. (2015: 15)
Subfamily ORTHOCLADIINAE
Antillocladius Saether, 1981
Antillocladius sp.—Silva et al. (2015: 15)
Compterosmittia Saether, 1981
Compterosmittia croizati Mendes, Andersen & Saether, 2004. Silva et al., 2015: 15
Compterosmittia sp. 1—Silva et al. (2015: 15)
Cricotopus Van der Wulp, 1874
Cricotopus sp. 1—Silva et al. (2015: 15)
Cricotopus sp. 2—Silva et al. (2015: 15)
Diplosmittia Saether, 1981
Diplosmittia caribensis Wiedenbrug & Silva, 2016: 71 *DR > TL: National Botanical Garden, Gran Cañada river
Lipurometriocnemus Saether, 1981
Lipurometriocnemus glabalus Saether, 1981. Silva et al., 2015: 15
Metriocnemus Van der Wulp, 1874
Metriocnemus sp.—Silva et al. (2015: 15)
Orthocladiinae sp. 1—Silva et al. (2015: 15)
Orthocladiinae sp. 2—Silva et al. (2015: 15)
Orthocladius Van der Wulp, 1874
Orthocladius sp.—Silva et al. (2015: 15)
Parametriocnemus Goetghebuer, 1932
Parametriocnemus sp.—Silva et al. (2015: 15)
Psectrocladius Kieffer, 1906

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Psectrocladius sp. 1—Silva et al. (2015: 15)
Pseudosmittia Goetghebuer, 1932
Pseudosmittia sp.—Silva et al. (2015: 16)
Subfamily TANYPODINAE
Tribe PENTANEURINI
Ablabesmyia Johannsen, 1905
Ablabesmyia electrohispaniolana† Grund, 2005: 32 [Fossil in Dominican amber]
Labrundinia Fittkau, 1962
Labrundinia sp. 1—Silva et al. (2015: 16)
Labrundinia sp. 2—Silva et al. (2015: 16)
Labrundinia sp. 3—Silva et al. (2015: 16)
Monopelopia Fittkau, 1962
Monopelopia sp.†—Grund (2006a: 412) [Fossil in Dominican amber]
Monopelopia sp.—Silva et al. (2015: 16)
Pentaneura Philippi, 1865
Pentaneura sp. 1—Silva et al. (2015: 16)
Pentaneura sp. 2—Silva et al. (2015: 16)
Thienemannimyia Fittkau, 1957
Thienemannimyia sp.—Silva et al. (2015: 16)
Tribe PROCLADIINI
Procladius Skuse, 1889
Procladius (Holotanypus) sp. 1—Silva et al. (2015: 16)
Tribe TANYPODINI
Tanypus Meigen, 1803
Tanypus sp.†—Grund (2006a: 412) [Fossil in Dominican amber]

Family CORETHRELLIDAE
Subfamily CORETHRELLINAE
Corethrella Coquillett, 1902
Corethrella appendiculata Grabham, 1906. Stone, 1966: 11.1; Belkin & Heinemann, 1972: 69
Corethrella (Corethrella) hispaniolensis Borkent, 2008: 165 *DR > TL: Rd. Constanza to Bonao, La Vega, 700 m
Corethrella (Corethrella) longituba Belkin, Heinemann & Page, 1970. Borkent, 2008: 171
Corethrella nudistyla† Borkent & Szadziewski, 1991: 460 [Fossil in Dominican amber]
Corethrella sp.—Belkin & Heinemann (1972: 70)
Sayomyia Coquillett, 1903
Sayomyia sp.—Belkin & Heinemann (1972: 70)
Sayomyia sp.†—Poinar & Poinar (1999: 207) [Misspelled Dayomyia] [Fossil in Dominican amber]

Family CULICIDAE
Generic classification follows the Mosquito Taxonomic Inventory (mosquito-taxonomic-inventory.info/).
Tribe AEDINI
Aedes Meigen, 1818
Aedes sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Gymnometopa Coquillett, 1905
Gymnometopa mediovittata (Coquillett, 1906). Dyar & Knab, 1906: 196; Belkin & Heinemann, 1972: 68
Howardina Theobald, 1903
Howardina albonotata (Coquillett, 1905: 183). Dyar & Knab, 1906: 196; Belkin & Heinemann, 1972: 68
Ochlerotatus Lynch Arribálzaga, 1891
Ochlerotatus pertinax Grabham, 1906. Belkin & Heinemann, 1972: 68
Ochlerotatus scapularis (Rondani, 1848). By synonymy of species below
=Aedes (Ochlerotatus) hemisurus (Dyar & Knab, 1906). Belkin & Heinemann, 1972: 67
Ochlerotatus sollicitans (Walker, 1856). Belkin & Heinemann, 1972: 68
Ochlerotatus taeniorhynchus (Weidemann, 1821). Belkin & Heinemann, 1972: 68
Ochlerotatus tortilis (Theobald, 1903). Belkin & Heinemann, 1972: 68

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Stegomyia Theobald, 1901
Stegomyia aegypti (Linnaeus, 1762). Wolcott, 1927: 32; Belkin & Heinemann, 1972: 68 [Introduced]
Stegomyia albopicta (Skuse, 1895). Peña, 1993: 4; Peña et al., 2003: 208 [Introduced]
Tribe ANOPHELINI
Anopheles Meigen, 1818
Anopheles (Anopheles) crucians Wiedemann, 1828. Belkin & Heinemann, 1972: 64
Anopheles (Anopheles) grabhami Theobald, 1901. Belkin & Heinemann, 1972: 64; Rodríguez-Sosa et al., 2019: 3
Anopheles (Anopheles) vestitipennis Dyar & Knab, 1906. Belkin & Heinemann, 1972: 64
Anopheles (Nyssorhynchus) albimanus Weidemann, 1820. Wolcott, 1927: 74; Belkin & Heinemann, 1972: 64; Rodríguez-Sosa
et al., 2019: 3
Anopheles (Nyssorhynchus) dominicanus† Zavortink & Poinar, 2000: 1231 [Fossil in Dominican amber]
Tribe CULICINI
Culex Linnaeus, 1758
Culex (Culex) bahamensis Dyar & Knab, 1906. Belkin & Heinemann, 1972: 64
Culex (Culex) duplicator Dyar & Knab, 1909: 258. Belkin & Heinemann, 1972: 65 *DR > TL: San Francisco Mountains, Santo
Domingo
Culex (Culex) garciai González Broche, 2000. Rueda et al., 2018: 96; Rodríguez-Sosa et al., 2019: 6
Culex (Culex) habilitator Dyar & Knab, 1906: 212. Belkin & Heinemann, 1972: 65 *DR > TL: small pool in a cave in coral
cliffs near the ocean in San Domingo
Culex (Culex) janitor Theobald, 1903. Belkin & Heinemann, 1972: 65
Culex (Culex) nigripalpus Theobald, 1901. Belkin & Heinemann, 1972: 65; Rodríguez-Sosa et al., 2019: 6 [Introduced]
=Culex carmodyae Dyar & Knab, 1906: 210
=Culex regulator Dyar & Knab, 1906: 213
Culex (Culex) quinquefasciatus Say, 1823. Belkin & Heinemann, 1972: 65; Rodríguez-Sosa et al., 2019: 8 [Introduced]
Culex (Culex) secutor Theobald, 1901. Belkin & Heinemann, 1972: 65; Rodríguez-Sosa et al., 2019: 10
=Culex lamentator Dyar & Knab, 1906: 219
Culex malariager† Poinar, 2005: 549 [Fossil in Dominican amber]
Culex (Melanoconion) atratus Theobald, 1901. Belkin & Heinemann, 1972: 66; Rodríguez-Sosa et al., 2019: 4
Culex (Melanoconion) carcinophilus Dyar & Knab, 1906: 220. Belkin & Heinemann, 1972: 66 *DR > TL: Crab holes
containing fresh water near Santo Domingo city
Culex (Melanoconion) erraticus (Dyar & Knab, 1906). Belkin & Heinemann, 1972: 66
Culex (Melanoconion) inhibitator Dyar & Knab, 1906: 216. Belkin & Heinemann, 1972: 66 *DR > TL: San Francisco Mts. of
San Domingo
Culex (Melanoconion) opisthopus Komp, 1927. Belkin & Heinemann, 1972: 66
Culex (Melanoconion) pilosus (Dyar & Knab, 1906). Belkin & Heinemann, 1972: 66
Culex (Micraedes) antillummagnorum Dyar, 1928. Belkin & Heinemann, 1972: 66
Culex (Micraedes) biscaynensis Zavortink & O’Meara, 1999. Rodríguez-Sosa et al., 2019: 4
Culex (Phenacomyia) corniger Theobald, 1903. Belkin & Heinemann, 1972: 65; Rodríguez-Sosa et al., 2019: 6
Tribe MANSONINI
Mansonia Blanchard, 1901
Mansonia (Mansonia) dyari Belkin et al., 1970. Peña & Chadee, 2004: 449
Mansonia (Mansonia) flaveola (Coquillett, 1906). Peña & Chadee, 2004: 449
Mansonia (Mansonia) titillans (Walker, 1948). Belkin & Heinemann, 1972: 67
Tribe ORTHOPODOMYINI
Orthopodomyia Theobald, 1904
Orthopodomyia signifera (Coquillett, 1896). Belkin & Heinemann, 1973: 219
Orthopodomyia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Tribe SABETHINI
Aedeomyia Theobald, 1801
Aedeomyia squamipennis Lynch-Arribálzaga, 1878. Peña & Chadee, 2004: 449
Culiseta Felt, 1904
Culiseta sp.†—Grimaldi (1993: 165) [Fossil in Dominican amber]
Deinocerites Theobald, 1901
Deinocerites cancer Theobald, 1901. Belkin & Heinemann, 1972: 67

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Limatus Theobald, 1901
Limatus hoffmani Root, 1927. Belkin & Heinemann, 1972: 69
Lutzomiops Lane, 1942
Lutzomiops? sp.—Belkin & Heinemann (1975: 381)
Psorophora Robineau-Desvoidy, 1827
Psorophora (Grabhamia) infinis (Dyar & Knab, 1906: 182). Belkin & Heinemann, 1972: 67
Psorophora (Grabhamia) insularia (Dyar & Knab, 1906). Belkin & Heinemann, 1972: 67
Psorophora (Grabhamia) jamaicensis (Theobald, 1901). Dyar & Knab, 1906: 183; Belkin & Heinemann, 1972: 67
Psorophora (Grabhamia) pygmaea (Theobald, 1903). Dyar & Knab, 1906: 182; Belkin & Heinemann, 1972: 67
Psorophora (Janthinosoma) ferox (Humboldt, 1819). Belkin & Heinemann, 1972: 67
Psorophora (Janthinosoma) johnstonii (Grabham, 1905). Belkin & Heinemann, 1972: 67
Sabethes Robineau-Desvoidy, 1827
Sabethes (Sabethes) bipartites Dyar & Knab, 1906. Belkin & Heinemann, 1972: 69
Toxorhynchites Theobald, 1901
Toxorhynchites (Lynchiella) portoricensis (von Röder, 1885). Belkin & Heinemann, 1972: 69; Rodríguez-Sosa et al., 2019: 13
Wyeomyia Theobald, 1901
Wyeomyia (Wyeomyia) mitchellii (Theobald, 1905). Belkin & Heinemann, 1972: 69; Rodríguez-Sosa et al., 2019: 13
=Wyeomyia abia Dyar & Knab, 1908. Belkin & Heinemann, 1972: 69
=Wyeomyia ochrura Dyar & Knab, 1906. Belkin & Heinemann, 1972: 69
=Wyeomyia glaucocephala Dyar & Knab, 1906. Belkin & Heinemann, 1972: 69
Wyeomyia (Wyeomyia) nigritubus Galindo et al., 1951. Belkin & Heinemann, 1972: 69
Wyeomyia (Wyeomyia) sororcula Dyar & Knab, 1906. Belkin & Heinemann, 1972: 69
Wyeomyia (Wyeomyia) sp. 1—Belkin & Heinemann (1972: 69)
Wyeomyia (Wyeomyia) sp. 2—Belkin & Heinemann (1972: 69)
Tribe URANOTAENINII
Uranotaenia Lynch & Arribálzaga, 1891
Uranotaenia cooki Root, 1937. Belkin & Heinemann, 1972: 64. Rodríguez-Sosa et al., 2018: 141; Rodríguez-Sosa et al., 2019:
13
Uranotaenia lowii Theobald, 1901. Belkin & Heinemann, 1972: 64
Uranotaenia socialis Theobald, 1901. Belkin & Heinemann, 1972: 64

Family DIXIDAE
Dixella Dyar & Shannon, 1924
Dixella sp.—Belkin & Heinemann (1972: 70)
Mesodixa Belkin et al., 1970
Mesodixa sp.—Belkin & Heinemann (1972: 70)

Family SIMULIIDAE
Simulium Latreille, 1902
Simulium (Ectemnaspis) ochraceum Walker, 1861. Coscarón & Coscarón Arias, 2007: 416
Simulium (Psilopelmia) bipunctatum Malloch, 1912. Crosskey & Howard, 1997: 58

Infraorder BIBIONOMORPHA
Family ANISOPODIDAE
Mesochria Enderlein, 1910
Mesochria neotropica† Grimaldi, 1991: 21 [Fossil in Dominican amber]
Mycetobia Meigen, 1818
Mycetobia antillea† Grimaldi, 1991: 13. Amorim et al., 2016: 17 [Fossil in Dominican amber]
Mycetobia cryptambra† Grimaldi, 1991: 17. Amorim et al., 2016: 17 [Fossil in Dominican amber]
Mycetobia sp.†—Grimaldi (1991: 20) [Fossil in Dominican amber]
Olbiogaster Osten Sacken, 1886
Olbiogaster perezi† Grimaldi & Amorim, 1995: 562 [Fossil in Dominican amber]

Family BIBIONIDAE

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Subfamily PLECIINAE
Dilophus Meigen, 1803
Dilophus matilei† Waller et al., 2000: 150 [Fossil in Dominican amber]
Plecia Wiedemann, 1828
Plecia fasciapenna Fitzgerald, 1998: 146
Plecia quasimaculata Hardy, 1961: 90 *H > TL: Kenscoff
Plecia pristina† Hardy, 1971. Skartveit & Bechly, 2013: 99 [Fossil in Dominican amber]
Plecia sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]

Family CECIDOMYIIDAE
Subfamily CECIDOMYIINAE
Asphondylia Loew, 1850
Asphondylia websteri Felt, 1917. Gagné, 1994: 127
Contarinia Rondani, 1860
Contarinia maculipennis Felt, 1933. Serra et al., 2003: 108 [Introduced]
Contarinia sp.†—Gagné (1994: 146) [Fossil in Dominican amber]
Diadiplosis Felt, 1911
Diadiplosis coccidarum Cockerell, 1892. Gagné, 1994: 184
Diadiplosis multifila (Felt, 1907). Gagné, 1994: 185
Diadiplosis pulvinariae (Felt, 1912). Gagné, 1994: 185
Feltiella Rübsaamen, 1910
Feltiella insularis (Felt, 1913). Etienne & Gagné, 2017: 455
Neolasioptera Felt, 1905
Neolasioptera portulacae (Cook, 1906). Etienne & Gagné, 2017: 455
Neolasioptera rostrata Gagné, 1989: 170. Gagné, 1994: 75
Stenodiplosis Reuter, 1895
Stenodiplosis sorghicola (Coquillett, 1899). Gagné, 1968: 23.29; SEA, 1999: 99
Xenasphondylia Felt, 1915
Xenasphondylia albipes Felt, 1915: 202. Callan, 1940: 755; Gagné, 1994: 198 *DR > TL: San Francisco Mountains, Santo
Domingo

Family KEROPLATIDAE
Keroplatus Bosc, 1792
Keroplatus sp.†—Poinar & Poinar (1999: 207) [Misspelled Keraplatus] [Fossil in Dominican amber]
Orfelia Costa, 1857
Orfelia sp.†—Peñalver (1995: 59) [Fossil in Dominican amber]
Proceroplatus Edwards, 1925
Proceroplatus hennigi† Schmalfuss, 1979: 4 [Fossil in Dominican amber]
Proceroplatus pictipennis Williston, 1896. Penney, Evenhuis & Green, 2013: 598
Proceroplatus preziosii† Evenhuis and Penney, 2013: 595 [Fossil in Dominican amber]

Family LYGISTORRHINIDAE
Lygistorrhina Skuse, 1890
Lygistorrhina caribbiana† Grund, 2012: 640 [Fossil in Dominican amber]

Family MYCETOPHILIDAE
Allodia Winnertz, 1863
Allodia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Aphrastomyia Coher & Lane, 1949
Aphrastomyia planistylus† Baxter & Poinar, 1994: 100 [Fossil in Dominican amber]
Boletina Staeger, 1840
Boletina sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Coelosia Winnertz, 1863
Coelosia sp.†—Poinar & Poinar (1999: 207) [Misspelled Celosia] [Fossil in Dominican amber]

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Cordyla Meigen, 1803
Cordyla sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Euceroplatus Edwards, 1829
Euceroplatus sp.†—Grimaldi & Mathis (1993: 401) [Fossil in Dominican amber]
Exechia Winnertz, 1863
Exechia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Exechiopsis Tuomikoski, 1966
Exechiopsis sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Leia Gimmerthal, 1832
Leia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Macrocera Meigen, 1803
Macrocera sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Megophthalmidia Dziedzicki, 1889
Megophthalmidia sp.†—Poinar & Poinar (1999: 207) [Misspelled Megophthalmedia] [Fossil in Dominican amber]
Metaleia Baxter, 1994
Metaleia maculicauda† Baxter & Poinar, 1994: 97) [Fossil in Dominican amber]
Mycetophila Meigen, 1803
Mycetophila sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Mycomya Rondani, 1856
Mycomya sp.†—Poinar & Poinar (1999: 207) [Misspelled Mycomyia] [Fossil in Dominican amber]
Orthellia Robineau-Desvoidy, 1863
Orthellia spp.†—Poinar et al. (1994: 94) [Fossil in Dominican amber]
Phronia Winnertz, 1863
Phronia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Platyura Meigen, 1803
Platyura sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Rymosia Winnertz, 1863
Rymosia sp.†—Poinar & Poinar (1999: 207) [Misspelled Rhimozia] [Fossil in Dominican amber]
Sciophila Meigen, 1818
Sciophila sp.—Poinar & Poinar (1999: 207) [Misspelled Sciophita] [Fossil in Dominican amber]
Synapha Meigen, 1818
Synapha sp.†—Poinar & Poinar (1999: 207) [Misspelled Synaphe] [Fossil in Dominican amber]
Tetragoneura Winnertz, 1846
Tetragoneura brevistipitis† Baxter & Poinar, 1994: 94 [Fossil in Dominican amber]
Trichonta Winnertz, 1863
Trichonta sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]

Family SCIARIDAE
Archicratyna† Mohrig, 2005
Archicratyna arcana† Mohrig & Röschmann, 2005: 331 [Fossil in Dominican amber]
Archicratyna subarcana† Mohrig & Röschmann, 2005: 333 [Fossil in Dominican amber]
Archicratyna trichoarcana† Mohrig & Röschmann, 2005: 332 [Fossil in Dominican amber]
Archicratyna zimbabweensis† Mohrig, 2005: 330 [Fossil in Dominican amber]
Bradysia Winnertz, 1867
Bradysia acerba Mohrig & Röschmann in Mohrig et al., 2004: 291 *DR > TL: Duarte Prov., 13 km N San Francisco de
Macorís, 800 ft.
Bradysia antiqua† Mohrig & Röschmann, 2005: 337 [Fossil in Dominican amber]
Bradysia cauta Mohrig & Röschmann in Mohrig et al., 2004: 296 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Bradysia ciliocera Mohrig & Röschmann in Mohrig et al., 2004: 290 *DR > TL: La Vega Prov., 12 km S Constanza, rd. to San
José de Ocoa, 5000 ft.
Bradysia clara Mohrig & Röschmann in Mohrig et al., 2004: 288 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Bradysia crassicera Mohrig & Röschmann in Mohrig et al., 2004: 293 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las

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Abejas, 1500 m
Bradysia dimidiata† Mohrig & Röschmann, 2005: 339 [Fossil in Dominican amber]
Bradysia exoleta† Mohrig & Röschmann, 2005: 338 [Fossil in Dominican amber]
Bradysia longioculosa Mohrig & Röschmann in Mohrig et al., 2004: 296 *DR > TL: Pedernales Prov., 20 km NE Pedernales,
Las Abejas, 1500 m
Bradysia mellea Mohrig & Röschmann in Mohrig et al., 2004: 294 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Bradysia obtusa Mohrig & Röschmann in Mohrig et al., 2004: 292 *DR > TL: La Vega Prov., 12 km S Constanza, rd. to San
José de Ocoa, 5000 ft.
Bradysia oculosa Mohrig & Röschmann in Mohrig et al., 2004: 295 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Bradysia pararufescens Mohrig & Röschmann in Mohrig et al., 2004: 307 *DR > TL: Duarte Prov., 13 km N San Francisco de
Macorís, 800 ft.
Bradysia reticulata Mohrig & Röschmann in Mohrig et al., 2004: 289 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Bradysia spinea Mohrig & Röschmann in Mohrig et al., 2004: 289 *DR > TL: Duarte Prov., 13 km N San Francisco de
Macorís, 800 ft.
Bradysia subspinea Mohrig & Röschmann in Mohrig et al., 2004: 290 *DR > TL: Barahona Prov., just S La Ciénaga, 100 ft.
Bradysia turgida Mohrig & Röschmann in Mohrig et al., 2004: 287 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Bradysiopsis Tuomikoski, 1960
Bradysiopsis repentina† Mohrig & Röschmann, 2005: 335 [Fossil in Dominican amber]
Chaetosciara Frey, 1942
Chaetosciara gilva Mohrig & Röschmann in Mohrig et al., 2004: 274 *DR > TL: Duarte Prov., 13 km N San Francisco de
Macorís, 800 ft.
Chaetosciara obsoleta† Mohrig & Röschmann, 2005: 334 [Fossil in Dominican amber]
Corynoptera Winnertz, 1867
Corynoptera iocosa† Mohrig & Röschmann, 2005: 343 [Fossil in Dominican amber]
Corynoptera prisca† Mohrig & Röschmann, 2005: 342 [Fossil in Dominican amber]
Corynoptera pristina† Mohrig & Röschmann, 2005: 341 [Fossil in Dominican amber]
Corynoptera sicca Mohrig & Röschmann in Mohrig et al., 2004: 302 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Corynoptera turbata Mohrig & Röschmann in Mohrig et al., 2004: 301 *DR > TL: Pedernales Prov., 20 km NE Pedernales,
Las Abejas, 1500 m
Cratyna Winnertz, 1867
Cratyna (Cratyna) fumoalata Mohrig & Röschmann in Mohrig et al., 2004: 275 *DR > TL: Duarte Prov., 13 km N San
Francisco de Macorís, 800 ft.
Cratyna (Cratyna) tempestiva† Mohrig & Röschmann, 2005: 326 [Fossil in Dominican amber]
Cratyna interposita† Mohrig & Röschmann, 2005: 328 [Fossil in Dominican amber]
Cratyna villosoantennata† Mohrig & Röschmann, 2005: 327 [Fossil in Dominican amber]
Epidapus Haliday, 1852
Epidapus adstrictosetus† Mohrig & Röschmann, 2005: 346 [Fossil in Dominican amber]
Epidapus (Clandestina) conciliatus Mohrig, 2003. Mohrig & Röschmann, 2004: 305
Epidapus (Epidapus) flavothoracicus Mohrig & Röschmann in Mohrig et al., 2004: 305 *DR > TL: Duarte Prov., 13 km N San
Francisco de Macorís, 800 ft.
Epidapus longisetus† Mohrig & Röschmann, 2005: 347 [Fossil in Dominican amber]
Epidapus macrospinatus† Mohrig & Röschmann, 2005: 345 [Fossil in Dominican amber]
Epidapus microspinus† Mohrig & Röschmann, 2005: 344 [Fossil in Dominican amber]
Epidapus (Pseudoaptanogyna) obstinatus Mohrig & Röschmann in Mohrig et al., 2004: 306 *DR > TL: Pedernales Prov., 20
km NE Pedernales, Las Abejas, 1500 m
Hyperlasion Schmitz, 1918
Hyperlasion multisetus Mohrig & Röschmann in Mohrig et al., 2004: 276 *DR > TL: Duarte Prov., 13 km N San Francisco de
Macorís, 800 ft.
Hyperlasion wasmanni Schmitz, 1919. Mohrig & Röschmann in Mohrig et al., 2004: 276

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Keilbachia Mohrig, 1987
Keilbachia paucipalpa Mohrig & Röschmann in Mohrig et al., 2004: 303 *DR > TL: Duarte Prov., 13 km N San Francisco de
Macorís, 800 ft.
Leptosciarella Tuomikoski, 1960
Leptosciarella (Leptosciarella) ampullocera Mohrig & Röschmann in Mohrig et al., 2004: 270 *DR > TL: Pedernales Prov., 20
km NE Pedernales, Las Abejas, 1500 m
Leptosciarella (Leptosciarella) manifesta† Mohrig & Röschmann, 2005: 324 [Fossil in Dominican amber]
Leptosciarella (Leptosciarella) nudinervosa Mohrig & Röschmann in Mohrig et al., 2004: 269 *DR > TL: Duarte Prov., 13 km
N San Francisco de Macoris
Pseudolycoriella Menzel & Mohrig, 1998
Pseudolycoriella barbata Mohrig & Rulik in Mohrig et al., 2004: 279 *DR > TL: Duarte Prov., 13 km N San Francisco de
Macorís, 800 ft.
Pseudolycoriella bicornis (Menzel, 1997). Mohrig & Röschmann, 2004: 278
Pseudolycoriella curvimedia Mohrig & Rulik in Mohrig et al., 2004: 286 *DR > TL: Pedernales Prov., 20 km NE Pedernales,
Las Abejas, 1500 m
Pseudolycoriella florentissima Mohrig & Rulik in Mohrig et al., 2004: 285 *DR > TL: Pedernales Prov., 20 km NE Pedernales,
Las Abejas, 1500 m
Pseudolycoriella fuscivenosa Mohrig & Rulik in Mohrig et al., 2004: 284 *DR > TL: Pedernales Prov., 20 km NE Pedernales,
Las Abejas, 1500 m
Pseudolycoriella indocera Mohrig & Rulik in Mohrig et al., 2004: 283 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Pseudolycoriella parilis (Johannsen, 1912). Mohrig et al., 2013: 227
Pseudolycoriella pulla Mohrig & Rulik in Mohrig et al., 2004: 278 *DR > TL: La Vega Prov., 12 km S Constanza, rd. to San
José de Ocoa, 5000 ft.
Pseudolycoriella rotundostyla Mohrig & Rulik in Mohrig et al., 2004: 282 *DR > TL: Monte Plata, 9 km N Bayaguana, 300 ft.
Pseudolycoriella torva Mohrig & Rulik in Mohrig et al., 2004: 277 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Pseudolycoriella virgata Mohrig & Rulik in Mohrig et al., 2004: 282 *DR > TL: Pedernales Prov., 20 km NE Pedernales, Las
Abejas, 1500 m
Pseudosciara Schiner, 1866
Pseudosciara aitkeni Lane, 1959. Mohrig et al., 2004: 271
Pseudosciara cariba Lane, 1959. Mohrig et al., 2004: 271
Pseudosciara fragistyla Mohrig & Röschmann in Mohrig et al., 2004: 273 *DR > TL: La Romana, sea level
Pseudosciara longicera Mohrig & Röschmann in Mohrig et al., 2004: 272 *DR > TL: Pedernales Prov., 20 km NE Pedernales,
Las Abejas, 1500 m
Scaptosciara Tonnor & Edwards,1927
Scaptosciara (Scatopsciara) ventosa Mohrig & Röschmann in Mohrig et al., 2004: 297 *DR > TL: La Vega Prov., 12 km S
Constanza, rd. to San José de Ocoa, 5000 ft.
Scaptosciara (Xenopygina) abiecta Mohrig & Röschmann in Mohrig et al., 2004: 299 *DR > TL: Peravia Prov., 30 km N San
José de Ocoa, 3000 ft.
Scaptosciara (Xenopygina) caribiana Mohrig & Röschmann in Mohrig et al., 2004: 298 *DR > TL: La Vega Prov., 21 km W
Bonao, 3400 ft.
Scaptosciara (Xenopygina) destituta Mohrig & Röschmann in Mohrig et al., 2004: 300 *DR > TL: La Romana, sea level
Scaptosciara (Xenopygina) terribilis Mohrig & Röschmann in Mohrig et al., 2004: 300 *DR > TL: Pedernales Prov., 20 km NE
Pedernales, Las Abejas, 1500 m
Trichosia Winnertz, 1867
Trichosia (Mouffetina) nova† Mohrig & Röschmann, 2005: 322 [Fossil in Dominican amber]

Suborder BRACHYCERA
Infraorder STRATIOMYOMORPHA
Family STRATIOMYIDAE
Key reference: Woodley (2001).
Subfamily CLITELLARIINAE
Cyphomyia Wiedemann, 1819

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Cyphomyia albomaculata (Macquart, 1838). Woodley, 2001: 158
Cyphomyia baoruco Woodley, 2014: 112 *DR > TL: Pedernales Prov., Parque Nacional Sierra de Bahoruco, Las Abejas, 1150
m
Cyphomyia chalybea (Wiedemann, 1824). Wolcott, 1927: 325; Smith & Audant, 1930: 978; Woodley, 2001: 159
Cyphomyia marginata Loew, 1866. Wolcott, 1927: 318; Curran, 1928: 17; James, 1940: 131; James, 1972: 146; Woodley,
2001: 161
Subfamily HERMETINAE
Hermetia Latreille, 1804
Hermetia illucens (Linnaeus, 1758). Wolcott, 1927: 317; James, 1972: 145; Woodley, 2001: 182
Hermetia sexmaculata Macquart, 1834. Woodley, 2001: 184
Subfamily NEMOTELINAE
Brachychara Sharp, 1883
Brachychara maculata (James, 1953). Woodley, 2001: 308
Subfamily PACHYGASTRINAE
Brachyodina Lindner, 1949
Brachyodina janestanleyae Woodley, 2014: 430 *DR > TL: Punta Cana near Ecological Reserve
Pachygaster Germar, 1817
Pachygaster hymenaea† Grund & Hauser, 2005: 30 [Fossil in Dominican amber]
Subfamily SARGINAE
Microchrysa Loew, 1855
Microchrysa bicolor (Wiedemann, 1830). James, 1972: 147; Woodley, 2001: 203
Ptecticus Loew, 1855
Ptecticus insularis James, 1972: 147. Woodley, 2001: 212 *DR > TL: Santo Domingo
Sargus Fabricius, 1798
Sargus fasciatus Fabricius, 1805. Woodley, 2001: 223
Sargus pleuriticus Loew, 1866. Woodley, 2001: 228
Subfamily STRATIOMYINAE
Hedriodiscus Enderlein, 1914
Hedriodiscus dorsalis (Fabricius, 1805). Woodley, 2001: 263
Hedriodiscus trivittatus (Say, 1829). James, 1972: 147; Woodley, 2001: 265
Nothomyia Loew, 1869
Nothomyia nigra James, 1942: 55. Woodley, 2001: 258 *DR > TL: Valle Nuevo, S.E. Constanza, 7000 ft.
Nothomyia sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]

Infraorder TABANOMORPHA
Family ACROCERIDAE
Ogcodes Latreille, 1796
Ogcodes exotica† Grimaldi, 1995: 2 [Fossil in Dominican amber]
Turbopsebius Schlinger, 1972
Turbopsebius brunnipennis (Sabrosky, 1948: 397). Schlinger, 1972: 425; Perez-Gelabert, 2011: 123 *H, *DR > TL: Furcy

Family RHAGIONIDAE
Chrysopilus Loew, 1840
Chrysopilus sp.†—Wu (1997: 516) [Fossil in Dominican amber]

Family TABANIDAE
Key reference: Fairchild and Burger (1994).
Subfamily CHRYSOPINAE
Tribe CHRYSOPINI
Chrysops Meigen, 1800
Chrysops frazari Williston, 1887. Bequaert, 1940b: 282; Fairchild, 1980: 171; Fairchild & Burger, 1994: 57
Chrysops variegatus (De Geer, 1776). Bequaert, 1940b: 276; Fairchild, 1980: 171
Subfamily TABANINAE
Tribe DIACHLORINI

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Chlorotabanus Lutz, 1909
Chlorotabanus inanis (Fabricius, 1787). Bequaert, 1940b: 318
=Tabanus sulphureus Palisot de Beauvois, 1819. Fairchild & Burger, 1994: 87
Lepiselaga Macquart, 1838
Lepiselaga crassipes (Fabricius, 1805: 108). Bequaert, 1940b: 285; Fairchild, 1980: 173; Fairchild & Burger, 1994: 125
Stenotabanus Lutz, 1913
Stenotabanus (Aegialomyia) barahona Fairchild, 1980: 173. Fairchild & Burger, 1994: 77 *DR > TL: Barahona, Barahona
Agricultural Experimental Station
Stenotabanus (Aegialomyia) jamaicensis (Newstead, 1909). Fairchild, 1980: 173; Fairchild & Burger, 1994: 77
Stenotabanus (Aegialomyia) picticornis (Bigot, 1892). Bequaert, 1940b: 344; Fairchild & Burger, 1994: 78 *H > TL: Haiti
Stenotabanus (Stenotabanus) alticolus Fairchild, 1980: 177. Fairchild & Burger, 1994: 72 *DR > TL: Constanza
Stenotabanus (Stenotabanus) batesi (Bequaert, 1940b: 321). Fairchild, 1980: 177. Fairchild & Burger, 1994: 72 *H, *DR > TL:
La Selle Range, La Visite
Stenotabanus (Stenotabanus) brodzinskyi† Lane et al., 1988: 594. Fairchild & Burger, 1994: 73 [Fossil in Dominican amber]
Stenotabanus (Stenotabanus) brunettii (Bequaert, 1940b: 319). Fairchild & Burger, 1994: 73
Stenotabanus (Stenotabanus) fenestra (Williston, 1887). Bequaert, 1940b: 336; Fairchild, 1980: 181; Fairchild & Burger, 1994:
74
Stenotabanus (Stenotabanus) hispaniolae (Bequaert, 1940b: 341). Fairchild, 1980: 182; Fairchild & Burger, 1994: 74 *DR >
TL: Mt. Quita Espuela
Stenotabanus (Stenotabanus) marcanoi Fairchild, 1980: 180. Fairchild & Burger, 1994: 74 *DR > TL: Río Cana, between
Rancho de la Guardia and Hondo Valle, Elías Piña Prov.
Stenotabanus (Stenotabanus) parvulus (Williston, 1887). Bequaert, 1940b: 343; Fairchild, 1980: 177; Fairchild & Burger,
1994: 75
Stenotabanus (Stenotabanus) woodruffi† Fairchild & Lane, 1989: 630. Fairchild & Burger, 1994: 76 [Fossil in Dominican
amber]
Tribe TABANINI
Tabanus Linnaeus, 1758
Tabanus claripennis Bigot, 1892. Fairchild, 1980: 186
Tabanus clenchi Bequaert, 1940b: 314. Fairchild, 1980: 183; Fairchild & Burger, 1994: 134 *DR, *H > TL: Barahona
Tabanus commixtus Walker, 1860. Fairchild, 1980: 187; Fairchild & Burger, 1994: 134
Tabanus darlingtoni Bequaert, 1940b: 302. Fairchild, 1980: 184; Fairchild & Burger, 1994: 135 *DR > TL: Valle Nuevo,
southeast of Constanza, about 7,000 ft.
Tabanus flocculus Bequaert, 1940b: 304. Fairchild, 1980: 185; Fairchild & Burger, 1994: 137 *DR
Tabanus gymnorhynchus Fairchild, 1980: 182. Fairchild & Burger, 1994: 139 *DR > TL: Prov. Estrelleta, Higüerito - Bánica
Tabanus haitiensis Krober, 1931: 297. Fairchild, 1980: 186; Fairchild & Burger, 1994: 139 *H > TL: Port-au-Prince
Tabanus monops Bequaert, 1940b: 307. Fairchild, 1980: 184; Fairchild & Burger, 1994: 141 *DR > TL: Valle Nuevo, southeast
of Constanza, about 7,000 ft.
Tabanus vittiger guatemalanus Hine, 1906. Fairchild, 1980: 187
Note: The record of Tabanus dominicanus Krober, 1931: 301 from “S. Domingos” is considered by Stone (1970) to be “an
obvious error”. This species is from Dominica, Lesser Antilles.

Superfamily ASILOIDEA
Family ASILIDAE
Key references: Scarbrough & Perez-Gelabert (2005, 2006).
Subfamily APOCLINAE
Efferia Coquillett, 1893
Efferia alia Scarbrough & Perez-Gelabert, 2009: 8 *DR > TL: Pedernales Prov., 25 km N of Cabo Rojo, 679 m
Efferia angusta Scarbrough & Perez-Gelabert, 2009: 11 *DR > TL: Azua Prov., La Furnia, Barreras
Efferia bullata Scarbrough & Perez-Gelabert, 2009: 13 *DR > TL: Parque Nacional Jaragua, trail to Carlitos, ca. 5 km S of
Highway 44, 106 m
Efferia clava Scarbrough & Perez-Gelabert, 2009: 15 *DR > TL: Azua Prov., Monte Río, ~4.5 km W of beach
Efferia exacta Scarbrough & Perez-Gelabert, 2009: 18 *DR,*H > TL: Santiago Prov., entrance to Sabana Iglesia, 372 m
Efferia fortis (Walker, 1855b: 623). Scarbrough & Perez-Gelabert, 2006: 7; Scarbrough & Perez-Gelabert, 2009: 20 *DR, *H >
TL: St. Domingo

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Efferia fulvibarbis (Macquart, 1848: 188). Hine, 1919: 129; Scarbrough & Perez-Gelabert, 2006: 8; Scarbrough & Perez-
Gelabert, 2009: 23 *DR, *H > TL: Haïti
=Efferia pachychaeta (Bromley, 1928: 1). Scarbrough & Perez-Gelabert, 2006: 8; Scarbrough & Perez-Gelabert, 2009: 23
Efferia haitensis (Macquart, 1848: 188). Scarbrough & Perez-Gelabert, 2009: 26; Scarbrough & Perez-Gelabert, 2009: 26 *DR,
*H > TL: Haïti
Efferia incisura Scarbrough & Perez-Gelabert, 2009: 29 *DR > TL: Isla Beata, trail to Cueva del Tubo
Efferia picea Scarbrough & Perez-Gelabert, 2009: 31 *DR > TL: La Vega Prov., Parque Nacional Armando Bermúdez, trail La
Ciénaga de Manabao-Los Tablones
Efferia serrula Scarbrough & Perez-Gelabert, 2009: 33 *DR > TL: Santiago Prov., Babosico, on way to Jánico, 515 m
Efferia sinuosa Scarbrough & Perez-Gelabert, 2009: 35 *DR > TL: Peravia Prov., El Maizal, near Presa de Valdesia, 97 m
Efferia spinula Scarbrough & Perez-Gelabert, 2009: 38 *DR > TL: Terrenos Zona Franca, E. of Duvergé, 12 m
Efferia suspiciosa Scarbrough & Perez-Gelabert, 2009: 40 *DR > TL: La Vega Prov., PN Armando Bermúdez, km 1–3 trail W
of La Ciénaga, 900–1100 m
Efferia woodleyi Scarbrough & Perez-Gelabert, 2009: 42 *DR > TL: Independencia Prov., PN Sierra de Bahoruco, Caseta 1, S.
of Puerto Escondido, 1215 m
Proctacanthella Bromley, 1934
Proctacanthella taina Scarbrough & Perez-Gelabert, 2006: 12 *DR > TL: Montecristi Prov., 4 km N of Botoncillo
Proctacanthus Macquart, 1838
Proctacanthus darlingtoni Curran, 1951: 6 *DR > TL: Barahona
Proctacanthus dominicana Curran, 1951: 9 *DR > TL: Sánchez
Proctacanthus vittatus (Olivier, 1789: 263). Scarbrough & Perez-Gelabert, 2006: 8 *DR, *H > TL: San Domingo
Wilcoxia James, 1941
Wilcoxia sp.†—Poinar & Poinar (1999: 206). [This could be a misspelling of Wilcoxius as Wilcoxia is a Nearctic genus] [Fossil
in Dominican amber]
Wilcoxius Martin, 1975
Wilcoxius planus Scarbrough & Perez-Gelabert, 2005: 45 *DR > TL: ~2 km road to San Juan, near bridge on Viajama river,
Azua Prov., 294 m
Wilcoxius tumidus Scarbrough & Perez-Gelabert, 2005: 47 *DR > TL: 2 km road Balneario Ingenito, Presa de Sabaneta, San
Juan Prov.
Subfamily LAPHRIINAE
Andrenosoma Rondani, 1856
Andrenosoma chalybeum Williston, 1885: 56. Scarbrough & Perez-Gelabert, 2006: 17
Andrenosoma ruficaudum (Williston, 1885: 55). Scarbrough & Perez-Gelabert, 2006: 18
*H, *DR > TL: San Domingo
Atomosia Macquart, 1838
Atomosia anacaona Scarbrough & Perez-Gelabert, 2006: 22 *DR > TL: Santiago Prov., Babosico, on way to Jánico, 515 m
Atomosia ciguaya Scarbrough & Perez-Gelabert, 2006: 25 *DR > TL: San Juan Prov., Road San Juan—Vallejuelo, near river,
449 m
Atomosia jagua Scarbrough & Perez-Gelabert, 2006: 28 *DR > TL: Independencia Prov., Caseta 1, Parque Nacional Sierra de
Bahoruco, 1,239 m
Atomosia jimagua Scarbrough & Perez-Gelabert, 2006: 31 *DR > TL: Parque Nacional Jaragua, trail to Carlitos, ca. 6 km S
Highway 44, 106 m
Atomosia yurabia Scarbrough & Perez-Gelabert, 2006: 33 *DR > TL: San Juan Prov., few km E. Vallejuelo, 704 m
Atoniomyia Hermann, 1912
Atoniomyia mikii (Williston, 1886: 290). Scarbrough & Perez-Gelabert, 2006: 36
*DR > TL: Santo Domingo
Atoniomyia sp.—Scarbrough & Perez-Gelabert (2006: 38)
Cerotainia Schiner, 1868
Cerotainia sola Scarbrough & Perez-Gelabert, 2006: 39
Dasyllis Loew, 1851
Dasyllis rufa (Bromley, 1931: 132). Scarbrough & Perez-Gelabert, 2006: 20 *H, *DR > TL: Santo Domingo
Eumecosoma Schiner, 1866
Eumecosoma caerulum Scarbrough & Perez-Gelabert, 2006: 42 *DR > TL: La Vega Prov., Parque Nacional Armando
Bermúdez, km 1–3 along trail W of Ciénaga, 900–1100 m

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Pilica Curran, 1931
Pilica sp.—Scarbrough & Perez-Gelabert (2006: 21)
Rhopalogaster Macquart, 1834
Rhopalogaster albidus Scarbrough & Perez-Gelabert, 2006: 45 *DR > TL: La Vega Prov., Parque Nacional Armando
Bermúdez, km 1–3 along trail W of Ciénaga, 900–1100 m
Subfamily LEPTOGASTRINAE
Key reference: Scarbrough and Perez-Gelabert (2005).
Apachekolos Martin, 1957
Apachekolos clavipes (Johnson, 1897). Scarbrough, Perez-Gelabert & Page, 2005: 32
Apachekolos flaventis Scarbrough & Perez-Gelabert in Scarbrough et al., 2005: 33 *DR > TL: Azua Prov., Loma El
Cucurucho, 814 m
Apachekolos invasus Scarbrough & Perez-Gelabert in Scarbrough et al., 2005: 34 *DR > TL: Barahona Prov., Cabral, Hotel
Rincón, 24 m
Apachekolos magnus Scarbrough & Perez-Gelabert in Scarbrough et al., 2005: 35 *DR > TL: Montecristi Prov., Rd. to Playa
Buen Hombre, 132 m
Apachekolos volubilis Scarbrough & Perez-Gelabert in Scarbrough et al., 2005: 37 *DR > TL: Peravia Prov., Matadero, start of
hills, 450 m
Apachekolos vultus Scarbrough & Perez-Gelabert in Scarbrough et al., 2005: 38 *DR > TL: Puerto Plata Prov., 8 km S
Bombita, beside karst mogote, 144 m
Beameromyia Martin, 1957
Beameromyia dicrana Scarbrough, 1997b: 241. Scarbrough et al., 2005: 39 *DR > TL: Prov. Pedernales, ca. 35 km N. Cabo
Rojo, 1250 m, Las Abejas
Beameromyia incisuralis Scarbrough & Page in Scarbrough et al., 2005: 41 *DR > TL: Pedernales Prov., Upper Las Abejas, 38
km NNW Cabo Rojo, 1350 m
Beameromyia melana Scarbrough, 1997b: 238. Scarbrough et al., 2005: 43 *DR > TL: Jarabacoa
Eurhabdus Aldrich, 1923
Eurhabdus sororius Scarbrough & Perez-Gelabert in Scarbrough et al., 2005: 44 *DR > TL: Pedernales Prov., 26 km N Cabo
Rojo, 730 m
Leptogaster Meigen, 1803
Leptogaster hyacinthina Scarbrough, 1996: 197 *DR > TL: Pedernales Prov., ca. 35 km NNW Cabo Rojo, 1430 m, El
Aceitillar
Leptogaster lineatus Scarbrough, 1996: 200 *DR > TL: Pedernales Prov., ca. 35 km NNW Cabo Rojo, 1430 m, El Aceitillar
Leptogaster sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]
Psilonyx Aldrich, 1923
Psilonyx zephyrus Scarbrough & Page in Scarbrough et al., 2005: 45 *DR > TL: Pedernales Prov., Upper Las Abejas, 38 km
NNW Cabo Rojo, 1350 m
Schildia Aldrich, 1923
Schildia martini† Dikow & Bayless, 2009: 278 [Fossil in Dominican amber]
Subfamily OMMATINAE
Key reference: Scarbrough and Perez-Gelabert (2003).
Ommatius Wiedemann, 1821
Ommatius atrosus Scarbrough, 1997a: 11 *DR > TL: Pedernales Prov., 26 km N Cabo Rojo, 730 m
Ommatius bastardoanus Scarbrough in Scarbrough & Perez-Gelabert, 2003: 43 *DR > TL: La Vega Prov., Arroyazo, Reserva
Científica Ebano Verde, 3,500 ft.
Ommatius cinnamomeus Scarbrough, 1984: 147 *H > TL: La Hotte
Ommatius elusivus Scarbrough in Scarbrough & Perez-Gelabert, 2003: 46 *DR > TL: Santiago Prov., Caseta Parques,
Diferencia, Parque Nacional Armando Bermúdez, 740 m
Ommatius fimbrillus† Scarbrough, 2000: 917 [Fossil in Dominican amber]
=Ommatius fimbriatus† Scarbrough, 1997a: 13 [Preoccupied name. Fossil in Dominican amber]
Ommatius flavescens Scarbrough in Scarbrough & Perez-Gelabert, 2003: 48 *DR > TL: Pedernales Prov., 23.5 km N Cabo
Rojo, 540 m
Ommatius geminus Scarbrough & Perez-Gelabert, 2006: 51 *DR > TL: La Vega Prov., Parque Nacional Armando Bermúdez,
km 1–3 along trail W of Ciénaga, 900–1100 m
Ommatius gwenae Scarbrough, 1984b: 139 *DR > TL: Constanza

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Ommatius haitiensis Scarbrough, 1984b: 132 *H > TL: Haiti
Ommatius hierroi Scarbrough in Scarbrough & Perez-Gelabert, 2003: 49 *DR > TL: Duarte Prov., 8 mi. N San Francisco de
Macorís, Loma Quita Espuela, 800 m
Ommatius hispaniolae Scarbrough, 1984b: 137 *DR > TL: Barahona
Ommatius inflatus Scarbrough in Scarbrough & Perez-Gelabert, 2003: 50 *DR > TL: Montecristi Prov., 9 km N Montecristi
Ommatius laticrus Scarbrough & Perez-Gelabert, 2006: 58 *DR > TL: Peravia Prov., southern slopes of Cerro Gordo, Baní
Ommatius lucidatus Scarbrough, 1997a: 13 *DR > TL: Pedernales Prov., 26 km N Cabo Rojo, 730 m
Ommatius maculosus Scarbrough & Perez-Gelabert, 2006: 62 *DR > TL: Pedernales Prov., ~2 km on trail to Carlitos, Parque
Nacional Jaragua, 124 m
Ommatius nigellus Scarbrough, 1984b: 144 *H > TL: Furcy
Ommatius praelongus Scarbrough & Perez-Gelabert, 2006: 66 *DR > TL: Pedernales Prov., Las Abejas, Parque Nacional
Sierra de Bahoruco, 1,310 m
Ommatius pulverius Scarbrough, 1997: 16 *DR > TL: Prov. Pedernales, Cabo Rojo, 0–10 m
Ommatius russelli Scarbrough, 1984b: 142 *DR > TL: La Palma
Ommatius stramineus Scarbrough, 1984: 246 *H, *DR > TL: Hispaniola
Ommatius subtus† Scarbrough & Poinar, 1992: 16 [Fossil in Dominican amber]
Ommatius vitreus Bigot, 1875: 246 *H, *DR > TL: Haiti
Ommatius vivus Scarbrough, 1997a: 18 *DR > TL: Pedernales Prov., ca. 35 km N Cabo Rojo, Las Abejas, 1250 m
Subfamily STENOPOGONINAE
Plesiomma Macquart, 1838
Plesiomma angustum (Macquart, 1848: 20). Scarbrough & Perez-Gelabert, 2006: 73 *H > TL: Haiti
Plesiomma inflatum (Hull, 1962: 219). Scarbrough & Perez-Gelabert, 2006: 76 *H > TL: Mount Basil, North Haiti, 4700 ft.
Plesiomma lineata (Fabricius, 1781). Scarbrough & Perez-Gelabert, 2006: 80
Plesiomma simile Scarbrough & Perez-Gelabert, 2006: 81 *DR > TL: Barahona Prov., nr. Filipinas, Larimar mine
Subfamily TRIGOMIMINAE
Holcocephala Jaennicke, 1867
Holcocephala indigena Scarbrough & Perez-Gelabert, 2006: 84 *DR > TL: La Vega Prov., La Compartición—Los Tablones,
Parque Nacional Armando Bermúdez, 2,450 –1,110 m

Family BOMBYLIIDAE
Subfamily ANTHRACINAE
Tribe ANTHRACINI
Anthrax Scopoli, 1763
Anthrax delicatulus Walker, 1849. Painter et al., 1978: 38.31; Evenhuis & Greathead, 1999: 299
Anthrax funebris Macquart, 1840. Painter et al., 1978: 38.31; Evenhuis & Greathead, 1999: 301
Anthrax insulanus Marston, 1970. Painter et al., 1978: 38.32; Evenhuis & Greathead, 1999: 305
Anthrax oedipus Fabricius, 1805. Perez-Gelabert, 2008: 159
Anthrax succini† Greenwalt & Evenhuis in Greenwalt et al., 2015: 3 [Fossil in Dominican amber]
Tribe EXOPROSOPINI
Exoprosopa Macquart, 1841
Exoprosopa cubana Loew, 1869. Perez-Gelabert, 2008: 159
Ligyra Newman, 1841
Ligyra cerberus Fabricius, 1794. Painter et al., 1978: 38.43; Evenhuis & Greathead, 1999: 402
Ligyra harpyia (Wiedemann, 1828). Painter et al., 1978: 38.43; Evenhuis & Greathead, 1999: 406
Tribe VILLINI
Chrysanthrax Osten-Sacken, 1886
Chrysanthrax fuscus (Fabricius, 1805). Painter et al., 1978: 38.46; Evenhuis & Greathead, 1999: 427
Chrysanthrax gorgon (Fabricius, 1805). Evenhuis & Greathead, 1999: 427
Hemipenthes Loew, 1869
Hemipenthes bidegradata Loew, 1869. Perez-Gelabert, 2008: 159
Hemipenthes dominicana† Evenhuis, 2013: 282 [Fossil in Dominican amber]
Neodiploclampta Curran, 1934
Neodiploclampta roederi Curran, 1931. Perez-Gelabert, 2008: 159
Poecilanthrax Osten Sacken, 1886

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Poecilanthrax lucifer (Fabricius, 1775). Painter et al., 1978: 38.38; Evenhuis & Greathead, 1999: 471
Villa Lioy, 1864
Villa lateralis (Say, 1823). Perez-Gelabert, 2008: 159
Subfamily BOMBYLIINAE
Tribe BOMBYLIINI
Heterostylum Macquart, 1848
Heterostylum haemorrhoicum (Loew, 1863). Painter et al., 1978: 38.5; Cunha et al., 2007: 17; Evenhuis & Greathead, 1999:
156
=Bombylius semirufus Loew, 1872: 78. Cunha et al., 2007: 17; Evenhuis & Greathead, 1999: 156
Heterostylum pallipes Bigot, 1892: 361. Painter et al., 1978: 38.5; Cunha et al., 2007: 19; Evenhuis & Greathead, 1999: 157
*H, *DR > TL: Haiti
Subfamily GERONTINAE
Geron Meigen, 1820
Geron (Geron) articulatus Scarbrough & Davidson, 1985: 1256; Evenhuis & Greathead, 1999: 38 *DR > TL: Palo Verde,
Monte Cristi
Geron senilis (Fabricius, 1794). Perez-Gelabert, 2008: 160
Subfamily PHTHIRINAE
Tribe POECILOGNATHINI
Poecilognathus Jaennicke, 1867
Poecilognathus sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]

Family MYDIDAE
Subfamily MYDINAE
Baliomydas Wilcox & Pimentel, 1989
Baliomydas gracilis (Macquart, 1834). Kondratieff & Perez-Gelabert, 2006: 382
Ceriomydas Williston, 1898
Ceriomydas darlingtoni Papavero & Wilcox, 1974: 47 *DR > TL: Sánchez
Nemomydas Curran, 1934
Nemomydas dominicanus Kondratieff & Perez-Gelabert, 2004: 380 *DR > TL: La Malena de Boca Chica, Santo Domingo
Prov., nr. sea level

Family MYTHICOMYIIDAE
Glabellula Loew, 1873
Glabellula brunnifrons† Evenhuis, 2002c: 3 [Fossil in Dominican amber]
Glabellula grimaldii† Evenhuis, 2002c: 5 [Fossil in Dominican amber]
Glabellula kuehnei† Schlüter, 1976: 358 [Fossil in Dominican amber]
Mythicomyia Coquillett, 1893
Mythicomyia dominicana† Evenhuis, 2002c: 8 [Fossil in Dominican amber]
Mythicomyia sp.†—Poinar & Poinar (1999: 206) [Misspelled Mythecomyia] [Fossil in Dominican amber]
Pieza Evenhuis, 2002
Pieza dominicana† Evenhuis, 2002a: 10. Evenhuis, 2002b: 55 [Fossil in Dominican amber]
Pieza sinclairi Evenhuis, 2002a: 22. Evenhuis, 2002b: 55

Family SCENOPINIDAE
Metatrichia Coquillett, 1900
Metatrichia pria† Yeates & Grimaldi, 1993: 2 [Fossil in Dominican amber]

Family THEREVIDAE
Ambradolon† Metz & Irwin, 2000
Ambradolon grimaldii† Metz & Irwin, 2000b: 991 [Fossil in Dominican amber]
Elcaribe Webb in Webb & Metz, 2006
Elcaribe bifidus Webb in Webb & Metz, 2006: 167 *DR > TL: Puerto Plata
Elcaribe elongatus Webb in Webb & Metz, 2006: 170
Elcaribe glabrus Webb in Webb & Metz, 2006: 172 *DR > TL: Independencia, 4 km S Los Pinos, Loma de Vientos, 475 m

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Elcaribe laticornis (Loew, 1869). Webb & Metz, 2006: 178
Elcaribe obscurus (Coquillett, 1893). Webb & Metz, 2006: 183
Elcaribe platycera Loew, 1872. Webb et al., 2012: 38
Elcaribe stellus Webb in Webb & Metz, 2006: 198 *DR > TL: Isla Saona, Mano Juan
Elcaribe vexans (Curran, 1926). Curran, 1928: 25; Webb & Metz, 2006: 158
Insulatitan Metz & Irwin, 2000
Insulatitan trishae Metz & Irwin, 2000b: 997 *DR > TL: 7 km WNW Barrero, east side of crest of Sierra Martín García, Azua
Prov.
Insulatitan youngi Metz & Irwin, 2000b: 1002
Nesonana Gaimari & Irwin, 2000
Nesonana xenoverpa Gaimari & Irwin, 2000a: 172 *DR > TL: Pedernales: 3.3 km NE Los Arroyos, 1450 m
Nesonana youngi Gaimari & Irwin, 2000a: 175 *DR > TL: Pedernales: 26 km N Cabo Rojo, 730 m

Superfamily EMPIDOIDEA
Family DOLICHOPODIDAE
Subfamily DIAPHORINAE
Chrysotus Meigen, 1824
Chrysotus runyoni Grichanov, 2017: 134 *DR > TL: Barahona env.
Subfamily DOLICHOPODINAE
Paraclius Loew, 1864
Paraclius pilifer Aldrich, 1896. Perez-Gelabert, 2008: 161
Subfamily HYDROPHORINAE
Diostracus Loew, 1861
Diostracus sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Subfamily MEDETERINAE
Thrypticus Gerstaecker, 1864
Thrypticus violaceus Van Duzee, 1927. Perez-Gelabert, 2008: 161
Subfamily SCIAPODINAE
Condylostylus Bigot, 1859
Condylostylus pilosus (Loew, 1861). Perez-Gelabert, 2008: 161
Condylostylus pruinosus (Coquillett, 1904). Pollet et al., 2004: 71
Condylostylus quadricolor (Walker, 1849). Perez-Gelabert, 2008: 161
Condylostylus sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Amblypsilopus Bigot, 1888
Amblypsilopus bellimanus (Van Duzee, 1927: 10). Robinson, 1970: 40.15; Bickel, 2002: 548 *DR > TL: Sánchez
=Psilopus spinimanus Van Duzee, 1927: 6. Curran, 1928: 32
Amblypsilopus unicinctus (Van Duzee, 1927: 6). Robinson, 1970: 40.16; Bickel, 2002: 548
=Sciapus angelicus Parent, 1930: 109. Robinson, 1970: 40.14; Bickel, 2002: 548
Notes: The Psilopodinus hirtulus Bigot, 1888: xxix. Bigot, 1890: 286; Bickel, 2002: 556, Psilopodinus pampoecilus Bigot,
1888: xxix. Robinson, 1970: 40.10; Bickel, 2002: 556 and Psilopodinus polychromus Bigot, 1890: 289. Robinson, 1970: 40.10;
Bickel, 2002: 556 are unidentifiable species considered nomina dubia. They were described early in the 19th century from Haiti
and have been removed from the list of valid species (Bickel, 2002).

Family EMPIDIDAE
Subfamily TACHYDROMIINAE
Chersodromia Bigot, 1891
Chersodromia dominicana† Solórzano et al., 2005: 42 [Fossil in Dominican amber]
Drapetis Meigen, 1822
Drapetis (Elaphropeza) flavida (Williston, 1896). Smith, 1967: 39.4
Porphyrochroa Melander, 1927
Porphyrochroa argentata Rafael & Ale-Rocha, 2002: 247 *DR > TL: Las Abejas, 38 km N Cabo Rojo, 1250 m
Porphyrochroa dominicanensis Rafael & Ale-Rocha, 2002: 242 *DR > TL: Independencia, Sierra de Neiba near crest, 5.5 km
NNW Angel Feliz, 1750 m
Porphyrochroa genalis Rafael & Ale-Rocha, 2002: 244 *DR > TL: Independencia, Sierra de Neiba near crest, 5.5 km NNW

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Angel Feliz, 1750 m
Porphyrochroa latifrons Rafael & Ale-Rocha, 2002: 242 *DR > TL: Las Abejas, 38 km N Cabo Rojo, 1250 m
Porphyrochroa pectinata Rafael & Ale-Rocha, 2002: 244 *DR > TL: Independencia, Sierra de Neiba near crest, 5.5 km NNW
Angel Feliz, 1750 m
Rhamphomyia Meigen, 1822
Rhamphomyia sp.†—Wu (1997: 192) [Fossil in Dominican amber]

Family HYBOTIDAE
Subfamily HYBOTINAE
Euhybus Coquillett, 1896
Euhybus martiniensis Ale-Rocha & Rafael, 2004: 496 *DR > TL: Azua: East side of crest, Sierra Martín García, 7 km WNW
Barrero, 860 m
Neohybos Ale-Rocha & De Carvalho, 2003
Neohybos cinereus Ale-Rocha & Rafael, 2004: 498 *DR > TL: Independencia: Sierra de Neiba, just south of crest, 5 km NNW
Angel Feliz, 1780 m
Neohybos longicornis Ale-Rocha & Rafael, 2004: 500 *DR > TL: Pedernales: Upper Las Abejas, 38 km NNW Cabo Rojo,
1350 m
Neohybos pruinosus Ale-Rocha & Rafael, 2004: 500 *DR > TL: Independencia: Sierra de Neiba, just south of crest, 5.5 km
NNW Angel Feliz, 1750 m
Neohybos setosus Ale-Rocha & Rafael, 2004: 502 *DR > TL: La Vega: 9 km SE Constanza near Valle Nuevo, 1930 m
Syneches Walker, 1852
Syneches dominicanus Ale-Rocha & Rafael, 2004: 506
Syneches inversus Curran, 1928. Ale-Rocha & Rafael, 2004: 507
Syneches pallidus Wilder, 1974. Ale-Rocha & Rafael, 2004: 507
Syneches vineus Wilder, 1974. Ale-Rocha & Rafael, 2004: 507

Infraorder MUSCOMORPHA
Family PHORIDAE
Subfamily AENIGMATIINAE
Diplonevra Lioy, 1864
Diplonevra sp.†—Brown (1999: 1570) [Fossil in Dominican amber]
Dohrniphora Dahl, 1898
Dohrniphora calvata† Solórzano Kraemer & Brown, 2018: 313 [Fossil in Dominican amber]
Dohrniphora cornuta (Bigot, 1857). Kung & Brown, 2006: 1932
Dohrniphora dispar (Enderlein, 1912). Kung & Brown, 2006: 1934
Dohrniphora divaricata (Aldrich, 1896). Kung & Brown, 2006: 1936
Dohrniphora kraemergabi† Solórzano Kraemer & Brown, 2018: 304 [Fossil in Dominican amber]
Dohrniphora minoris† Solórzano Kraemer & Brown, 2018: 312 [Fossil in Dominican amber]
Dohrniphora paleocircularis† Solórzano Kraemer & Brown, 2018: 317 [Fossil in Dominican amber]
Dohrniphora poinari† Disney, 1987: 377 [Fossil in Dominican amber]
Dohrniphora processa† Solórzano Kraemer & Brown, 2018: 316 [Fossil in Dominican amber]
Dohrniphora quasicornuta† Solórzano Kraemer & Brown, 2018: 314 [Fossil in Dominican amber]
Dohrniphora quinti† Solórzano Kraemer & Brown, 2018: 309 [Fossil in Dominican amber]
Dohrniphora saepis† Solórzano Kraemer & Brown, 2018: 306 [Fossil in Dominican amber]
Dohrniphora sexspinosa Kung & Brown, 2006: 1941 *DR > TL: Pedernales, 23.5 km N Cabo Rojo, 540 m
Dohrniphora sp.†—Solórzano Kraemer & Brown (2018: 320) [Fossil in Dominican amber]
Dohrniphora translucendi† Solórzano Kraemer & Brown, 2018: 308 [Fossil in Dominican amber]
Dohrniphora volkswageni† Solórzano Kraemer & Brown, 2018: 318 [Fossil in Dominican amber]
Subfamily CONICERINAE
Conicera Meigen, 1830
Conicera sp.†—Brown (1999: 1570) [Fossil in Dominican amber]
Subfamily HYPOCERINAE
Abaristophora Schmitz, 1927
Abaristophora domicambarae† Disney in Disney & Ross, 1997: 128 [Fossil in Dominican amber]

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Borophaga Enderlein, 1924
Borophaga sp.†—Brown (1999: 1570) [Fossil in Dominican amber]
Subfamily PHORINAE
Neopleurophora Brown, 1992
Neopleurophora sp.†—Brown (1999: 1570) [Fossil in Dominican amber]
Subfamily METOPININAE
Apocephalus Coquillett, 1901
Apocephalus (Mesophora) grandiflavus Brown, 1994: 4 *DR > TL: Pedernales: 3.3 km NE Los Arroyos, 1450 m
Apocephalus (Mesophora) brevicercus Brown, 1993: 223
Apocephalus (Mesophora) crassus Brown, 1996: 24 *DR > TL: Pedernales: 3.3 km NE Los Arroyos, 1450 m
Apocephalus (Mesophora) echinatus Brown, 1996: 25 *DR > TL: Pedernales, 5 km NE Los Arroyos
Apocephalus (Mesophora) micrepelis Brown, 1993: 224
Apocephalus (Mesophora) pilatus Brown, 1996: 23 *DR > TL: Pedernales, Las Abejas, 38 km NNW Cabo Rojo, 1250 m
Apocephalus (Mesophora) prolixus Brown, 1996: 24 *DR > TL: Pedernales, Las Abejas, 38 km NNW Cabo Rojo, 1250 m
Apocephalus (Mesophora) secundus Brown, 1996: 25 *DR > TL: Pedernales, Las Abejas, 38 km NNW Cabo Rojo, 1250 m
Apocephalus succineus† Brown, 2000: 17 [Fossil in Dominican amber]
Beckerina Malloch, 1910
Beckerina sp.†—Brown (1999: 1570) [Fossil in Dominican amber]
Chonocephalus Wandoleck, 1898
Chonocephalus fletcheri Schmitz, 1912. Disney, 2008: 18
Chonocephalus sp.†—Brown (1999: 1570) [Fossil in Dominican amber]
Megaselia Rondani, 1856
Megaselia amberae† Disney, 1987: 378 [Fossil in Dominican amber]
Megaselia bernsteinae† Disney, 1987: 378 [Fossil in Dominican amber]
Megaselia dominicana† Disney, 1987: 378 [Fossil in Dominican amber]
Metopina Macquart, 1835
Metopina sp.—Grimaldi (1989: 65); Brown (1999: 1569)
Myriophora Brown, 1992
Myriophora communis Hash & Brown, 2015: 17
Myriophora dolionatis Hash & Brown, 2015: 21
Myriophora sp.†—Brown (1999: 1570) [Fossil in Dominican amber]
Pseudacteon Coquillett, 1907
Pseudacteon grandis Greene, 1941. Plowes et al., 2009: 944
Puliciphora Dahl, 1897
Puliciphora rontaleri† Disney, 1997: 131 [Fossil in Dominican amber]
Rhyncophoromyia Malloch, 1923
Rhyncophoromyia sp.†—Brown (1999: 1570) [Fossil in Dominican amber]

Family PIPUNCULIDAE
Subfamily PIPUNCULINAE
Basileunculus Rafael, 1987
Basileunculus aliceae Rafael, 1987. Rafael, 1996: 371
Cephalops Fallén, 1810
Cephalops (Cephalops) pedernalensis Rafael, 1996: 368 *DR > TL: 23.5 km N. Cabo Rojo, Pedernales Prov.
Cephalops (Semicephalops) varius stygius (Hardy, 1948). Rafael, 1996: 369
Cephalops sp.—Rafael (1996: 370)
Cephalosphaera Enderlein, 1936
Cephalosphaera (Cephalosphaera) insularis Rafael, 1996: 367
Cephalosphaera (Cephalosphaera) mocaensis (Hardy, 1948). Rafael, 1996: 365
Cephalosphaera spp.—Rafael (1996: 368)
Eudorylas Aczel, 1940
Eudorylas curvicaudatus Rafael, 1996: 377 *DR > TL: 3 km W. Cueva de Arena, Parque Los Haitises, Hato Mayor Prov.
Eudorylas dominicanensis Rafael, 1996: 377 *DR > TL: 7 km E. Los Arroyos, Pedernales Prov.
Eudorylas sp. A†—De Meyer (1995: 208) [Fossil in Dominican amber]

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Eudorylas sp. B†—De Meyer (1995: 210) [Fossil in Dominican amber]
Metadorylas Rafael, 1987
Metadorylas antillensis Rafael, 1996: 375 *DR > TL: Pedernales, 23.5 km N Cabo Rojo, 540 m
Metadorylas cornutus Rafael, 1996: 374 *DR > TL: 3.3 km NE Los Arroyos, Pedernales Prov.
Metadorylas cressoni (Johnson, 1919). Rafael, 1996: 375
Metadorylas dominicensis Scarbrough & Knutson, 1989. Rafael, 1996: 376
Metadorylas pilosus Rafael, 1996: 376 *DR > TL: 23.5 km N. Cabo Rojo, Pedernales Prov.
Metadorylas youngi Rafael, 1996: 372 *DR > TL: 26 km N. Cabo Rojo, Pedernales Prov.
Microcephalops De Meyer, 1989
Microcephalops latifrons (Hardy, 1948). Rafael, 1996: 370
Microcephalops williamsi (Hardy, 1954). Rafael, 1996: 370
Tomosvaryella Aczel, 1939
Tomosvaryella mexicanensis Ale-Rocha & Rafael, 1995. Rafael, 1996: 380
Tomosvaryella tuberculata Hardy, 1948. Scarbrough & Knutson, 1989: 528; Rafael, 1996: 379

Family PLATYPEZIDAE
Bertamyia Kessel, 1970
Bertamyia notata (Loew, 1866). Woodley, 2005: 731
Lindneromyia Kessel, 1965
Lindneromyia dominicana† Grimaldi, 2018: 57 [Fossil in Dominican amber]
Lindneromyia neomedialis† Grimaldi, 2018: 53 [Fossil in Dominican amber]

Family SYRPHIDAE
Subfamily ERISTALINAE
Tribe VOLUCELLINI
Copestylum Macquart, 1846
Copestylum hispaniolae Thompson, 1981: 129 *H, *DR > TL: San Cristóbal
Copestylum purpurascens (Loew, 1869: 31). Thompson, 1981: 134 *H, *DR > TL: Hayti
=Volucella watsoni Curran, 1930a: 5
Copestylum sexmaculatum (Palisot de Beauvois, 1819). Thompson, 1981: 138
Copestylum sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Ornidia Lepeletier & Serville, 1828
Ornidia obesa (Fabricius, 1775). Wolcott, 1927: 319; Thompson, 1981: 120; Thompson, 1991: 256
Tribe ERISTALINI
Meromacrus Rondani, 1848
Meromacrus pinguis (Fabricius, 1775). Thompson, 1981: 161
Meromacrus pratorum (Fabricius, 1775). Thompson, 1981: 162
Palpada Macquart, 1834
Palpada agrorum (Fabricius, 1787). Thompson, 1981: 150
Palpada albifrons (Wiedemann, 1830). Thompson, 1981: 151
Palpada atrimana (Loew, 1866). Thompson, 1981: 151
Palpada hortorum (Fabricius, 1775). Thompson, 1981: 152
Palpada vinetorum (Fabricius, 1798). Wolcott, 1927: 319; Thompson, 1981: 153
Tribe MILESIINI
Neplas Porter, 1927
Neplas pretiosus (Loew, 1861). Thompson, 1981: 172
Subfamily MICRODONTINAE
Laetodon Reemer in Reemer and Ståhls, 2013
Laetodon laetus (Loew, 1864). Thompson, 1981: 115
Microdon Meigen, 1803
Microdon bruesi Hull, 1945: 77. Thompson, 1981: 114 *H > TL: Port-au-Prince and vicinity
Microdon pulcher (Williston, 1887: 5). Thompson, 1981: 115 *DR > TL: Saint Domingo
Subfamily SYRPHINAE
Tribe SYRPHINI

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Allograpta Osten Sacken, 1875
Allograpta (Allograpta) obliqua (Say, 1823). Fluke, 1942: 19; Thompson, 1981: 30
Allograpta (Antillus) ascita (Vockeroth, 1969: 131). Thompson, 1981: 26 *H, *DR > TL: Mt. La Hotte, Desbarrière, nr. 4000
ft.
Allograpta neotropica Curran, 1936. Perez-Gelabert, 2015: 70
Dioprosopa Hull, 1949
Dioprosopa clavata (Fabricius, 1794). Wolcott, 1927: 321; Thompson, 1981: 33; Kassebeer, 2000: 80
Ocyptamus Macquart, 1834
Ocyptamus antiphates (Walker, 1849). Thompson, 1981: 46
Ocyptamus cylindricus (Fabricius, 1781). Thompson, 1981: 46
Ocyptamus dimidiatus (Fabricius, 1781). Thompson, 1981: 47
Ocyptamus fasciatus Roeder, 1885. Thompson, 1981: 49
Ocyptamus megafemur Thompson in Mengual et al., 2018: 17
Ocyptamus oriel (Hull, 1943: 96). Thompson, 1981: 53 *DR > TL: Loma Rucilla and Mts., 5–8000 ft.
Syrphus Philippi, 1865
Syrphus vockerothi Thompson, 1981: 21
Rhinoprosopa Hull, 1942
Rhinoprosopa aenea (Hull, 1937). Thompson, 1981: 26; Mengual, 2014: 5 *H, *DR > TL: Desbarriere/Mt. La Hotte
Tribe TOXOMERINI
Toxomerus Macquart, 1855
Toxomerus arcifer (Loew, 1866). Thompson, 1981: 82
Toxomerus aurulentus (Williston, 1887). Thompson, 1981: 82
Toxomerus buscki Thompson, 1981: 83 *DR > TL: San Francisco Mts., [Prov. San Cristóbal]
Toxomerus dispar (Fabricius, 1794). Thompson, 1981: 86
Toxomerus floralis (Fabricius, 1798). Thompson, 1981: 89
Toxomerus lacrymosus (Bigot, 1884). Perez-Gelabert, 2015: 70
Toxomerus ornithoglyphus (Hull, 1942a: 17). Thompson, 1981: 93 *DR > Monte Diego de Ocampo, 3000–4000 ft.
Toxomerus pictus (Macquart, 1842). Perez-Gelabert, 2015: 70
Toxomerus politus (Say, 1823). Thompson, 1981: 94; Metz & Thompson, 2001: 240
Toxomerus pulchellus (Macquart, 1846). Thompson, 1981: 95
Toxomerus una (Hull, 1943: 23). Thompson, 1981: 97 *H > TL: Haiti
Toxomerus veve (Hull, 1942a: 23). Thompson, 1981: 99 *H > TL: La Selle Range, La Visite and vic., 5000–6000 ft.
Toxomerus violaceus Curran, 1926). Perez-Gelabert, 2015: 70
Toxomerus watsoni (Curran, 1930b: 3). Thompson, 1981: 99
Toxomerus species A—Thompson (1981: 100)
Tribe BACCHINI
Leucopodella Hull, 1949
Leucopodella gracilis (Williston, 1891). Thompson, 1981: 103
Xanthandrus Verrall, 1901
Xanthandrus (Xanthandrus) simplex (Loew, 1861). Thompson, 1981: 109

CALYPTRATAE
Family ANTHOMYIIDAE
Coenosopsites Michelsen, 1996
Coenosopsites poinari† Michelsen, 1996: 442 [Fossil in Dominican amber]
Grimalantha Rohacek, 1998
Grimalantha vulnerata† Rohácek, 1998: 147 [Fossil in Dominican amber]

Family CALLIPHORIDAE
Subfamily CALLIPHORINAE
Calliphora Robineau-Desvoidy, 1830
Calliphora maestrica Peris et al., 1998. Whitworth, 2010: 8
Subfamily CHRYSOMYINAE
Chloroprocta Van der Wulp, 1896

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Chloroprocta idioidea (Robineau-Desvoidy, 1830). Whitworth, 2010: 10
Chrysomya Robineau-Desvoidy, 1830
Chrysomya megacephala (Fabricius, 1794). Whitworth, 2010: 12
Cochliomyia Townsend, 1915
Cochliomyia hominivorax (Coquerel, 1858). Alvarez, 1984: 73; Dear, 1985: 138
=Callitroga americana Cushing & Patton, 1933. Laake et al., 1936: 3
Cochliomyia macellaria (Fabricius, 1775). Dear, 1985: 139
=Cochliomyia fontanai Garcia, 1952. Dear, 1985: 144
Cochliomyia minima Shannon, 1926: 124. James, 1970: 102.7; Dear, 1985: 136; Whitworth, 2010: 16
Subfamily LUCILIINAE
Lucilia Robineau-Desvoidy, 1830
Lucilia cuprina (Wiedemann, 1830). Alvarez, 1984: 43; Whitworth, 2010: 21; Whitworth, 2014: 25
Lucilia eximia (Wiedemann, 1819). Whitworth, 2010: 21
Lucilia retroversa (James, 1971). Whitworth, 2010: 26; Whitworth, 2014: 45
Lucilia rica Shannon, 1926. Whitworth, 2010: 30
=Somomyia soulouquina Bigot, 1877: 47
=Phaenicia rica (Shannon, 1926). James, 1970: 102.11
Lucilia sericata (Meigen, 1826: 53). Guimarães, Papavero & Pires do Prado, 1983: 256
Subfamily MELANOMYINAE
Glutoxys Aldrich, 1929
Glutoxys elegans Aldrich, 1929. Whitworth, 2010: 33
Pseudopsodexia Townsend, 1935
Pseudopsodexia sp.—Whitworth (2010: 33)

Family HIPPOBOSCIDAE
Subfamily ORNITHOMYIINAE
Olfersia Wiedemann, 1830
Olfersia sordida Bigot, 1885. Bequaert, 1940: 321; Bequaert, 1957: 469
Ornithoctona Speiser, 1902
Ornithoctona erythrocephala (Leach, 1817). Bequaert, 1940: 315; Bequaert, 1954: 254
Ornithoctona fusciventris (Wiedemann, 1830). Bequaert, 1940: 317; Bequaert, 1954: 210
Pseudolynchia Bequaert, 1925
Pseudolynchia canariensis (Macquart, 1840). Bequaert, 1940: 324; Bequaert, 1955: 390 [Introduced]
Stilbometopa Coquillett, 1899
Stilbometopa ramphastonis Ferris, 1930. Bequaert, 1935: 325; Bequaert, 1940: 319; Bequaert, 1954: 254

Family STREBLIDAE
Subfamily NYCTEROPHILIINAE
Enischnomyia Poinar & Brown, 2012
Enischnomyia stegosoma† Poinar & Brown, 2012: 82 [Fossil in Dominican amber]
Subfamily STREBLINAE
Strebla Wiedemann, 1824
Strebla avium (Macquart, 1854). Kessel, 1925: 30
Subfamily TRICHOBIINAE
Trichobius Gervais, 1844
Trichobius dominicanus Peterson & Hůrka, 1974: 1065 *DR > TL: Cueva Jugres, Monción, from Monophyllus sp.
Trichobius frequens Peterson & Hůrka, 1974: 1061. Guerrero, 1994: 9
Trichobius intermedius Peterson & Hůrka, 1974: 1049
Trichobius neotropicus Peterson & Hůrka, 1974: 1054 *DR > TL: Cueva sin nombre, Sánchez, from Macrotus waterhousii
waterhousii

Family FANNIIDAE
Subfamily FANNIINAE
Fannia Robineau-Desvoidy, 1830

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Fannia femoralis (Stein, 1898). Curran, 1928: 89
Fannia trimaculata (Stein, 1898). Seago, 1954: 9

Family MUSCIDAE
Key reference: Carvalho et al. (2005).
Subfamily ATHERIGONINAE
Atherigona Rondani, 1856
Atherigona orientalis Schiner, 1868. Carvalho et al., 2005: 12
Subfamily AZELIINAE
Tribe REINWARDTIINI
Philornis Meinert, 1890
Philornis pici (Macquart, 1853). Carvalho et al., 2005: 67; Löwenberg-Neto & De Carvalho, 2013: 33 ; Hayes et al., 2019: 157
Synthesiomyia Brauer & Bergenstamm, 1893
Synthesiomyia nudiseta (Wulp, 1883). Carvalho et al., 2005: 75
Subfamily COENOSIINAE
Tribe LIMNOPHORINI
Lispe Latreille, 1796
Lispe nasoni Stein, 1898. Vikhrev, 2016: 181
Lispe probohemica Speiser, 1914. Vikhrev, 2016: 182
Lispe sordida Aldrich, 1913. Vikhrev, 2016: 183
Subfamily MUSCINAE
Tribe MUSCINI
Archaeopolietes† Pont & Carvalho, 1997
Archaeopolietes tertiaria† Pont & Carvalho, 1997: 178. Carvalho et al., 2005: 14 [Fossil in Dominican amber]
Morellia Robineau-Desvoidy, 1830
Morellia basalis (Walker, 1853). Carvalho et al., 2005: 15; Löwenberg-Neto & De Carvalho, 2013: 33
=Pyrellia flora Bigot, 1878. Carvalho et al., 2005: 16
Morellia (Parapyrellia) maculipennis (Macquart, 1846). Medina Gaud, 2007: 144; Löwenberg-Neto & De Carvalho, 2013: 14
Musca Linnaeus, 1758
Musca domestica Linnaeus, 1758. Wolcott, 1927: 323; Alvarez, 1984: 27; Carvalho et al., 2005: 22
Tribe STOMOXYINI
Haematobia Lepeletier & Serville, 1828
Haematobia irritans (Linnaeus, 1758). Carvalho et al., 2005: 37
Stomoxys Geoffroy, 1762
Stomoxys calcitrans (Linnaeus, 1758). Wolcott, 1927: 324; Carvalho et al., 2005: 39
Subfamily MYDAEINAE
Brontaea Kowarz, 1873
Brontaea debilis (Williston, 1896). Carvalho et al., 2005: 135
=Gymnodia scatophaga Curran, 1938: 5. Carvalho et al., 2005: 136
Brontaea quadristigma (Thomson, 1869). Carvalho et al., 2005: 137
Graphomya Agassiz, 1846
Graphomya stipata (Walker, 1856). Medina Gaud, 2007: 123
Myospila Rondani, 1856
Myospila pallidicornis (Bigot, 1887). Carvalho et al., 2005: 146; Löwenberg-Neto & De Carvalho, 2013: 82
=Myospila obsoleta (Brauer & Bergenstamm, 1891). Curran, 1928: 90; Carvalho et al., 2005: 147
Subfamily PHAONIINAE
Phaonia Robineau-Desvoidy, 1830
Phaonia electrica† Pont & Carvalho, 1997: 176. Carvalho et al., 2005: 127 [Fossil in Dominican amber]
Phaonia succini† Pont & Carvalho, 1997: 174. Carvalho et al., 2005: 133 [Fossil in Dominican amber]

Family OESTRIDAE
Subfamily GASTEROPHILINAE
Gasterophilus Leach, 1817
Gasterophilus intestinalis (De Geer, 1776) [By synonymy of species below]

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=Gasterophilus equi (Clark, 1797). Wolcott, 1927: 72
Gasterophilus nasalis Linnaeus, 1758. Alvarez, 1984: 53; Marcano, 1964: 155
Gyrostigma Brauer, 1885
Gyrostigma rhinocerontis (Owen, 1830) [By synonymy of name below]
=Gyrostigma pavesii (Corti, 1895). Alvarez, 1984: 63 [In rhino at Santo Domingo zoo]
Subfamily HYPODERMATINAE
Hypoderma Latreille, 1818
Hypoderma (Hypoderma) lineatum (Villers, 1789). Guimarães, 1967: 106.2

Family MESEMBRINELLIDAE
Mesembrinella Giglio-Tos, 1893
Mesembrinella caenozoica† Cerreti et al., 2017: 4 [Fossil in Dominican amber]

Family SARCOPHAGIDAE
Key reference: Pape (1996).
Subfamily SARCOPHAGINAE
Argoravinia Townsend, 1917
Argoravinia aurea (Townsend, 1917). Pape, 1996: 175
Blaesoxipha Loew, 1861
Blaesoxipha (Acanthodotheca) acridophagoides (Lopes & Downs, 1951). Pape, 1996: 177
Blaesoxipha (Giganthoteca) plinthopyga (Wiedemann, 1830). Pape, 1994: 66; Pape, 1996: 201
Blaesoxipha (Tephromyia) blandita (Brèthes, 1920). Pape, 1996: 208
Boettcheria Parker, 1914
Boettcheria solo Pape, 1990: 431. Pape, 1996: 215 *H, *DR > TL: Prov. El Seibo
Dexosarcophaga Townsend, 1917
Dexosarcophaga varenna (Dodge, 1968). Pape, 1996: 220
Fletcherimyia Townsend, 1917
Fletcherimyia morionella (Aldrich, 1930). Pape, 1996: 227
Oxysarcodexia Townsend, 1917
Oxysarcodexia bakeri (Aldrich, 1930). Pape, 1996: 263
Peckia Robineau-Desvoidy, 1830
Peckia (Euboettcheria) percussa (Lopes, 1938: 337). Pape, 1996: 276 *H > TL: Port-au-Prince
Peckia (Peckia) imperialis Robineau-Desvoidy, 1830. Buenaventura & Pape, 2013: 57
=Sarcophaga fortipes Walker, 1861. Buenaventura & Pape, 2013: 57
Peckia (Peckia) praeceps (Wiedemann, 1830). Pape, 1996: 280
Peckia (Sarcodexia) dominicana (Lopes, 1982: 291). Pape, 1996: 275; Buenaventura & Pape, 2013: 73 *DR > TL: Prov.
Samaná
Peckia (Squamatodes) virgo (Pape, 1994). Pape, 1996: 276; Buenaventura & Pape, 2013: 81
Peckia plumipes Robineau-Desvoidy, 1830: 336. Pape, 1996: 282 *H > TL: L’Ile d’Haïti
Ravinia Robineau-Desvoidy, 1830
Ravinia effrenata (Walker, 1861). Pape, 1996: 286
Sarcodexia Townsend, 1892
Sarcodexia lambens (Wiedemann, 1830). Pape, 1996: 292
Sarcofahrtiopsis Hall, 1933
Sarcofahrtiopsis spathor de Mello & Pape, 2000: 189 *DR > TL: Azua (East side of crest), Sierra Martín García, 7 km WNW
Barrero, 860 m
Sarcophaga Meigen, 1826
Sarcophaga (Mehria) insularis (Lopes, 1946: 123). Pape, 1996: 362 *H > TL: Haiti, larva parasitic in egg-bags of gregarious
epeirid spider
Titanogrypa Townsend, 1917
Titanogrypa (Sarconeiva) fimbriata (Aldrich, 1916). Pape, 1996: 432
Tricharaea Thomson, 1869
Tricharaea (Sarcophagula) occidua (Fabricius, 1794). Pape, 1996: 434
Tricharaea (Sarothromyia) simplex (Aldrich, 1916). Pape, 1996: 435

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 219
Unrecognized nominal species-taxa assigned to Tricharaea
“Anapunaphyto” lopesi Thompson & Pont, 1993: 161. Pape, 1996: 436
=Musca pusilla Wiedemann, 1930
Unidentified nominal species-group taxa
Myophora Robineau-Desvoidy, 1830
Myophora fervida Robineau-Desvoidy, 1830: 341. Pape, 1996: 444 *H > TL: Haïti

Family SCATHOPHAGIDAE
Scathophaga Fabricius, 1805
Scathophaga stercoraria (Linnaeus, 1758). De Oliveira Albuquerque, 1984: 96B.3 [Introduced]

Family TACHINIDAE
Subfamily EXORISTINAE
Belvosia Robineau-Desvoidy, 1830
Belvosia vanderwulpi Williston, 1886: 303. Guimarães, 1971: 104.183 *DR > TL: San Domingo
Carcelia Robineau-Desvoidy, 1830
Carcelia rubrella Robineau-Desvoidy, 1830: 179. Guimarães, 1971: 104.203 *DR > TL: Saint-Domingue [“unrecognized”
species]
Leschenaultia Robineau-Desvoidy, 1830
Leschenaultia arnaudi Toma & Guimarães, 2002: 41 *H > TL: La Selle, La Visite & vic. (5–700 ft.)
Lixophaga Townsend, 1908
Lixophaga diatraeae (Townsend, 1916). Wolcott, 1927: 321; Santoro, 1960: 146
Subfamily PHASIINAE
Beskia Guimarães, (1971)
Beskia aelops (Walker, 1849). Guimarães, 1971: 104.14
=Beskia willistoni (Townsend, 1894: 796). Guimarães, 1971: 104.15
Cistogaster Latreille, 1829
Cistogaster dominica Curran, 1927: 14. Guimarães, 1971: 104.5 *DR > TL: Sánchez
Trichopoda Berthold, 1827
Trichopoda (Trichopoda) haitensis Robineau-Desvoidy, 1830: 285. Guimarães, 1971: 104.9 *H > TL: L’ile d’Haïti
Trichopoda (Trichopoda) subalipes Townsend, 1897: 30. Guimarães, 1971: 104.9 *DR > TL: Santo Domingo
Subfamily PROSENINAE
Chaetocalirrhoe Townsend, 1935
Chaetocalirrhoe grandis Townsend, 1935: 218. Guimarães, 1971: 104.28 *H > TL: Kenscoff
Euphasiopteryx Townsend, 1915
Euphasiopteryx dominicana (Townsend, 1919: 548). Guimarães, 1971: 104.21
Ormia Robineau-Desvoidy, 1830
Ormia punctata Robineau-Desvoidy, 1830. Sabrosky, 1953: 176
Ptilodexia Brauer & Bergenstamm, 1893
Ptilodexia spinosa (Bigot, 1888: 266). Guimarães, 1971: 104.34 *H > TL: Haïti [“unrecognized” species]
Ptilodexia strenua (Robineau-Devoidy, 1830: 315). Guimarães, 1971: 104.34 *H > TL: Port-au-Prince
Subfamily TACHININAE
Archytas Jaennicke, 1867
Archytas marmoratus (Townsend, 1915). Sabrosky, 1955: 78
Deopalpus Townsend, 1908
Deopalpus sp.—Woodley (2007: 857)
Jurinella Brauer & Bergenstamm, 1889
Jurinella baoruco Woodley, 2007: 857 *DR > TL: Pedernales Prov., Parque Nacional Sierra de Bahoruco, Las Abejas, 1150 m
Paradejeania Brauer & Bergenstamm, 1893
Paradejeania xenisma Woodley, 1993: 185 *DR > TL: Independencia Prov., 1 km E of El Aguacate, 950 m
Peleteria Robineau-Desvoidy, 1830
Peleteria sp.—Woodley (2007: 857)

ACALYPTRATAE

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Superfamily NERIOIDEA
Family CYPSELOSOMATIDAE
Latheticomyia Wheeler, 1956
Latheticomyia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]

Family MICROPEZIDAE
Subfamily TAENIAPTERINAE
Grallipeza Rondani, 1850
Grallipeza abeja Marshall, 2013: 53 *DR > TL: Pedernales Prov., P. N. Sierra de Bahoruco, Las Abejas, 1150 m
Grallipeza albiterga Marshall, 2013: 53 *DR > TL: Hato Mayor, Parque Los Haitises, 3 km W Cueva de Arena, 20 m
Grallipeza nigrivittata Marshall, 2013: 69 *DR > TL: Pedernales, Upper Las Abejas, 38 km NNW Cabo rojo, 1350 m
Grallipeza paraplacida Marshall, 2013: 70 *DR > TL: Hato Mayor, Parque Los Haitises, 3 km W Cueva de Arena, 20 m
Grallipeza perezi Marshall, 2013: 72 *DR > TL: Las Abejas, P. N. Sierra de Bahoruco, 1,310 m
Hoplocheiloma Cresson, 1926
Hoplocheiloma hispaniola Marshall, 2011: 7 *DR > TL: 3.3 km NE Los Arroyos, Pedernales Prov.
Hoplocheiloma maculosum (Loew, 1866). Marshall, 2011: 11
=Hoplocheiloma notitipenne Cresson, 1926: 272; Marshall, 2011: 11
Hoplocheiloma perforatum Enderlein, 1922. Marshall, 2011: 15
Rainieria Rondani, 1843
Rainieria sp.†—Poinar & Poinar (1999: 207) [Misspelled Raineria] [Fossil in Dominican amber]
Systellapha Enderlein, 1922
Systellapha placida (Loew, 1866). Cresson, 1930: 323
Taeniaptera Macquart, 1835
Taeniaptera lasciva (Fabricius, 1798). Wolcott, 1927: 328; Paes de Albuquerque, 1980: 663.
Note: The species Grallipeza nebulosa (Loew, 1866) has been restricted to the southeastern United States (Marshall, 2013).
Therefore the record by Steyskal, 1968a: 48.7 for Hispaniola and cited by Perez-Gelabert, 2008: 168 has to represent a different
species.

Family NERIIDAE
Subfamily NERIINAE
Nerius Fabricius, 1805
Nerius pilifer Fabricius, 1805. Cresson, 1930: 314
Nerius plurivittatus Bigot, 1886. Steyskal, 1968: 49.5
=Nerius robusta (Enderlein, 1922). Cresson, 1930: 315

Superfamily CONOPOIDEA
Family CONOPIDAE
Occemyia Marschall, 1873
Occemyia haitiensis Parsons, 1940: 35. Aczél, 1950: 80 *H > TL: Port-au-Prince
Physocephala Schiner, 1861
Physocephala venusta Parsons, 1940: 32 *H > TL: Manneville
Physoconops Szilady, 1926
Physoconops pictus (Fabricius, 1775). Parsons, 1940: 32; Camras, 1955: 166; Aczél, 1950: 60
Stylogaster Macquart, 1935
Stylogaster iviei Camras, 1992: 84 *DR > TL: Prov. Pedernales, 24 km N Cabo Rojo, 610 m
Stylogaster sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]

Superfamily DIOPSOIDEA
Family SYRINGOGASTRIDAE
Syringogaster Cresson, 1912
Syringogaster miocenecus† Grimaldi in Marshall, Buck, Skevington & Grimaldi, 2009: 71 [Fossil in Dominican amber]
Syringogaster craigi† Grimaldi in Marshall, Buck, Skevington & Grimaldi, 2009: 75 [Fossil in Dominican amber]

Family TANYPEZIDAE

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Neotanypeza Hendel, 1903
Neotanypeza dominicana† Lonsdale & Apigian, 2010: 215 [Fossil in Dominican amber]

Superfamily TEPHRITOIDEA
Family LONCHAEIDAE
Dasiops Rondani, 1856
Dasiops sp.—SEA (1999: 84)
Lonchaea Fallén, 1820
Lonchaea chalybea Wiedemann, 1830. Santoro, 1960: 389
Neosilba McAlpine, 1962
Neosilba perezi (Romero S. & Ruppel, 1973). McAlpine & Steyskal, 1982: 126

Family RICHARDIIDAE
Epiplatea Loew, 1868
Epiplatea erosa Loew, 1867. Steyskal, 1958: 306
Johnrichardia Perez-Gelabert & Thompson, 2006
Johnrichardia vockerothi Perez-Gelabert & Thompson, 2006: 28 *DR > TL: La Ciénaga—Los Tablones, Parque Nacional
Armando Bermúdez, La Vega Prov., 1100–1270 m

Family TEPHRITIDAE
Subfamily BLEPHARONEURINAE
Ceratodacus Hendel, 1914
Ceratodacus priscus† Norrbom & Condon, 2000: 141 [Fossil in Dominican amber]
Problepharoneura† Norrbom & Condon, 2000
Problepharoneura antiqua† Norrbom & Condon, 2000: 144 [Fossil in Dominican amber]
Subfamily DACINAE
Tribe CERATIDINI
Ceratitis MacLeay, 1829
Ceratitis capitata (Wiedemann, 1824). Valdez, Matos & Álvarez, 2016: 162 (Introduced, Erradicated]
Subfamily TACHINISCINAE
Tribe ORTALOTRYPETINI
Protortalotrypeta† Norrbom, 1994
Protortalotrypeta grimaldii† Norrbom, 1994: 4 [Fossil in Dominican amber]
Subfamily TEPHRITINAE
Tribe ACROTAENIINI
Acrotaenia Loew, 1873
Acrotaenia spadix Bates, 1934: 10. Aczél, 1949: 269
Acrotaenia testudinea (Loew, 1873). Foote, 1960: 107
Tetreuaresta Hendel, 1928
Tetreuaresta obscuriventris (Loew, 1873). Perez-Gelabert, 2008: 169
Tetreuaresta timida (Loew, 1862). Perez-Gelabert, 2008: 169
Tomoplagia Coquillett, 1910
Tomoplagia incompleta (Williston, 1896). Perez-Gelabert, 2008: 169
Tribe EUTETRINI
Polymorphomyia Snow, 1894
Polymorphomyia basilica Snow, 1894. Bates, 1933: 163; Aczél, 1949: 249
Tribe TEPHRITINI
Acinia Robineau-Desvoidy, 1830
Acinia picturata (Snow, 1894). Perez-Gelabert, 2008: 169
Dioxyna Frey, 1945
Dioxyna picciola (Bigot, 1857). Perez-Gelabert, 2008: 169
Dioxyna thomae (Curran, 1928). Perez-Gelabert, 2008: 169
Dyseuaresta Hendel, 1928
Dyseuaresta mexicana (Wiedemann, 1830). Perez-Gelabert, 2008: 169

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Euaresta Loew, 1873
Euaresta sp. poss. bella (Loew, 1862) [By synonymy of species below]
=Euaresta sp.—Norrbom (1993: 207)
Euarestoides Benjamin, 1934
Euarestoides acutangulus (Thomson, 1869). Savaris et al., 2019: 305
Trupanea Schrank, 1795
Trupanea eclipta Benjamin, 1934. Perez-Gelabert, 2008: 170
Tribe INCERTAE SEDIS
Xanthaciura Hendel, 1914
Xanthaciura chrysura (Thomson, 1869). Perez-Gelabert, 2008: 170
Xanthaciura connexionis Benjamin, 1934. Perez-Gelabert, 2008: 170
Xanthaciura insecta (Loew, 1862). Bates, 1933: 163; Aczél, 1951: 127
Subfamily TRYPETINAE
Tribe TOXOTRYPANINI
Anastrepha Schiner, 1868
Anastrepha antilliensis Norrbom, 1998: 173
Anastrepha curvicauda (Gerstaecker, 1860). Wolcott, 1927: 327; Smith & Audant, 1930: 976; Norrbom et al., 2018: 838
Anastrepha dissimilis Stone, 1942: 7. Aczél, 1949: 207; Hernández-Ortíz & Aluja, 1993: unpaginated appendix [Introduced]
Anastrepha insulae Stone, 1942. Hernández-Ortíz & Aluja, 1993: unpaginated appendix
Anastrepha obliqua (Macquart, 1835). Stone, 1942: 7 (as A. mombipraeoptans); Schmutterer, 1990: 238; Hernández-Ortíz &
Aluja, 1993: unpaginated appendix [Introduced]
Anastrepha ocresia (Walker, 1849). Stone, 1942: 7; Aczél, 1949: 221; Weems, 1968: 1; Hernández-Ortíz & Aluja, 1993:
unpaginated appendix [Introduced]
=Anastrepha tricincta (Loew, 1873). Bates, 1933: 161; Greene, 1934: 146
Anastrepha stonei Steyskal, 1977. Norrbom, 1998: 188
Anastrepha suspensa (Loew, 1862). Stone, 1942: 7; Aczél, 1949: 230; Meyer & Knight, 1998: 235; Hernández-Ortíz & Aluja,
1993: unpaginated appendix [Introduced]
Tribe TRYPETINI
Parastenopa Hendel, 1914
Parastenopa sp.—Perez-Gelabert (2008: 170)
Rhagoletotrypeta Aczél, 1950
Rhagoletotrypeta gelabertae Norrbom & Savaris in Norrbom et al., 2016: 552 *DR > TL: Boca de Yuma, Parque Nacional del
Este
Tribe INCERTAE SEDIS
Alujamyia Norrbom, 2006
Alujamyia isolata Norrbom, 2006: 48
Pseudophorellia Lima, 1934
Pseudophorellia flavida Norrbom, 2006: 100 *DR > TL: Pedernales, Las Abejas, 38 km NNW Cabo Rojo, 1160 m

Family ULIDIIDAE
Subfamily ULIDIINAE
Acrosticta Loew, 1867
Acrosticta apicalis (Williston, 1896). Steyskal, 1968: 54.13
Eumecosomyia Hendel, 1909
Eumecosomyia nubila (Wiedemann, 1830). Steyskal, 1968: 54.15
Euxesta Loew, 1868
Euxesta annonae (Fabricius, 1794). Steyskal, 1968: 54.16
Euxesta stigmatias Loew, 1868. App, 1938: 182
Neoacanthoneura Hendel, 1914
Neoacanthoneura trypetina Kameneva, 2012: 32 *DR > TL: Barahona, 9.2 km NW Paraiso, confluence of Río Nizao and Río
Cortico, 230 m

Superfamily LAUXANIOIDEA
Family LAUXANIIDAE

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Camptoprosopella Hendel, 1907
Camptoprosopella diversa Curran, 1926: 13. Curran, 1928: 82
Neogriphoneura Malloch, 1924
Neogriphoneura striatifrons Hendel or N. striga Curran (undetermined). De Mello & Silva, 2008: 45

Superfamily SCIOMYZOIDEA
Family SCIOMYZIDAE
Dominimyza† Poinar, 2013
Dominimyza tanyacaena† Poinar, 2013: 431 [Fossil in Dominican amber]
Sepedomerus Steyskal, 1973
Sepedomerus caeruleus (Melander, 1920: 332). Knutson et al., 1976: 64.10

Family SEPSIDAE
Subfamily SEPSINAE
Archisepsis Silva, 1993
Archisepsis armata (Schiner, 1868). Steyskal, 1968: 63.2; Silva, 1993: 131
=Sepsis hoplicnema Melander & Spuler, 1917: 17. Steyskal, 1968: 63.2; Ozerov, 2005: 11
Archisepsis discolor (Bigot, 1857). Ozerov, 2005: 12
Archisepsis ecalcarata (Thomson, 1869). Ozerov, 2005: 12
=Sepsis insularis Williston, 1896. Steyskal, 1968: 63.2
Archisepsis pusio (Schiner, 1868). Ozerov, 2005: 12
Meropliosepsis Duda, 1926
Meropliosepsis sexsetosa Duda, 1926. Ozerov, 2005: 20
Microsepsis Silva, 1993
Microsepsis armillata (Melander & Spuler, 1917: 18). Steyskal, 1968: 63.2; Silva, 1993: 141
Microsepsis furcata (Melander & Spuler, 1917). Ozerov, 2005: 23
Microsepsis mitis (Curran, 1927). Ozerov, 2005: 23
Microsepsis simplicula (Steyskal, 1968). Ozerov, 2005: 23

Superfamily OPOMYZOIDEA
Family AGROMYZIDAE
Subfamily AGROMYZINAE
Agromyza Fallén, 1810
Agromyza parvicornis Loew, 1869. Martinez et al., 1993: 167
Japanagromyza Sasakawa, 1958
Japanagromyza perpetua Spencer, 1973. Martinez et al., 1993: 168
Melanagromyza Hendel, 1920
Melanagromyza compositoides Spencer, 1963. Martinez et al., 1993: 168
Melanagromyza consueta Spencer, 1973. Martinez et al., 1993: 168
Melanagromyza diadema (Melander, 1913: 259). Spencer, 1963: 311
Melanagromyza floridensis Spencer, 1963: 312. Martinez et al., 1993: 168
Melanagromyza minima (Malloch, 1913). Martinez et al., 1993: 168
Melanagromyza obtusa (Malloch, 1914). Etienne et al., 2004: 105 [Introduced]
Melanagromyza occulta Spencer, 1973. Martinez et al., 1993: 170
Melanagromyza sp. 1—Martinez et al. (1993: 170)
Melanagromyza sp. 2—Martinez et al. (1993: 170)
Melanagromyza sp. 3—Martinez et al. (1993: 170)
Melanagromyza sp. 4—Martinez et al. (1993: 171)
Nemorimomyza Hendel, 1931
Nemorimomyza maculosa (Malloch, 1913). Etienne & Martinez, 2003: 94
Ophiomyia Braschnikov, 1897
Ophiomyia gentilis Spencer, 1973. Martinez et al., 1993: 171
Ophiomyia sp. 1—Martinez et al. (1993: 172)
Ophiomyia sp. 2—Martinez et al. (1993: 172)

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Ophiomyia texana (Malloch, 1913). Martinez et al., 1993: 171
Subfamily PHYTOMYZINAE
Amauromyza Hendel, 1931
Amauromyza (Annimyzella) maculosa (Malloch, 1913). Martinez et al., 1993: 176
Calycomyza Hendel, 1931
Calycomyza allecta (Melander, 1913). Spencer, 1963: 338
Calycomyza eupatorivora Spencer, 1973. Martinez et al., 1993: 177
Calycomyza ipomaea (Frost, 1931). Spencer, 1963: 345; Martinez et al., 1993: 177
Calycomyza sidae Spencer, 1973. Martinez et al., 1993: 177
Calycomyza stegmaieri Spencer, 1966. Martinez et al., 1993: 178
Liriomyza Mik, 1894
Liriomyza blechi Spencer, 1973. Martinez et al., 1993: 173
Liriomyza brassicae (Riley, 1885). SEA, 1999: 27
Liriomyza huidobrensis (Blanchard, 1926). Martinez et al., 1993: 173; Zuluaga & Mesa, 2000: 197 [Introduced]
Liriomyza marginalis (Malloch, 1913). Martinez et al., 1993: 174
Liriomyza sativae Blanchard, 1938. Martinez et al., 1993: 174
Liriomyza sp.—Martinez et al. (1993: 175)
Liriomyza trifolii (Burgess, 1880). Schmutterer, 1990: 470; Martinez et al., 1993: 175; Zuluaga & Mesa, 2000: 197
[Introduced]
Liriomyza tubifer Melander, 1913: 266. Spencer, 1963: 372

Family ANTHOMYZIDAE
Anthomyza Fallén, 1810
Anthomyza sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]

Family ASTEIIDAE
Asteia Meigen, 1830
Asteia expansa Sabrosky, 1957. Sabrosky, 1966: 92.3
Asteia sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]
Loewimyia Sabrosky, 1943
Loewimyia sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]

Family AULACIGASTRIDAE
Aulacigaster Macquart, 1835
Aulacigaster grimaldii Rung & Mathis, 2011: 49
Aulacigaster sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]

Family CLUSIIDAE
Subfamily CLUSIINAE
Chaetoclusia Coquillett, 1904
Chaetoclusia furva Lonsdale & Marshall, 2006a: 180
Chaetoclusia nigromaculata Melander & Argo, 1924. Lonsdale & Marshall, 2006a: 172 *H > TL: Hayti
Clusiodes Coquillett, 1904
Clusiodes sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Craspedochaeta Czerny, 1903
Craspedochaeta annulipes (Johnson, 1913). Lonsdale & Marshall, 2006b: 59
=Czernyola piceoflava (Soós, 1962: 146) [from Dominican Republic, Not Dominica]
Craspedochaeta apsilutea Lonsdale & Marshall, 2006b: 19
Craspedochaeta pleuralis (Williston, 1896). Lonsdale & Marshall, 2006b: 44
Subfamily CLUSIODINAE
Heteromeringia Czerny, 1903
Heteromeringia czernyi Kertesz, 1903. Lonsdale & Marshall, 2007: 54
Heteromeringia fumipennis Melander & Argo, 1924. Lonsdale & Marshall, 2007: 45
Subfamily SOBAROCEPHALINAE

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 225
Sobarocephala Czerny, 1903
Sobarocephala atrifacies Sabrosky & Steyskal, 1974. Lonsdale & Marshall, 2012: 161
Sobarocephala biovata Lonsdale & Marshall, 2012: 104
Sobarocephala bivittata Melander & Argo, 1924. Lonsdale & Marshall, 2012: 106
Sobarocephala bucki Lonsdale & Marshall, 2012: 41
Sobarocephala bulbipenis Lonsdale & Marshall, 2012: 107
Sobarocephala hyalina Lonsdale & Marshall, 2012: 73
Sobarocephala lucibilis Lonsdale & Marshall, 2012: 92
Sobarocephala panamaensis Soós, 1965. Lonsdale & Marshall, 2012: 175
Sobarocephala plumatella Melander & Argo, 1924. Lonsdale & Marshall, 2012: 50
Sobarocephala pruinosa Soós, 1962. Lonsdale & Marshall, 2012: 208

Family ODINIIDAE
Odinia Robineau-Desvoidy, 1830
Odinia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]

Family PERISCELIDIDAE
Periscelis Loew, 1858
Periscelis (Myodris) amberifera† Grimaldi & Mathis, 1993: 387 [Fossil in Dominican amber]
Periscelis (Myodris) brodzinskyi† Grimaldi & Mathis, 1993: 391 [Fossil in Dominican amber]
Periscelis (Myodris) fascianota† Grimaldi & Mathis, 1993: 389 [Fossil in Dominican amber]
Planinasus Cresson, 1914
Planinasus insularis Mathis, Rung & Kotrba, 2012: 52
Planinasus electrus† Grimaldi & Mathis, 1993: 396; Mathis, Rung & Kotrba, 2012: 79 [Fossil in Dominican amber]
Stenomicra Coquillett, 1900
Stenomicra anacrostichalis† Grimaldi & Mathis, 1993: 401 [Fossil in Dominican amber]
Stenomicra sabroskyi† Grimaldi & Mathis, 1993: 399 [Fossil in Dominican amber]

Superfamily CARNOIDEA
Family CANACIDAE
Subfamily CANACINAE
Canacea Malloch, 1924
Canacea macateei Malloch, 1924. Munari & Mathis, 2010: 14
Subfamily NOCTICANACINAE
Paracanace Mathis & Wirth, 1978
Paracanace aicen Mathis & Wirth, 1978. Mathis, 1997: 142
Subfamily TETHININAE
Tethina Haliday in Curtis, 1837
Tethina albula (Loew, 1869). Foster & Mathis, 2008: 745; Munari & Mathis, 2010: 50
Tethina cohiba Foster & Mathis, 1998: 625
Tethina willistoni (Melander, 1913). Foster & Mathis, 1998: 615
Tethina xanthopoda (Williston, 1896). Foster & Mathis, 1998: 20

Family CARNIDAE
Meoneura Rondani, 1856
Meoneura vieja† Grimaldi, 1997: 16 [Fossil in Dominican amber]

Family CHLOROPIDAE
Subfamily OSCINELLINAE
Apallates Sabrosky, 1980
Apallates convexus (Loew, 1866). Sabrosky & Paganelli, 1984: 81.5
Apallates lutzi (Curran, 1926). Sabrosky & Paganelli, 1984: 81.6
Elachiptera Macquart, 1835
Elachiptera attenuata (Adams, 1908). Sabrosky & Paganelli, 1984: 81.13

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Hippelates Loew, 1863
Hippelates brumpti Séguy, 1940. Sabrosky & Paganelli, 1984: 81.17
Liohippelates Duda, 1929
Liohippelates currani (Aldrich, 1931). Sabrosky & Paganelli, 1984: 81.20
Liohippelates flavipes (Loew, 1866). Wolcott, 1922: 36; Smith & Audant, 1930: 976; Santoro, 1960: 452; Sabrosky &
Paganelli, 1984: 81.20
Liohippelates peruanus (Becker, 1912). Sabrosky & Paganelli, 1984: 81.21
=Siphonella circumdata Becker, 1912: 185
Liohippelates pusio (Loew, 1872). Sabrosky & Paganelli, 1984: 81.21
Monochaetoscinella Duda, 1930
Monochaetoscinella anonyma (Williston, 1896). Sabrosky & Paganelli, 1984: 81.24
Olcella Enderlein, 1911
Olcella trilineata (Duda, 1930). Sabrosky & Paganelli, 1984: 81.26
Pseudogaurax Malloch, 1915
Pseudogaurax lancifer (Coquillett, 1900). Wolcott, 1927: 329; Sabrosky & Paganelli, 1984: 81.29
Rhodesiella Adams, 1905
Rhodesiella spp.—Sabrosky & Paganelli (1984: 81.31)
Rhopalopterum Duda, 1929
Rhopalopterum limitatum (Becker, 1912: 211). Curran, 1928: 53; Sabrosky & Paganelli, 1984: 81.31
Siphunculina Rondani, 1856
Siphunculina striolata (Wiedemann, 1830). Sabrosky & Paganelli, 1984: 81.31
Tricimba Lioy, 1864
Tricimba sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]

Family MILICHIIDAE
Subfamily MADIZINAE
Desmometopa Loew, 1866
Desmometopa tarsalis Loew, 1866. Sabrosky, 1983: 56
Subfamily MILICHIINAE
Milichiella Giglio-Tos, 1895
Milichiella archaia† Brake, 2006: 19 [Fossil in Dominican amber]
Milichiella cinerea (Coquilett, 1900). Sabrosky, 1973: 75.4; Brake, 2009: 65
Milichiella dolichosurstyla† Brake, 2006: 21 [Fossil in Dominican amber]
Milichiella dominicana† Brake, 2006: 21 [Fossil in Dominican amber]
Milichiella hennigi† Brake, 2006: 23 [Fossil in Dominican amber]
Milichiella lacteipennis (Loew, 1866). Sabrosky, 1973: 75.4
Milichiella margaretae† Brake, 2006: 24 [Fossil in Dominican amber]
Milichiella punctata Brake, 2009: 138 *DR > TL: La Ciénaga
Milichiella quadrisetosa† Brake, 2006: 25 [Fossil in Dominican amber]
Milichiella theodori† Brake, 2006: 25 [Fossil in Dominican amber]
Pholeomyia Bilimek, 1867
Pholeomyia myopa Melander, 1913: 238. Sabrosky, 1959: 325; Brake, 2000: 108
Pholeomyia politifacies Sabrosky, 1959: 323
Pholeomyia sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Subfamily PHYLLOMYZINAE
Phyllomyza Fallén, 1810
Phyllomyza sp.†—Poinar (1992: 186) [Fossil in Dominican amber]

Superfamily SPHAEROCEROIDEA
Family SPHAEROCERIDAE
Subfamily ARCHIBORBORINAE
Poecilantrops Kits & Marshall, 2013
Poecilantrops baorucensis Kits & Marshall, 2013: 96 *DR > TL: Las Abejas, 30 km N of Cabo Rojo, 1300 m
Poecilantrops dominicus Kits & Marshall, 2013: 99 *DR > TL: Independencia: La Descubierta, 32 km NW Sabana Real, 1800

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m
Subfamily LIMOSININAE
Aptilotella Duda, 1924
Aptilotella caerulea Luk & Marshall, 2014: 18 *DR > TL: Independencia, 32 km NW La Descubierta, Sabana Real, 1800 m
Bifronsina Roháček, 1983
Bifronsina bifrons (Stenhammar, 1855). Marshall et al., 2011: 235
Bromeloecia Spuler, 1924
Bromeloecia aculatus Yau and Marshall, 2018: 85
Bromeloecia ephippium Yau and Marshall, 2018: 95
Bromeloecia wolverinei Yau and Marshall, 2018: 74
Coproica Rondani, 1861
Coproica acutangula (Zetterstedt, 1847). Roháček et al., 2001: 135; Marshall et al., 2011: 238; Bergeron et al., 2015: 18
Coproica bispatha Bergeron et al., 2015: 20
Coproica brachystyla Bergeron et al., 2015: 22
Coproica diabolica Bergeron et al., 2015: 24
Coproica emarginata Bergeron et al., 2015: 27
Coproica rufifrons Hayashi, 1991. Bergeron et al., 2015: 37
Coproica setulosa (Duda, 1929). Roháček et al., 2001: 141; Bergeron et al., 2015: 38
Dahlimosina Roháček, 1983
Dahlimosina sp. 1†—Marshall et al. (1999: 299) [Fossil in Dominican amber]
Druciatus Marshall, 1995
Druciatus ovisternus Marshall, 1995: 287. Roháček et al., 2001: 143 *DR > TL: La Ciénaga
Lepidosina Marshall & Buck, 2007
Lepidosina angusticercus Marshall & Buck in Marshall et al., 2007: 581; Marshall et al., 2011: 244
Lepidosina gibba (Spuler, 1925). Marshall et al., 2011: 244
Leptocera Olivier, 1813
Leptocera erythrocera (Becker, 1920). Roháček et al., 2001: 153; Buck & Marshall, 2009: 34; Marshall et al., 2011: 246
Leptocera fulva (Malloch, 1912). Roháček et al., 2001: 154; Buck & Marshall, 2009: 104; Marshall et al., 2011: 247
Leptocera insularum Buck & Marshall, 2009: 109; Marshall et al., 2011: 247
Minilimosina Roháček, 1983
Minilimosina (Svarciella) niveipennis (Malloch, 1913). Roháček & Marshall, 1988: 257; Roháček et al., 2001: 169
Opacifrons Duda, 1918
Opacifrons quarta Marshall & Langstaff, 1998. Roháček et al., 2001: 178
Opalimosina Roháček, 1983
Opalimosina sp. 1†—Marshall et al. (1999: 300) [Fossil in Dominican amber]
Opalimosina sp. 2†—Marshall et al. (1999: 300) [Fossil in Dominican amber]
Phthitia Enderlein, 1938
Phthitia (Kimosina) antillensis Marshall 1992: 16. Roháček et al., 2001: 192 *DR > TL: La Ciénaga, at base of trail to Pico
Duarte
Poecilosomella Duda, 1925
Poecilosomella angulata (Thomson, 1869). Roháček et al., 2001: 200; Marshall et al., 2011: 259
Pterogramma Spuler, 1924
Pterogramma adustum Smith & Marshall, 2004: 36. Marshall et al., 2011: 265 *DR > TL: La Ciénaga
Pterogramma aquatile Smith & Marshall, 2004: 45; Marshall et al., 2011: 265
Pterogramma lobosternum Smith & Marshall, 2004: 47; Marshall et al., 2011: 266
Pterogramma substitutum (Richards, 1961). Roháček et al., 2001: 215; Marshall et al., 2011: 267
Pullimosina Roháček, 1983
Pullimosina (Pullimosina) zayensis Marshall, 1986. Roháček et al., 2001: 222; Marshall et al., 2011: 269
Rachispoda Lioy, 1864
Rachispoda amarilla Wheeler in Wheeler & Marshall, 1995: 1252. Roháček et al., 2001: 225 *DR > TL: Manabao, sweep near
creek
Rachispoda fuscinervis (Malloch, 1912). Wheeler & Marshall, 1995: 1260; Roháček et al., 2001: 231
Rachispoda m-nigrum (Malloch, 1912). Wheeler & Marshall, 1995: 1304; Roháček et al., 2001: 135
Rachispoda paludicola Wheeler in Wheeler & Marshall, 1995: 1225. Roháček et al., 2001: 241

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Rachispoda rhizophora Wheeler, 1995. Roháček et al., 2001: 243
Robustagramma Marshall & Cui, 2005
Robustagramma multiseta Cui & Marshall in Marshall & Cui, 2005: 92. Marshall et al., 2011: 275 *DR > TL: Cabo Rojo (of
Pedernales), Alcoa rd. km 26
Sclerocoelus Marshall, 1995
Sclerocoelus caribensis Marshall, 1997: 102. Roháček et al., 2001: 249
Spelobia Spuler, 1924
Spelobia (Bifronsina) bifrons (Stenhammar, 1855). Roháček et al., 2001: 253
Spelobia palaeobrevipteryx† Marshall in Marshall et al., 1999: 296 [Fossil in Dominican amber]
Spinilimosina Roháček, 1983
Spinilimosina brevicostata (Duda, 1918). Marshall et al., 2011: 279
Telomerina Roháček, 1983
Telomerina orpha Marshall & Roháček, 1984. Roháček et al., 2001: 270
Telomerina sp. 1—Marshall et al. (1999: 296)
Xenolimosina Roháček, 1983
Xenolimosina palaeospinosa† Marshall in Marshall et al., 1999: 297 [Fossil in Dominican amber]
Subfamily SPHAEROCERINAE
Parasphaerocera Spuler, 1924
Parasphaerocera varipes (Malloch, 1925). Roháček et al., 2001: 103
Sphaerocera Latreille, 1804
Sphaerocera flaviceps Malloch, 1925. Roháček et al., 2001: 107

Superfamily EPHYDROIDEA
Family CURTONOTIDAE
Curtonotum Macquart, 1843
Curtonotum electrodominicum† Grimaldi & Kirk-Spriggs, 2012: 3 [Fossil in Dominican amber]
Depressonotum† Grimaldi & Kirk-Spriggs, 2012
Depressonotum priscum† Grimaldi & Kirk-Spriggs, 2012: 10 [Fossil in Dominican amber]

Family DROSOPHILIDAE
Subfamily DROSOPHILINAE
Chymomyza Czerny, 1903
Chymomyza laevilimbata Duda, 1927. Wheeler, 1970: 79.7
Chymomyza primaeva† Grimaldi, 1987: 3 [Fossil in Dominican amber]
Cladochaeta Coquillett, 1900
Cladochaeta abeja Grimaldi & Nguyen, 1999: 92 *DR > TL: Pedernales: Upper Las Abejas, 38 km NNW Cabo Rojo, 1350 m
Cladochaeta abrupta Grimaldi & Nguyen, 1999: 93 *DR > TL: Pedernales Prov.: 20 km NE Pedernales, Las Abejas
Cladochaeta calvovis Grimaldi & Nguyen, 1999: 274 *DR > TL: La Vega, 9 km SE Constanza, near Valle Nuevo, 1930 m
Cladochaeta dejecta Grimaldi & Nguyen, 1999: 143 *DR > TL: La Vega Prov., 7.2 mi S Constanza, on road to San José de
Ocoa, 5000 ft.
Cladochaeta dominicana Grimaldi & Nguyen, 1999: 15 *DR > TL: Pedernales Prov.: 20 km NE Pedernales, 1500 m
Cladochaeta dominitica Grimaldi & Nguyen, 1999: 144 *DR > TL: Pedernales Prov.: 20 km NE Pedernales, 1500 m
Cladochaeta masneri Grimaldi & Nguyen, 1999: 22 *H > TL: Kenscoff
Cladochaeta nebulosa Coquillett, 1900. Wheeler, 1957: 103
Cladochaeta spira Grimaldi & Nguyen, 1999: 28 *DR > TL: Pedernales: 9.7 km NE Los Arroyos, 2070 m
Cladochaeta vapida Grimaldi & Nguyen, 1999: 217 *DR > TL: La Vega, 24 km E El Río
Clastopteromyia Malloch, 1924
Clastopteromyia (Clastopteromyia) aberrans Wheeler, 1957: 107. Wheeler, 1970: 79.8 *H > TL: Kenscoff (4000 ft.)
Drosophila Fallén, 1823
Drosophila (Drosophila) acutilabella Stalker, 1953. Heed, 1962: 174; Wheeler, 1970: 79.11
Drosophila (Drosophila) annularis Sturtevant, 1916. Markow & O’Grady, 2006: 156
Drosophila (Drosophila) bromeliae Sturtevant, 1921. Markow & O’Grady, 2006: 156
Drosophila (Drosophila) carcinophila Wheeler, 1960. Carson, 1967: 330; Markow & O’Grady, 2006: 156
Drosophila (Drosophila) cardini Sturtevant, 1916: 336. Sturtevant, 1921: 78; Stalker, 1953: 351

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Drosophila (Drosophila) lauta Wheeler & Takada in Wheeler et al., 1962: 403. Wheeler, 1970: 79.16 *H > TL: Kenscoff pines,
ca. 5500–6000 ft.
Drosophila (Drosophila) mayaguana Vilela, 1983. Heed & Grimaldi, 1991: 10
Drosophila (Drosophila) mediodiffusa Heed & Wheeler, 1957: 27. Wheeler, 1970: 79.17
Drosophila (Drosophila) micromettleri Heed, 1989: 276
Drosophila (Drosophila) mulleri Sturtevant, 1939. Vilela, 1983: 65
Drosophila (Drosophila) nesiota Wheeler & Takada in Wheeler et al., 1962: 409. Wheeler, 1970: 79.19 *H > TL: Near
Pétionville
Drosophila (Drosophila) ornatipennis Williston, 1896. Wheeler, 1957: 85; Wheeler, 1970: 79.20
Drosophila (Drosophila) paraguttata Thompson in Wheeler, 1957. Markow & O’Grady, 2006: 161
Drosophila (Drosophila) parisiena Heed & Grimaldi, 1991: 3
Drosophila (Drosophila) peninsularis Patterson & Wheeler, 1942. Ruiz & Wasserman, 1992: 585
Drosophila (Drosophila?) poinari† Grimaldi, 1987: 7 [Fossil in Dominican amber]
Drosophila (Drosophila) repleta Wollaston, 1858. Sturtevant, 1921: 99
Drosophila (Drosophila) similis Williston, 1896. Sturtevant, 1921: 79; Stalker, 1953: 352; Wheeler, 1970: 79.22
Drosophila (Drosophila) straubae Heed & Grimaldi, 1991: 9
Drosophila (Drosophila?) succini† Grimaldi, 1987: 5 [Fossil in Dominican amber]
Drosophila (Hirtodrosophila) palaeothoracis† Grimaldi, 1987: 9 [Fossil in Dominican amber]
Drosophila (Hirtodrosophila) prognatha Sturtevant, 1916. Sturtevant, 1921: 75; Wheeler, 1970: 79.24
Drosophila (Siphlodora) flexa Loew, 1865. Wheeler, 1957: 79; Wheeler, 1970: 79.25
Drosophila (Sophophora) ananassae Doleschall, 1858. Hernández Martich, 1982: 40/81
Drosophila (Sophophora) caribea Sturtevant, 1916: 335. Sturtevant, 1921: 92
Drosophila (Sophophora) equinoxialis Dobzhansky, 1946. Spassky et al., 1971: 132
Drosophila (Sophophora) lusaltans Magalhães, 1962: 140. Wheeler, 1970: 79.27 *H > TL: Pétionville
Drosophila (Sophophora) melanogaster Meigen, 1830. Yukilevich & True, 2008: 2809
Drosophila (Sophophora) simulans Sturtevant, 1919. Yoon et al., 1990: 18
Drosophila (Sophophora) nebulosa Sturtevant, 1916: 327. Sturtevant, 1921: 88
Drosophila (Sophophora) neoalagitans Wheeler & Magalhães, 1962: 162. Wheeler, 1970: 79.27
Drosophila (Sophophora) sturtevanti Duda, 1927. Magalhães, 1962: 143
Drosophila (Sophophora) tolteca Patterson & Mainland, 1944. Sulerud & Miller, 1966: 447
Drosophila (Sophophora) tropicalis Burla and da Cunha in Burla et al., 1949. Spassky et al., 1971: 132
Drosophila (Sophophora) tropicalis cubana Townsend, 1954.Wheeler, 1970: 79.28
Drosophila (Sophophora) willistoni Sturtevant, 1916: 327. Sturtevant, 1921: 89
Drosophila sp. A†—Grimaldi (1987: 20) [Fossil in Dominican amber]
Drosophila sp. B†—Grimaldi (1987: 20) [Fossil in Dominican amber]
Drosophila bromeliae Sturtevant, 1921. Grimaldi, 2016: 16
Drosophila penispina Grimaldi, 2016: 37
Drosophila stylipennis Grimaldi, 2016: 47
Drosophila starki Grimaldi, 2016: 48
Drosophila unnamed species 5—Wheeler et al. (1962: 412)
Mycodrosophila Oldenberg, 1914
Mycodrosophila projectans (Sturtevant, 1916: 342). Sturtevant, 1921: 63; Wheeler, 1970: 79.31
Miomyia Grimaldi, 1987
Miomyia io† Grimaldi, 1987: 12 [Fossil in Dominican amber]
Neotanygastrella Duda, 1925
Neotanygastrella tricoloripes Duda, 1925. Wheeler, 1970: 79.32
Neotanygastrella wheeleri† Grimaldi, 1987: 14 [Fossil in Dominican amber]
Paramycodrosophila Duda, 1924
Paramycodrosophila costaricana Duda, 1925. Wheeler, 1970: 79.32
Protochymomyza† Grimaldi, 1987
Protochymomyza miocena† Grimaldi, 1987: 15 [Fossil in Dominican amber]
Scaptomyza Hardy, 1849
Scaptomyza dominicana† Grimaldi, 1987: 16 [Fossil in Dominican amber]
Scaptomyza (Mesoscaptomyza) coquilletti Wheeler & Takada, 1967. Wheeler, 1970: 79.32

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Scaptomyza (Mesoscaptomyza) paravittata Wheeler, 1952. Wheeler, 1970: 79.32
Zygothrica Wiedemann, 1830
Zygothrica circumveha Grimaldi, 1987a: 175
Zygothrica (Zygothrica) bilineata (Williston, 1896). Wheeler, 1970: 79.35
Zygothrica (Zygothrica) vittatifrons (Williston, 1896). Wheeler, 1970: 79.35
Subfamily STEGANINAE
Hyalistata Wheeler, 1960
Hyalistata vitrea† Grimaldi, 1993: 14 [Fossil in Dominican amber]
Leucophenga Mik, 1886
Leucophenga (Leucophenga) elegans Duda, 1927. Wheeler, 1970: 79.3
Leucophenga (Leucophenga) frontalis (Williston, 1896). Wheeler, 1970: 79.3
Leucophenga maculosa (Coquillett, 1895). Sturtevant, 1916: 324; Sturtevant, 1921: 60
Stegana Meigen, 1830
Stegana sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]

Family EPHYDRIDAE
Subfamily DISCOMYZINAE
Tribe DISCOMYZINI
Clasiopella Hendel, 1914
Clasiopella uncinata Hendel, 1914. Perez-Gelabert, 2008: 176
Guttipsilopa Wirth, 1956
Guttipsilopa (Nesopsilopa) bahamaensis (Mathis & Wirth, 1977). Perez-Gelabert, 2008: 176
Guttipsilopa (Nesopsilopa) stonei (Mathis & Wirth, 1977). Perez-Gelabert, 2008: 177
Guttipsilopa (Nesopsilopa) umbrosa (Loew, 1862). Perez-Gelabert, 2008: 177
Paratissa Coquillett, 1900
Paratissa semilutea (Loew, 1869). Perez-Gelabert, 2008: 177
Tribe DISCOCERININI
Discocerina Macquart, 1835
Discocerina chalybea (Hendel, 1930). Perez-Gelabert, 2008: 177
Discocerina flavipes Cresson, 1841. Perez-Gelabert, 2008: 177
Discocerina obscura Williston, 1896. Perez-Gelabert, 2008: 177
Discocerina obscurella (Fallén, 1813). Perez-Gelabert, 2008: 177
Gymnoclasiopa Hendel, 1933
Gymnoclasiopa bella (Mathis, 1997: 700). Mathis & Zatwarnicki, 2012b: 21
Hydrochasma Hendel, 1936
Hydrochasma faciale (Williston, 1896). Mathis & Zatwarnicki, 2013: 23
Hydrochasma dolabrutum Mathis & Zatwarnicki, 2013: 58
Hydrochasma glochium Mathis & Zatwarnicki, 2013: 65
Hydrochasma incisum Mathis & Zatwarnicki, 2013: 68
Hydrochasma peniculum Mathis & Zatwarnicki, 2013: 91
Hydrochasma urnulum Mathis & Zatwarnicki, 2013: 98
Hydrochasma capsum Mathis & Zatwarnicki, 2013: 119
Hydrochasma edmistoni Mathis & Zatwarnicki, 2013: 125
Hydrochasma leucoproctum (Loew, 1861). Mathis & Zatwarnicki, 2013: 132
Hydrochasma robustum Mathis & Zatwarnicki, 2013: 144
Lamproclasiopa Hendel, 1933
Lamproclasiopa nana (Williston, 1896). Wirth, 1968a: 77.7; Costa et al., 2016: 54
Orasiopa Zatwarnicki & Mathis, 2001
Orasiopa mera (Cresson, 1939). Perez-Gelabert, 2008: 177; Mathis et al., 2016: 5/15
Pectinifer Cresson, 1944
Pectinifer aeneus (Cresson, 1918). Perez-Gelabert, 2008: 177; Mathis et al., 2016: 10/15
Polytrichophora Cresson, 1924
Polytrichophora adarca Mathis & Zatwarnicki, 2012a: 30
Polytrichophora agens Cresson, 1924. Perez-Gelabert, 2008: 177; Mathis & Zatwarnicki, 2012a: 37

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Polytrichophora barba Mathis & Zatwarnicki, 2012a: 46
Polytrichophora desmata (Williston, 1896). Perez-Gelabert, 2008: 177; Mathis & Zatwarnicki, 2012a: 74
Polytrichophora reginae Mathis, 1997. Perez-Gelabert, 2008: 177; Mathis & Zatwarnicki, 2012a: 108
Polytrichophora setulosa (Cresson, 1918). Perez-Gelabert, 2008: 177; Mathis & Zatwarnicki, 2012a: 66
Tribe GASTROPINI
Beckeriella Williston, 1897
Beckeriella sp.†—Poinar & Poinar (1999: 207) [Fossil in Dominican amber]
Tribe GYMNOMYZINI
Athyroglossa Loew, 1860
Athyroglossa (Athyroglossa) atra (Williston, 1896). Perez-Gelabert, 2008: 177
Athyroglossa (Athyroglossa) dubia (Williston, 1896). Perez-Gelabert, 2008: 177
Athyroglossa (Athyroglossa) glaphyropus Loew, 1878. Perez-Gelabert, 2008: 177
Athyroglossa (Athyroglossa) nitida Williston, 1896. Perez-Gelabert, 2008: 177
Mosillus Latreille, 1804
Mosillus stegmaieri Wirth, 1969. Perez-Gelabert, 2008: 177
Placopsidella Kertész, 1901
Placopsidella cynocephala Kertész, 1901. Perez-Gelabert, 2008: 177
Tribe HECAMEDINI
Allotrichoma Becker, 1896
Allotrichoma (Pseudohecamede) abdominale (Williston, 1896). Perez-Gelabert, 2008: 177
Allotrichoma (Pseudohecamede) adustum Mathis, 1991. Perez-Gelabert, 2008: 177
Allotrichoma (Pseudohecamede) jamaicense Mathis, 1991. Perez-Gelabert, 2008: 177
Diphuia Cresson, 1944
Diphuia flinti Mathis, 1997: 33 *DR > TL: El Seibo: near Rincón
Hecamede Haliday, 1837
Hecamede (Hecamede) brasiliensis Cresson, 1938. Perez-Gelabert, 2008: 177
Tribe LIPOCHAETINI
Glenanthe Haliday, 1839
Glenanthe caribea Mathis, 1995. Perez-Gelabert, 2008: 177
Glenanthe litorea Cresson, 1925. Perez-Gelabert, 2008: 177
Glenanthe ruetzleri Mathis, 1995. Perez-Gelabert, 2008: 177
Lipochaeta Coquillett, 1896
Lipochaeta slossonae Coquillett, 1896. Mathis & Trautwein, 2003: 39
Paraglenanthe Wirth, 1956
Paraglenanthe bahamensis Wirth, 1956. Perez-Gelabert, 2008: 178
Tribe OCHTHERINI
Ochthera Latreille, 1802
Ochthera exsculpta Loew, 1862. Perez-Gelabert, 2008: 178
Ochthera lauta Wheeler, 1896. Perez-Gelabert, 2008: 178
Ochthera loreta Cresson, 1931. Perez-Gelabert, 2008: 178
Tribe PSILOPINI
Ceropsilopa Cresson, 1917
Ceropsilopa coquilletti Cresson, 1922. Wirth, 1968a: 77.11
Cressonomyia Arnaud, 1958
Cressonomyia aciculata (Loew, 1862). Mathis & Zatwarnicki, 2004: 255
Cressonomyia skinneri (Cresson, 1922). Mathis & Zatwarnicki, 2004: 269
Leptopsilopa Cresson, 1922
Leptopsilopa nigrimana (Williston, 1896). Perez-Gelabert, 2008: 178
Leptopsilopa similis (Coquillett, 1900). Perez-Gelabert, 2008: 178
Mimapsilopa Cresson, 1941
Mimapsilopa bacoa Mathis & Zatwarnicki, 1998: 13
Mimapsilopa cressoni Lizarralde de Grosso, 1982. Mathis & Zatwarnicki, 1998: 12
Mimapsilopa dominicana Mathis & Zatwarnicki, 1998: 20 *DR > TL: La Vega: nr. Jarabacoa, Salto Guasara, 680 m
Psilopa Fallén, 1823

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Psilopa compta (Meigen, 1830). Perez-Gelabert, 2008: 178
Psilopa girschneri von Röder, 1889. Perez-Gelabert, 2008: 178
Psilopa pulchripes Loew, 1878. Perez-Gelabert, 2008: 178
Subfamily EPHYDRINAE
Tribe DAGINI
Brachydeutera Loew, 1862
Brachydeutera longipes Hendel, 1913. Perez-Gelabert, 2008: 178
Brachydeutera neotropica Wirth, 1964. Perez-Gelabert, 2008: 178
Dagus Cresson, 1935
Dagus dominicanus Mathis in Mathis & Zatwarnicki, 1988: 116. Stark, 1993: 3 *DR > TL: La Vega Prov.: Constanza (12 km
S)
Dagus grimaldii Stark, 1993: 6. Mathis & Zatwarnicki, 1995: 232 *DR > TL: Peravia: 18 km N San José de Ocoa, 830 m
Dagus hermani Stark, 1993: 7. Mathis & Zatwarnicki, 1995: 232 *DR > TL: La Vega: 12 W Bonao, 3100 ft.
Dagus mathisi Stark, 1993: 4. Mathis & Zatwarnicki, 1995: 232 *DR > TL: Peravia: 18 km N San José de Ocoa, 830 m
Dagus spanglerorum Mathis in Mathis & Zatwarnicki, 1988: 114. Stark, 1993: 3 *DR > TL: La Vega Prov.: Constanza (8.5 km
S)
Dagus trichocerus Mathis, 1983. Mathis & Zatwarnicki, 1995: 233
Dagus wirthi Mathis, 1983. Perez-Gelabert, 2008: 178
Physemops Cresson, 1934
Physemops panops Wirth, 1970: 175. Mathis & Zatwarnicki, 1995: 234
Tribe EPHYDRINI
Ephydra Fallén, 1810
Ephydra (Halephydra) gracilis Packard, 1871. Mathis & Marinoni, 2016: 95
Tribe PARYDRINI
Parydra Stenhammar, 1844
Parydra (Parydra) humilis Williston, 1897. Perez-Gelabert, 2008: 178
Tribe SCATELLINI
Limnellia Malloch, 1825
Limnellia huachuca Mathis, 1978. Perez-Gelabert, 2008: 178
Scatella Robineau-Desvoidy, 1830
Scatella (Scatella) obscura Williston, 1896. Perez-Gelabert, 2008: 178
Scatella (Scatella) triseta Coquillett, 1902. Perez-Gelabert, 2008: 178
Subfamily HYDRELLIINAE
Tribe ATISSINI
Atissa Haliday, 1839
Atissa luteipes Cresson, 1944. Perez-Gelabert, 2008: 178
Ptilomyia Coquillett, 1900
Ptilomyia lobiochaeta Sturtevant & Wheeler, 1954. Perez-Gelabert, 2008: 179
Ptilomyia mabelae (Cresson, 1926). Perez-Gelabert, 2008: 179
Ptilomyia parva (Williston, 1896). Perez-Gelabert, 2008: 179
Tribe DRYXIINI
Oedenops Becker, 1903
Oedenops nuda (Coquillett, 1902). Mathis & Zatwarnicki, 2002: 57
Paralimna Loew, 1862
Paralimna (Paralimna) maculata Ale-Rocha & Mathis, 2015: 42
Paralimna (Paralimna) multipunctata Williston, 1896. Perez-Gelabert, 2008: 179
Paralimna (Paralimna) pauca Ale-Rocha & Mathis, 2015: 60 *RD > TL: Independencia, La Descubierta, 0 m
Paralimna (Paralimna) plumbiceps Cresson, 1916. Wirth, 1968a: 77.15; Ale-Rocha & Mathis, 2015: 71
Paralimna (Paralimna) thomae (Wiedemann, 1830). Ale-Rocha & Mathis, 2015: 102
Paralimna (Phaiosterna) decipiens Loew, 1878. Mathis & Zatwarnicki, 2002: 72; Ale-Rocha & Mathis, 2015: 112
Paralimna (Phaiosterna) longiseta Mathis & Zatwarnicki, 2002: 81. Ale-Rocha & Mathis, 2015: 117
Paralimna (Phaiosterna) obscura Williston, 1896: 83. Ale-Rocha & Mathis, 2015: 119
Tribe HYDRELLINI
Hydrellia Robineau-Desvoidy, 1830

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Hydrellia calverti Cresson, 1918. Wirth, 1968a: 77.13; Mathis & Zatwarnicki, 1995: 66
Hydrellia tibialis Cresson, 1917. Perez-Gelabert, 2008: 179
Lemnaphila Cresson, 1933
Lemnaphila wirthi Lizarralde de Grosso, 1977. Perez-Gelabert, 2008: 179
Tribe NOTIPHILINI
Notiphila Fallén, 1810
Notiphila (Dichaeta) decorata Williston, 1896. Perez-Gelabert, 2008: 179
Notiphila (Notiphila) erythrocera Loew, 1878. Perez-Gelabert, 2008: 179
Tribe TYPOPSILOPINI
Typopsilopa Cresson, 1916
Typopsilopa archboldi Wirth, 1968. Perez-Gelabert, 2008: 179
Subfamily ILYTHEINAE
Tribe ILYTHEINI
Ilythea Haliday, 1839
Ilythea caniceps Cresson, 1918. Perez-Gelabert, 2008: 179
Lytogaster Becker, 1896
Lytogaster pallipes Cresson, 1914. Wirth, 1968a: 77.19
Zeros Cresson, 1943
Zeros calverti (Cresson, 1918). Perez-Gelabert, 2008: 179
Zeros fenestralis (Cresson, 1918). Perez-Gelabert, 2008: 179
Zeros flavipes (Williston, 1896). Perez-Gelabert, 2008: 179
Zeros obscurus (Cresson, 1918). Perez-Gelabert, 2008: 179
Tribe PHILYGRIINI
Nostima Coquillett, 1900
Nostima atriscuta Edmiston & Mathis, 2005: 15
Nostima duaguttata Edmiston & Mathis, 2005: 21
Nostima elegantula Hendel, 1930. Edmiston & Mathis, 2005: 23
Nostima flavida Cresson, 1947. Edmiston & Mathis, 2005: 25
Nostima franciscana Edmiston & Mathis, 2005: 29
Nostima gilvipes (Coquillett, 1900). Edmiston & Mathis, 2005: 31
Nostima giovannolii Wirth, 1956. Edmiston & Mathis, 2005: 34
Nostima lutea Edmiston & Mathis, 2005: 41
Nostima maculata Edmiston & Mathis, 2005: 43
Nostima niveivenosa Cresson, 1930. Mathis & Zatwarnicki, 1995: 192; Edmiston & Mathis, 2005: 52
Nostima pulchra (Williston, 1896). Wirth, 1968a: 77.17; Edmiston & Mathis, 2005: 60
Nostima schildi Cresson, 1941. Edmiston & Mathis, 2005: 60
Nostima slossonae Coquillett, 1900. Edmiston & Mathis, 2005: 69
Nostima spilogaster Cresson, 1947. Edmiston & Mathis, 2005: 72
Nostima xehohypopia Edmiston & Mathis, 2005: 84 *DR > TL: Pedernales Prov., Sierra de Bahoruco, Las Abejas, 1300 m

Order EMBIIDINA
Family ANISEMBIIDAE
Glyphembia Ross, 2003
Glyphembia amberica† Ross, 2003: 21 [Fossil in Dominican amber]
Glyphembia dominicana Ross, 2003: 24 *DR > TL: 13 km S of Azua. Azua Prov.
Glyphembia haitiana (Ross, 1940: 646). Ross, 2003: 24 *H > TL: Gros Morne
Glyphembia vetehae† (Szumik, 1998: 7). Ross, 2003: 27 [Fossil in Dominican amber]
Poinarembia† Ross, 2003
Poinarembia rota† Ross, 2003: 38 [Fossil in Dominican amber]

Family TERATEMBIIDAE
Oligembia Davis, 1939
Oligembia vetusta† Szumik, 1994: 71 [Fossil in Dominican amber]

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Order EPHEMEROPTERA
Family BAETIDAE
Subfamily BAETINAE
Baetis Leach, 1815
Baetis sp.†—Spahr (1992: 79) [Fossil in Dominican amber]
Cloeodes Traver, 1938
Cloeodes sp.†—Poinar (1992: 98) [Fossil in Dominican amber]
Cloeodes inferior Kluge, 1991. Naranjo & Peters, 2016: 84; Naranjo et al., 2019: 6
Cloeodes superior Kluge, 1991. Naranjo & Peters, 2016: 84; Naranjo et al., 2019: 6
Fallceon Waltz & McCafferty, 1987
Fallceon poeyi (Eaton, 1885). Naranjo et al., 2019: 8
=Fallceon nikitai McCafferty & Lugo-Ortiz, 1994. Naranjo & Peters, 2016: 85
Paracloeodes Day, 1955
Paracloeodes sp. – Naranjo & Peters (2016: 85)

Family CAENIDAE
Caenis Stephens, 1833
Caenis sp. – Naranjo & Peters (2016: 85

Family EUTYPHLOCIIDAE
Mesoplocia Demoulin, 1952
Mesoplocia inaccessibile (Kluge & Naranjo, 1994). Naranjo & Peters, 2016: 85; Naranjo et al., 2019: 16

Family LEPTOPHLEBIIDAE
Subfamily ATALOPHLEBIINAE
Borinquena Traver, 1938
Borinquena (?) caeciliana† Staniczek, 2003: 13 [Fossil in Dominican amber]
Borinquena maculata† Staniczek, 2003: 6 [Fossil in Dominican amber]
Borinquena parva† Staniczek, 2003: 9; Godunko & Krzeminski, 2009: 131; Penney et al., 2012: 3 [Fossil in Dominican amber]
Borinquena schawallfussi† Staniczek, Godunko & Krzeminski, 2017: 114 [Fossil in Dominican amber]
Careospina Peters, 1971
Careospina annulata Peters, 1971: 14 *H > TL: Mt. La Hotte, 1,000 ft., Roche Croix
Hagenulites† Staniczek, 2003
Hagenulites hitchingsi† Staniczek, 2003: 2 [Fossil in Dominican amber]
Hagenulus Eaton, 1882
Hagenulus eatoni Banks, 1924: 423 *H > TL: Diquini

Family LEPTOHYPHIDAE
Tricorythodes Ulmer, 1920
Tricorythodes rolstoni (Allen, 1973: 371). Naranjo & Peters, 2016: 86 *DR > TL: San Cristóbal

Order HEMIPTERA

Taxonomic summary:
Infraorder Enicocephalomorpha
Superfamily Enicocephaloidea
Family Enicocephalidae
Infraorder Leptopodomorpha
Superfamily Leptopodoidea
Family Leptopodidae
Family Saldidae
Infraorder Dipsocoromorpha
Superfamily Dipsocoroidea
Family Schizopteridae

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 235
Infraorder Gerromorpha
Superfamily Mesoveloidea
Family Hebridae
Family Mesoveliidae
Superfamily Hydrometroidea
Family Hydrometridae
Superfamily Gerroidea
Family Gerridae
Family Veliidae
Infraorder Nepomorpha
Superfamily Naucoroidea
Family Naucoridae
Superfamily Nepoidea
Family Belostomatidae
Superfamily Notonectoidea
Family Notonectidae
Superfamily Ochteroidea
Family Ochteridae
Superfamily Pleoidea
Family Pleidae
Superfamily Corixoidea
Family Corixidae
Infraorder Cimicomorpha
Superfamily Thaumastocoroidea
Family Thaumastocoridae
Superfamily Tingoidea
Family Tingidae
Superfamily Miroidea
Family Miridae
Superfamily Cimicoidea
Family Anthocoridae
Family Cimicidae
Family Nabidae
Superfamily Reduvoidea
Family Reduviidae
Infraorder Pentatomomorpha
Superfamily Aradoidea
Family Aradidae
Superfamily Lygaeoidea
Family Berytidae
Family Blissidae
Family Cymidae
Family Geocoridae
Family Lygaeidae
Family Ninidae
Family Oxycarenidae
Family Pachygronthidae
Family Rhyparochromidae
Superfamily Pyrrhocoroidea
Family Largidae
Family Pyrrhocoridae
Superfamily Coreoidea
Family Alydidae
Family Coreidae

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 Family Rhopalidae
Superfamily Pentatomoidea
Family Cydnidae
Family Corimelaenidae
Family Megarididae
Family Pentatomidae
Family Tessaratomidae
Family Plataspidae
Family Scutelleridae
Suborder Auchenorrhyncha
Infraorder Fulgoromorpha
Superfamily Fulgoroidea
Family Acanaloniidae
Family Achilidae
Family Caliscelidae
Family Cixiidae
Family Delphacidae
Family Derbidae
Family Flatidae
Family Issidae
Family Kinnaridae
Family Nogodinidae
Family Tropiduchidae
Infraorder Cicadomorpha
Superfamily Cercopoidea
Family Cercopidae
Family Clastopteridae
Superfamily Cicadoidea
Family Cicadidae
Superfamily Cicadelloidea
Family Cicadellidae
Family Membracidae
Suborder Sternorrhyncha
Superfamily Psylloidea
Family Aphalaridae
Family Liviidae
Family Phacopteronidae
Family Psyllidae
Family Triozidae
Superfamily Aleyrodoidea
Family Aleyrodidae
Superfamily Aphidoidea
Family Aphididae
Family Drepanosiphidae
Family Hormaphididae
Family Termitaphididae
Superfamily Coccoidea
Family Cerococcidae
Family Coccidae
Family Conchaspididae
Family Dactylopiidae
Family Asterolecaniidae
Family Diaspididae
Family Eriococcidae

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 Family Monophlebidae
Family Pseudococcidae
Family Ortheziidae
Family Margarodidae

Suborder HETEROPTERA
Infraorder ENICOCEPHALOMORPHA
Superfamily ENICOCEPHALOIDEA
Family ENICOCEPHALIDAE
Subfamily ENICOCEPHALINAE
Enicocephalus Westwood, 1838
Enicocephalus cubanus Bruner, 1924. Wygodzinsky & Schmidt, 1991: 107
=Enicocephalus marimutti Kritsky, 1979: 95. Wygodzinsky & Schmidt, 1991: 108
Enicocephalus exophthalmus Wygodzinsky & Schmidt, 1991: 116 *H > TL: Desbarriere, Mt. La Hotte, near 4000 ft.
Enicocephalus omen† Grimaldi et al., 1993: 19 [Fossil in Dominican amber]
Enicocephalus prius† Grimaldi et al., 1993: 23 [Fossil in Dominican amber]
Enicocephalus seniculus† Grimaldi et al., 1993: 15 [Fossil in Dominican amber]
Enicocephalus subvitreus† Grimaldi et al., 1993: 17 [Fossil in Dominican amber]
Subfamily ALIENATINAE
Alienates Barber, 1953
Alienates brodzinskyi† Grimaldi et al., 1993: 7 [Fossil in Dominican amber]
Alienates electrodominicus† Grimaldi et al., 1993: 8 [Fossil in Dominican amber]
Alienates sp.†—Grimaldi et al. (1993: 10) [Fossil in Dominican amber]
Paralienates† Maldonado Capriles et al., 1996
Paralienates hyalinus† Maldonado Capriles et al., 1996: 48 [Fossil in Dominican amber]

Infraorder LEPTOPODOMORPHA
Superfamily LEPTOPODOIDEA
Family LEPTOPODIDAE
Subfamily LEPTOPODINAE
Leptosalda Cobben, 1971
Leptosalda dominicana† Grimaldi & Engel in Grimaldi et al., 2013: 232 [Fossil in Dominican amber]
Leptosalda niarchos† Grimaldi & Engel in Grimaldi et al., 2013: 233 [Fossil in Dominican amber]
Leptosalda sp.†—Wu (1997: 159); Grimaldi & Engel (2005: 326) [Fossil in Dominican amber]

Family SALDIDAE
Subfamily CHILOXANTHINAE
Pentacora Reuter, 1912
Pentacora signoreti (Guérin-Méneville, 1856). Drake, 1949: 189
Pentacora sphacelata (Uhler, 1877). Drake, 1949: 189
Subfamily SALDIINAE
Saldula Van Duzee, 1914
Saldula lomata Polhemus, 1985: 186
Saldula pallipes (Fabricius, 1803). Kirkaldy & Torre-Bueno, 1908: 177; Barber, 1939: 415
=Acanthia interstitialis Say, 1825. Kirkaldy & Torre-Bueno, 1908: 176
Saldula sulcata (Barber, 1939). Drake, 1949: 189

Infraorder DIPSOCOROMORPHA
Superfamily DIPSOCOROIDEA
Family SCHIZOPTERIDAE
Hypselosoma Reuter, 1891
Hypselosoma dominicana† Poinar & Brown, 2015: 373 [Fossil in Dominican amber]
Hypselosomops Hoey-Chamberlain & Weirauch, 2016
Hypselosomops pecki Hoey-Chamberlain & Weirauch, 2016: 93

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Schizoptera Fieber, 1860
Schizoptera dominicana† Poinar & Brown, 2015: 374 [Fossil in Dominican amber]
Schizoptera hispaniolae† Poinar & Brown, 2015: 375 [Fossil in Dominican amber]

Infraorder GERROMORPHA
Superfamily MESOVELOIDEA
Family HEBRIDAE
Miohebrus Garrouste & Nel, 2010
Miohebrus anderseni† Garrouste & Nel, 2010: 97 [Fossil in Dominican amber]

Family MESOVELIIDAE
Mesovelia Mulsant & Rey, 1852
Mesovelia amoena Uhler, 1894. Spangler, 1990: 93
Mesovelia dominicana† Garrouste & Nel, 2010: 94 [Fossil in Dominican amber]
Mesovelia mulsanti White, 1879. Horváth, 1915: 548; Drake & Maldonado Capriles, 1956: 55

Superfamily HYDROMETROIDEA
Family HYDROMETRIDAE
Hydrometra Latreille, 1796
Hydrometra australis Say, 1832. Nieser, 1970: 147
=Hydrometra myrae Torre-Bueno, 1926. Hungerford & Evans, 1934: 105
Hydrometra caraiba Guérin-Méneville, 1856. Drake, 1954: 63; Drake & Lauck, 1959: 51; Nieser, 1970: 143
=Hydrometra championana Torre-Bueno, 1926. Hungerford & Evans, 1934: 94; Drake, 1954: 63
Hydrometra consimilis Barber, 1934. Drake & Hottes, 1952: 110

Superfamily GERROIDEA
Family GERRIDAE
Subfamily CHARMATOMETRINAE
Brachymetroides† Andersen, 2000
Brachymetroides atra† Andersen, 2000: 414 [Fossil in Dominican amber]
Subfamily ELECTROBATINAE
Electrobates Andersen & Poinar, 1992
Electrobates spinipes† Andersen & Poinar, 1992: 261. Andersen, 2000: 419 [Fossil in Dominican amber]
Subfamily GERRINAE
Aquarius Schellenberg, 1800
Aquarius remigis (Say, 1832). Barber, 1939: 408
Eobates Drake & Harris, 1934
Eobates morrisoni Drake & Harris, 1934a: 176 *DR > TL: S. Francisco Mts.
Eobates vittatus (Shaw, 1933: 224). Drake & Maldonado Capriles, 1956: 53
Eurygerris Hungerford & Matsuda, 1958
Eurygerris cariniventris (Champion, 1898). Drake & Harris, 1934b: 197; Barber, 1939: 408
Eurygerris dominicus (Drake & Maldonado Capriles, 1956: 54)
Limnogonus Stål, 1868
Limnogonus franciscanus (Stål, 1859). Barber, 1939: 407. Kort-Gommers & Nieser, 1969: 7
=Gerris guerini Lethierry & Severin, 1896. Drake & Maldonado Capriles, 1956: 53
Subfamily RHAGADOTARSINAE
Rheumatobates Bergroth, 1892
Rheumatobates bergrothi Meinert, 1895. Drake & Maldonado Capriles, 1956: 53
Rheumatobates imitator (Uhler, 1894). Barber, 1939: 407
Rheumatobates meinerti Schroeder, 1931: 70. Spangler et al., 1985: 198
Subfamily TREPOBATINAE
Metrobates Uhler, 1871
Metrobates tumidus Anderson, 1932. Drake & Harris, 1937: 358
Trepobates Uhler, 1894

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Trepobates taylori (Kirkaldy, 1899). Drake & Maldonado Capriles, 1956: 53

Family VELIIDAE
Subfamily HALOVELIINAE
Halovelia Bergroth, 1893
Halovelia electrodominica† Andersen & Poinar, 1998: 1. Andersen, 2000b: 426 [Fossil in Dominican amber]
Subfamily MICROVELIINAE
Microvelia Westwood, 1834
Microvelia albonotata Champion, 1898. Drake & Maldonado Capriles, 1956: 55
Microvelia cubana Drake, 1951. Drake & Hussey, 1955: 113
Microvelia electra† Andersen, 2000b: 422 [Fossil in Dominican amber]
Microvelia grimaldii† Andersen, 2000b: 420 [Fossil in Dominican amber]
Microvelia hinei Drake, 1920. Drake & Maldonado Capriles, 1956b: 55
Microvelia longipes Uhler, 1894. Drake & Maldonado Capriles, 1956b: 55; Kort-Gommers & Nieser, 1969: 84
Microvelia polhemi† Andersen, 2000a: 136. Andersen, 2000b: 424 [Fossil in Dominican amber]
Microvelia pulchella Westwood, 1834. Drake & Maldonado Capriles, 1956b: 55
Subfamily RHAGOVELIINAE
Rhagovelia Mayr, 1865
Rhagovelia aestiva J. Polhemus, 1980: 314 *H > TL: NE Foothills, La Hotte, 2400 ft.
Rhagovelia collaris (Burmeister, 1835). Kirkaldy & Torre-Bueno, 1908: 205; Gould, 1931: 21; Barber, 1939: 413
Rhagovelia elegans Uhler, 1894. Padilla-Gil & Figuereido, 2013: 414
Rhagovelia plumbea Uhler, 1894. Drake & Van Doesburg, 1966: 68
Rhagovelia secluda Drake & Maldonado Capriles, 1956a: 14. Drake & Maldonado Capriles, 1956b: 55 *DR > TL: Valle
Nuevo, La Vega Prov.
Rhagovelia vegana Drake & Maldonado Capriles, 1956b: 55 *DR > TL: La Vega Prov., Valle Nuevo, 2200 m
Subfamily VELIINAE
Platyvelia J. Polhemus & D. Polhemus, 1993
Platyvelia brachialis (Stål, 1860). Drake and Maldonado-Capriles, 1956: 55; Perez-Gelabert & Floriano, 2016: 471
Steinovelia J. Polhemus & D. Polhemus, 1993
Steinovelia permista (Drake, 1951). J. Polhemus & D. Polhemus, 1993: 397

Infraorder NEPOMORPHA
Superfamily NAUCOROIDEA
Family NAUCORIDAE
Decarloa La Rivers, 1969
Decarloa darlingtoni La Rivers, 1969: 1. Ludwick & Sites, 2015: 14 *H, *DR > TL: La Hotte (NE foothills), 2400 ft. el.
Pelocoris Stål, 1876
Pelocoris poeyi (Guérin-Méneville, 1844). García Avila & Peña, 1993: 214

Superfamily NEPOIDEA
Family BELOSTOMATIDAE
Subfamily BELOSTOMATINAE
Belostoma Latreille, 1807
Belostoma anurum (Herrich-Schäffer, 1848). Uhler, 1894: 223; Ribeiro, 2005: 255
Belostoma elongatum Montandon, 1908. Ribeiro, 2007: 28
Belostoma minor (Mayr, 1871). Kirkaldy & Torre-Bueno, 1908: 192; Lauck, 1964: 121; García Avila & Peña, 1993: 214
Belostoma subspinosum subspinosum (Palisot de Beauvois, 1820). Kirkaldy & Torre-Bueno, 1908: 192; Lauck, 1962: 66
Subfamily LETHOCERINAE
Lethocerus Mayr, 1853
Lethocerus (Lethocerus) annulipes (Herrich-Schäffer, 1848). Menke, 1963: 264
Note: Belostoma boscii (Lepeletier & Serville, 1825) cited for Hispaniola by Kirkaldy & Torre-Bueno, 1908: 190 and Barber,
1939: 425 is a Nomen dubium.

Superfamily NOTONECTOIDEA

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Family NOTONECTIDAE
Subfamily ANISOPINAE
Buenoa Kirkaldy, 1904
Buenoa antigone (Kirkaldy, 1899). Kirkaldy, 1904: 122; Kirkaldy & Torre-Bueno, 1908: 200; Truxal, 1953: 1376; Nieser,
1969: 92
Buenoa macrophthalma (Fieber, 1851). Barber, 1939: 420; Truxal, 1953: 1480
Buenoa mutabilis Truxal, 1953: 1432
Buenoa pallipes (Fabricius, 1803). Barber, 1939: 421
Buenoa platycnemis (Fieber, 1851). Truxal, 1953: 1421; Nieser, 1969: 96
Subfamily NOTONECTINAE
Notonecta Linnaeus, 1758
Notonecta indica Linnaeus, 1771. Nieser, 1969: 89; García Avila & Peña, 1993: 214

Superfamily OCHTEROIDEA
Family OCHTERIDAE
Riegerochterus† Popov & Heiss, 2014 [Fossil in Dominican amber]
Riegerochterus baehri† Popov & Heiss, 2014: 187 [Fossil in Dominican amber]

Superfamily PLEOIDEA
Family PLEIDAE
Neoplea Esaki & China, 1928
Neoplea punctifer (Barber, 1923). Drake & Maldonado Capriles, 1956: 53; Nieser, 1969: 62
Paraplea Esaki & China, 1928
Paraplea puella (Barber, 1923). Drake & Maldonado Capriles, 1956: 53; Nieser, 1969: 61

Superfamily CORIXOIDEA
Family CORIXIDAE
Key reference: Hungerford (1948).
Subfamily CORIXINAE
Tribe CORIXINI
Centrocorisa Lundblad, 1928
Centrocorisa nigripennis (Fabricius, 1803). Hungerford, 1948: 437; Nieser, 1969: 145
Ramphocorixa Abbott, 1912
Ramphocorixa rotundocephala Hungerford, 1927: 1. Hungerford, 1948: 452; Nieser, 1969: 147
Trichocorixa Kirkaldy, 1908
Trichocorixa louisianae Schwinghammer, 1905. Sailer, 1948: 324
Trichocorixa reticulata (Guérin-Méneville, 1857). Sailer, 1948: 343; Nieser, 1969: 153
Trichocorixa verticalis verticalis (Fieber, 1851). Sailer, 1948: 358; Nieser, 1969: 156

Infraorder CIMICOMORPHA
Superfamily THAUMASTOCOROIDEA
Family THAUMASTOCORIDAE
Discocoris Bergh, 1877
Discocoris dominicanus† Slater & Baranowski, 2000: 350 [Fossil in Dominican amber]
Paleodoris† Poinar & Santiago-Blay, 1997
Paleodoris lattini† Poinar & Santiago-Blay, 1997: 308 [Fossil in Dominican amber]

Superfamily TINGOIDEA
Family TINGIDAE
Subfamily CANTACADERINAE
Phatnoma Fieber, 1844
Phatnoma mattijoae† Jepson, Penney & Green, 2011: 61 [Fossil in Dominican amber]
Phatnoma pulchra† Golub & Heiss, 2018: 334 [Fossil in Dominican amber]
Subfamily TINGINAE

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 241
Acanthocheila Stål, 1858
Acanthocheila armigera (Stål, 1858). Drake & Poor, 1937: 306; Drake & Ruhoff, 1965: 56; Froeschner, 1995: 336
Acanthocheila spinicosta Van Duzee, 1907. Barber, 1939: 370; Drake & Ruhoff, 1965: 59; Froeschner, 1995: 336
Allotingis Drake, 1930
Allotingis insulicola Drake & Poor, 1939: 31. Drake & Ruhoff, 1965: 78 *H > TL: Fond des Negres
Amberobyrsa Heiss, 2009
Amberobyrsa brandti† Heiss, 2009: 10 [Fossil in Dominican amber]
Amblystira Stål, 1873
Amblystira amica Drake & Ruhoff, 1960. Drake & Ruhoff, 1965: 79 *H > TL: Hinche
=Amblystira solida Drake, 1942: 17
=Amblystira dozieri Drake & Hambleton, 1944: 94
Amblystira fuscitarsis Champion, 1897. Drake & Hambleton, 1945: 358; Drake & Ruhoff, 1965: 80
Amblystira morrisoni Drake, 1922: 360. Drake & Bruner, 1924: 148; Drake & Ruhoff, 1965: 81 *DR > TL: San Domingo City
Corythaica Stål, 1873
Corythaica carinata Uhler, 1894. Drake & Bruner, 1924: 151; Barber, 1939: 369; Hurd, 1945: 89; Drake & Cobben, 1960: 87;
Drake & Ruhoff, 1965: 136
Corythaica passiflorae Berg, 1884. By synonymy of species below.
=Corythaica cyathicollis (Costa, 1864). Santoro, 1960: 80; SEA, 1999: 56
=Corythaica planaris (Uhler, 1893). SEA, 1999: 44
Corythucha Stål, 1873
Corythucha gossypii (Fabricius, 1794). Wolcott, 1927: 180; Leonard & Mills, 1931: 309; Drake & Ruhoff, 1965: 149
Dicrotingis Drake & Ruhoff, 1960
Dicrotingis digitalis (Drake, 1928a: 22). Drake & Ruhoff, 1965: 180 *H > TL: Croix des Missiones, Plaine Cul de Sac
Dictyla Stål, 1874
Dictyla balli (Drake, 1922: 355). Drake & Bruner, 1924: 144; Drake, 1943: 142; Drake & Ruhoff, 1965: 182
Dictyla c-nigrum (Champion, 1898). Drake, 1922: 355; Barber, 1939: 372; Drake & Ruhoff, 1965: 183
Dictyla haitiensis (Drake & Poor, 1938: 28). Drake & Ruhoff, 1965: 187
Dictyla monotropidia (Stål, 1858). Drake, 1922: 354; Drake & Poor, 1937: 301; Barber, 1939: 371; Drake & Ruhoff, 1965: 190
Eocader Drake & Hambleton, 1934
Eocader babyrussus† Golub & Popov, 2000a: 166 [Fossil in Dominican amber]
Gargaphia Stål, 1862
Gargaphia vanduzei Gibson, 1919. SEA, 1999: 52
Leptodictya Stål, 1873
Leptodictya (Hanuala) bambusae Drake, 1918. Barber, 1939: 370; Drake & Ruhoff, 1965: 261
Leptopharsa Stål, 1873
Leptopharsa antica† Golub & Heiss, 2018: 336 [Fossil in Dominican amber]
Leptopharsa elegantula Stål, 1873. Drake & Poor, 1937: 308
Leptopharsa evsyunini† Golub & Popov, 2000b: S292 [Fossil in Dominican amber]
Leptopharsa frater† Golub & Popov, 2003: 104 [Fossil in Dominican amber]
Leptopharsa hoffmani Drake, 1928a: 21. Drake & Ruhoff, 1965: 277 *H > TL: Haiti
Leptopharsa poinari† Golub & Popov, 2000b: S290 [Fossil in Dominican amber]
Leptopharsa vicina Drake & Poor, 1939: 33. Drake & Ruhoff, 1965: 287 *H > TL: Williamson
Leptoypha Stål, 1873
Leptoypha morrisoni Drake, 1922: 43. Drake & Ruhoff, 1965: 290
Phymacysta Monte, 1942
Phymacysta mcelfreshi (Drake, 1918: 175). Drake & Bruner, 1924: 149; Drake & Ruhoff, 1965: 327 *H > TL: Port-au-Prince
Phymacista stysi† Golub, Popov & Guilbert, 2008: 486 [Fossil in Dominican amber]
Phymacysta tumida (Champion, 1897). Drake & Cobben, 1960: 85; Drake & Ruhoff, 1965: 327; Marcano & Abud, 1994: 11/
94
=Leptopharsa walcotti Drake, 1928b: 55. Drake and Ruhoff, 1965: 327
Pseudacysta Blatchley, 1926
Pseudacysta perseae (Heidemann, 1908). Abud & Marcano, 1990: 4/90; Abud, 1991: 4; Zuluaga & Mesa, 2000: 197
[Introduced]
Stephanitis Stål, 1873

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Stephanitis rozanovi† Golub & Popov, 2003: 106 [Fossil in Dominican amber]
Teleonemia Costa, 1864
Teleonemia adelphe Drake & Maldonado Capriles, 1965: 317 *H > TL: Kenscoff
Teleonemia lustrabilis Drake, 1953: 151. Drake & Ruhoff, 1965: 378 *DR > TL: Constanza, 3000–4000 ft. elevation
Teleonemia scrupulosa Stål, 1873. Drake & Ruhoff, 1965: 383
Teleonemia scrupulosa haytiensis Drake, 1920. Drake & Bruner, 1924: 145; Drake & Frick, 1939: 201; Drake & Ruhoff, 1965:
384
Vatiga Drake & Hambleton, 1946
Vatiga illudens (Drake, 1922: 370). Barber, 1939: 370; Drake & Cobben, 1960: 85; Drake & Ruhoff, 1965: 424; Froeschner,
1993: 458

Superfamily MIROIDEA
Family MIRIDAE
Key reference: Schuh (1995).
Subfamily BRYOCORINAE
Tribe DICYPHINI
Engytatus Reuter, 1875
Engytatus modestus (Distant, 1893). Schmutterer, 1990: 577
=Engytatus geniculatus Reuter, 1876. Uhler, 1894: 193
=Dicyphus luridus (Gibson, 1917). SEA, 1999: 121
Macrolophus Fieber, 1858
Macrolophus praeclarus (Distant, 1884). Marcano, 1964: 30
=Dicyphus prasinus (Gibson, 1917). SEA, 1999: 121
Nesidiocoris Kirkaldy, 1902
Nesidiocoris tenuis (Reuter, 1895). Smith & Audant, 1930: 978; Schmutterer, 1990: 577; Alvarez et al., 1993: 36
Tupiocoris China & Carvalho, 1952
Tupiocoris notatus (Distant, 1893). SEA, 1999: 121
Tribe ECCRITOTARSINI
Cyrtocapsus Reuter, 1875
Cyrtocapsus caligineus (Stål, 1859). Henry & Carvalho, 1987: 292
=Hemisphaerodella mirabilis (Reuter, 1908). Henry & Carvalho, 1987: 292
Cyrtocapsus haitianus Carvalho, 1954: 16. Henry & Carvalho, 1987: 292 *H > TL: Camp Perrin
Pycnoderes Guérin-Méneville, 1857
Pycnoderes incurvus (Distant, 1884). Russo, 1927: 44; Marcano, 1964: 29
Subfamily DERAEOCORINAE
Tribe CLIVINEMATINI
Dominicanocoris Ferreira, 1996
Dominicanocoris barahonensis Ferreira, 1996: 264 *DR > TL: Barahona, 6 km NW Fundación
Tribe DERAEOCORINI
Diplozona Van Duzee, 1915
Diplozona mexicana Carvalho & Gomes, 1972. Carvalho & Costa, 1990: 186
Tribe HYALODINI
Antias Distant, 1884
Antias dominicana Carvalho, 1982: 329 *DR > TL: 3 miles W Haina, San Cristóbal Prov.
Antias maldonadoi Carvalho, 1982: 330 *DR > TL: Constanza
Fennahiella Carvalho, 1955
Fennahiella singularis (Carvalho, 1952: 181) *H > TL: Haiti
Paracarniella Henry & Ferreira, 2003
Paracarniella dominicanensis Henry & Ferreira, 2003: 109 *DR > TL: La Vega Prov., Rt. 12, 20 km SW of jct. 1 above Bonao,
3,400 ft.
Paracarnus Distant, 1884
Paracarnus haitianus Carvalho, 1952: 182 *H > TL: Cap Haitien
Subfamily ISOMETOPINAE
Tribe ISOMETOPINI

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Diphleps Bergroth, 1924
Diphleps yenli† Santiago-Blay & Poinar, 1993: 71 [Fossil in Dominican amber]
Subfamily MIRINAE
Tribe MIRINI
Dagbertus Distant, 1904
Dagbertus fasciatus (Reuter, 1876). Maldonado Capriles, 1986: 125
Derophthalma Berg, 1883
Derophthalma dominicana Carvalho & Gomes, 1980: 106 *DR > TL: Prov. La Vega, 8 milhas a oeste de Jayaco
Phytocoris Fallén, 1814
Phytocoris eximius Reuter, 1875. Uhler, 1894: 191
Stomatomiris† Maldonado Capriles & Poinar, 1995
Stomatomiris funebris† Maldonado Capriles & Poinar, 1995: 282 [Fossil in Dominican amber]
Tribe RESTHENINI
Prepops Reuter, 1905
Prepops maldonadoi Carvalho & Fontes, 1973: 543
Tribe STENODEMINI
Collaria Provancher, 1872
Collaria oleosa (Distant, 1883). Schmutterer, 1990: 41; Morales et al., 2016: 228
Subfamily ORTHOTYLINAE
Tribe CERATOCAPSINI
Ceratocapsidea Henry, 2015
Ceratocapsidea dominicanensis Henry, 2015: 30 *DR > TL: Santo Domingo City, 43 m
Tribe ORTHOTYLINI
Falconia Distant, 1884
Falconia maculipennis Maldonado, 1969. Henry et al., 2019: 180
Hyalochloria Reuter, 1907
Hyalochloria unicolor Reuter, 1907. Henry, 2001: 258
Jobertus Distant, 1863
Jobertus chryselectrus Distant, 1893. Maldonado Capriles, 1980: 307
Subfamily PHYLINAE
Tribe LEUCOPHOROPTERINI
Tytthus Fieber, 1864
Tytthus parviceps (Reuter, 1890). Schuh, 1995: 249
=Cylloceps pellicia Uhler, 1893: 712. Schuh, 1995: 249
Tribe PILOPHORINI
Sthenaridea Reuter, 1884
Sthenaridea araguaiana (Carvalho, 1948). Schuh & Schwartz, 1988: 186
Sthenaridea vulgaris (Distant, 1893). Schuh & Schwartz, 1988: 192
Tribe PHYLINI
Gonoporomiris Henry & Schuh, 2002
Gonoporomiris hispaniolae Schuh & Schwartz, 2004: 22 *DR > TL: Santo Domingo
Gonoporomiris mirificus (Distant, 1893). Henry & Schuh, 2002: 213
Hyalopsallus Carvalho & Schaffner, 1973
Hyalopsallus diaphanus (Reuter, 1907). Carvalho & Schaffner, 1973: 19
Platyscytus Reuter, 1907
Platyscytus binotatus Reuter, 1907. Maldonado Capriles & Carvalho, 1981: 386
Pseudatomoscelis Poppius, 1911
Pseudatomoscelis nubila Henry, 2002: 102 *DR > TL: Prov. Pedernales, 5 km N Cabo Rojo, nr. sea level
Pseudatomoscelis seriata (Reuter, 1876). Henry, 1991: 389
Ranzovius Distant, 1893
Ranzovius moerens (Reuter, 1905). Henry, 1999: 103
Reuteroscopus Kirkaldy, 1905
Reuteroscopus carvalhoi† Maldonado Capriles & Poinar, 1995: 347 [Fossil in Dominican amber]
Reuteroscopus hamatus Kelton, 1964: 146

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Reuteroscopus ornatus (Reuter, 1876). Uhler, 1894: 195; Kelton, 1964: 1422
Rhinacloa Reuter, 1875
Rhinacloa puertoricensis Schuh & Schwartz, 1985: 426
Rhinacloa schaffneri Schuh & Schwartz, 1985: 406 *DR, *H > TL: La Vega Prov.: 2 mi. north Valle Nuevo
Subfamily PSALLOPINAE
Psallops Usinger, 1946
Psallops popovi† Herczek, 2011: 691 [Fossil in Dominican amber]
Note: The record of Engytatus varians (Distant, 1884) by Marcano, 1964: 30 needs confirmation. For the moment, I’m
discounting this record from the Hispaniolan fauna.

Superfamily CIMICOIDEA
Family ANTHOCORIDAE
Subfamily LASIOCHILINAE
Lasiochilus Reuter, 1871
Lasiochilus (Lasiochilus) hirtellus Drake & Harris, 1926. Carpintero, 2014: 54
Subfamily LYCTOCORINAE
Cardiastethus Fieber, 1860
Cardiastethus rugicollis Champion, 1900. Barber, 1939: 402
Orius Wolff, 1811
Orius insidiosus (Say, 1832). Wolcott, 1927: 178

Family CIMICIDAE
Cimex Linnaeus, 1758
Cimex hemipterus (Fabricius, 1803). Barber, 1939: 398
Cimex lectularius Linnaeus, 1758. Wolcott, 1927: 178; Barber, 1939: 398

Family NABIDAE
Subfamily NABINAE
Arachnocoris Scott, 1881
Arachnocoris darlingtoni Santiago-Blay & Mercado in Mercado et al., 2016: 44 *DR > TL: Sánchez, Samaná Prov.
Hoplistocelis Reuter, 1890
Hoplistocelis confusa Kerzhner & Henry, 2008: 989; Volpi & Coscaron, 2010: 55
Hoplistocelis sordida (Reuter, 1872). Barber, 1939: 395
Lasiomerus Reuter, 1890
Lasiomerus signatus (Uhler, 1894). Kerzhner, 2007: 232
Lasiomerus spinicrus (Reuter, 1890). Barber, 1939: 395; Volpi & Coscaron, 2010: 57
Nabis Latreille, 1802
Nabis (Tropiconabis) capsiformis Germar, 1837. Barber, 1939: 395; Kerzhner, 2007: 233

Superfamily REDUVOIDEA
Family REDUVIIDAE
Key reference: Maldonado Capriles (1990).
Subfamily ECTRICHODIINAE
Pseudopothea Wygodzinsky, 1951
Pseudopothea luacesi (Bruner and Fracker, 1926). Dougherty, 1995: 213
Subfamily EMESINAE
Alumeda† Popov, 1990
Alumeda antilliana† Popov, 1990: 10 [Fossil in Dominican amber]
Alumeda dominicana† Popov, 1990: 7 [Fossil in Dominican amber]
Alumeda nigricans† Popov, 1990: 4 [Fossil in Dominican amber]
Emesa Fabricius, 1803
Emesa tenerrima (Dohrn, 1860). Barber, 1939: 387; Maldonado Capriles, 1990: 84
=Emesa marmorata McAtee & Malloch, 1925. Maldonado Capriles, 1990: 84
Emesopsis Uhler, 1894

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 245
Emesopsis nubilus Uhler, 1894. Wygodzinsky, 1966: 365
Empicoris Hahn, 1834
Empicoris copal† Popov, 1987b: 4 [Fossil in Dominican amber]
Empiploiariola† Popov, 1993
Empiploiariola inermis† Popov, 1993: 437 [Fossil in Dominican amber]
Ghilianella Spinola, 1852
Ghilianella (Ghilianella) apiculata McAtee & Malloch, 1925: 180. Wygodzinsky, 1966: 475; Maldonado Capriles, 1990: 124
*DR > TL: Blanton Mine, north of San Cristóbal
Ghilianella gerstaeckeri (Dohrn, 1860: 223). Wygodzinsky, 1966: 484; Maldonado Capriles, 1990: 128 *H > TL: Haiti
[Species Incertae sedis]
Ghinallelia Wygodzinsky, 1966
Ghinallelia haitiana (Maldonado Capriles, 1960: 427). Wygodzinsky, 1966: 491; Maldonado Capriles, 1990: 129 *H > TL:
Kenscoff, near Port-au-Prince, 4–6000 ft. elev.
Ghinallelia zomata (Maldonado Capriles, 1963: 64). Maldonado Capriles, 1990: 131 *H > TL: Five miles south of Cap Haitien
along the road to Grande Rivière-du-Nord
Malacopus Stål, 1858
Malacopus wygodzinskyi† Popov, 1987a: 11 [Fossil in Dominican amber]
Paleoploiariola† Maldonado Capriles et al., 1993
Paleoploiariola venosa† Maldonado Capriles et al., 1993a: 97 [Fossil in Dominican amber]
Stenolemus Signoret, 1858
Stenolemus haitianus Maldonado Capriles, 1957: 457. Maldonado Capriles, 1990: 94 *H > TL: Furcy
Subfamily HARPACTORINAE
Apicrenus† Maldonado Capriles et al., 1993
Apicrenus fossilis† Maldonado Capriles et al., 1993b: 140 [Fossil in Dominican amber]
Apiomerus Hahn, 1831
Apiomerus sp.†—Poinar & Poinar (1999: 203) [Fossil in Dominican amber]
Heza Amyot & Serville, 1843
Heza clavata (Guérin-Méneville, 1857). Maldonado Capriles, 1976: 414
Heza haitiana Maldonado Capriles, 1976: 418. Maldonado Capriles, 1990: 216 *H > TL: Diquini
Heza ornata Maldonado Capriles, 1976. Perez-Gelabert, 2008: 187
Zelus Fabricius, 1803
Zelus longipes (Linnaeus, 1767). Barber, 1939: 391; Maldonado Capriles, 1990: 328
=Reduvius rubidus Lepeletier & Serville, 1825. Maldonado Capriles, 1990: 328
=Evagoras speciosus Burmeister, 1835. Maldonado Capriles, 1990: 328
Zelus puertoricensis Hart, 1987. Maldonado Capriles, 1990: 330
Zelus renardii (Kolenati, 1856). Perez-Gelabert, 2008: 187
Zelus subimpressus Stål, 1872. Barber, 1939: 392; Maldonado Capriles, 1990: 331
Subfamily HOLOPTILINAE
Praecoris† Disney, 1991
Praecoris dominicana† Poinar, 1991: 196 [Fossil in Dominican amber]
Subfamily PEIRATINAE
Melanolestes Stål, 1866
Melanolestes degener (Walker, 1873: 100). Maldonado Capriles, 1990: 361
Melanolestes picipes (Herrich-Schäffer, 1848). Maldonado Capriles, 1990: 361
Rasahus Amyot & Serville, 1843
Rasahus hamatus (Fabricius, 1781). Maldonado Capriles, 1990: 370
Rasahus sp.†—Poinar & Poinar (1999: 203) [Fossil in Dominican amber]
Sirthenea Spinola, 1840
Sirthenea amazona Stål, 1866. Maldonado Capriles, 1990: 372
Subfamily PHYMATINAE
Key reference: Froeschner and Kormilev (1989).
Capricephala Masonic, 2019
Capricephala chiaroscuro Masonic, 2019: 2 *DR > TL: Pedernales Province, Parque Nacional Sierra de Bahoruco, Las Abejas,
1150 m

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Kormilevida Van Doesburg, 1997
Kormilevida iviei (Kormilev & Van Doesburg, 1991: 280) *DR > TL: Pedernales Prov., 24 km N Cabo Rojo, 610 m
Lophoscutus Kormilev, 1951
Lophoscutus confusus Kormilev, 1989: 61 *DR > TL: S. of Jarabacoa, Pinar Quemado, 500–800 m
Lophoscutus dominicanus (Kormilev, 1963: 224) *DR > TL: Valle Nuevo
Lophoscutus haitiensis Kormilev, 1987: 703 *H > TL: Kenscoff
Lophoscutus hispaniolensis Kormilev & Van Doesburg, 1991: 278 *DR > TL: Pedernales Prov., 13.5 km N Cabo Rojo, 140 m
Lophoscutus leucographus (Westwood, 1843: 25). Barber, 1939: 379 *H > TL: Port-au-Prince
Lophoscutus pulcher (Kormilev, 1981: 218) *DR > TL: Foothills Cord. Centr. S. of Santiago
Lophoscutus testaceus (Kormilev, 1963: 223) *DR > TL: Puerto Plata
Lophoscutus ypsilon Kormilev, 1990: 223 *DR > TL: Pedernales Prov., “Minas Canote”, 35 km NE Cabo Rojo, ca. 1500 m
Paraphymata Kormilev, 1962
Paraphymata saileri Kormilev, 1960: 464 *H > TL: Acc. Hinche
Phymata Katreille, 1802
Phymata (Euryphymata) superba Kormilev, 1960: 452 *H > TL: Port-au-Prince
Phymata interjecta Dudich, 1922: 164. Kormilev, 1960: 452 *H, *DR > TL: Port-au-Prince
Phymata marginata (Fabricius, 1803). Barber, 1939: 373
Subfamily REDUVIINAE
Leogorrus Stål, 1859
Leogorrus litura (Fabricius, 1787). Maldonado Capriles, 1990: 412; Melo, 2007: 17
Subfamily SAICINAE
Kiskeyana Weirauch & Forero, 2007
Kiskeyana palassaina (Weirauch & Forero, 2007a: 63). Weirauch & Forero, 2007b: 68 *DR > TL: Santiago: Parque Nacional
Armando Bermúdez, along road to Mata Grande station, 914 m
Oncerotrachelus Stål, 1868
Oncerotrachelus cubanus Barber & Bruner, 1937: 182
Subfamily STENOPODAINAE
Oncocephalus Fischer, 1813
Oncocephalus ventralis Walker, 1873: 24. Maldonado Capriles, 1990: 521 *H, *DR > TL: St. Domingo
Stenopoda Laporte, 1832
Stenopoda cinerea Laporte, 1833. Barber, 1939: 388; Maldonado Capriles, 1990: 540
Subfamily TRIATOMINAE
Panstrongylus Berg, 1879
Panstrongylus hispaniolae† Poinar, 2014: 4 [Fossil in Dominican amber]
Triatoma Laporte, 1832
Triatoma dominicana† Poinar, 2005: 76 [Fossil in Dominican amber]
Triatoma rubrofasciata (De Geer, 1773). Neiva & Lent, 1941: 84; Lent & Wygodzinsky, 1979: 312 [Introduced]

Infraorder PENTATOMOMORPHA
Superfamily ARADOIDEA
Family ARADIDAE
Key reference: Kormilev and Froeschner (1987).
Subfamily ANEURINAE
Aneurus Curtis, 1825
Aneurus haitiensis Kormilev, 1968: 6. Kormilev & Froeschner, 1987: 27 *H > TL: Morne Guimby, 22 km southeast of Fond
Verrettes, 6500’
Aneurus hispaniolensis Picchi, 1977: 273. Kormilev & Froeschner, 1987: 27 *H, *DR > TL: La Vega Prov., 14 km north of
Jarabacoa, on road to La Vega
Subfamily CALISIINAE
Calisiopsis Champion, 1898
Calisiopsis brodzinskyorum† Froeschner, 1992: 34 [Fossil in Dominican amber]
Subfamily CARVENTINAE
Acaricoris Harris & Drake, 1944
Acaricoris haitiensis Kormilev, 1968: 2. Kormilev & Froeschner, 1987: 66 *H > TL: Morne Guimby, 22 km southeast of Fond

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 247
Verrettes, 6500 ft.
Acaricoris lattini Heiss, 2008: 948 *DR > TL: road La Vega - Jarabacoa
Acaricoris robertae† Heiss & Poinar, 2012: 51 [Fossil in Dominican amber]
Aglaocoris Drake & Maldonado Capriles, 1955
Aglaocoris drakei Kormilev, 1968: 5. Kormilev & Froeschner, 1987: 67 *H > TL: Port-au-Prince
Aglaocoris rectangulatus Usinger & Matsuda, 1959: 141. Kormilev & Froeschner, 1987: 67 *DR > TL: San Domingo
Carventius Stål, 1865
Carventius bechlyi† Heiss in Heiss & Poinar, 2012: 50 [Fossil in Dominican amber]
Eretmocoris Harris & Drake, 1944
Eretmocoris longicornis Usinger & Matsuda, 1959: 138. Kormilev & Froeschner, 1987: 78 *H > TL: Haiti
Eretmocoris sp.†—Poinar & Poinar (1999: 203) [Fossil in Dominican amber]
Neoproxius Usinger & Matsuda, 1959
Neoproxius latocanus† Heiss & Poinar, 2012: 48 [Fossil in Dominican amber]
Proneoproxius† Heiss & Poinar, 2012
Proneoproxius cornutus† Heiss & Poinar, 2012: 53 [Fossil in Dominican amber]
Rhysocoris Usinger & Matsuda, 1959
Rhysocoris rugosus Usinger & Matsuda, 1959: 130. Kormilev & Froeschner, 1987: 90 *H > TL: N. E. Foothills, La Hotte,
2,400 ft.
Subfamily MEZIRINAE
Brevisensoria Poinar in Poinar & Heiss, 2012
Brevisensoria incrustata† Poinar in Poinar & Heiss, 2012: 57 [Fossil in Dominican amber]
Mezira Amyot & Serville, 1843
Mezira abdominalis (Stål, 1873). Barber, 1939: 330
Mezira amberdominica† Heiss, 2012: 76 [Fossil in Dominican amber]
Mezira jamaicensis Bergroth, 1906. Perez-Gelabert, 2008: 189
Mezira legorskyi† Heiss, 2012: 74 [Fossil in Dominican amber]
Mezira placida haitiensis Kormilev, 1968: 15. Kormilev & Froeschner, 1987: 156
Mezira scheveni† Heiss, 2000: 7 [Fossil in Dominican amber]
Notapictinus Usinger & Matsuda, 1959
Notapictinus dominicus (Usinger, 1936: 512). Kormilev & Froeschner, 1987: 178 *DR > TL: La Cavite, St. Domingo

Superfamily LYGAEOIDEA
Key reference: Baranowski and Slater (2005).

Family BERYTIDAE
Subfamily METACANTHINAE
Gampsocoris Fuss, 1852
Gampsocoris decorus (Uhler, 1893). Henry, 1997: 25
Jalysus Stål, 1860
Jalysus reductus Barber, 1939: 331
Metacanthus Hawle & Corda, 1847
Metacanthus tenellus Stål, 1859. Henry, 1997: 129

Family BLISSIDAE
Blissus Burmeister, 1835
Blissus antillus Leonard, 1968. Baranowski & Slater, 2005: 59
Ischnodemus Fieber, 1837
Ischnodemus sallei (Signoret, 1857). Baranowski & Slater, 2005: 62
Ischnodemus variegatus (Signoret, 1857). Baranowski & Slater, 2005: 63
Toonglasa Distant, 1893
Toonglasa discalis (Barber, 1947). Baranowski & Slater, 2005: 64

Family CYMIDAE
Cymodema Spinola, 1837

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Cymodema breviceps (Stål, 1874). Baranowski & Slater, 2005: 54
Cymoninus Breddin, 1907
Cymoninus notabilis (Distant, 1882). Baranowski & Slater, 2005: 56
Neoninus Distant, 1882
Neoninus illustris Distant, 1882. Baranowski & Slater, 2005: 55

Family GEOCORIDAE
Subfamily BLEDIONOTINAE
Neopamphantus Barber & Bruner, 1933
Neopamphantus hispaniolus Slater, 1965: 188. Baranowski & Slater, 2005: 75 *H, *DR > TL: Kenscoff
Pamphantus Stål, 1874
Pamphantus atrohumeralis atrohumeralis Barber & Bruner, 1933: 537. Baranowski & Slater, 2005: 76 *H > TL: Le Trou
Pamphantus atrohumeralis dominicanus Slater, 1981. Baranowski & Slater, 2005: 76 *DR > TL: Sabana Abajo, Puerto Plata
Prov.
Pamphantus barberi Slater in Baranowski & Slater, 2005: 76 *DR > TL: Prov. Pedernales, km 24.5 N Cabo Rojo, elev. 750 m
Pamphantus pallidoides Slater, 1981: 83. Baranowski & Slater, 2005: 78 *DR > TL: Puerto Plata
Pamphantus trimaculatus Slater, 1981: 85. Baranowski & Slater, 2005: 79 *DR > TL: 3 miles W. Haina, San Cristóbal Prov.
Subfamily GEOCORINAE
Geocoris Fallén, 1814
Geocoris floridanus Blatchley, 1926. Baranowski & Slater, 2005: 65
Geocoris lividipennis Stål, 1862. Baranowski & Slater, 2005: 66
Geocoris punctipes (Say, 1832). Baranowski & Slater, 2005: 66

Family LYGAEIDAE
Subfamily ISCHNORHYNCHINAE
Kleidocerys Stephens, 1829
Kleidocerys hispaniola Baranowski in Baranowski & Slater, 2005: 51 *DR, *H > TL: Ebano Verde Nat. Park
Kleidocerys virescens (Fabricius, 1794). Baranowski & Slater, 2005: 51
Subfamily LYGAEINAE
Anachrostomus A. Slater, 1992
Anachrostomus formosoides Baranowski in Baranowski & Slater, 2005: 12 *DR > TL: Río Chavón
Anachrostomus formosus (Blanchard, 1840). Baranowski & Slater, 2005: 12
Hadrosomus A. Slater, 1992
Hadrosomus confraternus (Uhler, 1869). Baranowski & Slater, 2005: 15
Hadrosomus nigrocoxalis Baranowski & Brailovsky, 1999: 596. Baranowski & Slater, 2005: 15 *DR > TL: Río Chavón
Lygaeus Fabricius, 1794
Lygaeus alboornatus (Blanchard, 1852). Baranowski & Slater, 2005: 17
Lygaeus argutus Brailovsky, 1982: 271. Baranowski & Slater, 2005: 18
Lygaeus bahamensis Barber & Ashlock, 1960. Baranowski & Slater, 2005: 18
Melanopleuroides A. Slater & Baranowski, 2001
Melanopleuroides dominicanus A. Slater & Baranowski, 2001: 131. Baranowski & Slater, 2005: 20 *DR > TL: Parque
Nacional Sierra de Bahoruco, Pedernales Prov.
Melanopleurus Stål, 1874
Melanopleurus aureus Distant, 1882: 188. Baranowski & Slater, 2005: 21
Melanopleurus inflatus Brailovsky, 1979. Baranowski & Slater, 2005: 22 *DR > TL: Prov. of Pedernales, 36 km N of Cabo
Rojo
Melanopleurus maculicorium Maldonado Capriles, 1974: 24. Baranowski & Slater, 2005: 22 *DR > TL: Sección La Palma,
Constanza, 900 m
Ochrimnus Stål, 1874
Ochrimnus (Ochrimnus) collaris (Fabricius, 1803). Baranowski & Slater, 2005: 26
Ochrimnus (Ochrimnus) laevus Brailovsky, 1982. Baranowski & Slater, 2005: 28
Ochrimnus (Orthochrimnus) marginatus Baranowski in Baranowski & Slater, 2005: 29 *DR > TL: 20 km N San José de Ocoa
Ochrimnus (Ochrimnus) pseudocollaris Brailovsky, 1982. Baranowski & Slater, 2005: 30
Ochrostomus Stål, 1874

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Ochrostomus pulchellus (Fabricius, 1794). Baranowski & Slater, 2005: 31
Oncopeltus Stål, 18768
Oncopeltus aulicus (Fabricius, 1775). Baranowski & Slater, 2005: 33
Oncopeltus cingulifer Stål, 1874. Baranowski & Slater, 2005: 35
Oncopeltus fasciatus (Dallas, 1852). Baranowski & Slater, 2005: 36
Oncopeltus femoralis Stål, 1874. Baranowski & Slater, 2005: 37
Oncopeltus pictus Van Duzee, 1907. Baranowski & Slater, 2005: 38
Oncopeltus semilimbatus Stål, 1874. Baranowski & Slater, 2005: 39
Oncopeltus varicolor (Fabricius, 1794). Baranowski & Slater, 2005: 41
Subfamily ORSILLINAE
Neortholomus Hamilton, 1983
Neortholomus jamaicensis (Dallas, 1852). Baranowski & Slater, 2005: 48
Nysius Dallas, 1852
Nysius raphanus Howard, 1872. Baranowski & Slater, 2005: 46
Nysius scutellatus Dallas, 1852. Baranowski & Slater, 2005: 47
Nysius tenellus Barber, 1947. Baranowski & Slater, 2005: 47
Xyonysius Ashlock & Lattin, 1963
Xyonysius basalis (Dallas, 1852). Baranowski & Slater, 2005: 43
Xyonysius californicus (Stål, 1859). Baranowski & Slater, 2005: 44

Family NINIDAE
Ninyas Distant, 1882
Ninyas deficiens (Lethierry, 1881). Baranowski & Slater, 2005: 68
Ninyas dominicanus Baranowski in Baranowski & Slater, 2005: 70 *DR > TL: 33 km N San José de Ocoa
Ninyas woodruffi Baranowski in Baranowski & Slater, 2005: 73 *DR > TL: 33 km N San José de Ocoa

Family OXYCARENIDAE
Oxycarenus Fieber, 1837
Oxycarenus hyalinipennis (Costa, 1847). Baranowski & Slater, 2005: 81 [Introduced]

Family PACHYGRONTHIDAE
Cymophyes Fieber, 1870
Cymophyes nesocoris Baranowski & Slater, 1997. Baranowski & Slater, 2005: 92
Oedancala Amyot & Serville, 1843
Oedancala bimaculata (Distant, 1893). Baranowski & Slater, 2005: 83
Pachygrontha Germar, 1837
Pachygrontha compacta Distant, 1893. Baranowski & Slater, 2005: 87

Family RHYPAROCHROMIDAE
Tribe ANTILLOCORINI
Antillocoris Kirkaldy, 1874
Antillocoris pallidus (Uhler, 1894). Baranowski & Slater, 2005: 95
Antillocoris pilosulus (Stål, 1874). Baranowski & Slater, 2005: 96
Bathydema Uhler, 1893
Bathydema hispaniola Slater & Baranowski in Slater et al., 1977: 351 *DR > TL: Loma Rucilla and mts. N., 5–8000 ft.
Cligenes Distant, 1893
Cligenes distinctus Distant, 1893. Baranowski & Slater, 2005: 105
Tribe CLERADINI
Clerada Signoret, 1863
Clerada apicicornis Signoret, 1863. Baranowski & Slater, 2005: 110
Tribe LETHAEINI
Bubaces Distant, 1893
Bubaces uhleri (Distant, 1901). Baranowski & Slater, 2005: 112
Neopetissius O’Donnell, 2001

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Neopetissius variegatus O’Donnell, 2001. Baranowski & Slater, 2005: 120
Paragonatas Barber, 1939
Paragonatas divergens (Distant, 1882). Baranowski & Slater, 2005: 121
Valtissius Barber, 1918
Valtissius distinctus (Distant, 1901). Baranowski & Slater, 2005: 126
Tribe MYODOCHINI
Catenes Distant, 1893
Catenes spiculus Slater & Baranowski, 1994: 489. Baranowski & Slater, 2005: 131 *DR > TL: Guanuma, Finca Goya
Froeschneria Harrington, 1980
Froeschneria multispinus (Stål, 1874). Baranowski & Slater, 2005: 133
Froeschneria piligera (Stål, 1862). Baranowski & Slater, 2005: 135
Heraeus Stål, 1862
Heraeus caliginosus Slater & Baranowski, 1994: 491. Baranowski & Slater, 2005: 137; Dellapé et al., 2016: 50 *DR, *H > TL:
21 km N Cabo Rojo, Pedernales Prov.
Heraeus concolor Slater & Baranowski, 1994: 493. Baranowski & Slater, 2005: 137; Dellapé et al., 2016: 88
Heraeus dominicanus Dellapé et al., 2016: 51 *DR > TL: San Francisco Mountains [San Cristóbal Prov.]
Heraeus guttatus (Dallas, 1852). Baranowski & Slater, 2005: 139; Dellapé et al., 2016: 52
Heraeus hollyae Baranowski, 2005. Dellapé et al., 2016: 55
Heraeus plebejus Stål, 1874. Baranowski & Slater, 2005: 142
Heraeus sp. 1—Baranowski & Slater (2005: 145)
Heraeus sp. 2—Baranowski & Slater (2005: 145)
Heraeus triguttatus (Guérin-Méneville, 1857). Baranowski & Slater, 2005: 144
Ligyrocoris Stål, 1872
Ligyrocoris litigiosus (Stål, 1862). Baranowski & Slater, 2005: 145
Myodocha Ltreille, 1807
Myodocha froeschneri A. Slater, 1998: 10. Baranowski & Slater, 2005: 147
Neopamera Harrington, 1980
Neopamera albocincta (Barber, 1953). Baranowski & Slater, 2005: 150
Neopamera bilobata (Say, 1831). Baranowski & Slater, 2005: 152
Neopamera intermedia (Barber, 1923). Baranowski & Slater, 2005: 157
Neopamera neotropicalis (Kirkaldy, 1909). Baranowski & Slater, 2005: 158
Neopamera vicarius (Barber, 1954). Baranowski & Slater, 2005: 160
Paromius Fieber, 1861
Paromius dohrnii (Guérin-Méneville, 1857). Baranowski & Slater, 2005: 171
Paromius longulus (Dallas, 1852). Baranowski & Slater, 2005: 172
Prytanes Distant, 1893
Prytanes brunnea Baranowski in Baranowski & Slater, 2005: 175 *DR > TL: Prov. Pedernales, km 12 N Cabo Rojo
Prytanes dissimilis (Barber, 1953). Baranowski & Slater, 2005: 176
Prytanes formosa (Distant, 1882). Baranowski & Slater, 2005: 177
Prytanes minima (Guérin-Méneville, 1857). Baranowski & Slater, 2005: 179
Prytanes oblonga (Stål, 1862). Baranowski & Slater, 2005: 179
Pseudopachybrachius Malipatil, 1978
Pseudopachybrachius concepcioni Zheng & Slater, 1984: 98. Baranowski & Slater, 2005: 164
Pseudopachybrachius vinctus (Say, 1831). Uhler, 1894: 186; Baranowski & Slater, 2005: 166
Tribe OZOPHORINI
Ozophora Uhler, 1871
Ozophora atropicta Barber, 1939. Baranowski & Slater, 2005: 189
Ozophora auroseta Baranowski in Baranowski & Slater, 2005: 192 *DR > TL: La Vega, Res. Center Ebano Verde
Ozophora burmeisteri (Guérin-Méneville, 1857). Baranowski & Slater, 2005: 193
Ozophora cacumena Slater in Baranowski & Slater, 2005: 195 *DR > TL: La Vega-Monseñor Nouel, Loma El Casabito
(summit), 1390 m
Ozophora caliginosa Slater, 1990: 143. Baranowski & Slater, 2005: 196 *H, *DR > TL: Furcy
Ozophora caribbee Baranowski & Slater, 1984: 444. Baranowski & Slater, 2005: 196
Ozophora caroli Slater & Baranowski, 1983. Baranowski & Slater, 2005: 197

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Ozophora darlingtoni Slater, 1990: 144. Baranowski & Slater, 2005: 204 *H, *DR > TL: La Visite & vic. La Selle Range, 5–
7,000 ft.
Ozophora divaricata Barber, 1954. Baranowski & Slater, 2005: 204
Ozophora fuscipennis Baranowski in Baranowski & Slater, 2005: 206 *DR > TL: Prov. La Altagracia, Nisibón (Papagallo)
Ozophora helenae Baranowski & Slater, 1984: 457. Baranowski & Slater, 2005: 208
Ozophora heydoni Barber & Ashlock, 1960. Baranowski & Slater, 2005: 210
Ozophora hispaniola Slater, 1990: 140. Baranowski & Slater, 2005: 212 *H, *DR > TL: Port-au-Prince
Ozophora insulanus Slater in Baranowski & Slater, 2005: 215 *DR > TL: Pedernales, along Río Mulito, 13 km N Pedernales,
230 m
Ozophora laticephala Slater & O’Donnell, 1979. Baranowski & Slater, 2005: 216
Ozophora pallescens (Distant, 1893). Baranowski & Slater, 2005: 224
Ozophora parimpicta Baranowski in Baranowski & Slater, 2005: 225
Ozophora psarocoris Slater in Baranowski & Slater, 2005: 226 *DR > TL: Pedernales, 5 km NE Los Arroyos, 1680 m
Ozophora pusilla Slater, 1990: 145. Baranowski & Slater, 2005: 229 *DR > TL: Peravia, 16 km E San José de Ocoa
Ozophora quinquemaculata quinquemaculata Barber, 1939. Baranowski & Slater, 2005: 230
Ozophora rawlinsi Slater in Baranowski & Slater, 2005: 233 *DR > TL: Independencia, 4 km S Los Pinos, Loma de Vientos,
455 m
Ozophora reperta Blatchley, 1926. Baranowski & Slater, 2005: 234
Ozophora rubrolinea Slater, 1987. Baranowski & Slater, 2005: 235
Ozophora testacea Slater, 1990: 142. Baranowski & Slater, 2005: 238 *H, *DR > TL: Manneville, about 60 ft.
Ozophora triangularis Slater in Baranowski & Slater, 2005: 239 *DR, *H > TL: 5 km NE Los Arroyos, 1680 m
Ozophora umbrosa Slater, 1987b: 142. Baranowski & Slater, 2005: 241
Ozophora venusta Slater in Baranowski & Slater, 2005: 244 *DR > TL: Pedernales, 9.5 km N Cabo Rojo,
Ozophora woodruffi Slater in Baranowski & Slater, 2005: 245; Woodruff & Slater, 2012: 1 *DR > TL: Prov. La Vega, La
Ciénaga de Manabao, Park Hdqt.
Note: Originally described without type data, this species was validated in 2012.
Tribe RHYPAROCHROMINI
Dieuches Dohrn, 1860
Dieuches armatipes (Walker, 1872). Baranowski & Slater, 2005: 247 [Introduced]

Superfamily PYRRHOCOROIDEA
Family LARGIDAE
Tribe LARGULINI
Largus Hahn, 1831
Largus obovatus (Barber, 1923: 5). Barber, 1939: 364
Neolargulus Stehlík & Brailovsky, 2011
Neolargulus excavatus Stehlík & Brailovsky, 2011: 452 *DR > TL: 4 km E. of La Ciénaga de Manabao, 3050 ft.

Family PYRRHOCORIDAE
Subfamily PYRRHOCORINAE
Dysdercus Audinet-Serville, 1835
Dysdercus andreae (Linnaeus, 1758). Barber, 1923: 24; Barber, 1939: 366; Van Doesburg, 1968: 53
Dysdercus jamaicensis jindrai Stehlik in Schaefer & Stehlik, 2013: 375
Dysdercus mimulus mimulus Hussey, 1929. Van Doesburg, 1968: 104
Dysdercus ochreatus ochreatus (Say, 1832). Van Doesburg, 1968: 45
=Dysdercus fervens Walker, 1872: 182
Dysdercus sanguinarius sanguinarius Stål, 1870. Barber, 1939: 366; Van Doesburg, 1968: 48

Superfamily COREOIDEA
Family ALYDIDAE
Subfamily ALYDINAE
Burtinus Stål, 1860
Burtinus luteomarginatus Maldonado Capriles, 1953. Schaefer & Ahmad, 2007: 835
Neomegalotomus Schaffner & Schaefer, 1998

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Neomegalotomus rufipes (Westwood, 1842). Barber, 1923: 21; Barber, 1939: 323
Subfamily MICRELYTRINAE
Tribe LEPTOCORISINI
Leptocorisa Latreille, 1829
Leptocorisa filiformis (Fabricius, 1775). Barber, 1939: 324
Stenocoris Burmeister, 1839
Stenocoris furcifera (Westwood, 1842). Abud, 1993: 9/93

Family COREIDAE
Key reference: Brailovsky & Perez-Gelabert (2019).
Subfamily MEROPACHYINAE
Tribe MEROCORINI
Merocoris Perty, 1930
Merocoris (Corynocoris) distinctus distinctus Dallas, 1852. Brailovsky & Perez-Gelabert, 2019: 206
Merocoris (Corynocoris) typhaeus (Fabricius, 1798). Brailovsky & Perez-Gelabert, 2019: 206
Subfamily COREINAE
Tribe ACANTHOCERINI
Acanthocerus Palisot de Beauvois, 1818
Acanthocerus crucifer Palisot de Beauvois, 1818. Wolcott, 1927: 183; Brailovsky & Perez-Gelabert, 2019: 207
=Acanthocerus cruciger O’Shea, 1980: 61 [Unnecessary replacement name]
Tribe ANISOSCELINI
Chondrocera Laporte, 1832
Chondrocera mirabilis Brailovsky & Barrera, 1994: 54. Brailovsky & Perez-Gelabert, 2019: 210 *DR > TL: San Pedro de
Macorís
Leptoglossus Swainson, 1837
Leptoglossus balteatus (Linnaeus, 1771). Barber, 1939: 311; Allen, 1969: 97; Schmutterer, 1990: 293; Brailovsky & Perez-
Gelabert, 2019: 211
Leptoglossus concolor (Walker, 1871). Allen, 1969: 118; Brailovsky & Perez-Gelabert, 2019: 212
Leptoglossus confusus Alayo & Grillo, 1977. Brailovsky & Perez-Gelabert, 2019: 212
Leptoglossus gonagra (Fabricius, 1775). Allen, 1969: 84; Brailovsky & Perez-Gelabert, 2019: 213
Mamurius Stål, 1862
Mamurius cubanus Barber & Bruner, 1947. Brailovsky & Perez-Gelabert, 2019: 214
Phthia Stål, 1862
Phthia rubropicta (Westwood, 1842). Brailovsky & Perez-Gelabert, 2019: 215
Phthiacnemia Brailovsky, 2009
Phthiacnemia picta (Drury, 1770). Uhler, 1894: 179; Barber, 1923: 17; Brailovsky & Perez-Gelabert, 2019: 215
Rhytidophthia Brailovsky, 2009
Rhytidophthia splendida (Valdes, 1910). Brailovsky & Perez-Gelabert, 2019: 216
Tribe CHARIESTERINI
Chariesterus Laporte, 1832
Chariesterus gracilicornis Stål, 1870. Barber, 1923: 18; Barber, 1939: 316; Brailovsky & Perez-Gelabert, 2019: 217
Tribe HYPSELONOTINI
Althos Kirkaldy, 1904
Althos obscurator (Fabricius, 1803). Brailovsky & Perez-Gelabert, 2019: 220
Tribe COREINI
Anasa Amyot & Serville, 1843
Anasa acutangula Stål, 1870. Brailovsky & Perez-Gelabert, 2019: 222
Anasa bellator (Fabricius, 1787). Brailovsky, 1985: 90; Brailovsky & Perez-Gelabert, 2019: 222
Anasa pisina Brailovsky, 2001: 43. Brailovsky & Perez-Gelabert, 2019: 223 *DR > TL: La Vega, vic. Salto de Aguas Blancas
Anasa scorbutica Fabricius, 1775. Barber, 1923: 19; Brailovsky & Perez-Gelabert, 2019: 223
Anasa tristis (De Geer, 1773). Brailovsky & Perez-Gelabert, 2019: 225
Catorhintha Stål, 1859
Catorhintha guttula (Fabricius, 1794). Barber, 1923: 19; Packauskas, 2010: 115; Brailovsky & Perez-Gelabert, 2019: 225
Catorhintha selector Stål, 1859. Brailovsky & García, 1987: 55; Brailovsky & Perez-Gelabert, 2019: 226

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Sethenira Spinola, 1837
Sethenira ferruginea Stål, 1870. Brailovsky & Perez-Gelabert, 2019: 227
Zicca Amyot & Serville, 1843
Zicca gloriosa Brailovsky & Perez-Gelabert, 2019: 228 *DR > TL: Santo Domingo Prov., Sierra Prieta, lower hill, 140 m
Zicca rubricator rubricator (Fabricius, 1803). Brailovsky & Perez-Gelabert, 2019: 228
Zicca taeniola (Dallas, 1852). Brailovsky & Cadena, 1992: 90; Brailovsky & Perez-Gelabert, 2019: 229
Tribe SPARTHOCERINI
Eubule Stål, 1867
Eubule spartocerana Brailovsky, 1992. Brailovsky & Perez-Gelabert, 2019: 231
Spartocera Laporte, 1832
Spartocera batatas (Fabricius, 1798). Barber, 1939: 314; Halbert & Brambila, 2005: 135; Brailovsky & Perez-Gelabert, 2019:
232
Spartocera fusca (Thunberg, 1783). Perez-Gelabert, 2008: 194; Brailovsky & Perez-Gelabert, 2019: 233
Notes: The species Leptoglossus cinctus (Herrich-Schäffer, 1836) cited by Wolcott, 1927: 184 and Leptoglossus stigma
(Herbst, 1784) cited by Barber, 1939: 311 are based on erroneous identifications. These species are not part of the Hispaniolan
fauna (Brailovsky & Perez-Gelabert, 2019).

Family RHOPALIDAE
Subfamily RHOPALINAE
Corizus Fallén, 1814
Corizus punctatus Signoret, 1859. Hambleton, 1908: 144
Harmostes Burmeister, 1835
Harmostes serratus (Fabricius, 1794). Barber, 1923: 21; Barber, 1939: 325
Niesthrea Spinola, 1837
Niesthrea louisianica Sailer, 1961 [By synonymy of species below]
=Corizus hyalinus (Fabricius, 1794). Hambleton, 1908: 137; Barber, 1923: 22
Niesthrea sidae (Fabricius, 1794). Hambleton, 1908: 142; Barber, 1923: 22; Barber, 1939: 327
Subfamily SERINETHINAE
Jadera Stål, 1860
Jadera aeola (Dallas, 1852) [By synonymy of species below]
=Jadera antica (Walker, 1872). Barber, 1954: 2

Superfamily PENTATOMOIDEA
Family CYDNIDAE
Subfamily CYDNINAE
Amnestus Dallas, 1851
Amnestus electricus† Thomas, 1994: 305 [Fossil in Dominican amber]
Amnestus priscus† Thomas, 1994: 307 [Fossil in Dominican amber]
Amnestus pusio (Stål, 1860). Barber, 1939: 274; Froeschner, 1960: 654
Amnestus radialis Froeschner, 1960. Perez-Gelabert, 2008: 194
Amnestus subferrugineus (Westwood, 1837). Barber, 1939: 273
Amnestus trimaculatus Froeschner, 1960. Perez-Gelabert, 2008: 194
Dallasiellus Derg, 1901
Dallasiellus (Dallasiellus) succineus† Thomas, 1994: 304 [Fossil in Dominican amber]
Dallasiellus (Ecarinoceps) laevis Froeschner, 1960: 589 *DR, *H > TL: Constanza
Dallasiellus (Ecarinoceps) scitus (Walker, 1868). Froeschner, 1960: 587
Melanaethus Uhler, 1875
Melanaethus spinolae (Signoret, 1863). Froeschner, 1960: 449
Rhytidoporus Uhler, 1877
Rhytidoporus (Rhytidoporus) indentatus (Uhler, 1877). Barber, 1939: 271; Froeschner, 1960: 387
Rhytidoporus (Rhytidoporus) obsoletus Froeschner, 1960: 389 *H > TL: La Visite & vic., La Selle Range, 5–7,000 ft.
Tominotus Mulsant & Rey, 1866
Tominotus communis (Uhler, 1877). Froeschner, 1960: 551

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Family CORIMELAENIDAE
Corimelaena White, 1839
Corimelaena (Corimelaena) minuta (Uhler, 1863). Barber, 1939: 269
Corimelaena (Parapora) cognata (Van Duzee, 1907). Perez-Gelabert, 2008: 195
Galgupha Amyot & Serville, 1843
Galgupha (Gyrocnemis) guttiger (Stål, 1862). Perez-Gelabert, 2008: 195

Family MEGARIDIDAE
Megaris Stål, 1860
Megaris sp. 1—Perez-Gelabert (2008: 195)
Megaris sp. 2—Perez-Gelabert (2008: 195)
Minysporus† Poinar & Heiss, 2013
Minisporus dominicanus† Poinar & Heiss, 2013: 95 [Fossil in Dominican amber]

Family PENTATOMIDAE
Key reference: Perez-Gelabert and Thomas (2005).
Subfamily ASOPINAE
Alcaeorrhynchus Bergroth, 1891
Alcaeorrhynchus phymatophorus (Palisot de Beauvois, 1811: 112). Perez-Gelabert & Thomas, 2005: 323
Andrallus Bergroth, 1905
Andrallus spinidens (Fabricius, 1787). Perez-Gelabert & Thomas, 2005: 324
Podisus Herrich-Schäffer, 1851
Podisus maculiventris (Say, 1831). Perez-Gelabert & Thomas, 2005: 324
Podisus mucronatus Uhler, 1897. Perez-Gelabert & Thomas, 2005: 324
Podisus sagitta (Fabricius, 1794). Perez-Gelabert & Thomas, 2005: 325
Podisus subferrugineus Barber & Bruner, 1932. De los Santos & Bastardo, 2012: 121
Stiretrus Laporte, 1832
Stiretrus quinquepunctatus Germar, 1839: 20. Schumacher, 1912: 94; Perez-Gelabert & Thomas, 2005: 325 *H > TL: By error
in original description “Isle de Prince (Guinea)”. Type bears a label for Port-au-Prince, Haiti (Perez-Gelabert & Thomas 2005:
325).
Tylospilus Stål, 1870
Tylospilus acutissimus (Stål, 1870). Perez-Gelabert & Thomas, 2005: 325
Tyrannocoris Thomas, 1992
Tyrannocoris jole (Stål, 1862). Perez-Gelabert & Thomas, 2005: 325
Subfamily DISCOCEPHALINAE
Tribe OCHLERINI
Alathetus Dallas, 1851
Alathetus haitiensis Rolston, 1982: 156. Perez-Gelabert & Thomas, 2005: 325 *H > TL: Roche Croix, Mt. La Hotte, 5000 ft.
Subfamily EDESSINAE
Edessa Fabricius, 1803
Edessa bifida (Say, 1832). Perez-Gelabert & Thomas, 2005: 345
Edessa chlorophyla Barber & Bruner, 1932. Perez-Gelabert & Thomas, 2005: 346
Edessa cornuta Burmeister, 1835. Perez-Gelabert & Thomas, 2005: 346
Edessa excoriata Barber & Bruner, 1932. Perez-Gelabert & Thomas, 2005: 346
Edessa meditabunda (Fabricius, 1794). Perez-Gelabert & Thomas, 2005: 346
Edessa rawlinsi Thomas in Perez-Gelabert & Thomas, 2005: 346 *DR > TL: Barahona Prov., Eastern Sierra de Bahoruco,
Reserva Cachote, 1230 m
Subfamily PENTATOMINAE
Tribe CARPOCORINI
Euschistus Dallas, 1851
Euschistus acuminatus Walker, 1867. Perez-Gelabert & Thomas, 2005: 333
Euschistus bifibulus (Palisot de Beauvois, 1805). Perez-Gelabert & Thomas, 2005: 333
Euschistus crassus Dallas, 1851. Perez-Gelabert & Thomas, 2005: 333
Euschistus crenator (Fabricius, 1794). Perez-Gelabert & Thomas, 2005: 333

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Euschistus obscurus (Palisot de Beauvois, 1817). Perez-Gelabert & Thomas, 2005: 334
Tribe PENTATOMINI
Agonoscelis Spinola, 1837
Agonoscelis puberula Stål, 1857. Thomas et al., 2003: 151; Perez-Gelabert & Thomas, 2005: 327 [Introduced]
Arocera Spinola, 1837
Arocera (Euopta) placens (Walker, 1867). Perez-Gelabert & Thomas, 2005: 328
Arvelius Spinola, 1837
Arvelius albopunctatus (De Geer, 1773). Perez-Gelabert & Thomas, 2005: 328
Arvelius haitianus Brailovsky, 1981: 271. Perez-Gelabert & Thomas, 2005: 328 *H > TL: Le Trox
Arvelius porrectispinus Breddin, 1909. Perez-Gelabert & Thomas, 2005: 328
Banasa Stål, 1860
Banasa flavosa Thomas in Perez-Gelabert & Thomas, 2005: 329 *DR > TL: La Vega Prov., 4 km E of La Ciénaga de Manabao,
3050 ft.
Banasa herbacea (Stål, 1872). Perez-Gelabert & Thomas, 2005: 328
Banasa lenticularis Uhler, 1894. Perez-Gelabert & Thomas, 2005: 329
Banasa punctata Thomas in Perez-Gelabert & Thomas, 2005: 331 *DR > TL: La Vega Prov., El Convento, Constanza
Banasa punctatissima Barber & Bruner, 1932. Perez-Gelabert & Thomas, 2005: 329
Banasa zeteki Sailer, 1959. Perez-Gelabert & Thomas, 2005: 329
Caribo Rolston, 1984
Caribo fasciatus Rolston, 1984. Perez-Gelabert & Thomas, 2005: 332
Chinavia Orian, 1965
Chinavia insulani (Rolston, 1983: 113). Perez-Gelabert & Thomas, 2005: 325 *DR > TL: Constanza to Jarabacoa, 4000 ft.
Chinavia marginata (Palisot de Beauvois, 1817). Perez-Gelabert & Thomas, 2005: 326
Chinavia montivaga (Distant, 1890). Perez-Gelabert & Thomas, 2005: 327
Chinavia ubica (Rolston, 1983). Perez-Gelabert & Thomas, 2005: 327
Chinavia wygodzinskyi (Rolston, 1983). Perez-Gelabert & Thomas, 2005: 327
Chroantha Stål, 1872
Chroantha ornatula (Herrich-Schäffer, 1842). Perez-Gelabert & Thomas, 2005: 332 [Introduced]
Cyptocephala Berg, 1883
Cyptocephala antiguensis (Westwood, 1837). Perez-Gelabert & Thomas, 2005: 332
Cyptocephala bimini (Ruckes, 1952). Perez-Gelabert & Thomas, 2005: 332
Cyptocephala pallida Rolston, 1986. Perez-Gelabert & Thomas, 2005: 332
Fecelia Stål, 1872
Fecelia biorbis Eger, 1980: 29. Perez-Gelabert & Thomas, 2005: 334 *H, *DR > TL: Furcy, 4000 ft.
Fecelia nigridens (Walker, 1867). Perez-Gelabert & Thomas, 2005: 334
Fecelia proxima Grazia, 1980: 265. Perez-Gelabert & Thomas, 2005: 335
Grazia Rolston, 1981
Grazia tincta (Distant, 1880). Perez-Gelabert & Thomas, 2005: 335
Loxa Amyot & Serville, 1843
Loxa nesiotes Horvath, 1925. Perez-Gelabert & Thomas, 2005: 335
Loxa pallida Van Duzee, 1907. Perez-Gelabert & Thomas, 2005: 335
Loxa viridis (Palisot de Beauvois, 1805). Perez-Gelabert & Thomas, 2005: 335
Mediocampus Thomas, 1994
Mediocampus dominicanus Thomas, 1994: 259. Perez-Gelabert & Thomas, 2005: 336 *DR > TL: Pedernales Prov., Río
Pedernales, near Fuerte Banano, 280 m
Mediocampus perezi Thomas in Perez-Gelabert & Thomas, 2005: 336 *DR > TL: Independencia Prov. 4–5 km S Puerto
Escondido, on trail to Caseta 1, ~950 m
Mediocampus woodruffi Thomas in Perez-Gelabert & Thomas, 2005: 338 *DR > TL: La Altagracia Prov., 2.7 km E Batey
Papagayo, 150 ft.
Menudo Thomas, 1990
Menudo femoralis Thomas, 1994. Perez-Gelabert & Thomas, 2005: 339
Mormidea Amyot & Serville, 1843
Mormidea albisignis Stål, 1872. Perez-Gelabert & Thomas, 2005: 339
Mormidea angustata Stål, 1862. Perez-Gelabert & Thomas, 2005: 339

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Mormidea cubrosa (Dallas, 1851). Perez-Gelabert & Thomas, 2005: 339
Murgantia Stål, 1862
Murgantia thomasi McPherson & Ahmad, 2011: 149 *DR > TL: Boca de Yuma, La Altagracia Prov.
=Murgantia varicolor (Westwood, 1837). Perez-Gelabert & Thomas, 2005: 340 [Misidentification]
Nezara Amyot & Serville, 1843
Nezara viridula (Linnaeus, 1758). Santoro, 1960: 351; Schmutterer, 1990: 345; Perez-Gelabert & Thomas, 2005: 340
Oebalus Stål, 1862
Oebalus insularis (Stål, 1872). Perez-Gelabert & Thomas, 2005: 340
Oebalus linki (Heinemann, 1917). De los Santos & Bastardo, 2012: 122
Oebalus magnus Thomas in Perez-Gelabert & Thomas, 2005: 340 *DR > TL: Pedernales Prov., Mirador del Hoyo de
Pelempito, 1,250 m
Oebalus ornatus (Sailer, 1944). Schmutterer, 1990: 56; Perez-Gelabert & Thomas, 2005: 341
Oebalus poecilus (Dallas, 1851). Perez-Gelabert & Thomas, 2005: 341
Oebalus pugnax (Fabricius, 1775). Schmutterer, 1990: 56; Perez-Gelabert & Thomas, 2005: 342
Oebalus ypsilongriseus (De Geer, 1773). Perez-Gelabert & Thomas, 2005: 342
Piezodorus Fieber, 1860
Piezodorus guildinii (Westwood, 1837). Perez-Gelabert & Thomas, 2005: 342
Proxys Spinola, 1837
Proxys punctulatus (Palisot de Beauvois, 1805). Perez-Gelabert & Thomas, 2005: 342
Proxys victor (Fabricius, 1775). Perez-Gelabert & Thomas, 2005: 342
Thyanta Stål, 1862
Thyanta (Argosoma) obsoleta (Dallas, 1851). Perez-Gelabert & Thomas, 2005: 342
Thyanta (Argosoma) testacea (Dallas, 1851). Perez-Gelabert & Thomas, 2005: 343
Thyanta (Thyanta) perditor (Fabricius, 1794). Perez-Gelabert & Thomas, 2005: 343
Tibraca Stål, 1860
Tibraca limbativentris Stål, 1860. Zuluaga & Mesa, 2000: 197; Perez-Gelabert & Thomas, 2005: 343 [Introduced]
Vulsirea Spinola, 1837
Vulsirea nigrorubra Spinola, 1837. Perez-Gelabert & Thomas, 2005: 344
Tribe MECIDIINI
Mecidea Dallas, 1851
Mecidea longula Stål, 1854. Perez-Gelabert & Thomas, 2005: 344
Tribe PROCLETINI
Brepholoxa Van Duzee, 1904
Brepholoxa heidemanni Van Duzee, 1904. Perez-Gelabert & Thomas, 2005: 344
Brepholoxa rotundifrons Barber, 1939. Perez-Gelabert & Thomas, 2005: 344
Tribe SCIOCORINI
Antillosciocoris Thomas, 2005
Antillosciocoris palisoti Thomas in Perez-Gelabert & Thomas, 2005: 344 *DR > TL: Galeón, Baní, Prov. Peravia
Subfamily PODOPINAE
Amaurochrous Stål, 1872
Amaurochrous dubius (Palisot de Beauvois, 1805). Perez-Gelabert & Thomas, 2005: 347

Family TESSARATOMIDAE
Piezosternum Amyot & Serville, 1843
Piezosternum subulatum (Thunberg, 1783). Perez-Gelabert & Thomas, 2005: 347; De los Santos & Bastardo, 2012: 121

Family PLATASPIDAE
Subfamily PLATASPINAE
Brachyplatys Boisduval, 1835
Brachyplatys subaeneus (Westwood, 1838). Perez-Gelabert et al., 2019: 135 [Introduced]

Family SCUTELLERIDAE
Subfamily PACHYCORINAE
Diolcus Mayr, 1864

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 257
Diolcus cassidoides (Walker, 1867: 48). Eger & Lattin, 1995: 417 *DR, *H > TL: Haiti
Diolcus irroratus (Fabricius, 1775). Barber, 1939: 280
Diolcus variegatus (Herrich-Schäffer, 1836). Barber, 1939: 279; Eger & Baranowski, 2002: 267
Nesogenes Horváth, 1921
Nesogenes boscii (Fabricius, 1798). Barber, 1939: 280
Pachycoris Burmeister, 1835
Pachycoris fabricii (Linnaeus, 1771). Barber, 1939: 282
Sphyrocoris Mayr, 1864
Sphyrocoris obliquus (Germar, 1839). Kirkaldy, 1909: 279; Eger, 2012: 247
Symphylus Dallas, 1851
Symphylus caribbeanus Kirkaldy, 1909. Barber, 1939: 278
Tetyra Fabricius, 1803
Tetyra antillarum Kirkaldy, 1909. Barber, 1939: 282
Subfamily SCUTELLERINAE
Augocoris Burmeister, 1835
Augocoris illustris (Fabricius, 1781). Abud, 1994: 9/94

Suborder AUCHENORRHYNCHA
Infraorder FULGOROMORPHA
Superfamily FULGOROIDEA
Family ACANALONIIDAE
Acanalonia Spinola, 1839
Acanalonia phorcys Fennah, 1971: 337 *H > TL: Port-au-Prince
Acanalonia servillei Spinola, 1839. Wolcott, 1927: 191; Dozier, 1931: 13; Ball, 1933: 149; Metcalf, 1954: 31
Acanalonia sublinea (Walker, 1858e: 114). Metcalf, 1954: 34
Acanalonia umbraculata (Fabricius, 1803). Metcalf, 1954: 35
Acanalonia viridis Melichar, 1901: 187. Metcalf, 1954: 38 *H > TL: Haïti

Family ACHILIDAE
Catonia Uhler, 1895
Catonia haitiensis Dozier, 1931: 15. Metcalf, 1948: 29
Catonia intricata Uhler, 1895. Wolcott, 1922: 33; Metcalf, 1948: 31
Haitiana Dozier, 1936
Haitiana nigrita Dozier, 1936: 2. Fennah, 1950: 119 *H > TL: Hinche

Family CALISCELIDAE
Quizqueiplana† Burgoin & Wang in Burgoin et al., 2016
Quizqueiplana alexbrowni† Burgoin & Gnezdilov in Burgoin et al., 2016: 212 [Fossil in Dominican amber]

Family CIXIIDAE
Bothriocera Burmeister, 1835
Bothriocera emelyanovi O’Brien, 2006: 312 *DR > TL: Prov. La Vega, 15 km E El Río
Bothriocera ferruginea Fennah, 1971: 310 *H > TL: Port-au-Prince
Bothriocera haitiana Fennah, 1971: 311 *H > TL: Kenscoff
Bothriocera hispaniolae Caldwell, 1950: 1 *DR > TL: Samaná
Bothriocera latens Fennah, 1971: 309 *H > TL: Barr. Peligre
Bothriocera lua Fennah, 1971: 304 *H > TL: Plaisance
Bothriocera venosa Fowler, 1904. Dozier, 1931: 14
Cubanella Fennah, 1948
Cubanella haitiensis Caldwell, 1950: 2 *H > TL: Port-au-Prince
Cyclopoliarus Fennah, 1945
Cyclopoliarus hispaniolae Fennah, 1971: 319 *H > TL: Kenscoff
Haplaxius Fowler, 1904
Haplaxius cabrerensis Ferreira, McKamey & Martínez, 2010: 66 *DR > TL: Prov. María Trinidad Sánchez, Cabrera, Payita,

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Baoba del Pinar
=Haplaxius sp. nr. crudus—Howard et al. (1981: 44) [As Myndus]
Haplaxius crudus (Van Duzee, 1907). Howard et al. (1981: 39)
Haplaxius jamaicae (Kramer, 1979). Ferreira, McKamey & Martínez, 2010: 67
Mnemosyne Stål, 1866
Mnemosyne consoleae van Stalle, 1987: 128 *DR > TL: San Juan, 28 km SE San Juan
Mnemosyne dominicensis van Stalle, 1987: 126 *DR > TL: La Vega, 15 km NE Jarabacoa
Mnemosyne granulata van Stalle, 1987: 129 *DR > TL: La Vega, 19 km SE Constanza
Mnemosyne oblongostriata van Stalle, 1987: 127 *DR > TL: La Estrelleta, 11 km E. Hondo Valle
Mnemosyne vegensis van Stalle, 1987: 129 *DR > TL: La Vega, 23 km SE Jarabacoa
Nymphocixia Van Duzee, 1923
Nymphocixia caribbea Fennah, 1971. Howard et al., 1981: 44
Oecleus Stål, 1862
Oecleus acutus Ball, 1902: 157. Kramer, 1977: 419 *H > TL: Port-au-Prince
Oecleus sp. nr. acutus—Howard et al. (1981: 44)
Oliarus Stål, 1862
Oliarus complectus Ball, 1902. Caldwell, 1950: 1; Howard et al., 1981: 44
=Oliarus franciscanus Stål, 1859. Metcalf, 1936: 61
Oliarus kulickae† Szwedo, 2000: 162 [Fossil in Dominican amber]
Oligocixia Gebicki & Wegierek, 1993
Oligocixia electrina† Gebicki & Wegierek, 1993: 122 [Fossil in Dominican amber]
Pintalia Stål, 1860
Pintalia procellata (Uhler, 1901: 511) *H > TL: Near Port-au-Prince

Family DELPHACIDAE
Subfamily ASIRACINAE
Tribe ASIRACINI
Copicerus Swartz, 1802
Copicerus irroratus irroratus Swartz, 1802. Metcalf, 1943: 31
Subfamily DELPHACINAE
Tribe DELPHACINI
Caenodelphax Fennah, 1965
Caenodelphax teapae (Fowler, 1905). Bartlett, 2011: no pagination.
Delphacodes Fieber, 1866
Delphacodes floridae puertoricensis Caldwell, 1951. Bartlett, 2011: no pagination.
Delphacodes humilis (Van Duzee, 1907). Caldwell, 1950: 1
Euides Fieber, 1866
Euides magnistylus (Crawford, 1914). Caldwell, 1950: 1
Megamelus Fieber, 1866
Megamelus electrae Muir, 1926. Bartlett, 2011: no pagination
Metadelphax Wagner, 1963
Metadelphax propinqua (Fieber, 1866). Caldwell, 1950: 1; Gonzon & Bartlett, 2007: 242
Nilaparvata Distant, 1906
Nilaparvata caldwelli Metcalf, 1955. Bartlett, 2007: 54
Nothodelphax Fennah, 1963
Nothodelphax slossonae (Ball, 1903). Caldwell, 1950: 1
Peregrinus Kirkaldy, 1904
Peregrinus maidis (Ashmead, 1890). Wolcott, 1927: 394; Schmutterer, 1990: 15
Pissonotus Van Duzee, 1894
Pissonotus abdominalis (Crawford, 1914). Bartlett & Dietz, 2000: 102
Pissonotus albovenosus (Osborn, 1935). Bartlett & Dietz, 2000: 38
Sogatella Fennah, 1956
Sogatella kolophon (Kirkaldy, 1907). Bartlett, 2011: no pagination
Sogatella molina (Fennah, 1963). Bartlett, 2011: no pagination

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 259
Stobaera Stål, 1859
Stobaera concinna (Stål, 1859). Wilson & Hilburn, 1991: 417
Tagosodes Asche & Wilson, 1990
Tagosodes orizicola (Muir, 1926). Schmutterer, 1990: 43; Medina, 2002a: 61
Tribe SACCHAROSYDNINI
Saccharosydne Kirkaldy, 1907
Saccharosydne saccharivora (Westwood, 1833). Wolcott, 1922: 33; Wolcott, 1927: 191; Metcalfe, 1971: 565; Schmutterer,
1990: 77

Family DERBIDAE
Subfamily DERBINAE
Anticedusa Emeljanov, 2008
Anticedusa dominicana† Emeljanov, 2008: 793 [Fossil in Dominican amber]
Cedusa Fowler, 1904
Cedusa colona Caldwell in Caldwell & Martorell, 1950. Flynn & Kramer, 1983: 244
Cedusa credula† Emeljanov & Shcherbakov, 2000: 445 [Fossil in Dominican amber]
Cedusa inflata (Ball, 1902: 262). Howard et al., 1981: 44; Flynn & Kramer, 1983: 241; Howard, 2001: 119
Cedusa quinteca Kramer, 1986: 278 *DR > TL: Nisibón, El Cedro
Cedusa santaclara Myers, 1928. Metcalf, 1945: 130
Dawnarioides Dozier, 1929
Dawnarioides hispaniolus O’Brien, 1982: 313 *DR > TL: La Vega, 23 km SE Jarabacoa
Dysimia Muir, 1924
Dysimia imprudens† Emeljanov & Shcherbakov, 2000: 447 [Fossil in Dominican amber]
Dysimia maculata Muir, 1924. Dozier, 1931: 13; Broomfield, 1985: 89
Neocenchrea Metcalf, 1923
Neocenchrea sp.—Howard et al. (1981: 44)
Omolicna Fennah, 1945
Omolicna sp.—Howard et al. (1981: 44)

Family FLATIDAE
Subfamily FLATINAE
Cryptoflata Melichar, 1901
Cryptoflata dominicensis (Spinola, 1839: 439). Metcalf, 1957: 270 *DR, *H > TL: Saint-Domingue
Cyarda Stål, 1866
Cyarda acuminipennis acuminipennis (Spinola, 1839: 445). Melichar, 1902: 134; Fennah, 1965: 112 *DR, *H > TL: Saint-
Domingue
Cyarda acuminipennis haitensis Metcalf & Bruner, 1948. Wolcott, 1927: 191; Fennah, 1965: 113 *H > TL: Port-au-Prince
=Cyarda haitensis Metcalf & Bruner, 1948: 103
Cyarda difformis Walker, 1858e: 121. Melichar, 1902: 134; Dozier, 1931: 14; Fennah, 1965: 114
Cyarda regina Fennah, 1965: 117 *H > TL: Haiti
Cyclopterum Gnezdilov & O’Brien, 2014
Cyclopterum hispaniolum Gnezdilov & O’Brien, 2014: 146 *DR > TL: San Juan, 28 km SE San Juan
Eurocalia Van Duzee, 1907
Eurocalia punctifera (Walker, 1858e: 71). Metcalf, 1957: 431
Euhyloptera Fennah, 1945
Euhyloptera haitiana Fennah, 1965: 110 *H > TL: Haiti
Euhyloptera maculipennis Fennah, 1965: 108 *H > TL: Limbé
Euhyloptera priapus Fennah, 1965: 109 *H > TL: Haiti
Euhyloptera simillima Fennah, 1965: 111 *H > TL: Haiti
Exoma Melichar, 1902
Exoma robusta (Uhler, 1901: 515). Fennah, 1965: 108 *H > TL: Port-au-Prince
=Exoma viduata Melichar, 1902: 158. Medler, 2003: 355
Flatormenis Melichar, 1923
Flatormenis glaucescens (Walker, 1858e: 118). Melichar, 1902: 97; Metcalf, 1957: 317 *DR > TL: St. Domingo

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Melormenis Metcalf, 1938
Melormenis infuscata domingensis (Russo, 1930: 509). Metcalf, 1957: 333
Ormenana Metcalf & Bruner, 1948
Ormenana sp.—Schmutterer (1990: 157)
Ormenaria Metcalf & Bruner, 1948
Ormenaria sp.—Abud (1982: 2/82)
Ormenina Fennah, 1947
Ormenina coffeacola (Dozier, 1931: 17). Fennah, 1947a: 114 *H > TL: Fond-des-Nègres
Ormenina paupera (Walker, 1858e: 119). Melichar, 1902: 99; Fennah, 1947a: 114 *DR > TL: St. Domingo
Petrusa Stål, 1860
Petrusa epilepsis (Kirkaldy, 1906) [New name for Petrusa marginata Linnaeus, 1767]. Caldwell, 1950: 1; Metcalf, 1957: 337
Subfamily FLATOIDINAE
Flatoides Guérin-Méneville, 1844
Flatoides brunnea Muir, 1924: 472. Metcalf, 1957: 502
Flatoides cervina Walker, 1851. Metcalf, 1957: 502
Flatoides guttata (Uhler, 1901: 513)
Flatoides lichenosus Melichar, 1902: 222. Medler, 2003: 350 *H > TL: Port-au-Prince
Flatoidinus Melichar, 1923
Flatoidinus acutus (Uhler, 1901: 514). Metcalf, 1954: 14; Metcalf, 1957: 528
Flatoidinus punctatus (Walker, 1851). Metcalf, 1957: 533
Phalaenomorpha Amyot & Serville, 1843
Phalaenomorpha corticina (Burmeister, 1835). Metcalf, 1957: 514
Planodascalia Metcalf & Bruner, 1948
Planodascalia sp.—Howard et al. (1981: 45)
Tetraceratium Muir, 1924
Tetraceratium wolcotti Muir, 1924: 468 *DR > TL: Haina, Santo Domingo

Family ISSIDAE
Colpoptera Burmeister, 1835
Colpoptera clerodendri Dozier, 1931: 19 *H > TL: Fond-des-Nègres
Colpoptera elevans (Walker, 1858e). Metcalf, 1958: 205
Colpoptera insularis Dozier, 1931: 20 *H > TL: Fond-des-Nègres
Colpoptera nana Dozier, 1931: 20 *DR, *H > TL: Kenscoff
Thionia Stål, 1859
Thionia gibbicollis Dozier, 1931: 17 *H > TL: Hinche
Thionia douglundbergi† Stroinski & Szwedo, 2008: 531 [Fossil in Dominican amber]

Family KINNARIDAE
Atopocixius Muir, 1926
Atopocixius collaris Fennah, 1945: 129 *DR > TL: Constanza, 3,000–4,000 ft.
Atopocixius major Fennah, 1945: 131 *DR > TL: 7,000 ft., Valle Nuevo, S. E. Constanza
Atopocixius ornatus Muir, 1926: 336 *H > TL: Carrefour, near Port-au-Prince
Lomagenes Fennah, 1945
Lomagenes delphacoides Fennah, 1945: 133 *DR > TL: 6,000 ft., at Loma Vieja, S. Constanza
Microissus Fennah, 1947
Microissus psychoda Fennah, 1947b: 94 *H > TL: La Gonave Island
Oeclidius Van Duzee, 1914
Oeclidius browni† Bourgoin & Lefebvre, 2002: 583 [Fossil in Dominican amber]
Oeclidius conopa Fennah, 1980: 693 *H > TL: Port-au-Prince
Oeclidius princeps Fennah, 1980: 691 *H > TL: Port-au-Prince
Oeclidius salaco† Emeljanov & Shcherbakov, 2000: 439 [Fossil in Dominican amber]
Quilessa Fennah, 1942
Quilessa stolida† Emeljanov & Shcherbakov, 2000: 442 [Fossil in Dominican amber]

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Family NOGODINIDAE
Tainosia† Szwedo & Stroinski, 2001
Tainosia quisqueyae† Szwedo & Stroinski, 2001: 29 [Fossil in Dominican amber]

Family TROPIDUCHIDAE
Subfamily TAMBINIINAE
Cyphoceratops Uhler, 1901
Cyphoceratops furcatus Uhler, 1901: 511. Dozier, 1931: 15
Parahydriena Muir, 1923
Parahydriena hyalina acuta Caldwell & Martorell, 1951. Fennah, 1945b: 173
Pelitropis Van Duzee, 1908
Pelitropis haitiana Fennah, 1965: 97 *H > TL: Morne à Cabrits
Remosa Distant, 1906
Remosa cultellator (Walker, 1858e: 62). Fennah, 1945b: 145
Tangiopsis Uhler, 1901
Tangiopsis tetrastichus Uhler, 1901: 513. Dozier, 1931: 15 *H, *DR > TL: Port-au-Prince
Subfamily TROPIDUCHINAE
Tangella Metcalf & Bruner, 1930
Tangella dictys Fennah, 1971: 330 *H > TL: Grand Goave
Subfamily INCERTAE SEDIS
Tangyria Uhler, 1901
Tangyria frontalis Uhler, 1901: 512. Fennah, 1971: 331 *H > TL: Port-au-Prince

Infraorder CICADOMORPHA
Superfamily CERCOPOIDEA
Family CERCOPIDAE
Subfamily APHROPHORINAE
Tribe LEOCOMIINI
Enocomia Ball, 1919
Enocomia (Enocomia) ampliata Ball, 1919: 147 *H > TL: Port-au-Prince
Leocomia Ball, 1919
Leocomia (Leocomia) parabolocrata Ball, 1919: 149 *H > TL: Port-au-Prince
Lepyronura Hamilton, 2014
Lepyronura furcata Hamilton, 2014: 38 *DR > TL: Blanton Mine, N of San Cristóbal
Subfamily CERCOPINAE
Prosapia Fennah, 1949
Prosapia bicincta (Say, 1830). Ramos, 1988: 65; Rowland, 2001: 77
Subfamily TOMASPIDINAE
Tribe TOMASPIDINI
Aeneolamia Fennah, 1949
Aeneolamia varia (Fabricius, 1787). Marcano & Abud, 1980: 10/80

Family CLASTOPTERIDAE
Clastoptera Germar, 1839
Clastoptera proteus Fitch, 1851. Metcalf, 1962: 34
Prisciba† Poinar & Brown in Poinar et al., 2014
Prisciba dominicana† Poinar & Brown in Poinar et al., 2014: 3 [Fossil in Dominican amber]
Prisciba serrata† Poinar & Brown in Poinar et al., 2014: 3 [Fossil in Dominican amber]

Superfamily CICADOIDEA
Family CICADIDAE
Subfamily CICADETTINAE
Tribe CICADETTINI
Cicadettana Marshall & Hill, 2017

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Cicadettana ramosi (Sanborn, 2009: 1). Marshall & Hill, 2017: 543 *DR, *H > TL: Peravia Prov., 15 km E. San José de Ocoa
=Cicadetta calliope (Walker, 1850). Ramos, 1983: 66 [Misidentification]
Tribe TAPHURINI
Minyscapheus† Poinar & Kritsky, 2012
Minyscapheus dominicanus† Poinar & Kritsky, 2012: 330. Moulds, 2018: 457 [Fossil in Dominican amber]
Psallodia Uhler, 1903
Psallodia espinii Uhler, 1903: 66. Ramos, 1983: 66 *H, *DR > TL: Vicinity of Port de Paix
Subfamily CICADINAE
Tribe FIDICININI
Ollanta Distant, 1905
Ollanta melvini Ramos, 1983: 65 *DR > TL: Baní
Proarna Stål, 1864
Proarna hilaris (Germar, 1843). Ramos, 1983: 64
Proarna palisoti (Metcalf, 1963). Ramos, 1983: 62 *DR > TL: Saint Domingue
Tribe ZAMMARINI
Chinaria Davis, 1934
Chinaria vivianae Ramos, 1983: 63 *DR > TL: La Estrelleta Prov., 4 km SE Río Limpio, ca. 760 m
Odopoea Stål, 1862
Odopoea cariboea Uhler, 1892: 169. Ramos, 1983: 62 *DR > TL: Near the city of San Domingo
Odopoea dilatata (Fabricius, 1775). Goemans, 2010: 5
=Odopoea domingensis Uhler, 1892: 172
=Odopoea degiacomii Distant, 112. Ramos, 1983: 62 [Misidentification]
Odopoea strigipennis (Walker, 1858d: 3). Ramos, 1983: 62 *H, *DR > TL: Haiti
Odopoea suffusa (Walker, 1850: 37). Ramos, 1983: 62 *DR > TL: St. Domingo
Odopoea vacillans (Walker, 1858e: 3). Ramos, 1983: 63 *DR > TL: St. Domingo
Uhleroides Distant, 1912
Uhleroides hispaniolae Davis, 1939: 292 *DR > TL: San Francisco Mountains, St. Domingo
Uhleroides samanae Davis, 1939: 294 *DR > TL: San Francisco Mountains, St. Domingo
Tribe INCERTAE SEDIS
Dominicicada† Poinar, Kritsky & Brown, 2011
Dominicicada youngi† Poinar & Kritsky, 2011: 463. Moulds, 2018: 453 [Fossil in Dominican amber]

Superfamily CICADELLOIDEA
Family CICADELLIDAE
Subfamily AGALLIINAE
Aceratagallia Kirkaldy, 1907
Aceratagallia grandis (Nielson & Maes, 1993). Perez-Gelabert, 2008: 204
Agallia Curtis, 1833
Agallia albidula Uhler, 1895. Oman, 1933: 35
Agallia configurata Oman, 1933: 36
Agallia reticulata Ball, 1900: 127
Agallia (?) sp.†—Dietrich & Vega (1995: 268) [Fossil in Dominican amber]
Agalliana Oman, 1933
Agalliana sticticollis (Stål, 1859). Oman, 1933: 70
Agalliopsis Kirkaldy, 1907
Agalliopsis acrita Freytag, 2009: 272 *DR > TL: San Juan, Sierra de Neiba, Sabana del Silencio, 10.1 km SSW El Cercado,
2017 m
Agalliopsis acuticauta (Oman, 1933: 28). Freytag, 2009: 261 *H > TL: Port-au-Prince
Agalliopsis ancyra Freytag, 2009: 263 *DR > TL: Sierra de Neiba, south slope near summit, 4 km N Angel Feliz, 1825 m
Agalliopsis balli (Baker, 1901). Oman, 1933: 10. Freytag, 2009: 260 *H, *DR > TL: Haiti
Agalliopsis bidigita Freytag, 2009: 264 *DR > TL: San Juan, 8 km NE Vallejuelo, 690 m
Agalliopsis clitellaria (Ball, 1900: 127). Oman, 1933: 13. Freytag, 2009: 260 *H > TL: Port-au-Prince
Agalliopsis conformis Oman, 1933: 11. Freytag, 2009: 260 *DR > TL: Blanton mine, north of San Cristóbal
Agalliopsis davidsoni Freytag, 2009: 274 *DR > TL: Barahona, Eastern Sierra de Bahoruco, Reserva Cachote, 12.8 km NE

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Paraiso, 1219 m
Agalliopsis dichroma Freytag, 2009: 270 *DR > TL: San Juan, Sierra de Neiba, Sabana del Silencio, 10 km SSW El Cercado,
2009 m
Agalliopsis extenda Freytag, 2009: 274 *DR > TL: Duarte, Reserva Loma Quita Espuela, El Cadillar, 6.7 km NE San Francisco
de Macoris, 280 m
Agalliopsis extrema Freytag, 2009: 268 *DR Independencia, Sierra de Bahoruco, north slope, 13.5 km SE Puerto Escondido,
1789 m
Agalliopsis intera Freytag, 2009: 268 *DR > TL: Sierra de Neiba, south slope near summit, 4 km N Angel Feliz, 1825 m
Agalliopsis lathripa Freytag, 2009: 262 *DR > TL: Pedernales, 26 km N Cabo Rojo, 730 m
Agalliopsis longistyla Freytag, 2009: 271 *DR > TL: La Vega, Cordillera Central, Loma Casabito, 16 km NW Bonao, 1487 m
Agalliopsis meiosa Freytag, 2009: 273 *DR > TL: La Vega, Cordillera Central, Reserva Valle Nuevo, La Nevera, 15.1 km SE
Valle Nuevo, 2252 m
Agalliopsis pepino (DeLong & Wolcott, 1923). Oman, 1933: 12. Freytag, 2009: 260
Agalliopsis treptona Freytag, 2009: 269 *DR > TL: Pedernales, 9.7 km NE Los Arroyos, 2070 m
Agalliopsis uncula Freytag, 2009: 275 *DR > TL: Santiago Rodriguez, Gurabo, Sabaneta
Brasa Oman, 1937
Brasa dozieri Nielson & Freytag, 1998: 132 *H > TL: Fond des Negres
Brasa media Nielson & Freytag, 1998: 133 *DR > TL: Pedernales Prov., ca. 35 km N. Cabo Rojo, 1250 m, Las Abejas
Brasa rugicollis (Dozier, 1927: 264). Kramer, 1964: 153; Nielson & Freytag, 1998: 132 *DR > TL: Cerro del Cacique, Santo
Domingo
Cubrasa Young, 1977
Cubrasa brevida Freytag, 2008a: 34 *DR > TL: 3.3 km NE Los Arroyos, Pedernales Prov.
Cubrasa longula Freytag, 2008a: 34 *DR > TL: 3.3 km NE Los Arroyos, Pedernales Prov.
Subfamily CICADELLINAE
Apogonalia Evans, 1947
Apogonalia angusta Freytag, 2004: 67. McKamey, 2007: 31 *DR > TL: Azua, East side of crest, Sierra Martin Garcia, 7 km
WNW Barrero, 860 m
Apogonalia histrio (Fabricius, 1794). Young, 1977: 919; McKamey, 2007: 32
Apogonalia interrupta (Signoret, 1855). Dozier, 1931: 5; Young, 1977: 919; McKamey, 2007: 32 *H, *DR > TL: Port-au-
Prince
Apogonalia loxa Freytag, 2004: 67. McKamey, 2007: 32 *DR > TL: Pedernales, 20 km N Cabo Rojo, 1070 m
Apogonalia noda Freytag, 2004: 69. McKamey, 2007: 32 *DR > TL: Hato Mayor, Parque Los Haitises, near Cueva de Arena,
10 m
Apogonalia pinguis Young, 1977: 925. McKamey, 2007: 33 *DR, *H > TL: Paso de la Vaca, Mons. Nouel—Constanza Rd.
Apogonalia robusta (Walker, 1851). Young, 1977: 920; McKamey, 2007: 33
=Tettigella fausta (Walker, 1858e: 198). McKamey, 2007: 33
Caribovia Young, 1977
Caribovia aperta Freytag, 2005b: 350. McKamey, 2007: 66 *DR > TL: La Vega, 57 km SE Constanza
Caribovia bifasciata Freytag, 2005b: 359. McKamey, 2007: 66 *DR > TL: Sierra de Neiba, just south of crest, 5 km WNW
Angel Feliz, 1780 m
Caribovia constans (Walker, 1858e: 198). Dozier, 1931: 6; Young, 1977: 913; Freytag, 2005: 345; McKamey, 2007: 67 *DR >
TL: St. Domingo
Caribovia divisa Freytag, 2005b: 353. McKamey, 2007: 67 *DR > TL: La Vega, Reserva Cientifica Valle Nuevo, Sector La
Nevera, 3 km WNW La Nuez, 2200 m
Caribovia doloma Freytag, 2005b: 349. McKamey, 2007: 67 *DR > TL: 14 km SE Constanza
Caribovia dorsolinea Freytag, 2005b: 355. McKamey, 2007: 67 *DR > TL: Barahona, 4 km NE Polo, 1260 m
Caribovia furcata Freytag, 2005b: 350. McKamey, 2007: 67 *DR > TL: La Vega, 18 kkm SE Constanza
Caribovia glabra Young, 1977: 913. Freytag, 2005: 361; McKamey, 2007: 67 *DR > TL: Colonia Ramfis, Prov. Trujillo
Caribovia hamata Freytag, 2005b: 355. McKamey, 2007: 67 *DR > TL: Independencia, Sierra de Neiba, just south of crest, 5
km NNW Angel Feliz, 1780 m
Caribovia nesiotica Young, 1977: 914. Freytag, 2005: 358 *DR > TL: Paso de la Vaca, Mons. Nouel—Constanza Rd., Prov. La
Vega, alt. 1500 m
Caribovia obrienae Freytag, 2005b: 359. McKamey, 2007: 68 *DR > TL: La Vega, 24 km E El Río
Caribovia ovata Freytag, 2005b: 353. McKamey, 2007: 68 *DR > TL: Pedernales, 5 km NE Los Arroyos, 1680 m

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Caribovia plagata Freytag, 2005b: 352. McKamey, 2007: 68 *DR > TL: La Vega, 14 km SE Constanza
Caribovia rubida Freytag, 2005b: 361. McKamey, 2007: 68 *DR > TL: La Vega, 18 km E El Río
Caribovia trimaculata Freytag, 2005b: 359. McKamey, 2007: 68 *DR > TL: La Vega, 18 km SE Constanza
Caribovia vittata Freytag, 2005b: 349. McKamey, 2007: 68 *DR, *H > TL: La Vega, 6 km SE Constanza
Chlorogonalia Young, 1977
Chlorogonalia coeruleovittata (Signoret, 1855). Perez-Gelabert, 2008: 203
Ehagua Melichar, 1926
Ehagua dubiosa (Dozier, 1931: 5). Young, 1977: 1084; McKamey, 2007: 109 *DR, *H > TL: Port-Margot
Hadria Metcalf & Bruner, 1937
Hadria dominicana (Dozier, 1931: 10). Young, 1977: 970; McKamey, 2007: 137
Hadria haitiensis (Dozier, 1931: 9). Young, 1977: 970; McKamey, 2007: 137 *DR, *H > TL: Mt. Pilboreau
Hadria hispaniola (Dozier, 1931: 8). Young, 1977: 970; McKamey, 2007: 137 *DR, *H > TL: Mt. Pilboreau
Hadria maldonadoi Young, 1977: 974. McKamey, 2007: 138 *DR > TL: Blanton Mine, north of San Cristóbal
Hortensia Metcalf & Bruner, 1937
Hortensia similis (Walker, 1851). Howard et al., 1981: 43; Medina, 2002: 63; McKamey, 2007: 141; Lozada, 2010: 105
Neohadria Freytag, 2007
Neohadria amaurota Freytag, 2007: 394 *DR > TL: Independencia, Sierra de Bahoruco, Loma del Toro, 5.3 km SW El
Aguacate, 2316 m
Neohadria delecta Freytag, 2007: 395 *DR > TL: Barahona, Filipinas, Larimar Mine, 1006 m
Neohadria diversa Freytag, 2007: 393 *DR > TL: Sierra de Neiba, near crest, 5.5 km NNW Angel Feliz, 1750 m
Neohadria grisea Freytag, 2007: 395 *DR > TL: Barahona, Eastern Sierra de Bahoruco, Reserva Cachote, 12.8 km NE Paraiso,
1230 m
Neohadria maculata Freytag, 2007: 393 *DR > TL: Barahona, Eastern Sierra de Bahoruco, Reserva Cachote, 12.8 km NE
Paraiso, 1198 m
Parahadria Freytag, 2007
Parahadria octolineata Freytag, 2007: 386 *DR > TL: Duarte, Reserva Loma Quita Espuela, La Canela, 13.1 km NNE San
Francisco de Macoris, 512 m
Tylozygus Fieber, 1866
Tylozygus fasciatus (Walker, 1851). Young, 1977: 589; McKamey, 2007: 256
Xyphon Hamilton, 1985
Xyphon reticulata (Signoret, 1854). Perez-Gelabert, 2008: 203
Xyphon sagittifera (Uhler, 1895). Nottingham, 1932: 111
Uncertain species
Correct genus unknown
“Tettigonia” marginata Palisot de Beauvois, 1820: 170. McKamey, 2007: 95 *DR, *H > TL: Saint-Domingue
“Tettigonia” tripartita Walker, 1858d: 96. Metcalf, 1965: 147; McKamey, 2007: 79 *DR > TL: St. Domingo
Subfamily COELIDIINAE
Tribe COELIDIINI
Kramerolidia Nielson, 1982
Kramerolidia mera (Van Duzee, 1907). Nielson, 1982: 241
=Jassus wolcotti (Dozier, 1931: 12). Nielson, 1982: 241
Kramerolidia serrata Nielson, 1982: 238
Subfamily DELTOCEPHALINAE
Tribe ACINOPTERINI
Acinopterus Van Duzee, 1892
Acinopterus angulatus Lawson, 1922. Dozier, 1931: 4; Zanol, 2006: 89
Tribe ATHYSANINI
Bandara Ball, 1931
Bandara hyalina (Osborn, 1923). Metcalf, 1967: 496
Chlorotettix Van Duzee, 1892
Chlorotettix minimus Baker, 1898. Howard et al., 1981: 43
Chlorotettix viridius Van Duzee, 1892. Perez-Gelabert, 2008: 204
Cicadulina China, 1926
Cicadulina tortilla Caldwell, 1952. Perez-Gelabert, 2008: 204

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Exitianus Ball, 1929
Exitianus atratus Linnavuori, 1959. Perez-Gelabert, 2008: 204
Exitianus exitiosus (Uhler, 1880). Perez-Gelabert, 2008: 204
Ileopeltus Cwikla, 1988
Ileopeltus tethys (Van Duzee, 1907). Perez-Gelabert, 2008: 204
Ollarianus Ball, 1936
Ollarianus sp.—Perez-Gelabert (2008: 204)
Stirellus Osborn & Ball, 1902
Stirellus bicolor (Van Duzee, 1892). Perez-Gelabert, 2008: 204
Tribe BALCLUTHINI
Balclutha Kirkaldy, 1900
Balclutha abdominalis abdominalis (Van Duzee, 1892). Zanol, 2006: 102
Balclutha aridula Linnavuori, 1959. Perez-Gelabert, 2008: 204
Balclutha caldwelli Blocker, 1967: 36. Zanol, 2006: 102
Balclutha flavescens (Baker, 1903). Perez-Gelabert, 2008: 204
Balclutha incisa (Matsumura, 1902). Zanol, 2006: 104
Balclutha lucida (Butler, 1877) [By synonymy of species below]
=Balclutha floridana (DeLong & Davidson, 1933). Zanol, 2006: 103
Balclutha neglecta (DeLong & Davidson, 1933). Perez-Gelabert, 2008: 204
Balclutha rosea (Scott, 1876) [By synonymy of species below]
=Balclutha guajanae (DeLong, 1923). Zanol, 2006: 104
=Balclutha hebe (Kirkaldy, 1906). Blocker, 1967: 34; Zanol, 2006: 104
Balclutha saltuella (Kirschbaum, 1868). Perez-Gelabert, 2008: 204
Balclutha youngi Blocker, 1967. Perez-Gelabert, 2008: 204
Tribe DELTOCEPHALINI
Amplicephalus DeLong, 1926
Amplicephalus maculellus (Osborn, 1926). Perez-Gelabert, 2008: 204
Graminella DeLong, 1950
Graminella bipunctella Linnavuori, 1959. Perez-Gelabert, 2008: 204
Graminella cognita Caldwell, 1952. Perez-Gelabert, 2008: 204
Graminella lambda Kramer, 1965: 23
Graminella nigrifrons (Forbes, 1885). Kramer, 1967: 613
Graminella punctata Caldwell, 1952. Perez-Gelabert, 2008: 204
Graminella sonora (Ball, 1900). Perez-Gelabert, 2008: 204
Haldorus Oman, 1938
Haldorus australis (DeLong, 1926). Perez-Gelabert, 2008: 204
Laevicephalus DeLong, 1926
Laevicephalus nigripennis (DeLong, 1923). Dozier, 1931: 4
Planicephalus Linnavuori, 1954
Planicephalus flavicosta (Stål, 1860). Perez-Gelabert, 2008: 205
Sanctanus Ball, 1932
Sanctanus fasciatus (Osborn, 1900: 190). Osborn, 1935: 147; Zanol, 2007: 9
Unerus DeLong, 1936
Unerus colonus (Uhler, 1895). Metcalf, 1967: 859; Freytag, 1986: 81
Tribe HECALINI
Spangbergiella Signoret, 1879
Spangbergiella vulnerata (Uhler, 1877). Perez-Gelabert, 2008: 205
Tribe KRISNINI
Archiokrisna Freytag, 2012
Archiokrisna garciamarquezi† (Dietrich & Vega, 1995: 263). Freytag, 2012: 2 [Fossil in Dominican amber]
Neokrisna Freytag, 2012
Neokrisna breva Freytag, 2012: 5 *DR > TL: Sierra de Neiba, just South of crest
Neokrisna decliva Freytag, 2012: 4 *DR > TL: Loma El Casabito
Neokrisna libera Freytag, 2012: 7 *DR > TL: 15 km N Jarabacoa

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Neokrisna longula Freytag, 2012: 5 *DR > TL: Loma El Casabito
Neokrisna oncora Freytag, 2012: 4 *DR > TL: 26 km N. Cabo Rojo
Neokrisna stena Freytag, 2012: 7 *DR > TL: La Cienaga de Manabao
Tribe MACROSTELINI
Circulifer Zachvatkin, 1935
Circulifer tenellus (Baker, 1896). Perez-Gelabert, 2008: 205
Dalbulus DeLong, 1950
Dalbulus maidis (DeLong & Wolcott, 1923). Osborn, 1935: 169; Oman, 1948: 34; Howard et al., 1981: 43; Zanol, 2006: 108
Macrosteles Fieber, 1866
Macrosteles divisus (Uhler, 1877). Perez-Gelabert, 2008: 205
=Macrosteles fascifrons (Stål, 1858). Zanol, 2006: 109 [Misidentification]
Nephotettix Matsumura, 1902
Nephotettix bipunctatus (Fabricius, 1803). SEA, 1999: 90 [Introduced]
Tribe SCAPHOIDEINI
Osbornellus Ball, 1932
Osbornellus (Nereius) arcus Freytag, 2008b: 139 *DR > TL: 18 km E El Río, La Vega Prov.
Osbornellus (Nereius) bellus Freytag, 2008b: 133 *DR > TL: Reserva Cachote, 12.8 km NE Paraiso, Eastern Sierra de
Bahoruco, Barahona Prov.
Osbornellus (Nereius) bimarginatus (DeLong, 1932). Perez-Gelabert, 2008: 205
Osbornellus (Nereius) brevitubus Freytag, 2008b: 134 *DR > TL: Larimar Mine, Filipinas, Barahona Prov.
Osbornellus (Nereius) coloritubus Freytag, 2008b: 137 *DR > TL: El Cadillar, Reserva Loma Quita Espuela, 6.7 km NE San
Francisco de Macoris, Duarte Prov.
Osbornellus (Nereius) dicellus Freytag, 2008b: 138 *DR > TL: Nisibón, La Altagracia Prov.
Osbornellus (Nereius) dipilus Freytag, 2008b: 133 *DR > TL: Alcoa Headquarters, Cabo Rojo, Pedernales Prov.
Osbornellus (Nereius) eccritus Freytag, 2008b: 137 *DR > TL: Loma Casabito, La Vega Prov.
Osbornellus (Nereius) eratus Freytag, 2008b: 143 *DR > TL: 24 km N Cabo Rojo, Pedernales Prov.
Osbornellus (Nereius) fasciatus (DeLong, 1932). Perez-Gelabert, 2008: 205
Osbornellus (Nereius) hispanus Freytag, 2008b: 141 *DR > TL: 2.9 km SW Boca de Yuma, Parque del Este, La Altagracia
Prov.
Osbornellus (Nereius) parvus Freytag, 2008b: 142 *DR > TL: 2 km N La Romana, La Romana Prov.
Osbornellus (Nereius) proximus Freytag, 2008b: 138 *DR > TL: 40 km N Angel Felix, Sierra de Neiba, south slope near
summit, Independencia Prov.
Osbornellus (Nereius) sagarus Freytag, 2008b: 142 *DR > TL: 13 km NW Baní, Peravia Prov.
Osbornellus (Nereius) tetrus Freytag, 2008b: 143 *DR > TL: 15 km N Cabo Rojo, Pedernales Prov.
Osbornellus (Nereius) tubus Freytag, 2008b: 136 *DR > TL: Reserva Cachote, 12.8 km NE Paraiso, Eastern Sierra de
Bahoruco, Barahona Prov.
Osbornellus (Nereius) venustus Freytag, 2008b: 141 *DR > TL: 24 km N Cabo Rojo, Pedernales Prov.
Subfamily EVACANTHINAE
Tribe NIRVANINI
Antillonirvana Dietrich, 2004
Antillonirvana freytagi Dietrich, 2004: 465 *DR > TL: Pedernales, Las Abejas, 38 km NNW Cabo Rojo, 1250 m
Antillonirvana turnbowi Dietrich, 2004: 465 *DR > TL: Payaso, Barahona
Jassosqualus Kramer, 1964
Jassosqualus hispaniolensis† Dietrich & Vega, 1995: 267 [Fossil in Dominican amber]
Subfamily GYPONINAE
Curtara DeLong & Freytag, 1972
Curtara (Curtara) acroschismata Freytag & Carlo, 1991: 11. Freytag, 2005a: 324 *DR > TL: Peravia, 13 km NW Baní
Curtara (Curtara) albimaculata (Osborn, 1926). DeLong & Freytag, 1976: 40; Freytag, 2005a: 330
=Phlepsius smithi Dozier, 1931: 11
Curtara (Curtara) barrera Freytag, 2005a: 331 *DR > TL: Azua, East side of crest, Sierra Martin Garcia, 7 km WNW Barrero,
860 m
Curtara (Curtara) blancoi Freytag & Carlo, 1991: 12. Freytag, 2005a: 324 *DR > TL: Dajabon Prov., 13 km S Loma de
Cabrera, ca. 400 m
Curtara (Curtara) carloorum Freytag & Carlo, 1991: 11. Freytag, 2005a: 324 *DR > TL: Peravia, 13 km NW Baní

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Curtara (Curtara) cavera DeLong & Freytag, 1976. Freytag, 2005a: 324
Curtara (Curtara) cornuta Freytag, 2005a: 327 *DR > TL: Pedernales, 5 km NE Los Arroyos, 1680 m
Curtara (Curtara) minima Freytag, 2005a: 331 *DR > TL: Hato Mayor, Parque Los Haitises, near Cueva de Arena, 10 m
Curtara (Curtara) nunezi Freytag, 2005: 330 *DR > TL: Elias Piña, Sierra de Neiba, 9.3 km WSW Hondo Valle, 1901 m
Curtara (Curtara) sata DeLong & Freytag, 1976: 9. Freytag & Carlo, 1991: 10; Freytag, 2005a: 324 *H, *DR > TL: Port-au-
Prince
Curtara (Curtara) youngi Freytag, 2005a: 327 *DR > TL: Pedernales, along Río Mulito, 13 km N Pedernales, 230 m
Gypona Germar, 1821
Gypona (Marganalana) rawlinsi Freytag, 2005a: 338 *DR > TL: Pedernales, 9.7 km NE Los Arroyos, 2070 m
Gypona (Marganalana) spina Freytag, 2005a: 340 *DR > TL: Pedernales, 3.3 km NE Los Arroyos, 1450 m
Gypona (Marganalana) woodruffi Freytag, 2005a: 338 *DR > TL: Barahona, Filipinas, Larimar Mine, 3300 ft.
Gypona (Obtusana) curta Freytag, 2005a: 333 *DR > TL: Pedernales, Cabo Rojo
Gypona (Obtusana) ileota Freytag, 2005a: 334 *DR > TL: Peravia, 15 km E San José de Ocoa
Gypona (Obtusana) imita Freytag, 2005a: 336 *DR > TL: La Vega, 23 km SE Constanza, 2225 m
Gypona (Obtusana) toxum Freytag, 2005a: 336 *DR > TL: Pedernales, Cabo Rojo, Alcoa Headquarters
Prairiana Ball, 1920
Prairiana dualis DeLong, 1942. Freytag, 2005a: 323
Rugosana DeLong, 1936
Rugosana carpa Freytag, 2005a: 332 *DR > TL: La Vega, 46 km SE Constanza
Rugosana reta Freytag, 2005a: 332 *DR > TL: La Vega, 24 km SE Constanza
Subfamily IASSINAE
Tribe IASSINI
Comanopa Blocker, 1979
Comanopa puertoricensis (Caldwell, 1952). Perez-Gelabert, 2008: 206
Penestragania Beamer & Lawson, 1945
Penestragania confusa Linnavuori, 1956. Perez-Gelabert, 2008: 206
Subfamily IDIOCERINAE
Idiocerus Lewis, 1835
Idiocerus parvulus Osborn, 1935. Perez-Gelabert, 2008: 206
Subfamily LEDRINAE
Tribe XEROPHLOEINI
Xerophloea Germar, 1839
Xerophloea viridis (Fabricius, 1794). Dozier, 1931: 4
Subfamily NEOCOELIDIINAE
Krocarites Dietrich & Vega, 1995
Krocarites reflexa† Dietrich & Vega, 1995: 266. Dietrich, 2003: 701 [Fossil in Dominican amber]
Subfamily TYPHLOCYBINAE
Tribe ALEBRINI
Elabra Young, 1952
Elabra parallela (Osborn, 1928). Perez-Gelabert, 2008: 206
Eualebra Baker, 1898
Eualebra sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]
Omegalebra Young, 1957
Omegalebra cordiae (Osborn, 1929). Perez-Gelabert, 2008: 206
Plagalebra McAtee, 1926
Plagalebra similis (Baker, 1899). Dozier, 1931: 4; Young, 1957: 146; Metcalf, 1968: 23
Protalebra Baker, 1899
Protalebra sp.—Perez-Gelabert (2008: 206)
Protalebrella Young, 1952
Protalebrella brasiliensis (Baker, 1899). Young, 1957: 257
Rabela Young, 1952
Rabela tabebuiae (Dozier, 1927). Perez-Gelabert, 2008: 206
Trypanalebra Young, 1952
Trypanalebra maculata (Baker, 1903). Perez-Gelabert, 2008: 206

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Tribe DIKRANEURINI
Alconeura Ball & DeLong, 1925
Alconeura (Hyloidea) pistola (Caldwell, 1952). Perez-Gelabert, 2008: 206
Alconeura (Hyloidea) portoricensis Metcalf, 1968. Perez-Gelabert, 2008: 206
Hybla McAtee, 1932
Hybla maculata McAtee, 1932. Metcalf, 1968: 285
Idona DeLong, 1931
Idona minuenda (Moznette, 1919). Dozier, 1931: 4
Kunzeana Oman, 1949
Kunzeana sp.—Perez-Gelabert (2008: 206)
Typhlocybella Baker, 1903
Typhlocybella minima Baker, 1903. Perez-Gelabert, 2008: 206
Youngszella Freytag, 2000
Youngszella pseudomarginata (Caldwell, 1952). Perez-Gelabert, 2008: 206
Tribe EMPOASCINI
Empoasca Walsh, 1862
Empoasca canavalia DeLong, 1932: 114. Metcalf, 1968: 309
Empoasca canda Ross & Moore, 1957: 120
Empoasca cypha Caldwell, 1952. Perez-Gelabert, 2008: 207
Empoasca dicyrta Caldwell, 1952. Perez-Gelabert, 2008: 207
Empoasca dilitara DeLong & Davidson, 1935. Perez-Gelabert, 2008: 207
Empoasca fabalis DeLong, 1930: 92. DeLong, 1932: 113; Osborn, 1935: 185; Metcalf, 1968: 320
Empoasca gongyla Caldwell, 1952. Perez-Gelabert, 2008: 207
Empoasca gossypii DeLong, 1932: 114. Osborn, 1935: 186; Metcalf, 1968: 329
Empoasca hastosa Ross & Moore, 1957. SEA, 1999: 32
Empoasca kraemeri Ross & Moore, 1957. Schmutterer, 1990: 346
Empoasca millsi Ross, 1959: 315
Empoasca montana Caldwell, 1952. Perez-Gelabert, 2008: 207
Empoasca obliqua Caldwell, 1952. Perez-Gelabert, 2008: 207
Empoasca panisea Ross & Moore, 1957. Perez-Gelabert, 2008: 207
Empoasca papayae Oman, 1937. Martorell & Adsuar, 1952: 866; Metcalf, 1968: 343; Schmutterer, 1990: 267
Empoasca plebeia DeLong & Davidson, 1935. Metcalf, 1968: 536
Empoasca prona DeLong & Davidson, 1940. Perez-Gelabert, 2008: 207
Empoasca zapoides DeLong, 1930. Perez-Gelabert, 2008: 207
Tribe JORUMINI
Joruma McAtee, 1924
Joruma (Joruma) neascripta Oman, 1937. Perez-Gelabert, 2008: 207
Subfamily XESTOCEPHALINAE
Xestocephalus Van Duzee, 1892
Xestocephalus agassizi Van Duzee, 1912. Perez-Gelabert, 2008: 207
Xestocephalus bipunctatus Van Duzee, 1907. Perez-Gelabert, 2008: 207
Xestocephalus desertorum (Berg, 1879). Cwikla, 1985: 202
Xestocephalus sp.†—Dietrich & Vega (1995: 269) [Fossil in Dominican amber]
Xestocephalus subtessellatus Linnavuori, 1959. Cwikla, 1985: 199
Note: The citations for Hispaniola of Empoasca sativae Poos, 1959 by Metcalf, 1968: 542; Joruma ascripta McAtee, 1926 by
Metcalf, 1968: 602 and Joruma pisca McAtee, 1924 by Metcalf, 1968: 605 seem to be erroneous records (Freytag, pers.
comm.).

Family MEMBRACIDAE
Key reference: Ramos (1979).
Subfamily CENTROTINAE
Brachycentrotus Metcalf & Bruner, 1926
Brachycentrotus intermedius Ramos, 1979: 9 *DR > TL: Constanza, Prov. La Vega
Brachycentrotus rufinervis Ramos, 1957. Ramos, 1979: 10

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 269
Daimon Buckton, 1903
Daimon serricornis (Walker, 1858d: 77). Wolcott, 1927: 188; Ramos, 1979: 12
Monobeloides Ramos, 1979
Monobeloides cuneatus Ramos, 1979: 58 *DR > TL: Foothills of Cordillera Central, S. of Santiago
Monobeloides stuarti Ramos, 1979: 56
Monobelus Stål, 1866
Monobelus abudi Ramos, 1979: 23 *DR > TL: Constanza, 3–4000 ft.
Monobelus biguttatus (Fabricius, 1803). Ramos, 1979: 22
=Monobelus fasciatus Coquebert, 1899. Wolcott, 1927: 189
Monobelus flavidus (Fairmaire, 1846). Metcalf & Wade, 1965: 158
Monobelus majusculus Ramos, 1979: 25 *DR > TL: Jarabacoa
Monobelus obtusiceps Stål, 1869. Metcalf & Wade, 1965: 160
Orthobelus Stål, 1869
Orthobelus furcatus Ramos, 1979: 18 *DR > TL: San José de las Matas, 1000–2000 ft.
Orthobelus gomezmenori Peláez, 1936: 279. Ramos, 1979: 15
Orthobelus marcanoi Ramos, 1979: 17 *DR > TL: San Lorenzo
Orthobelus poeyi (Fairmaire, 1846). Peláez, 1936: 278; Metcalf & Wade, 1965: 152
Orthobelus truncaticornis Ramos, 1979: 19 *DR > TL: Without type locality data
Orthobelus urus (Fairmaire, 1846). Wolcott, 1927: 188; Peláez, 1936: 278; Ramos, 1979: 14
=Orthobelus megaceros Walker, 1851. Metcalf & Wade, 1965: 153
Orthobelus verrucosus Ramos, 1979: 20 *DR > TL: El Seibo
Orthobelus wolcotti Goding, 1928. Ramos, 1979: 16
Subfamily MEMBRACINAE
Platycotis Stål, 1869
Platycotis (Lophopelta) tuberculata (Fairmaire, 1846). Dozier, 1931: 2; Ramos, 1979: 60
Ramosella McKamey & Deitz, 1996
Ramosella dominicensis (Ramos, 1979: 62). McKamey & Deitz, 1996: 306 *DR > TL: La Laguna, Elias Piña
Ramosella thalli McKamey & Deitz, 1996: 306 *H > TL: Kenscoff
Umbonia Burmeister, 1835
Umbonia crassicornis Amyot & Serville, 1843. Ramos, 1979: 59 [Introduced]
Subfamily NESORRHINAE
Callicentrus Stål, 1869
Callicentrus callicerus Ramos, 1979: 50 *DR > TL: Loma Rucilla and Mts. N.
Callicentrus castaneus Ramos, 1979: 49 *H > TL: Furcy
Callicentrus ferrugineus Ramos, 1979: 47 *DR > TL: La Vega, La Palma, Constanza
Callicentrus multidentatus Ramos, 1979: 45 *DR > TL: Constanza
Callicentrus paucidentatus Ramos, 1979: 26 *DR > TL: Los Yayales, Nagua, Prov. María T. Sánchez
Callicentrus pluridentatus Ramos, 1979: 46 *DR > TL: Valle Nuevo, Prov. La Vega
Marshallella Goding, 1927
Marshallella padovanii Ramos, 1979: 54 *DR > TL: Loma Rucilla & Mts. N., 5–8000 ft.
Nessorhinus Amyot & Serville, 1843
Nessorhinus vulpes Amyot & Serville, 1843. Dozier, 1931: 2; Ramos, 1979: 52
Orekthophora Funkhouser, 1930
Orekthophora bicornata Ramos, 1990: 161 *DR > TL: Valle Nuevo, Constanza, Prov. La Vega
Orekthophora cornuta Funkhouser, 1930. Ramos, 1979: 52
Subfamily SMILIINAE
Antianthe Fowler, 1896
Antianthe foliacea (Stål, 1864). Ramos, 1979: 67
Antianthe viridissima (Walker, 1858). Wolcott, 1927: 188; Dozier, 1931: 2
Entylia Germar, 1835
Entylia carinata (Forster, 1771) [By synonymy of species below]
=Entylia areolata Walker, 1858d: 71. Metcalf & Wade, 1965: 902
Idioderma Van Duzee, 1909
Idioderma virescens Van Duzee, 1909. Howard et al., 1981: 45

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Micrutalis Fowler, 1895
Micrutalis calva (Say, 1830). Ramos, 1979: 68
Spissistilus Caldwell, 1949
Spissistilus festinus (Say, 1830). Ramos, 1979: 64; Abud, 1986: 4/86
Spissistilus rotundatus (Stål, 1869). McKamey, 1998: 256
Vanduzea Goding, 1893
Vanduzea segmentata (Fowler, 1895). Ramos, 1979: 65
Subfamily STEGASPIDINAE
Togotolania Cryan & Dietz, 2002
Togotolania longicorna Cryan & Dietz, 2002: 879. Dietrich, 2017: 64 *DR > TL: 2 km east of Boca Chica

Suborder STERNORRHYNCHA
Superfamily PSYLLOIDEA
Follows revised classification of Burckhardt & Ouvrard (2012).

Family APHALARIDAE
Subfamily RHINOCOLINAE
Leurolophus Tuthill, 1942
=Vicinilura† Klimaszewski, 1997
Leurolophus sp.†—Burckhardt & Basset (2000: 128) [Fossil in Dominican amber]
Tainarys Brèthes, 1920
Tainarys reposta† (Klimaszewski, 1996: 26). Burckhardt & Ouvrard, 2012: 24; Burckhardt & Queiroz, 2017: 562 [Fossil in
Dominican amber]

Family LIVIIDAE
Subfamily EUPHYLLURINAE
Tribe DIAPHORININI
Diaphorina Loew, 1880
Diaphorina citri Kuwayama, 1908. Halbert & Nuñez, 2004: 401 [Introduced]
Subfamily LIVIINAE
Diclidophlebia Crawford, 1920
=Primascena† Klimaszewski, 1998
Diclidophlebia subita† (Klimaszewski, 1998: 21) [Fossil in Dominican amber]

Family PHACOPTERONIDAE
Sulciana† Klimaszewski, 1998
Sulciana macroconi† Klimaszewski, 1998: 22 [Fossil in Dominican amber]

Family PSYLLIDAE
Subfamily CIRIACREMINAE
Euceropsylla Boselli, 1929
Euceropsylla russoi Boselli, 1929: 71. Maia, 2006: 8 *DR > TL: San Domingo su Inga vera
Heteropsylla Crawford, 1914
Heteropsylla cubana Crawford, 1914. Muddiman et al., 1992: 84
Heteropsylla fusca Crawford, 1914: 48. Muddiman et al., 1992: 87
Heteropsylla huasache Caldwell, 1941. Muddiman et al., 1992: 88
Heteropsylla propinqua Muddiman et al., 1992: 91
Heteropsylla reducta Caldwell & Martorell, 1952. Muddiman et al., 1992: 92
Subfamily MACROCORSINAE
Euryconus Aulmann, 1912
=Otroacizzia† Klimaszewski, 1996
Euryconus muta† (Klimaszewski, 1996: 32) [Fossil in Dominican amber]
Euryconus prosapia† (Klimaszewski, 1996: 33) [Fossil in Dominican amber]
Euryconus soriae† (Peñalver & García-Gimeno, 2006: 200) [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 271
Euryconus tertia† (Klimaszewski, 1996: 34) [Fossil in Dominican amber]
Subfamily PSYLLINAE
Limbopsylla Brown & Hodkinson, 1988
=Trioacantha† Klimaszewski, 1998
=Indepsylla† Klimaszewski, 1996
Limbopsylla indocilia† (Klimaszewski, 1998: 24) [Fossil in Dominican amber]
Limbopsylla mortua† (Klimaszewski, 1996: 28) [Fossil in Dominican amber]
Subfamily ACIZZIINAE
Platycorypha Tuthill, 1945
=Paropsylla† Klimaszewski, 1996
=Parapsyllopsis† Klimaszewski, 1996
Platycorypha carabeus† (Klimaszewski, 1996: 30) [Fossil in Dominican amber]
Platycorypha olima† (Klimaszewski, 1996: 36) [Fossil in Dominican amber]

Family TRIOZIDAE
Calinda Blanchard, 1852
Calinda graciliforceps Olivares & Burckardt, 1997: 276
Ceropsylla Riley, 1883
Ceropsylla sideroxyli Riley, 1884. Abud, 1993: 8/93
Heterotrioza Dobreanu & Manolache, 1960
Heterotrioza grimaldii† Klimaszewski, 1996: 36 [Fossil in Dominican amber]

Superfamily ALEYRODOIDEA
Family ALEYRODIDAE
Key reference: Mound and Halsey (1978).
Subfamily ALEYRODINAE
Aleurocanthus Quaintance & Baker, 1914
Aleurocanthus spiniferus (Quaintance, 1903). Evans, 2007: 86
Aleurocanthus woglumi Ashby, 1915. Wolcott, 1927: 198; Schmutterer, 1990: 208 [Introduced]
Aleuroglandulus Bondar, 1923
Aleuroglandulus malangae Russell, 1944. Schmutterer, 1990: 517; Alvarez et al., 1993: 35
Aleuroglandulus subtilis Bondar, 1923. Martin, 2005: 15
Aleurolobus Quaintance & Baker, 1914
Aleurolobus marlatti (Quaintance, 1903). Evans, 2007: 139
Aleuroplatus Quaintance & Baker, 1914
Aleuroplatus cococolus Quaintance & Baker, 1917. Evans, 2007: 158
Aleuroplatus validus Quaintance & Baker, 1917. Evans, 2007: 169
Aleuropleurocelus Drews & Sampson, 1956
Aleuropleurocelus abnormis (Quaintance & Baker, 1900). Evans, 2007: 171
Aleurotrachelus Quaintance & Baker, 1914
Aleurotrachelus trachoides (Back, 1912). Russo, 1927: 45; Schmutterer, 1990: 499; Alvarez et al., 1993: 35
Aleurothrixus Quaintance & Baker, 1914
Aleurothrixus floccosus (Maskell, 1895). Mound & Halsey, 1978: 62; Alvarez et al., 1993: 35
Aleurothrixus myrtacei Bondar, 1923. Evans, 2007: 183
Aleurotuberculatus Takahashi, 1932
Aleurotuberculatus minutus (Singh, 1931). Abud, 1996: 1/96; SEA, 1999: 111
Bellitudo Russell, 1943
Bellitudo hispaniolae Russell, 1943: 136. Mound & Halsey, 1978: 110 *DR, *H > TL: Santo Domingo
Bemisia Quaintance & Baker, 1914
Bemisia tabaci (Gennadius, 1889). Schmutterer, 1990: 399; Alvarez et al., 1993: 34; Zuluaga & Mesa, 2000: 197 [Introduced]
=Bemisia argentifolii Bellows & Perring, 1994. Zuluaga & Mesa, 2000: 197
Crenidorsum Russell, 1945
Crenidorsum aroidephagus Martin & Aguiar, 2001: 274. Evans, 2007: 274
Crenidorsum commune Russell, 1945: 60. Mound & Halsey, 1978: 127

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Crenidorsum debordae Russell, 1945: 64. Mound & Halsey, 1978: 128 *H > TL: Petit Gonave Island, from Coccoloba
Crenidorsum diaphanum Russell, 1945: 65. Mound & Halsey, 1978: 128 *H > TL: West of Cabaret, from Coccoloba
Crenidorsum magnisetae Russell, 1945: 63. Mound & Halsey, 1978: 128
Crenidorsum marginale Russell, 1945: 63. Mound & Halsey, 1978: 128 *DR > TL: from Coccoloba pubescens L., near
Barahona
Dialeurodes Cockerell, 1902
Dialeurodes citri (Ashmead, 1885). Evans, 2007: 290
Minutaleyrodes Jesudasan & David, 1990
Minutaleyrodes minuta (Singh, 1931). Evans, 2007: 369
Parabemisia Takahashi, 1952
Parabemisia myricae (Kuwana, 1927). Evans, 2007: 388
Paraleyrodes Quaintance, 1909
Paraleyrodes urichii Quaintance & Baker, 1913. Evans, 2007: 54
Singhiella Sampson, 1943
Singhiella citrifolii (Morgan, 1893). Schmutterer, 1990: 200; Evans, 2007: 436
Singhiella simplex (Singh, 1931). Serra et al., 2011: 148 [Introduced]
Tetraleurodes Cockerell, 1902
Tetraleurodes acaciae (Quaintance, 1900). Nakahara, 1995: 108
Tetraleurodes pseudacaciae Nakahara, 1995: 119
Trialeurodes Cockerell, 1902
Trialeurodes floridensis (Quaintance, 1900). Evans, 2007: 476
Trialeurodes vaporariorum (Westwood, 1856). Schmutterer, 1990: 395; Alvarez et al., 1993: 34; Zuluaga & Mesa, 2000: 197
[Introduced]
Trialeurodes variabilis (Quaintance, 1900). Evans, 2007: 487
Subfamily ALEYRODICINAE
Aleurodicus Douglas, 1892
Aleurodicus cocois (Curtis, 1846). Schmutterer, 1990: 125; Alvarez et al., 1993: 35
Aleurodicus dispersus Russell, 1965. Mound & Halsey, 1978: 230
Aleurodicus ornatus Cockerell, 1893. Evans, 2007: 22
Metaleurodicus Quaintance & Baker, 1913
Metaleurodicus cardini (Back, 1912). Evans, 2007: 43

Superfamily APHIDOIDEA
Family APHIDIDAE
Key reference: Etienne (2005).
Subfamily LACHNINAE
Tribe CINARINI
Cinara Curtis, 1835
Cinara carolina Tissot, 1944. Abud, 1981: 146
Subfamily CHAITOPHORINAE
Tribe SIPHINI
Sipha Passerini, 1860
Sipha flava (Forbes, 1884). Wolcott, 1922: 33; Smith & Cermeli, 1979: 60; Schmutterer, 1990: 77
Subfamily GREENIDEINAE
Quisqueyaaphis† Wegierek, 2001
Quisqueyaaphis heiei† Wegierek, 2001: 410 [Fossil in Dominican amber]
Subfamily APHIDINAE
Tribe APHIDINI
Subtribe APHIDINA
Aphis Linnaeus, 1758
Aphis craccivora Koch, 1854. Schmutterer, 1990: 347; Zuluaga & Mesa, 2000: 126; Etienne, 2005: 459
Aphis citricola Van der Goot, 1912. Smith & Cermeli, 1979: 8
Aphis fabae Scopoli, 1763. Smith & Cermeli, 1979: 10; Etienne, 2005: 459
Aphis forbesi Weed, 1889. SEA, 1999: 84

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 273
Aphis gossypii Glover, 1877. Wolcott, 1927: 195; Smith & Cermeli, 1979: 11; Schmutterer, 1990: 434; Etienne, 2005: 459
Aphis illinoisensis Shimer, 1866. Schmutterer, 1990: 263; Etienne, 2005: 459
Aphis nerii Boyer de Fonscolombe, 1841. Smith & Cermeli, 1979: 13; Etienne, 2005: 459
Aphis spiraecola Patch, 1914. Etienne, 2005: 459
Toxoptera Koch, 1856
Toxoptera aurantii (Boyer, 1841). Wolcott, 1927: 195; Smith & Cermeli, 1979: 64; Etienne, 2005: 460
Toxoptera citricidus (Kirkaldy, 1907). Leclant et al., 1992: 162; Stoetzel, 1994: 75; Zuluaga & Mesa, 2000: 197; Etienne, 2005:
460 [Introduced]
Subtribe RHOPALOSIPHINA
Hysteroneura Davis, 1919
Hysteroneura setariae (Thomas, 1878). SEA, 1999: 99
Melanaphis van der Goot, 1917
Melanaphis sacchari (Zhentner, 1897). Schmutterer, 1990: 72; Etienne, 2005: 459
Rhopalosiphum Koch, 1854
Rhopalosiphum maidis (Fitch, 1856). Smith & Cermeli, 1979: 58; Etienne, 2005: 460
Rhopalosiphum pseudobrassicae (Davis, 1914). SEA, 1999: 27
Rhopalosiphum rufiabdominale (Sasaki, 1899). Abud & Marcano, 1988: 9/88; SEA, 1999: 90; Etienne, 2005: 460
Schizaphis Boerner, 1931
Schizaphis graminum (Rondani, 1852). Etienne, 2005: 460
Tribe MACROSIPHINI
Aulacorthum Mordvilko, 1914
Aulacorthum solani (Kaltenbach, 1834). Schmutterer, 1990: 455; Etienne, 2005: 459
Brevicoryne van der Goot, 1915
Brevicoryne brassicae (Linnaeus, 1758). Santoro, 1960: 212; Smith & Cermeli, 1979: 20; Etienne, 2005: 459
Cavariella Del Guercio, 1911
Cavariella sp.—SEA (1999: 14)
Chaetosiphon Mordvilko, 1914
Chaetosiphon fragaefolii (Cockerell, 1901). Abud, 1996: 4/96
Lipaphis Mordvilko, 1928
Lipaphis erysimi (Kaltenbach, 1843). Etienne, 2005: 459
Macrosiphoniella Del Guercio, 1911
Macrosiphoniella sanborni (Gillette, 1908). SEA, 1999: 23
Macrosiphum Passerini, 1860
Macrosiphum euphorbiae (Thomas, 1878). Schmutterer, 1990: 456; Etienne, 2005: 459
Macrosiphum martorelli Smith, 1960. Smith & Cermeli, 1979: 43; Schmutterer, 1990: 163
Macrosiphum mesosphaeri Tissot, 1934. SEA, 1999: 64
Macrosiphum rosae (Linnaeus, 1758). Abud, 1994b: 10/94
Macrosiphum luteum (Buckton, 1876). Abud, 1981: 91
Microparsus Patch, 1909
Microparsus brasiliensis (Moreira, 1925). Etienne, 2005: 459
Myzus Passerini, 1860
Myzus (Nectarosiphon) persicae (Sulzer, 1776). Beinhart, 1952: 713; Smith & Cermeli, 1979: 48; Schmutterer, 1990: 425;
Etienne, 2005: 460
Pentalonia Coquerel, 1859
Pentalonia nigronervosa Cockerell, 1859. Santoro, 1960: 330; Etienne, 2005: 460
Picturaphis Blanchard, 1922
Picturaphis brasiliensis Moreira, 1925. Abud, 1981b: 95; Schmutterer, 1990: 394
Sitobium Mordvilko, 1914
Sitobium martorelli (Smith, 1960). Etienne, 2005: 460
Uroleucon Mordvilko, 1914
Uroleucon ambrosiae (Thomas, 1878). Schmutterer, 1990: 457; SEA, 1999: 24
Subfamily HORMAPHIDINAE
Cerataphis Lichtenstein, 1882
Cerataphis brasiliensis (Hempel, 1901). Etienne, 2005: 459

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Cerataphis lataniae (Boisduval, 1867). Smith & Cermeli, 1979: 23; Etienne, 2005: 459
Cerataphis orchidearum (Westwood, 1879). SEA, 1999: 84
Cerataphis palmae (Ghesquière, 1934). SEA, 1999: 18
Cerataphis variabilis Hille Ris Lambers, 1953. Abud, 1994b: 3/94
Subfamily PEMPHIGINAE
Tribe ERIOSOMATINI
Eriosoma Leach, 1818
Eriosoma lanigerum (Hausmann, 1802). Russo, 1927: 45; Smith & Cermeli, 1979: 31; Etienne, 2005: 459
Tetraneura Hartig in Germar, 1841
Tetraneura nigriabdominalis (Sasaki, 1899). Valdez, Matos & Álvarez, 2016: 50
Tribe FORDINI
Geopemphigus Hille Ris Lambers, 1933
Geopemphigus floccosus (Moreira, 1925). Abud, 1993: 7/93

Family DREPANOSIPHIDAE
Mindazerius† Heie & Poinar, 1988
Mindazerius dominicanus† Heie & Poinar, 1988: 155 [Fossil in Dominican amber]
Tinocallis Matsumura, 1919
Tinocallis kahawaluokalani (Kirkaldy, 1907). Marcano & Abud, 1994: 2/94

Family HORMAPHIDIDAE
Dominicaphis† Heie & Poinar, 1999
Dominicaphis succini† Heie & Poinar, 1999: 816 [Fossil in Dominican amber]

Family TERMITAPHIDIDAE
Termitaradus Myers, 1924
Termitaradus avitinquilinus† Grimaldi & Engel, 2008: 2 [Fossil in Dominican amber]
=Termitaradus sp.†—Grimaldi & Engel (2005: 329) [Fossil in Dominican amber]
Termitaradus dominicanus† Poinar in Poinar & Heiss, 2012: 53 [Fossil in Dominican amber]
Termitaradus mitnicki† Engel, 2009: 63 [Fossil in Dominican amber]

Superfamily COCCOIDEA
Family CEROCOCCIDAE
Cerococcus Comstock, 1982
Cerococcus deklei Kostarab & Vest, 1966. Matile-Ferrero & Etienne, 2006: 164

Family COCCIDAE
Ceroplastes Gray, 1828
Ceroplastes cirripediformis Comstock, 1881. Santoro, 1960: 183; SEA, 1999: 20
Ceroplastes floridensis Comstock, 1881. Gómez-Menor, 1941: 129
Ceroplastes giganteus Dozier, 1931: 2. Ben-Dov, 1993: 37 *H > TL: St. Marc
Ceroplastes rubens Maskell, 1893. Dekle, 1971: 1
Ceroplastes rusci (Linnaeus, 1751). SEA, 1999: 83
Ceroplastes utilis Cockerell, 1893. Gimpel et al., 1974: 67; Ben-Dov, 1993: 59
=Ceroplastes dozieri Cockerell & Bueker, 1930: 7. Ben-Dov, 1993: 59
Coccus Linnaeus, 1758
Coccus acutissimus (Green, 1896). SEA, 1999: 9
Coccus hesperidum (Linnaeus, 1758). Gómez-Menor, 1941: 131
Coccus moestus De Lotto, 1959. Gill et al., 1977: 28; Ben-Dov, 1993: 84
Coccus viridis (Green, 1889). Gómez-Menor, 1941: 131; Schmutterer, 1990: 147; Gill et al., 1977: 37; Matile-Ferrero &
Etienne, 2006: 165
Eucalymnatus Cockerell, 1901
Eucalymnatus tessellatus (Signoret, 1873). Panis & Martin, 1976: 7; Ben-Dov, 1993: 120; Matile-Ferrero & Etienne, 2006: 165
Kilifia De Lotto, 1965

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Kilifia acuminata (Signoret, 1873). Panis & Martin, 1976: 7; Schmutterer, 1990: 226; Ben-Dov, 1993: 150
Milviscutulus Williams & Watson, 1990
Milviscutulus mangiferae (Green, 1889). Gómez-Menor, 1941: 131; Panis & Martin, 1976: 7; Schmutterer, 1990: 226; Ben-
Dov, 1993: 186; Matile-Ferrero & Etienne, 2006: 165
Parasaissetia Takahashi, 1955
Parasaissetia nigra (Nietner, 1861). Ben-Dov, 1993: 209
Philephedra Cockerell, 1899
Philephedra tuberculosa Nakahara & Gill, 1985. SEA, 1999: 30
Protopulvinaria Cockerell, 1894
Protopulvinaria longivalvata (Green, 1909). Schmutterer, 1990: 258
Protopulvinaria pyriformis (Cockerell, 1894). Panis & Martin, 1976: 7; Ben-Dov, 1993: 243; Matile-Ferrero & Etienne, 2006:
166
Pulvinaria Targioni-Tozzetti, 1867
Pulvinaria elongata Newstead, 1917. Ben-Dov, 1993: 257
Pulvinaria psidii Maskell, 1893. Gómez-Menor, 1941: 140; Panis & Martin, 1976: 8; Ben-Dov, 1993: 278; Stocks, 2013: 349
Pulvinaria urbicola Cockerell, 1893. Gómez-Menor, 1941: 140; Panis & Martin, 1976: 8; Ben-Dov, 1993: 286; Matile-Ferrero
& Etienne, 2006: 167
Saissetia Déplanche, 1858
Saissetia coffeae (Walker, 1852) [By synonymy of species below]
=Saissetia hemisphaerica (Targioni-Tozetti, 1867). Gómez-Menor, 1941: 140
Saissetia neglecta De Lotto, 1969. Malumphi, 2017: 266 [Intercepted in England on Annona muricata fruit from Dominican
Republic; Introduced]
Saissetia oleae (Olivier, 1791). Gómez-Menor, 1941: 141; Schmutterer, 1990: 192
Vinsonia Signoret, 1872
Vinsonia stellifera (Westwood, 1871). Gómez-Menor, 1941: 142; Schmutterer, 1990: 223 [Introduced]

Family CONCHASPIDIDAE
Conchaspis Cockerell, 1893
Conchaspis cordiae Mamet, 1954. Panis & Martin, 1976: 8; Ben-Dov, 1981: 147; Howard et al., 2006: 102

Family DACTYLOPIIDAE
Dactylopius Costa, 1836
Dactylopius confusus (Cockerell, 1893). Miller, 1996: 71

Family ASTEROLECANIIDAE
Asterolecanium Targioni-Tozzetti, 1868
Asterolecanium arabidis (Signoret, 1876). Matile-Ferrero & Etienne, 2006: 162
Asterolecanium epidendri (Bouché, 1833). Russell, 1941: 83
Bambusaspis Cockerell, 1902
Bambusaspis bambusae (Boisduval, 1869). Russell, 1941: 47; Gómez-Menor, 1941: 129; Hamon, 1980: 2
Mycococcus Ferris, 1952
Mycococcus coperniciae Ferris, 1952. Matile-Ferrero & Etienne, 2006: 163
Russellaspis Bodenheimer, 1951
Russellaspis pustulans (Cockerell, 1892). Dozier, 1933: 96; Gómez-Menor, 1941: 128; Matile-Ferrero & Etienne, 2006: 163

Family DIASPIDIDAE
Subfamily ASPIDIOTINAE
Aonidiella Berlese & Leonardi, 1895
Aonidiella aurantii (Maskell, 1879). Nakahara, 1982: 6
Aonidiella comperei McKenzie, 1937. McKenzie, 1946: 31
Aonidiella inornata (McKenzie, 1938). Nakahara, 1982: 7
Aonidiella orientalis (Newstead, 1894). Schmutterer, 1990: 121
Aspidiella Leonardi, 1898
Aspidiella hartii (Cockerell, 1895). Gómez-Menor, 1941: 128

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Aspidiella sacchari (Cockerell, 1893). Gómez-Menor, 1941: 128
Aspidiotus Bouché, 1833
Aspidiotus destructor Signoret, 1869. Wolcott, 1927: 203; Gómez-Menor, 1941: 126; Santoro, 1960: 201; Schmutterer, 1990:
123; Matile-Ferrero & Etienne, 2006: 168
Aspidiotus excisus Green, 1896. Nakahara, 1982: 13; Matile-Ferrero & Etienne, 2006: 169
Aspidiotus lataniae Signoret, 1869. Dozier, 1933: 94; Gómez-Menor, 1941: 128
Aspidiotus nerii Bouché, 1833. Nakahara, 1982: 14
Aspidiotus persearum Cockerell, 1898. Nakahara, 1982: 15
Clavaspis MacGillivray, 1921
Clavaspis herculeana (Cockerell & Hadden, 1909). Malumphi, 2017: 266 [Intercepted in England on Annona muricata fruit
from Dominican Republic; Introduced]
Chrysomphalus Ashmead, 1880
Chrysomphalus aonidium (Linnaeus, 1758). Wolcott, 1927: 203; Smith & Audant, 1930: 978; Gómez-Menor, 1941: 130
Chrysomphalus dictyospermi (Morgan, 1889). Gómez-Menor, 1941: 130; Matile-Ferrero & Etienne, 2006: 170
Chrysomphalus pinnulifer (Maskell, 1891). Smith-Pardo, Evans & Dooley, 2012: 15
Crenulaspidiotus MacGillivray, 1921
Crenulaspidiotus portoricensis (Lindinger, 1910). Miller & Davidson, 1981: 572
Crenulaspidiotus rusellae Miller & Davidson, 1981: 576
Crenulaspidiotus truncus Miller & Davidson, 1981: 581
Diaspidiotus Berlese, 1896
Diaspidiotus viticola (Leonardi, 1913). Gómez-Menor, 1941: 128
Duplaspidiotus MacGillivray, 1921
Duplaspidiotus tesseratus (Grandpré & Charmoy, 1899). Gómez-Menor, 1941: 139
Dynaspidiotus Thiem & Gerneck, 1934
Dynaspidiotus californicus (Coleman, 1903) [By synonymy of species below]
=Aspidiotus pini Comstock, 1881. Gómez-Menor, 1941: 128
Furcaspis Lindinger, 1908
Furcaspis biformis (Cockerell, 1893). SEA, 1999: 66
Hemiberlesia Leonardi, 1897
Hemiberlesia lataniae (Signoret, 1869). Schmutterer, 1990: 263
Hemiberlesia palmae (Cockerell, 1893). Malumphi, 2017: 266 [Intercepted in England on Annona muricata fruit from Haiti;
Introduced]
Hemigymnaspis Lindinger, 1934
Hemigymnaspis pimentae Davidson & Miller, 1977: 514 *DR > TL: Dominican Republic, ex Pimenta sp. leaf
Lindingaspis MacGillivray, 1921
Lindingaspis floridana Ferris, 1942. Nakahara, 1982: 51
Melanaspis Cockerell, 1897
Melanaspis bromiliae (Leonardi, 1899). Nakahara, 1982: 53; Deitz & Davidson, 1986: 18
Melanaspis calura (Cockerell, 1898). Deitz & Davidson, 1986: 20; Miller, 1996: 75
Melanaspis coccolobae Ferris, 1943. Nakahara, 1982: 54; Matile-Ferrero & Etienne, 2006: 172
Melanaspis tenax McKenzie, 1944. Deitz & Davidson, 1986: 80
Morganella Cockerell, 1897
Morganella longispina (Morgan, 1889). Dekle, 1964: 1; Nakahara, 1982: 57; Matile-Ferrero & Etienne, 2006: 172
Mycetaspis Cockerell, 1897
Mycetaspis personata (Comstock, 1883). Gómez-Menor, 1941: 139; Matile-Ferrero & Etienne, 2006: 172
Oceanaspidiotus Takagi, 1984
Oceanaspidiotus spinosus (Comstock,1883). Nakahara, 1982: 15
Parlatoria Targioni-Tozzetti, 1868
Parlatoria cinerea Hadden, 1909. Morrison, 1939b: 10; Matile-Ferrero & Etienne, 2006: 172
Parlatoria crotonis Douglas, 1887. Morrison, 1939b: 12; Nakahara, 1982: 66
Parlatoria pergandii Comstock, 1881. Morrison, 1939b: 18; Gómez-Menor, 1941: 138
Parlatoria proteus (Curtis, 1843). Morrison, 1939b: 22
Parlatoria ziziphi (Lucas, 1853). Morrison, 1939: 27b; Matile-Ferrero & Etienne, 2006: 173
Pseudaonidia Cockerell, 1897

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 277
Pseudaonidia trilobitiformis (Green, 1896). Schmutterer, 1990: 228
Pseudischnaspis Hempel, 1900
Pseudischnaspis bowreyi (Cockerell, 1893). Gómez-Menor, 1941: 139; Nakahara, 1982: 75
Radionaspis Ferris, 1942
Radionaspis indica (Marlatt, 1908). Nakahara, 1982: 80
Selenaspidus Cockerell, 1897
Selenaspidus articulatus (Morgan, 1889). Gómez-Menor, 1941: 142
Subfamily COMSTOCKIELLINAE
Comstockiella Cockerell, 1896
Comstockiella sabalis (Comstock, 1883). Gómez-Menor, 1941: 132; Nakahara, 1982: 26
Subfamily DIASPIDINAE
Andaspis MacGillivray, 1921
Andaspis punicae (Laing, 1929). Matile-Ferrero & Etienne, 2006: 167
Aonidomytilus Leonardi, 1903
Aonidomytilus albus (Cockerell, 1893). Gómez-Menor, 1941: 138
Aulacaspis Cockerell, 1893
Aulacaspis rosae (Bouché, 1834). Nakahara, 1982: 15
Aulacaspis tubercularis (Newstead, 1906). Abud & Marcano, 1987: 11/87; Schmutterer, 1990: 229; Stocks, 2013: 354
[Introduced]
Aulacaspis yasumatsui Takagi, 1977. Perez-Gelabert, 2019: 525 [Introduced]
Chionaspis Signoret, 1869
Chionaspis pinifoliae (Fitch, 1855). Gómez-Menor, 1941: 130; SEA, 1999: 87
Diaspis Costa, 1828
Diaspis boisduvallii Signoret, 1869. Gómez-Menor, 1941: 132; Santoro, 1960: 327
Diaspis bromeliae (Kerner, 1778). Gómez-Menor, 1941: 132; Nakahara, 1982: 30
Diaspis echinocacti (Bouché, 1833). Gómez-Menor, 1941: 132
Fiorinia Targioni-Tozzetti, 1868
Fiorinia fioriniae (Targioni-Tozetti, 1867). Nakahara, 1982: 36
Howardia Berlese & Leonardi, 1896
Howardia biclavis (Comstock, 1883). Gómez-Menor, 1941: 137
Ischnaspis Douglas, 1887
Ischnaspis longirostris (Signoret, 1882). Gómez-Menor, 1941: 138; Matile-Ferrero & Etienne, 2006: 171
Lepidosaphes Shimer, 1868
Lepidosaphes beckii (Newman, 1869). Wolcott, 1927: 203; Gómez-Menor, 1941: 138; Santoro, 1960: 186
Lepidosaphes gloverii (Packard, 1869). Gómez-Menor, 1941: 138
Lepidosaphes tokionis (Kuwana, 1902). Nakahara, 1982: 50
Pinnaspis Cockerell, 1892
Pinnaspis aspidistrae (Signoret, 1869). SEA, 1999: 11
Pinnaspis buxi (Bouché, 1851). Gómez-Menor, 1941: 139
Pinnaspis strachani (Cooley, 1899). Schmutterer, 1990: 246; Malumphi, 2017: 266 [Introduced]
Pseudaulacaspis MacGillivray, 1921
Pseudaulacaspis major (Cockerell, 1893). Nakahara, 1982: 74
Pseudaulacaspis pentagona (Targioni-Tozetti, 1886). Wolcott, 1927: 203; Gómez-Menor, 1941: 129; Schmutterer, 1990: 361;
Matile-Ferrero & Etienne, 2006: 173
Pseudoparlatoria Cockerell, 1892
Pseudoparlatoria ostreata Cockerell, 1892. Gómez-Menor, 1941: 140; Nakahara, 1982: 76; Schmutterer, 1990: 271; Matile-
Ferrero & Etienne, 2006: 174
Rutherfordia MacGillivray, 1921
Rutherfordia major (Cockerell, 1894). Gómez-Menor, 1941: 129
Unaspis MacGillivray, 1921
Unaspis citri (Comstock, 1883). Gómez-Menor, 1941: 130; Santoro, 1960: 184; Schmutterer, 1990: 200
Subfamily LEUCASPIDINAE
Lopholeucaspis Balachowsky, 1953
Lopholeucaspis cockerelli (Grandpré & Charmoy, 1899). Nakahara, 1982: 52

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Family ERIOCOCCIDAE
Eriococcus Targioni-Tozzetti, 1868
Eriococcus amomidis Gómez Menor, 1935: 2152. Gómez-Menor, 1941: 132; Miller et al., 2000: 124 *DR > TL: San Cristóbal

Family MONOPHLEBIDAE
Tribe ICERYINI
Crypticerya Cockerell, 1895
Crypticerya rosae (Riley & Howard, 1890). Wolcott, 1927: 205; Gómez-Menor, 1941: 142
Icerya Signoret, 1875
Icerya montserratensis Riley & Howard, 1890. Gómez-Menor, 1941: 142
Icerya purchasi Maskell, 1878. Wolcott, 1927: 206; Gómez-Menor, 1941: 142; Schmutterer, 1990: 350; Matile-Ferrero &
Etienne, 2006: 175 [Introduced]
Tribe LLAVEIINI
Protortonia Towsend, 1898
Protortonia sp.—Williams & Gullan (2008: 87)

Family PSEUDOCOCCIDAE
Antonina Signoret, 1875
Antonina graminis (Maskell, 1897). Abud & Marcano, 1981: 104
Dysmicoccus Ferris, 1950
Dysmicoccus boninsis (Kuwana, 1909). Wolcott, 1922: 34; Gómez-Menor, 1941: 139; Schmutterer, 1990: 84; Ben-Dov, 1994:
122; Watson & Chandler, 2000: 30; Granara de Willink, 2009: 21
Dysmicoccus brevipes (Cockerell, 1893). Gómez-Menor, 1941: 140; Schmutterer, 1990: 444; Ben-Dov, 1994: 124; Watson &
Chandler, 2000: 30; Matile-Ferrero & Etienne, 2006: 177; Granara de Willink, 2009: 24
Dysmicoccus grassii (Leonardi, 1913). Ben-Dov, 1994: 130; Watson & Chandler, 2000: 30; Granara de Willink, 2009: 40
=Dysmicoccus alazon Williams, 1960. Ben-Dov, 1994: 130
Dysmicoccus neobrevipes Beardsley, 1959. Ben-Dov, 1994: 137; Watson & Chandler, 2000: 30; Granara de Willink, 2009: 51
Dysmicoccus texensis (Tinsley, 1900). Watson & Chandler, 2000: 30; Granara de Willink, 2009: 69 [Introduced]
=Dysmicoccus bispinosus Beardsley, 1965. Ben-Dov, 1994: 122; Granara de Willink, 2009: 69
Electromyrmococcus† Williams, 2001
Electromyrmococcus abductus† Williams in Johnson et al., 2001: 17 [Fossil in Dominican amber]
Electromyrmococcus inclusus† Williams & Agosti in Johnson et al., 2001: 17 [Fossil in Dominican amber]
Electromyrmococcus reginae† Williams in Johnson et al., 2001: 18 [Fossil in Dominican amber]
Ferrisia Fullaway, 1923
Ferrisia cristinae Kaydan & Gullan, 2012: 16
Ferrisia uzinuri Kaydan & Gullan, 2012: 48
Ferrisia virgata (Cockerell, 1893). SEA, 1999: 9
Geococcus Green, 1902
Geococcus coffeae Green, 1933. Ben-Dov, 1994: 172; Watson & Chandler, 2000: 30
Hypogeococcus Rau, 1938
Hypogeococcus pungens Granara de Willink, 1981. German-Ramirez et al., 2014: 320
Lantanacoccus Williams & Granara de Willink, 1992
Lantanacoccus sauroides Williams & Granara de Willink, 1992: 224. Ben-Dov, 1994: 212; Watson & Chandler, 2000: 31;
Matile-Ferrero & Etienne, 2006: 178
Maconellicoccus Ezzat, 1958
Maconellicoccus hirsutus (Green, 1908). Serra, 2006: 37 [Introduced]
Nipaecoccus Šulc, 1945
Nipaecoccus filamentosus (Cockerell, 1893). Ben-Dov, 1994: 250; Miller, 1996: 82
Nipaecoccus nipae (Maskell, 1893). Gómez-Menor, 1941: 139; Panis & Martin, 1976: 8; Schmutterer, 1990: 119; Ben-Dov,
1994: 252; Watson & Chandler, 2000: 30; Matile-Ferrero & Etienne, 2006: 180; Malumphi, 2017: 267
Paracoccus Ezzat & McConnell, 1956
Paracoccus marginatus Williams & Granara de Willink, 1992. Miller et al., 1999: 179; Watson & Chandler, 2000: 30; Walker
et al., 2003: 1; Matile-Ferrero & Etienne, 2006: 180; Stocks, 2013: 346 [Introduced]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 279
Paraputo Laing, 1929
Paraputo guatemalensis (Ferris, 1953). Ben-Dov, 1994: 70; Miller, 1996: 81; Watson & Chandler, 2000: 30
Phenacoccus Cockerell, 1893
Phenacoccus gossypii Townsend & Cockerell, 1898. Gómez-Menor, 1941: 138
Phenacoccus madeirensis Green, 1923. Ben-Dov, 1994: 329; Watson & Chandler, 2000: 30
Phenacoccus sisalanus Granara de Willink in Granara de Willink et al., 2007: 101 *H, *DR > TL: Rosario
Phenacoccus solenopsis Tinsley, 1898. Ben-Dov, 1994: 344; Miller, 1996: 83; Watson & Chandler, 2000: 30; Matile-Ferrero &
Etienne, 2006: 182 [Introduced]
Planococcus Ferris, 1950
Planococcus citri (Risso, 1813). Gómez-Menor, 1941: 139; Panis & Martin, 1976: 8; Ben-Dov, 1994: 355; Watson & Chandler,
2000: 30
Planococcus ficus (Signoret, 1875). Ezzat & McConnell, 1956: 80
Planococcus halli Ezzat & McConnell, 1956. Ben-Dov, 1994: 361; Watson & Chandler, 2000: 30; Matile-Ferrero & Etienne,
2006: 182
Planococcus lilacinus (Cockerell, 1905). Ben-Dov, 1994: 363; Watson & Chandler, 2000: 30; Stocks, 2013: 348 [Introduced]
Planococcus minor (Maskell, 1897). Gómez-Menor, 1941: 137; Ben-Dov, 1994: 365; Watson & Chandler, 2000: 31;
Malumphi, 2017: 267 [Introduced]
Pseudococcus Westwood, 1840
Pseudococcus boninensis Lindinger, 1914. SEA, 1999: 96
Pseudococcus cryptus (Hempel, 1918). Malumphi, 2017: 267 [Intercepted in England on Annona muricata fruit from
Dominican Republic; Introduced]
Pseudococcus elisae Borchsenius, 1947. Ben-Dov, 1994: 389
Pseudococcus jackbeardsleyi Gimpel & Miller, 1996: 66. Watson & Chandler, 2000: 30
Pseudococcus longispinus (Targioni-Tozetti, 1867). Schmutterer, 1990: 520; Ben-Dov, 1994: 397; Watson & Chandler, 2000:
30
=Pseudococcus adonidum (Linnaeus, 1767). Valdéz et al., 2016: 27
Pseudococcus microcirculus McKenzie, 1960. Gimpel & Miller, 1996: 83; Watson & Chandler, 2000: 30
Puto Signoret, 1876
Puto barberi (Cockerell, 1895). Ben-Dov, 1994: 419; Matile-Ferrero & Etienne, 2006: 183; Watson & Chandler, 2000: 30
Rhizoecus Künckel d’Herculais, 1878
Rhizoecus sp.—Gómez-Menor (1941: 140)
Saccharicoccus Ferris, 1950
Saccharicoccus sacchari (Cockerell, 1895). Gómez-Menor, 1941: 140; Ben-Dov, 1994: 478; Watson & Chandler, 2000: 30

Family ORTHEZIIDAE
Insignorthezia Kozár, 2004
Insignorthezia insignis (Browne, 1887). Gómez-Menor, 1941: 138
Mixorthezia Morrison, 1925
Mixorthezia dominicana† Vea & Grimaldi, 2012: 769 [Fossil in Dominican amber]
Mixorthezia kozari† Vea & Grimaldi, 2012: 769 [Fossil in Dominican amber]
Praelongorthezia Kozár, 2004
Praelongorthezia praelonga (Douglas, 1891). Abud & Marcano, 1993: 2/93 [Introduced]

Family MARGARODIDAE
Matsucoccus Cockerell, 1909
Matsucoccus subdegeneratus Morrison, 1939a: 17. Foldi, 2004: 166 *DR > TL: Monción, Monte Cristi Prov.
Stigmacoccus Hempel, 1900
Stigmacoccus sp.—Latta et al. (2001: 253)

Order HYMENOPTERA

Taxonomic summary:
Suborder Symphyta
Superfamily Orussoidea

280 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Superfamily Siricoidea
Superfamily Tenthredinoidea
Parasitica
Superfamily Trigonaloidea
Family Trigonalidae
Superfamily Stephanoidea
Family Stephanidae
Superfamily Platygastroidea
Family Platygastridae
Family Scelionidae
Superfamily Ichneumonoidea
Family Braconidae
Family Ichneumonidae
Superfamily Evanioidea
Family Evanidae
Family Gasteruptiidae
Superfamily Diaprioidea
Family Diapriidae
Superfamily Cynipoidea
Family Figitidae
Superfamily Chalcidoidea
Family Agaonidae
Family Aphelinidae
Family Chalcididae
Family Encyrtidae
Family Eucharitidae
Family Eulophidae
Family Eupelmidae
Family Eurytomidae
Family Leucospidae
Family Mymaridae
Family Ormyridae
Family Pteromalidae
Family Signiphoridae
Family Tanaostigmatidae
Family Torymidae
Family Trichogrammatidae
Superfamily Ceraphronoidea
Family Ceraphronidae
Family Megaspilidae
Suborder Apocrita
Aculeata
Superfamily Vespoidea
Family Formicidae
Family Mutiliidae
Family Pompilidae
Family Rhopalosomatidae
Family Scoliidae
Family Tiphiidae
Family Vespidae
Superfamily Chrysidoidea
Family Bethylidae
Family Chrysididae
Family Dryinidae

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 281
 Family Embolemidae
Family Sclerogibbidae
Family Scolebythidae
Superfamily Apoidea
Family Crabronidae
Family Sphecidae
Family Colletidae
Family Andrenidae
Family Halictidae
Family Megachilidae
Family Apidae

Suborder SYMPHYTA
Superfamily ORUSSOIDEA
Family ORUSSIDAE
Ophrynopus Konow, 1897
Ophrynopus peritus† Engel, 2008: 40 [Fossil in Dominican amber]

Superfamily SIRICOIDEA
Family SIRICIDAE
Sirex Linnaeus, 1761
Sirex hispaniola Goulet in Schiff et al., 2012: 142 *DR > TL: La Sal, Res. Ebano Verde
Teredon Norton, 1869
Teredon cubensis (Cresson, 1880). Goulet et al., 2015: 522

Superfamily TENTHRENOIDEA
Family ARGIDAE
Subfamily ERIGLENINAE
Sericoceros Konow, 1905
Sericoceros krugii (Cresson, 1880). Marcano, 1964: 153; Smith, 1992: 64
Subfamily STERICTIPHORINAE
Didymia Lepeletier & Serville, 1828
Didymia davisi† Smith & Poinar, 1992: 118 [Fossil in Dominican amber]
Didymia dominicana† Smith & Poinar, 1992: 118 [Fossil in Dominican amber]
Didymia ebena† Smith & Poinar, 1992: 120 [Fossil in Dominican amber]
Didymia poinari† Smith & Poinar, 1992: 120 [Fossil in Dominican amber]
Didymia protera† Smith & Poinar, 1992: 120 [Fossil in Dominican amber]

PARASITICA
Superfamily TRIGONALOIDEA
Family TRIGONALIDAE
Subfamily TRIGONALINAE
Taeniogonalos Schulz, 1906
Taeniogonalos estebani Genaro, 2017: 2 *DR > TL: Prov. Pedernales, 26 km N Cabo Rojo, 730 m

Superfamily STEPHANOIDEA
Family STEPHANIDAE
Subfamily STEPHANINAE
Megischus Brullé, 1846
Megischus brunneus Cresson, 1865. Genaro, 2012: 1

Superfamily PLATYGASTROIDEA
Family PLATYGASTRIDAE
Subfamily PLATYGASTRINAE

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Acerotella Masner, 1964
Acerotella sp.†—Poinar & Poinar (1999: 210) [Misspelled Acerotellas] [Fossil in Dominican amber]
Allostemma Masner & Huggert, 1989
Allostemma sp.†—Talamas & Buffington (2015: 6) [Fossil in Dominican amber]
Euxestonotus Fouts, 1925
Euxestonotus callisto Buhl, 1998: 242 *DR > TL: Baní, 10 m
Inostemma Haliday, 1833
Inostemma sp.†—Talamas & Buffington (2015: 6) [Fossil in Dominican amber]
Platygaster Latreille, 1809
Platygaster sp.†—Talamas & Buffington (2015: 7) [Fossil in Dominican amber]
Subfamily SCELIOTRACHELINAE
Allotropa Förster, 1856
Allotropa sp.—Serra (2006: 51)
Amitus Haldeman, 1850
Amitus fuscipennis MacGown & Nebeker, 1978. Evans & Serra, 2002: 210
Fidiobia Ashmead, 1894
Fidiobia sp.—Serra (2006: 50)
Fidiobia sp.†—Talamas & Buffington (2015: 8) [Fossil in Dominican amber]
Parabaeus Kieffer, 1910
Parabaeus sp.†—Talamas & Buffington (2015: 9) [Fossil in Dominican amber]

Family SCELIONIDAE
Subfamily SCELIONINAE
Aradophagus Ashmead, 1893
Aradophagus sp.†—García & Masner (1994: 69); Talamas & Buffington (2015: 11) [Fossil in Dominican amber]
Baeus Haliday, 1833
Baeus latrodecti Dozier, 1931: 27. Loiácono & Margaría, 2004: 86 *H > TL: Reared from egg sacs of Lactrodectus mactans
Fab., at Damien
Baeus sp.†—Poinar & Poinar (1999: 210) [Fossil in Dominican amber]
Baryconus Förster, 1856
Baryconus sp.†—Poinar & Poinar (1999: 210) [Misspelled Boryconis] [Fossil in Dominican amber]
Calliscelio Ashmead, 1893
Calliscelio alcoa Chen & Masner in Chen et al., 2017: 22 *DR > TL: Pedernales Prov., 25 km Alcoa Rd., 700 m
Calliscelio armila Chen & Masner in Chen et al., 2017: 30 *H > TL: Sud Dept., Hotte Mts., S. slope of Mt. Formon, 1405 m
Calliscelio glaber Chen & Masner in Chen et al., 2017: 66
Calliscelio pararemigio Chen & Masner in Chen et al., 2017: 91
Calliscelio remigio Chen & Masner in Chen et al., 2017: 98 *DR > TL: Barahona Prov., Baoruco Mts, cloud forest, Remigio
Knoll, 800 m
Calliscelio rubriclavus (Ashmead, 1887). Chen et al., 2017: 100
Calliscelio sp.†—Wu (1997: 209); Talamas & Buffington (2015: 12) [Fossil in Dominican amber]
Calliscelio virga Chen & Johnson in Chen et al., 2017: 128
Calotelea Westwood, 1837
Calotelea sp.†—Poinar & Poinar (1999: 210); Talamas & Buffington (2015: 12) [Fossil in Dominican amber]
Ceratobaeus Ashmead, 1893
Ceratobaeus sp.†—Poinar & Poinar (1999: 210) [Fossil in Dominican amber]
Duta Nixon, 1933
Duta sp.† –Talamas & Buffington (2015: 13) [Fossil in Dominican amber]
Embidobia Ashmead, 1896
Embidobia sp.†—Talamas & Buffington (2015: 16) [Fossil in Dominican amber]
Embioctonus Masner, 1986
Embioctonus sp.†—Talamas & Buffington (2015: 21) [Fossil in Dominican amber]
Gryon Haliday, 1833
Gryon sp.†—Wu (1997: 210); Talamas & Buffington (2015: 21) [Fossil in Dominican amber]
Holoteleia Kieffer, 1908

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 283
Holoteleia sp.†—Poinar & Poinar (1999: 210) [Fossil in Dominican amber]
Idris Förster, 1856
Idris sp.†—Wu (1997: 209); Talamas & Buffington (2015: 23) [Fossil in Dominican amber]
Leptoteleia Kieffer, 1908
Leptoteleia arndti Dozier in Arndt & Dozier, 1931: 7 *H > TL: Fond-des-Negres
Leptoteleia ferdinandi Masner, 1978: 374 *DR > TL: Colonia, Cordillera Central, 1000 m
Leptoteleia josephi Masner, 1978: 376 *DR > TL: Boca Chica, 10 m
Leptoteleia lubomiri Masner, 1978: 374 *DR > TL: Colonia, Cordillera Central, 1000 m
Leptoteleia monicae Masner, 1978: 372 *DR > TL: Colonia, Cordillera Central, 1000 m
Leptoteleia radeki Masner, 1978: 375 *DR > TL: Colonia, Cordillera Central, 1000 m
Leptoteleia sp.†—Talamas & Buffington (2015: 25) [Fossil in Dominican amber]
Leptoteleia stani Masner, 1978: 375 *DR > TL: Colonia, Cordillera Central, 1000 m
Macroteleia Westwood, 1835
Macroteleia carinata Ashmead, 1894. Muesebeck, 1977: 10
Macroteleia rufithorax Muesebeck, 1977: 37 *DR > TL: Cambita, Trujillo Prov. [San Cristóbal]
Macroteleia rugosa (Provancher, 1881). Muesebeck, 1977: 39
Macroteleia sp.†—Talamas & Buffington (2015: 25) [Fossil in Dominican amber]
Odontacolus Kieffer, 1910
Odontacolus sp.†—Talamas & Buffington (2015: 25) [Fossil in Dominican amber]
Oethecoctonus Ashmead, 1900
Oethecoctonus sp.†—Wu (1997: 209) [Fossil in Dominican amber]
Opistacantha Ashmead, 1893
Opistacantha sp.†—Talamas & Buffington (2015: 25) [Fossil in Dominican amber]
Paridris Kieffer, 1908
Paridris aenea (Ashmead, 1894). Talamas et al., 2012: 38
Paridris isabelicae Talamas et al., 2012: 60
Paridris yumai† Talamas in Talamas & Buffington, 2015: 25 [Fossil in Dominican amber]
Plaumannion Masner & Johnson, 2007
Plaumannion fistulosum† Talamas in Talamas & Buffington, 2015: 35 [Fossil in Dominican amber]
Probaryconus Kieffer, 1908
Probaryconus sp.†—Wu (1997: 210); Talamas & Buffington (2015: 38) [Fossil in Dominican amber]
Psilanteris Kieffer, 1916
Psilanteris sp.†—Talamas & Buffington (2015: 39) [Fossil in Dominican amber]
Scelio Latreille, 1804
Scelio sp.†—Poinar & Poinar (1999: 210) [Misspelled Scelia] [Fossil in Dominican amber]
Scelio sp.—Navarro Morales (2014: 143)
Spiniteleia Masner, 1980
Spiniteleia sp.†—Talamas & Buffington (2015: 40) [Fossil in Dominican amber]
Synoditella Muesebeck, 1972
Synoditella bisulcata (Kieffer, 1904). Perez-Gelabert & Bastardo, 2004: 932; Bastardo & Perez-Gelabert, 2013: 92
Thoronidea Masner & Huggert, 1979
Thoronidea taino Masner & Huggert, 1979: 914 *DR > TL: Cordillera Central, 600 m, 15 km SW Piedra Blanca, (Prov. La
Vega, road between Piedra Blanca and Rancho Arriba)
Triteleia Kieffer, 1906
Triteleia sp.†—Talamas & Buffington (2015: 44) [Fossil in Dominican amber]
Subfamily TELENOMINAE
Telenomus Haliday, 1833
Telenomus connectans Ashmead, 1895. Johnson, 1990: 306
Telenomus minutissimus Ashmead, 1895. Andrews, 1988: 638
Telenomus monilicornis Ashmead, 1894. Wolcott, 1927: 345; Johnson, 1990: 307
Telenomus remus Nixon, 1937. Cave, 2000: 21N; Molina-Ochoa et al., 2003: 274
Telenomus sphingis (Ashmead, 1887). Agudelo, 1980a: 233; Agudelo, 1980b: 341
Telenomus xanthosoma Johnson, 1980: 781 *DR > TL: Colonia, 15 km W San Cristóbal, 1000 m
Telenomus sp.†—Talamas & Buffington (2015: 9) [Fossil in Dominican amber]

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Trissolcus Ashmead, 1893
Trissolcus brochymenae (Ashmead, 1881). Johnson, 1987: 298
Trissolcus elimatus Johnson, 1987: 300
Subfamily INCERTAE SEDIS
Janzenella Masner & Johnson, 2007
Janzenella innupta† Masner & Johnson, 2007: 2 [Fossil in Dominican amber. Specimen presumably conspecific with extant
species in Costa Rica]

Superfamily ICHNEUMONOIDEA
Family BRACONIDAE
Subfamily ACAMPSOHELCONINAE
Urosigalphus Ashmead, 1889
Urosigalphus (Microurosigalphus) neoschwarzi Gibson, 1972: 141
Subfamily AGATHIDINAE
Alabagrus Enderlein, 1920
Alabagrus stigma (Brullé, 1846). Sharkey, 1988: 401
=Microdus stigmaterus Cresson, 1865. Myers, 1935: 221; Santoro, 1960: 1453
Alabagrus varipes (Cresson, 1865). Sharkey, 1988: 409
Crassomicrodus Ashmead, 1900
Crassomicrodus nigriceps (Cresson, 1872). Figueroa et al., 2011: 57
Cremnops Förster, 1862
Cremnops guanicanus Wolcott, 1924. Berta de Fernández, 1998: 244; Tucker et al., 2015: 53
Subfamily ALYSIINAE
Aphaereta Förster, 1862
Aphaereta subtropicalis Wharton, 1977: 800 *DR > TL: Prov. La Vega: La Guardarraya
Gnathopleura Fischer, 1975
Gnathopleura unicarinata Fischer, 2006: 618
Subfamily APHIDIINAE
Lysiphlebus Förster, 1862
Lysiphlebus testaceipes (Cresson, 1880). Wolcott, 1927: 340
Subfamily BRACONINAE
Bracon Fabricius, 1805
Bracon mellitor Say, 1836. Annand, 1942: 50 [Purposely introduced]
Bracon sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Habrobracon Johnson, 1895
Habrobracon sp.—Segura et al. (2005: unpaginated)
Subfamily CHELONINAE
Chelonus Jurine, 1807
Chelonus insularis (Cresson, 1865). Molina-Ochoa et al., 2003: 264
Phanerotoma Wesmael, 1838
Phanerotoma sp.†—Poinar (1992: 193) [Fossil in Dominican amber]
Subfamily DORYCTINAE
Aivalykus Nixon, 1938
Aivalykus dominicanus† Zuparko & Poinar, 1997: 744 [Fossil in Dominican amber]
Doryctopambolus Nunes & Zaldívar-Riverón in Nunes et al., 2012
Doryctopambolus clebschi Zaldívar-Riverón & Martínez in Nunes et al., 2012: 61 *DR > TL: A. Bermúdez NP, La Ciénaga,
Los Tablones
Doryctopambolus dominicanus Zaldívar-Riverón & Martínez in Nunes et al., 2012: 62 *DR > TL: Punta Cana
Holcobracon Cameron, 1905
Holcobracon limatus (Cresson, 1865). Marsh, 1988: 464
=Liobracon manni Brues, 1919: 69. Marsh, 1988: 464
=Liobracon bruesi Roman, 1924. Marsh, 1988: 464
Liobracon Szépligeti, 1901
Liobracon cressoni (Dalla Torre, 1898). Brues, 1919: 71

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 285
Odontobracon Cameron, 1887
Odontobracon armatus (Cresson, 1865). Marsh, 1988: 447
Subfamily GNAMPTODONTINAE
Pseudognaptodon Fischer, 1965
Pseudognaptodon whartoni Williams, 2004: 198
Subfamily MACROCENTRINAE
Macrocentrus Curtis, 1833
Macrocentrus sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Subfamily MASONINAE
Anoblepsis† Engel & Bennett, 2008
Anoblepsis konokeraia† Engel & Bennett, 2008: 369 [Fossil in Dominican amber]
Masona van Achterberg, 1995
Masona pyriceps† van Achterberg, 2001: 393 [Fossil in Dominican amber]
Subfamily MICROGASTRINAE
Apanteles Förster, 1862
Apanteles diatraeae Muesebeck, 1920. Shenefelt, 1972: 489
Apanteles disputabilis (Ashmead, 1900). Shenefelt, 1972: 494
Apanteles sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Choeras Mason, 1981
Choeras sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Cotesia Cameron, 1891
Cotesia americana (Lepeletier, 1825). Agudelo-Silva, 1980: 233
Deuterixys Mason, 1981
Deuterixys bennetti Whitfield, 1985. Whitfield & Oltra Moscardó, 2004: 136
Deuterixys erythrocephala Whitfield & Oltra Moscardó, 2004: 140
Diolcogaster Ashmead, 1900
Diolcogaster sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Glyptapanteles Ashmead, 1904
Glyptapanteles sp.—Serra (1992d: Tab. A13)
Hypomicrogaster Ashmead, 1898
Hypomicrogaster largus Valerio in Valerio & Whitfield, 2015: 66
Pseudapanteles Ashmead, 1898
Pseudapanteles dignus (Muesebeck, 1938). Serra, 1992d: Tab. A13
Subfamily OPIINAE
Doryctobracon Enderlein, 1920
Doryctobracon areolatus (Szépligeti, 1911). Serra, 2005: 114; Serra et al., 2011: 809 [Purposely introduced]
Opius Wesmael, 1835
Opius (Opius) hirtus Fischer, 1963: 376. Fisher, 1977: 865; Wharton & Marsh, 1978: 163; Wharton, 1983: 328
Opius (Pleurosema) utinanus Fischer, 2001: 312 *DR > TL: Puerto Plata, Sosua, S of El Batey
Opius (Rhogadopsis) dominicanus Fischer, 1963. Fischer, 2001: 311
Opius (Rhogadopsis) lateroareatus Fischer, 2001: 308 *DR > TL: Puerto Plata, Sosua, S of El Batey
Opius (Rhogadopsis) puertoplatanus Fischer, 2001: 311 *DR > TL: Puerto Plata, Sosua, El Batey
Opius rufoflavus (Fischer, 2001: 317) *DR > TL: Puerto Plata, Sosua, El Batey
Utetes Förster, 1862
Utetes anastrephae (Viereck, 1913). Serra, 2006: 50
Utetes sp.—Ortíz (2005: 13)
Subfamily ORGILINAE
Stantonia Ashmead, 1904
Stantonia pallida (Ashmead, 1894). Braet & Quicke, 2004: 1565
Subfamily ROGADINAE
Conobregma van Achterberg, 1995
Conobregma brevistigmus van Achterberg, 1995: 37 *DR > TL: Paraiso Valley, 700 m
Conobregma masneri van Achterberg, 1995: 39 *DR > TL: La Cumbre, 600 m
Yelicones Cameron, 1887

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Yelicones delicatus (Cresson, 1872). Quicke et al., 1996: 57
Yelicones nigrigaster Quicke et al., 1996: 46 *DR > TL: La Cumbre, 600 m

Family ICHNEUMONIDAE
Subfamily ANOMALINAE
Anomalon Jurine, 1807
Anomalon ejuncidum Say, 1836. Townes & Townes, 1966: 197
Subfamily CREMASTINAE
Temelucha Förster, 1869
Temelucha sp.—Serra (1992d: Tab. A13)
Subfamily CRYPTINAE
Messatoporus Cushman, 1929
Messatoporus dominicanus Santos in Santos & Aguiar, 2013: 115 *DR > TL: Puerto Plata, La Cumbre.
Messatoporus lunatus Santos in Santos & Aguiar, 2013: 144 *DR > TL: La Altagracia, Nisibon, near cacao plantation.
Nesolinoceras Ashmead, 1906
Nesolinoceras laluzbrillante Santos, 2016: 14 *DR > TL: Punta Cana Ecological Foundation, La Altagracia Prov.
Subfamily EPHIALTINAE
Neotheronia Krieger, 1899
Neotheronia nigrolineata tolteca (Cresson, 1874). Townes & Townes, 1966: 35
Neotheronia schoenachii (Dalla Torre, 1902). Townes & Townes, 1966: 36
Pimpla Fabricius, 1805
Pimpla marginella Brullé, 1846. Townes & Townes, 1966: 26
Tromatobia Förster, 1868
Tromatobia notator (Fabricius, 1804). Townes & Townes, 1966: 15
Subfamily GELINAE
Acroricnus Ratzeburg, 1852
Acroricnus tricolor Mitchell, 1950. Townes & Townes, 1966: 124
Chirotica Förster, 1868
Chirotica phrixonota Townes, 1983: 85 *DR > TL: La Cumbre, Cordillera Septentrional, 300 m
Compsocryptus Ashmead, 1900
Compsocryptus stangei Porter, 1989: 666 *DR > TL: 9 km N Villa Elisa, Monte Cristi Prov.
Dichrogaster Doumerc, 1855
Dichrogaster zonata Townes, 1983: 97 *DR > TL: La Cumbre, 300 m
Lymeon Förster, 1868
Lymeon fuscipennis (Brullé, 1846). Ashmead, 1900: 351
Polycyrtus Spinola, 1840
Polycyrtus lituratus (Brullé, 1846). Myers, 1931: 267; Townes & Townes, 1966: 111
Polycyrtus trilineatus (Brullé, 1846). Ashmead, 1900: 351
Subfamily ICHNEUMONINAE
Tricholabus Thomson, 1894
Tricholabus serricornis (Cresson, 1865). Townes & Townes, 1966: 254
Subfamily OPHIONINAE
Enicospilus Stephens, 1829
Enicospilus cressoni Hooker, 1912. Townes & Townes, 1966: 176; Gauld, 1988: 198
Enicospilus dajaboni Gauld, 1988: 186 *DR > TL: Dajabón Prov., 13 km S. of Loma de Cabrera, 400 m
Enicospilus dispilus (Szépligeti, 1906). Gauld, 1988: 296
Enicospilus fernaldi Hooker, 1912: 63. Townes & Townes, 1966: 177; Gauld, 1988: 228
Enicospilus flavoscutellatus (Brullé, 1846). Townes & Townes, 1966: 177; Gauld, 1988: 272
=Henicospilus trimaculatus Morley, 1912. Townes & Townes, 1966: 178; Gauld, 1988: 272
Enicospilus flavostigma Hooker, 1912. González-Moreno & Bordera, 2012: 31
Enicospilus flavus (Fabricius, 1775). Townes & Townes, 1966: 178; Gauld, 1988: 230
Enicospilus gallegosi Gauld, 1988: 285
Enicospilus glabratus (Say, 1836). Gauld, 1988: 128; González-Moreno & Bordera, 2012: 31
Enicospilus guatemalensis (Cameron, 1886). Gauld, 1988: 206; González-Moreno & Bordera, 2012: 31

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 287
Enicospilus howdenorum Gauld, 1988: 282
Enicospilus masneri Gauld, 1988: 195 *DR > TL: La Cumbre, P. Plata Prov., 300 m
Enicospilus monticola (Cameron, 1886). Townes & Townes, 1966: 182
Enicospilus neotropicus Hooker, 1912. Townes & Townes, 1966: 182; Gauld, 1988: 172
Enicospilus purgatus (Say, 1836). Gauld, 1988: 175
Enicospilus sondrae Gauld, 1988: 283
Enicospilus trilineatus (Brullé, 1846). Gauld, 1988: 189
=Enicospilus appendiculatus Felt, 1902. Gauld, 1988: 189
=Henicospilus brevinervis Morley, 1912. Gauld, 1988: 189
Ophion Fabricius, 1798
Ophion flavidus Brullé, 1846. Gauld, 1988: 37; González-Moreno & Bordera, 2012: 31
Ophion sp.—Townes & Townes (1966: 170); Abud (1980: 8/80)
Thyreodon Brullé, 1846
Thyreodon atriventris armstrongi (Hooker, 1912: 105). Townes & Townes, 1966: 185 *DR > TL: San Francisco Mountains,
Santo Domingo
Thyreodon ornatipennis Morley, 1912. Townes & Townes, 1966: 190
Subfamily CAMPOPLEGINAE
Diadegma Förster, 1868
Diadegma insulare (Cresson, 1865). Santos et al., 2000: 77
Subfamily CREMASTINAE
Eiphosoma Cresson, 1865
Eiphosoma haitiense Brues, 1917: 453. Townes & Townes, 1966: 163 *H > TL: Cape Haitien

Superfamily EVANIOIDEA
Family EVANIIDAE
Evania Fabricius, 1775
Evania appendigaster (Linnaeus, 1758). Wolcott, 1927: 341; Deans, 2005: 27 [Introduced]
Hyptia Illiger, 1807
Hyptia manni (Brues, 1916: 719). Deans, 2005: 62 *H > TL: Manneville
Hyptia servillei (Guérin-Méneville, 1843). Ashmead, 1901: 349; Deans, 2005: 66
Hyptia weithi Ashmead, 1901: 302. Deans, 2005: 69
Semaeomyia Bradley, 1908
Semaeomyia dominicana† (Nel, et al., 2002: 217). Deans, 2005: 164 [Fossil in Dominican amber]

Family GASTERUPTIIDAE
Gasteruption Latreille, 1796
Gasteruption masneri Cruz Macedo, 2011: 40 *DR > TL: Pedernales

Superfamily DIAPRIOIDEA
Family DIAPRIIDAE
Subfamily DIAPRIINAE
Basalys Westwood, 1833
Basalys sp.†—Wu (1997: 207) [Fossil in Dominican amber]
Monelata Förster, 1856
Monelata sp.†—Poinar & Poinar (1999: 209) [As Corynopria] [Fossil in Dominican amber]
Peckidium Masner & García, 2002
Peckidium enigmaticum Masner & García, 2002: 119
Trichopria Ashmead, 1893
Trichopria sp.†—Wu (1997: 207) [Fossil in Dominican amber]

Superfamily CYNIPOIDEA
Family FIGITIDAE
Subfamily ANACHARITINAE
Acanthaegilips Ashmead, 1897

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Acanthaegilips alienus Ros-Farre & Pujade-Villar in Ros-Farre et al., 2003: 16 *DR > TL: Prov. Pedernales, Sra. Bahoruco
(1350 m)
Xyalaspis Hartig, 1843
Xyalaspis dominicana Mata-Casanova & Pujade-Villar, 2015: 102 *DR > TL: 28 km NW La Descubierta, Sierra de Neiba
Subfamily ASPICERINAE
Prosaspicera Kieffer, 1907
Prosaspicera optivus Quinlan, 1979. Ross-Farre & Pujade-Villar, 2006: 39
Subfamily EUCOILINAE
Cothonaspis Hartig, 1840
Cothonaspis sp.—Serra (1992d: Tab. A13)
Disorygma Forster, 1869
Disorygma pacifica (Yoshimoto, 1962). Serra, 1992d: Tab. A13
Ganaspidium Weld, 1952
Ganaspidium utilis Beardsley, 1988. Serra, 1992d: Tab. A13
Subfamily FIGITINAE
Neralsia Cameron, 1883
Neralsia gibbosa Jiménez & Pujade-Villar, 2008: 50

Superfamily CHALCIDOIDEA
Family AGAONIDAE
Subfamily AGAONINAE
Pegoscapus Cameron, 1906
Pegoscapus peritus† Peñalver & Engel in Peñalver et al., 2006: 11 [Fossil in Dominican amber]
Tetrapus Mayr, 1885
Tetrapus apopnus† Peñalver & Engel in Peñalver et al., 2006: 5 [Fossil in Dominican amber]
Tetrapus delclosi† Peñalver & Engel in Peñalver et al., 2006: 3 [Fossil in Dominican amber]
Subfamily SYCOPHAGINAE
Idarnes Walker, 1843
Idarnes thanatos† Farache, Rasplus, Pereira & Compton, 2016: 4 [Fossil in Dominican amber]

Family APHELINIDAE
Subfamily APHELININAE
Aphelinus Dalman, 1820
Aphelinus sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Aphytis Howard, 1900
Aphytis chrysomphali (Mercet, 1912). Wolcott, 1927: 208; Dozier, 1933: 90; Rosen & DeBach, 1979: 593; De Santis, 1979:
311; De Santis, 1981: 24
Aphytis diaspidis (Howard, 1881). De Santis, 1989: 57
=Aphytis fuscipennis (Howard, 1880). Dozier, 1933: 89
Aphytis hispanicus (Mercet, 1912). De Santis, 1979: 312
Aphytis holoxanthus DeBach, 1960. Rosen & DeBach, 1979: 548
Aphytis limonus (Rust, 1915). Dozier, 1933: 89
Aphytis maculicornis (Masi, 1911). Dozier, 1933: 90
Aphytis maculatipennis (Dozier, 1933: 88). Compere, 1936: 312; De Santis, 1979: 317 *H > TL: Reared from a diaspine scale,
Aspidiotus (Diaspidiotus) sp., incrusting the trunk and branches of young mahogany, Swietenia mahagoni, in nursery at Cote
Plage
Marietta Motschulsky, 1863
Marietta dozieri Hayat, 1986: 9 *H > TL: Pétionville, ex Coccus mangifera on mango
=Marietta buscki (Howard, 1907). Dozier, 1933: 87; Compere, 1936: 286 [Misidentification in part]
Subfamily CALESINAE
Cales Howard, 1907
Cales noacki Howard, 1907. Dozier, 1933: 98; De Santis, 1979: 340; Evans & Serra, 2002: 204
Subfamily COCCOPHAGINAE
Coccobius Ratzeburg, 1852

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Coccobius uvae (Dozier, 1933: 94) *H > TL: Reared from Aspidiotus lataniae on grape vine at St. Marc
Coccophagoides Girault, 1915
Coccophagoides pulchellus (Dozier, 1937: 127). De Santis, 1979: 337
Coccophagus Westwood, 1833
Coccophagus asterolecanii (Dozier, 1932g: 99). Compere, 1936: 290; De Santis, 1979: 328 *H > TL: Reared from
Asterolecanium aureum (Boisd.) on wild Annona sp. on Morne-a-Cabrits
Coccophagus ceroplastae (Howard, 1895). De Santis, 1979: 327; De Santis, 1989: 59
Coccophagus cubaensis Compere, 1931. De Santis, 1979: 322
Coccophagus hispaniolae (Dozier, 1932g: 98). Compere, 1936: 292; De Santis, 1979: 328 *H > TL: Reared from a giant wax
scale, Ceroplastes giganteus Dozier, on branches of wild fig tree, Ficus rubricosta Warb. at Source Cazeau
Coccophagus mangiferae (Dozier, 1932: 95). Compere, 1936: 286; De Santis, 1979: 327
Coccophagus scutellaris (Dalman, 1825). De Santis, 1989: 60
Coccophagus sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]
Encarsia Förster, 1878
Encarsia berlesei (Howard, 1906). De Santis, 1989: 61
Encarsia catherinae (Dozier, 1933: 92). Evans & Serra, 2002: 205 *H > TL: Reared from an undescribed whitefly,
Aleuroplatus sp., on Haitian oak, Catalpa longissima, at Damien
Encarsia citrina (Craw, 1891). Dozier, 1933: 89; De Santis, 1979: 330; De Santis, 1989: 61
Encarsia cubensis Gahan, 1931. Dozier, 1933: 92; De Santis, 1979: 329; Evans & Serra, 2002: 205
Encarsia diaspidicola (Silvestri, 1909). Dozier, 1933: 93; De Santis, 1979: 334
Encarsia dominicana Evans in Evans & Serra, 2002: 205
Encarsia formosa (Gahan, 1924). Evans & Serra, 2002: 206 [Introduced]
Encarsia haitiensis Dozier, 1932a: 118. De Santis, 1979: 328; Evans & Serra, 2002: 206
Encarsia hispida De Santis, 1948. Evans & Serra, 2002: 206
=Prospaltella brasiliensis (Hempel, 1904). Dozier, 1932: 121; De Santis, 1979: 333
Encarsia lanceolata Evans & Polaszek, 1997. Evans & Serra, 2002: 206
Encarsia lounsburyi (Berlese & Paoli, 1916). Dozier, 1933: 89; De Santis, 1979: 331
Encarsia meritoria Gahan, 1927. Serra et al., 1996: 654
Encarsia nigricephala Dozier, 1937. Evans & Serra, 2002: 206
Encarsia pergandiella Howard, 1907. Serra et al., 1996: 654
Encarsia perplexa Huang & Polaszek, 1997. Evans & Serra, 2002: 206
Encarsia portoricensis Howard, 1907. Evans & Serra, 2002: 207
Encarsia protransvena Viggiani, 1985. Serra et al., 2011: 148 [Introduced]
Encarsia sophia (Girault & Dodd, 1915). Evans & Serra, 2002: 207
Encarsia tabacivora Viggiani, 1985. Evans & Serra, 2002: 207
Encarsia telemachusi Evans in Evans & Serra, 2002: 207 *H > TL: Source, Cazeau
Encarsia variegata Howard, 1908. Dozier, 1933: 92; De Santis, 1979: 330; Evans & Serra, 2002: 209
Pteroptrix Westwood, 1833
Pteroptrix albifemur (Girault, 1915) [By synonymy of species below]
=Pteroptrix imitatrix (Fullaway, 1989). De Santis, 1989: 66
Pteroptrix howardi (Dozier, 1937). De Santis, 1979: 340
Pteroptrix orientalis (Silvestri, 1909). De Santis, 1989: 66
Subfamily ERETMOCERINAE
Eretmocerus Haldeman, 1850
Eretmocerus pallidus Dozier, 1932f: 116. Compere, 1936: 320; De Santis, 1979: 337; Evans & Serra, 2002: 209
Eretmocerus paulistus Hempel, 1904. Dozier, 1932f: 117; Compere, 1936: 321; De Santis, 1979: 337
Eretmocerus portoricensis Dozier, 1932. De Santis, 1989: 66; Evans & Serra, 2002: 209
Eretmocerus serius Silvestri, 1927. De Santis, 1979: 338; Evans & Serra, 2002: 209
Eretmocerus spp.—Evans & Serra (2002: 209)

Family CHALCIDIDAE
Subfamily CHALCIDINAE
Brachymeria Westwood, 1829
Brachymeria annulata (Fabricius, 1793). De Santis, 1979: 57

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Brachymeria flavipes (Fabricius, 1793) [By synonymy of species below]
=Brachymeria robusta (Cresson, 1865). De Santis, 1989: 13
Brachymeria incerta (Cresson, 1865). De Santis, 1979: 58; De Santis, 1989: 13
Brachymeria mnestor (Walker, 1841). Bouček, 1992: 90
=Chalcis pendator Walker, 1862. Bouček, 1992: 90
Brachymeria ovata (Say, 1824). De Santis, 1979: 59
Brachymeria podagrica (Fabricius, 1787). De Santis, 1979: 61
Brachymeria sp.†—Poinar (1992: 195) [Fossil in Dominican amber]
Chalcis Fabricius, 1787
Chalcis arapha Burks, 1939: 1. De Santis, 1979: 39 *DR > TL: Sánchez
Conura Spinola, 1837
Conura eubule (Cresson, 1865). De Santis, 1979: 43
Conura femorata (Fabricius, 1775). De Santis, 1979: 44
Conura hirtifemora (Ashmead, 1885). De Santis, 1979: 46; Santos et al., 2000: 77
Conura immaculata (Cresson, 1865). De Santis, 1989: 12
Conura petioliventris (Cameron, 1884). Santos et al., 2000: 77
Conura pseudofulvovariegata (Becker, 1989). Santos et al., 2000: 77
Conura side (Walker, 1843) [By synonymy of species below]
=Spilochalcis flavopicta (Cresson, 1865). De Santis, 1989: 11
Conura sp.†—Poinar & Poinar (1999: 208) [As Spilochalcis] [Fossil in Dominican amber]
Subfamily DIRHININAE
Dirhinus Dalman, 1818
Dirhinus (Dirhinus) giffardi Silvestri, 1913. De Santis, 1979: 68 [Purposely introduced]
Subfamily HALTICHELLINAE
Psilochalcis Kieffer, 1905
Psilochalcis sp.—Bouček (1992: 70)
Psilochalcis sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]

Family ENCYRTIDAE
Subfamily ENCYRTINAE
Acerophagus F. Smith, 1880
Acerophagus nubilipennis Dozier, 1926. De Santis, 1989: 40
Acerophagus papayae Noyes & Schauff, 2003. Kairo et al. (2003: 18N); Serra, 2006: 50 [Purposely introduced]
Adelencyrtus Ashmead, 1900
Adelencyrtus moderatus (Howard, 1897). Panis, 2006: 288
Ageniaspis Dahlbom, 1857
Ageniaspis sp.—Navarro Morales (2014: 143)
Anicetus Howard, 1896
Anicetus annulatus Timberlake, 1919. Trjapitzin et al., 2004: 48; Trjapitzin & Cancino, 2009: 255
Apsilophrys De Santis, 1964
Apsilophrys sp.†—Poinar & Poinar (1999: 209) [Fossil in Dominican amber]
Carabunia Waterston, 1928
Carabunia myersi Waterston, 1928. Myers, 1931: 276; De Santis, 1979: 239
Comperiella Howard, 1906
Comperiella bifasciata Howard, 1906. Trjapitzin et al., 2004: 68
Copidosoma Ratzeburg, 1844
Copidosoma archeodominica† Zuparko & Trjapitzin, 2013: 230 [Fossil in Dominican amber]
=Copidosoma sp.†—Poinar & Poinar (1999: 209) [Fossil in Dominican amber]
Copidosoma sp.—Serra (1992d: Tab. A13)
Homalopoda Howard, 1894
Homalopoda cristata Howard, 1894. De Santis, 1979: 230
Homalotylus Mayr, 1875
Homalotylus terminalis (Say, 1829). Myers, 1931: 276; De Santis, 1979: 206
Metaphycus Mercet, 1917

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 291
Metaphycus eruptor (Howard, 1881) [By synonymy of species below]
=Coccidoctonus ceroplastae Dozier, 1935: 184. De Santis, 1989: 185; Trjapitzin et al., 2004: 95
Metaphycus portoricensis (Dozier, 1926). Dozier, 1933: 96
Pseudaphycus Clausen, 1915
Pseudaphycus angelicus (Howard, 1898). Kairo et al. (2003: 18N)
Pseudaphycus utilis Timberlake, 1923. De Santis, 1989: 41; Trjapitzin et al., 2004: 117
Pseudectroma Girault, 1915
Pseudectroma europaeum (Mercet, 1921). Trjapitzin et al., 2004: 118
Pseudhomalopoda Girault, 1915
Pseudhomalopoda prima Girault, 1915. De Santis, 1979: 230; Trjapitzin et al., 2004: 119
Pseudleptomastix Girault, 1915
Pseudleptomastix sp.—Trjapitsin et al. (2004: 120)
Psyllaephagus Ashmead, 1900
Psyllaephagus yaseeni Noyes, 1990: 39
Subfamily TETRACNEMINAE
Anagyrus Howard, 1896
Anagyrus ananatis Gahan, 1949. De Santis, 1979: 185
Anagyrus californicus (Compere, 1947). Kairo et al. (2003: 18N) [Purposely introduced]
Anagyrus coccidivorus Dozier, 1932c: 8. De Santis, 1979: 185; De Santis, 1989: 30
Anagyrus greeni Howard, 1896. De Santis, 1989: 31
Anagyrus kamali Moursi, 1948. Serra, 2006: 50 [Purposely introduced]
Anagyrus loecki Noyes & Menezes, 2000. Kairo et al. (2003: 18N); Serra, 2006: 50 [Purposely introduced]
Gyranusoidea Compere, 1947
Gyranusoidea indica Shafee et al., 1975. Serra et al., 2003: 9; Serra 2005: 112 [Purposely introduced]
Hambletonia Compere, 1936
Hambletonia pseudococcina Compere, 1936. De Santis, 1989: 35
Leptomastidea Mercet, 1917
Leptomastidea abnormis (Girault, 1915). Gómez-Menor, 1941: 126; De Santis, 1979: 191 [Purposely introduced]
Leptomastix Förster, 1856
Leptomastix dactylopii Howard, 1885. De Santis, 1979: 184

Family EUCHARITIDAE
Subfamily EUCHARITINAE
Kapala Cameron, 1884
Kapala terminalis Ashmead, 1892. De Santis, 1979: 104
Subfamily ORASEMINAE
Orasema Cameron, 1884
Orasema costaricensis Wheeler & Wheeler, 1937. Burks et al., 2018: 26
Orasema evansi Burks, Heraty & Dominguez, 2018: 33
Orasema minutissima Howard, 1894. Burks et al., 2018: 62

Family EULOPHIDAE
Subfamily ENTEDONINAE
Achrysocharoides Girault, 1913
Achrysocharoides sp.†—Gumovsky (2001: 136) [Fossil in Dominican amber]
Apleurotropis Girault, 1913
Apleurotropis assis Hansson, 2010: 45
Ametallon Ashmead, 1904
Ametallon dominicanense Hansson, 2004: 23 *DR > TL: Barahona, Sierra de Bahoruco, Alcoa Road, km 25
Ceranisus Walker, 1841
Ceranisus menes (Walker, 1839). De Santis, 1979: 284
Ceranisus sp. “A”—Triapitsyn & Morse (2005: 79)
Chrysocharis Förster, 1856
Chrysocharis oscinidis Ashmead, 1888 [By synonymy of species below]

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 =Chrysocharis parksi Crawford, 1912. De Santis, 1989: 51
Chrysonotomyia Ashmead, 1904
Chrysonotomyia dominicana† Gumovsky, 2001: 134 [Fossil in Dominican amber]
Closterocerus Westwood, 1833
Closterocerus lividus (Ashmead, 1894). De Santis, 1989: 51
Diglyphus Walker, 1844
Diglyphus ?begini (Ashmead, 1904). Serra (1992: Tab. A13)
Diglyphus isaea (Walker, 1838). Serra (1992: Tab. A13)
Diglyphus sp.—Serra (1992: Tab. A13)
Emersonella Girault, 1916
Emersonella lemae Girault, 1916. Hansson, 2002: 44
Emersonella rotunda (Ashmead, 1894). Hansson, 2002: 51
Entedononecremnus Girault, 1915
Entedononecremnus fulgens Hansson & LaSalle, 2003: 765 *DR > TL: Duarte, 20 km NE San Francisco de Macorís, Loma
Quita Espuela, 300 m
Eprhopalotus Girault, 1916
Eprhopalotus hansoni (Schauff, 1991). Hansson, 2004: 142
Goetheana Girault, 1920
Goetheana shakespearei Girault, 1920. Triapitzyn, 2005: 266
Horismenus Walker, 1843
Horismenus abagus Hansson, 2009: 72
Horismenus alas Hansson, 2009: 83
Horismenus albiscapus Hansson, 2009: 388
Horismenus alnus Hansson, 2009: 85
Horismenus apantelivorus Crawford, 1911. Hansson, 2009: 97
Horismenus chydaeus Hansson, 2009: 391
Horismenus citrus Hansson, 2009: 151
Horismenus bennetti Schauff, 1987: 39. De Santis, 1989: 50
Horismenus eudami (Girault, 1918). De Santis, 1989: 50
Horismenus graciliventris (Girault, 1916). De Santis, 1989: 50
Horismenus malus Hansson, 2009: 247
Inti Hansson, 2010
Inti levis Hansson, 2010: 59
Neochrysocharis Kurdjumov, 1912
Neochrysocharis sp.—Serra (1992d: Tab. A13)
Neopomphale LaSalle & Schauff, 1994
Neopomphale aleurothrixi (Dozier, 1932: 120). De Santis, 1989: 52; Evans & Serra, 2002: 209; Hansson & LaSalle, 2003: 720
Omphale Haliday, 1830
Omphale divina (Girault, 1917). Hansson, 2004: 180
Omphale huggerti (Hansson, 1988). Hansson, 2004: 198
Omphale safranota Hansson, 2004: 186
Pediobius Walker, 1830
Pediobius dolichops Hansson, 2002: 154
Pediobius cajanus Taveras & Hansson, 2015: 45 *DR > TL: San Juan de la Maguana
Tropicharis Hansson, 1998
Tropicharis cecivora Hansson, 1998: 691
Subfamily EUDERINAE
Euderus Walker, 1830
Euderus sp.—Serra (1992d: Tab. A13)
Euderus sp.†—Poinar & Poinar (1999: 209) [Fossil in Dominican amber]
Subfamily EULOPHINAE
Cirrospilus Westwood, 1832
Cirrospilus sp.—Evans (1999: 449)
Elasmus Westwood, 1833

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Elasmus sp.—Macarrulla (2000: 111)
Euplectrus Westwood, 1832
Euplectrus cinctiventris Ferrière, 1941. Zhu & Huang, 2003: 148
Euplectrus flavipes (Fonscolombe, 1832). Zhu & Huang, 2003: 150
Euplectrus leucostomus Rohwer, 1921. Zhu & Huang, 2003: 154
Grotiusomyia Girault, 1917
Grotiusomyia nigricans (Howard, 1894). De Santis, 1989: 49
Zagrammosoma Ashmead, 1904
Zagrammosoma multilineatum (Ashmead, 1888). De Santis, 1989: 50
Subfamily TETRASTICHINAE
Aprostocetus Westwood, 1833
Aprostocetus gala (Walker, 1847). Etienne et al., 1993: 105
Aprostocetus vaquitarum (Wolcott, 1924). De Santis, 1989: 53
Baryscapus Förster, 1856
Baryscapus fennahi (Schauff, 1987). Etienne et al., 1993: 105
Ceratoneura Ashmead, 1894
Ceratoneura petiolata Ashmead, 1894. Ikeda, 2001: 40
Eriastichus LaSalle, 1994
Eriastichus masneri LaSalle, 1994: 207 *DR > TL: Barahona, Sierra de Bahoruco, Loma Remigio, cloud forest, 800 m
Eriastichus nakos LaSalle, 1994: 208
Melittobia Westwood, 1848
Melittobia australica Girault, 1912. Gonzalez et al., 2004: 619
Oomyzus Rondani, 1870
Oomyzus sokolowskii (Kurdjumov, 1912). Santos et al., 2000: 77
Quadrastichus Girault, 1913
Quadrastichus haitiensis (Gahan, 1929). Myers, 1931: 276; De Santis, 1979: 294; Schauff, 1987: 34; De Santis, 1989: 53;
Etienne et al., 1993: 105
Tamarixia Mercet, 1924
Tamarixia radiata (Waterston, 1922). Serra et al., 2011: 151

Family EUPELMIDAE
Subfamily EUPELMINAE
Anastatus Motschulsky, 1860
Anastatus (Anastatus) masneri Gibson, 1995: 109 *DR > TL: Prov. Pedernales, Sierra Bahoruco, 1450 m
Calosota Curtis, 1836
Calosota setosa Gibson, 2010: 52
Eupelmus Dalman, 1820
Eupelmus coccidivorus Gahan, 1934. De Santis, 1989: 29
Lecaniobius Ashmead, 1896
Lecaniobius cockerellii Ashmead, 1896. De Santis, 1989: 30
Psomizopelma Gibson, 1995
Psomizopelma fuscum Gibson, 2018 *DR > TL: La Vega, around Manabao, 900 m
Uropelma Sharkov, 1988
Uropelma sp.—Gibson (1995: 293)
Zaischnopsis Ashmead, 1904
Zaischnopsis sp.†—Wu (1997: 206) [Fossil in Dominican amber]

Family EURYTOMIDAE
Subfamily EURYTOMINAE
Bephratelloides Girault, 1913
Bephratelloides cubensis (Ashmead, 1894). Santoro, 1960: 229; De Santis, 1989: 16; Peña & Bennett, 1995: 346
Bephratelloides pomorum (Fabricius, 1804). Peña & Bennett, 1995: 346
Eurytoma Illiger, 1807
Eurytoma insularis Ashmead, 1894. De Santis, 1989: 16

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Subfamily RILEYINAE
Neorileya Ashmead, 1904
Neorileya ashmeadi Crawford, 1913. Gates, 2008: 55
Rileya Ashmead, 1888
Rileya bicolor Gates, 2008: 103 *DR > TL: Sierra de Bahoruco, Alcoa Rd., km 25
Rileya megastigma (Ashmead, 1894). Gates, 2008: 170
Rileya pulchra (Ashmead, 1894). Gates, 2008: 208

Family LEUCOSPIDAE
Leucospis Fabricius, 1775
Leucospis glaesaria† Engel, 2002: 437 [Fossil in Dominican amber]
Leucospis gomezi Genaro, 2012: 83 *DR > TL: La Descubierta, Lago Enriquillo, Prov. Barahona

Family MYMARIDAE
Acmopolynema Ogloblin, 1946
Acmopolynema vittatipenne (Dozier, 1932b: 83). De Santis, 1979: 376
Alaptus Westwood, 1839
Alaptus minutus Dozier, 1932b: 89. De Santis, 1979: 362 *H > TL: Reared from lime foliage, infested with Parlatoria zizyphus
scale, etc., at Port-au-Prince
Anagrus Haliday, 1833
Anagrus (Anagrus) empoascae Dozier, 1932b: 86
Anagrus (Anagrus) flaveolus Waterhouse, 1913. Dozier, 1932b: 86; De Santis, 1979: 361
Anagrus (Anagrus) raygilli Triapitsyn, 2000. Triapitsyn, 2002: 220
Anagrus (Anagrus) yawi Fullaway, 1944. Triapitsyn, 2002: 217
Camptoptera Förster, 1856
Camptoptera minutissima Dozier, 1932b: 89 *H > TL: Reared from avocado foliage infested with Empoasca minuenda Ball at
Pétionville
Camptoptera sp.†—Wu (1997: 204) [Fossil in Dominican amber]
Camptopteroides Viggiani, 1974
Camptopteroides sp.—Huber & Naiquan (1999: 32)
Cosmocomoidea Howard, 1908
Cosmocomoidea portoricensis (Dozier, 1937). Triapitsyn et al., 2010: 205
Cosmocomoidea quadrivittatus (Dozier, 1932b: 81). De Santis, 1979: 366; Triapitsyn et al., 2010: 207 *H > TL: Reared from
foliage and stem material of Malpighia glabra, at Port-au-Prince
Erythmelus Enock, 1909
Erythmelus (Erythmelus) noeli (Dozier, 1932b: 87). De Santis, 1979: 369; Triapitsyn et al., 2007: 43 *H > TL: On leaf the Sea
Grape, Coccoloba uvifera, at Port-au-Prince
Lymaenon Walker, 1846
Lymaenon cubensis (Dozier, 1932b: 83). De Santis, 1979: 365; Triapitsyn et al., 2010: 46
Lymaenon flaviventris (Dozier, 1932b: 82). De Santis, 1979: 365; Triapitsyn et al., 2010: 47
Mymar Curtis, 1829
Mymar taprobanicus Ward, 1875. Navarro Morales, 2014: 145
Polynema Haliday, 1833
Polynema haitianum Dozier, 1932b: 85. De Santis, 1979: 376 *H > TL: Grass and weeds between coffee and bananas at Fond-
des-Negres
Polynema phaseoli (Dozier, 1932b: 84). De Santis, 1979: 353 *H > TL: Reared from red bean foliage at Damien
Palaeopatasson† Witsack, 1986
Palaeopatasson grollei† Witsack, 1986: 263 [Fossil in Dominican amber]
Schizophragma Ogloblin, 1949
Schizophragma bicolor (Dozier, 1932b: 88). De Santis, 1979: 370
Yoshimotoana Huber, 2015
Yoshimotoana masneri (Yoshimoto, 1990: 88); Triapitsyn et al., 2010: 177; Huber, 2015: 62 *DR > TL: El Río, La Vega Prov.
=Yoshimotoana hispaniola (Triapitsyn & Huber in Triapitsyn et al., 2010: 164); Huber, 2015: 62

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 295
Family ORMYRIDAE
Ormyrus Westwood, 1832
Ormyrus sp.—Serra et al. (2011: 148)

Family PTEROMALIDAE
Subfamily CEINAE
Spalangiopelta Masi, 1926
Spalangiopelta albigena Darling, 1991: 24
Spalangiopelta georgei† Darling, 1996: 248 [Fossil in Dominican amber]
Subfamily CEROCEPHALINAE
Dominocephala† Krogmann, 2013
Dominocephala vibrissae† Krogmann, 2013: 47 [Fossil in Dominican amber]
Tenuicornus† Bläser, Krogmann & Peters, 2015
Tenuicornus dominicus† Bläser, Krogmann & Peters, 2015: 96 [Fossil in Dominican amber]
Subfamily EUNOTINAE
Scutellista Motschulsky, 1859
Scutellista caerulea (Fonscolombe, 1832) [By synonymy of species below]
=Scutellista cyanea Motschulsky, 1859. De Santis, 1989: 22
Subfamily LEPTOFOENINAE
Leptofoenus F. Smith, 1862
Leptofoenus pittfieldae† Engel, 2009: 61 [Fossil in Dominican amber]
Subfamily MISCOGASTERINAE
Halticoptera Spinola, 1811
Halticoptera circulus (Walker, 1833). De Santis, 1989: 23; Serra, 1992d: Tab. A13
Subfamily PTEROMALINAE
Pachyneuron Walker, 1833
Pachyneuron albutius Walker, 1843. De Santis, 1979: 163
Subfamily SPALANGIINAE
Spalangia Latreille, 1805
Spalangia cameroni Perkins, 1910. Gibson, 2009: 35
Spalangia drosophilae Ashmead, 1887. Gibson, 2009: 50
Spalangia endius Walker, 1839. Gibson, 2009: 56
Spalangia flavicrus Gibson, 2009: 70 *DR > TL: La Vega, National Park Armando Bermúdez, La Ciénaga, 1100 m
Spalangia haematobiae Ashmead, 1894. Gibson, 2009: 81
Spalangia imitator Gibson, 2009: 85
Spalangia masneri Gibson, 2009: 106
Spalangia nigroaenea Curtis, 1839. De Santis, 1979: 118
Spalangia simplex Perkins, 1910. Gibson, 2009: 141
Subfamily SYCORYCTINAE
Philotrypesis Förster, 1878
Philotrypesis sp.—Ramírez Pérez (2018: 139)

Family SIGNIPHORIDAE
Signiphora Ashmead, 1880
Signiphora aleyrodis Ashmead, 1900. Evans & Serra, 2002: 210; Woolley & Molin, 2017: 11
Signiphora dozieri Woolley & Molin, 2017: 40
Signiphora fax Girault, 1913. Woolley & Molin, 2017: 53
Signiphora flava Girault, 1913. De Santis, 1989: 47
Signiphora insularis (Dozier, 1933: 98)
Signiphora maculata Girault, 1913. Dozier, 1933: 99; Woolley & Molin, 2017: 77
Signiphora magniclavus (Dozier, 1933: 99) *H > TL: Taken sweeping grass, weeds and shrubbery at Port-au-Prince
Signiphora perpauca Girault, 1915. Woolley & Molin, 2017: 83
Signiphora townsendi Ashmead, 1900. Evans & Serra, 2002: 210
Signiphora unifasciata Ashmead, 1900. Dozier, 1933: 98

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Family TANAOSTIGMATIDAE
Tanaostigma Howard, 1890
Tanaostigma coursetiae Howard, 1890. LaSalle, 1987: 108
Tanaostigmodes Ashmead, 1896
Tanaostigmodes haematoxyli (Dozier, 1932: 103). De Santis, 1979: 167; LaSalle, 1987: 26; Cicero, 1992: 6/92

Family TORYMIDAE
Gummilumpus† Grissell, 1995
Gummilumpus bouceki† (Grissell, 1980: 255). Grissell, 1995: 135 [Fossil in Dominican amber]
Zophodetus† Grissell, 1980
Zophodetus woodruffi† Grissell, 1980: 253 [Fossil in Dominican amber]

Family TRICHOGRAMMATIDAE
Bloodiella Nowicki, 1935
Bloodiella ormenidis (Dozier, 1932d: 35). De Santis, 1979: 354 *H > TL: Reared from cottony covered eggs of the common
fulgorid, Ormenis sp., on red-flowered ornamental shrub on the Ecole Centrale grounds at Damien
Brachyufens Viggiani, 1968
Brachyufens osborni (Dozier, 1932). De Santis, 1979: 358; Schauff, 1987: 32
Ceratogramma De Santis, 1957
Ceratogramma etiennei Delvare, 1988. Etienne et al., 1993: 105 [Purposely introduced]
Ittys Girault, 1911
Ittys platycotis Dozier, 1932d: 32 *H > TL: Reared from eggs of the tree hopper Platycotis (Lophopelta) tuberculatus Fairm., in
avocado twigs in the mountains at Kenscoff
Megaphragma Timberlake, 1924
Megaphragma mymaripenne Timberlake, 1924. Dozier, 1932d: 29. De Santis, 1979: 341
Paracentrobia Howard, 1897
Paracentrobia ducassei (Dozier, 1932d: 30). De Santis, 1979: 356
Pseudoligosita Girault, 1913
Pseudoligosita plebeia (Perkins, 1912). Viggiani, 1981: 109
=Chaetostricha phaseoli Dozier, 1932: 31
Szelenyia Nowicki, 1940
Szelenyia masneri (Pinto & Viggiani, 1991: 720)
Trichogramma Westwood, 1833
Trichogramma fuentesi Torre, 1980. Pinto, 1999: 142
Trichogramma minutum (Riley, 1871). Wolcott, 1927: 342; Dozier, 1932d: 29; De Santis, 1979: 349; De Santis, 1989: 68
Trichogramma pretiosum Riley, 1879. Pinto, 1999: 1884
Trichogramma zeta Pinto, 1999. Pinto, 1999: 108
Ufens Girault, 1911
Ufens beneficus Dozier, 1932d: 33. De Santis, 1979: 359
Ufens niger (Ashmead, 1888). Owen, 2011: 112

Superfamily CERAPHRONOIDEA
Family CERAPHRONIDAE
Ceraphron Jurine, 1807
Ceraphron ceuthonymus† Engel, 2013: 2 [Fossil in Dominican amber]
Ceraphron sp.†—Poinar & Poinar (1999: 208) [Fossil in Dominican amber]

Family MEGASPILIDAE
Subfamily MEGASPILINAE
Conostigmus Dahlbom, 1857
Conostigmus sp.†—Poinar & Poinar (1999: 209) [Fossil in Dominican amber]
Dendrocerus Ratzeburg, 1852
Dendrocerus sp.†—Wu (1997: 208) [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 297
Suborder APOCRITA
ACULEATA
Superfamily VESPOIDEA
Family FORMICIDAE
Key reference: Lubertazzi (2019). General classification follows Bolton (2003).
Subfamily AMBLYOPONINAE
Tribe AMBLYOPONINI
Fulakora Mann, 1919
Fulakora falcata (Lattke, 1991). Lubertazzi & Alpert, 2014: 3; Lubertazzi, 2019: 113
Prionopelta Mayr, 1866
Prionopelta sp.†—Baroni Urbani (1995: 20); Wu (1997: 214) [Fossil in Dominican amber]
Subfamily CERAPACHYINAE
Tribe ACANTHOSTICHINI
Acanthostichus Mayr, 1887
Acanthostichus hispaniolicus† De Andrade, 1998b: 20 [Fossil in Dominican amber]
Tribe CYLINDROMYRMECINI
Cylindromyrmex Mayr, 1870
Cylindromyrmex darlingtoni Wheeler, 1937. Lubertazzi, 2019: 108
Cylindromyrmex inopinatus† De Andrade, 2001: 52 [Fossil in Dominican amber]
Cylindromyrmex sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Subfamily DOLICHODERINAE
Tribe BOTHRIOMYRMECINI
Bothriomyrmex Dubovikoff, 2005
Bothriomyrmex enigmaticus Prebus & Lubertazzi, 2016: 1; Lubertazzi, 2019: 77 *DR > TL: 7 km WSW El Factor, 195 m
Tribe DOLICHODERINI
Azteca Forel, 1878
Azteca alpha† Wilson, 1985c: 25 [Fossil in Dominican amber]
Azteca eumeces† Wilson, 1985c: 31 [Fossil in Dominican amber]
Dolichoderus Lund, 1831
Dolichoderus caribbaeus† (Wilson, 1985c: 19) [Fossil in Dominican amber]
Dolichoderus dibolius† Wilson, 1985c: 18 [Fossil in Dominican amber]
Dolichoderus intermedius† MacKay, 1993: 64 [Fossil in Dominican amber]
Dolichoderus primitivus† (Wilson, 1985c: 22) [Fossil in Dominican amber]
Dolichoderus prolaminatus† (Wilson, 1985c: 20) [Fossil in Dominican amber]
Dorymyrmex Mayr, 1866
Dorymyrmex antillana Snelling, 2005. Lubertazzi & Alpert, 2014: 3; Lubertazzi, 2019: 111
Gracilidris Wild & Cuezzo, 2006
Gracilidris humiloides† (Wilson, 1985c: 33). Heterick & Shattuck, 2011: 170 [Fossil in Dominican amber]
Linepithema Mayr, 1866
Linepithema dispertitum (Forel, 1885). Wild, 2007: 43; Lubertazzi, 2019: 124
Linepithema flavescens (Wheeler & Mann, 1914: 43). Wild, 2007: 49; Lubertazzi, 2019: 125 *H > TL: Cape Haitien
=Linepithema keiteli flavescens (Wheeler & Mann, 1914: 43). Wheeler, 1936: 204
Linepithema keiteli (Forel, 1907). Wild, 2007: 76; Lubertazzi, 2019: 126 *DR, *H > TL: Port-au-Prince
=Linepithema keiteli subfasciatus (Wheeler & Mann, 1914: 43)
Linepithema hispaniolae† (Wilson, 1985c: 32). Shattuck, 1992: 16 [Fossil in Dominican amber]
Leptomyrmex Mayr, 1862
Leptomyrmex neotropicus† (Baroni Urbani, 1980c: 4). Baroni Urbani & Wilson, 1987: 1; Boudinot et al., 2016: 667; Barden et
al., 2017: 766 [Fossil in Dominican amber]
Tapinoma Förster, 1850
Tapinoma litorale Wheeler, 1905. Menozzi & Russo, 1930: 164; Wetterer & Guerrero, 2017: 145; Lubertazzi, 2019: 183
Tapinoma melanocephalum (Fabricius, 1793). Wheeler & Mann, 1914: 43; Menozzi & Russo, 1930: 165; Lubertazzi, 2019:
184 [Introduced]
Tapinoma opacum Wheeler & Mann, 1914: 43; Lubertazzi, 2019: 185 *H > TL: Furcy

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Tapinoma rasenum Smith & Lavigne, 1973. Lubertazzi, 2019: 186
Tapinoma troche† Wilson, 1985c: 34 [Fossil in Dominican amber]
Technomyrmex Mayr, 1872
Technomyrmex caritatis† Brandão et al., 1999: 416 [Fossil in Dominican amber]
Technomyrmex hispaniolae† (Wilson, 1985c: 32); Heterick & Shattuck, 2011: 171 [Fossil in Dominican amber]
Subfamily DORYLINAE
Syscia Roger, 1861
Syscia sp. 1—Lubertazzi (2019: 125)
Subfamily ECITONINAE
Tribe ECITONINI
Neivamyrmex Borgmeier, 1940
Neivamyrmex ectopus† Wilson, 1985b: 11 [Fossil in Dominican amber]
Subfamily ECTATOMMINAE
Tribe ECTATOMMINI
Gnamptogenys Roger, 1863
Gnamptogenys brunoi† Lattke, 2002: 137 [Fossil in Dominican amber]
Gnamptogenys casca† Lattke, 2002: 138 [Fossil in Dominican amber]
Gnamptogenys falcaria Lattke, 2002: 139. Lubertazzi, 2019: 115
Gnamptogenys haytiana (Wheeler & Mann, 1914: 11). Lubertazzi, 2019: 115 *H > TL: Mountains north of Jacmel
Gnamptogenys insularis Lattke, 2002: 140. Lubertazzi, 2019: 115 *DR > TL: 16 km ENE Pedernales, 800 m
Gnamptogenys levinates† Baroni Urbani, 1980a: 5 [Fossil in Dominican amber]
Gnamptogenys lineolata Brown, 1992: 275. Lubertazzi, 2019: 116 *DR > TL: Prov. La Vega, Parque Nacional [Armando
Bermúdez], La Ciénaga, 1100 m
Gnamptogenys pristina† Baroni Urbani, 1980d: 2 [Fossil in Dominican amber]
Gnamptogenys schmitti (Forel, 1901: 334). Wheeler & Mann, 1914: 8; Lubertazzi, 2019: 117 *H > TL: Haïti
Gnamptogenys semiferox Brown, 1958: 324. Lubertazzi, 2019: 118 *DR > TL: Mt. Diego de Ocampo, 3000–4000 ft. altitude
Gnamptogenys striatula Mayr, 1884. Lubertazzi, 2019: 119
Subfamily FORMICINAE
Tribe CAMPONOTINI
Camponotus Mayr, 1861
Camponotus (Dendromyrmex) sp.†—Baroni Urbani (1995: 20) [Fossil in Dominican amber]
Camponotus (Myrmeurynota) albistramineus Wheeler, 1936: 208. Lubertazzi, 2019: 80 *H > TL: Mt. Trou d’Eau, 4000 ft.
Camponotus (Myrmeurynota) altivagans Wheeler, 1936: 207. Lubertazzi, 2019: 81 *H > TL: Between La Visite and Kenscoff
Camponotus (Myrmeurynota) augustei Wheeler & Mann, 1914: 59. Wheeler, 1936: 207; Lubertazzi, 2019: 82 *H > TL:
Pétionville
Camponotus (Myrmeurynota) christophei Wheeler & Mann, 1914: 57. Wheeler, 1936: 207; Lubertazzi, 2019: 83 *H > TL:
Near Milot on blades of a tall grass outside the citadel of Henri Christophe
Camponotus (Myrmeurynota) saussurei Forel, 1879. Lubertazzi, 2019: 90
Camponotus (Myrmeurynota) toussainti Wheeler & Mann, 1914: 60. Wheeler, 1936: 207; Lubertazzi, 2019: 91 *H > TL:
Pétionville in the mountains north of Jacmel
Camponotus (Myrmosphincta) sexguttatus (Fabricius, 1793). Wheeler & Mann, 1914: 56; Menozzi & Russo, 1930: 167
Camponotus (Pseudocolobopsis) claviscapus occultus Wheeler & Mann, 1914: 56. Menozzi & Russo, 1930: 167; Lubertazzi,
2019: 84 *H, *DR > TL: Pétionville
Camponotus sp.†—Wu (1997: 214) [Fossil in Dominican amber]
Camponotus (Tanaemyrmex) fumidus fraterculus Wheeler & Mann, 1914: 50. Lubertazzi, 2019: 85 *H > TL: Furcy
Camponotus (Tanaemyrmex) fumidus haytianus Wheeler & Mann, 1914: 48. Wheeler, 1936: 205; Lubertazzi, 2019: 86 *H >
TL: Diquini
Camponotus (Tanaemyrmex) fumidus illitus Wheeler & Mann, 1914: 51. Wheeler, 1936: 205; Lubertazzi, 2019: 86 *H > TL:
Port-au-Prince
Camponotus (Tanaemyrmex) fumidus imbecillus Wheeler & Mann, 1914: 52. Lubertazzi, 2019: 87 *H > TL: Grande Rivière
Camponotus (Tanaemyrmex) fumidus soulouquei Wheeler & Mann, 1914: 50. Lubertazzi, 2019: 87 *H > TL: Gonaives
Camponotus (Tanaemyrmex) lucayanus Wheeler, 1936. Lubertazzi, 2019: 88
Camponotus (Tanaemyrmex) plombyi Wheeler & Mann, 1914: 47. Wheeler, 1936: 205; Lubertazzi, 2019: 88 *H > TL: Furcy
Camponotus (Tanaemyrmex) ramulorum vernulus Wheeler, 1936: 206. Lubertazzi, 2019: 89 *H > TL: Northeastern foothills of

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the Massif de la Hotte, at an altitude of 3000–4000 ft.
Camponotus ustus Forel, 1879. Lubertazzi, 2019: 92
=Camponotus (Tanaemyrmex) larvigerus maculifrons Menozzi in Menozzi & Russo, 1930: 167. Wheeler, 1936: 205 *DR >
TL: Pueblo Viejo, Azua
=Camponotus (Tanaemyrmex) ustus depolitus Wheeler, 1936: 205
=Camponotus (Tanaemyrmex) ustus furnissi Wheeler & Mann, 1914: 55
=Camponotus (Tanaemyrmex) ustus sublautus Wheeler & Mann, 1914: 55
=Camponotus (Tanaemyrmex) ustus ulysses Forel, 1907. Wheeler & Mann, 1914: 54
Tribe PLAGIOLEPIDINI
Brachymyrmex Mayr, 1868
Brachymyrmex heeri Forel, 1874. Wheeler & Mann, 1914: 44; Menozzi & Russo, 1930: 166; Lubertazzi, 2019: 78
Brachymyrmex obscurior Forel, 1893. Menozzi & Russo, 1930: 166; Lubertazzi, 2019: 78
Brachymyrmex sp.†—Baroni Urbani (1995: 20) [Fossil in Dominican amber]
Myrmelachista Roger, 1863
Myrmelachista (Myrmelachista) gagates Wheeler, 1936: 204; Lubertazzi, 2019: 133 *H, *DR > TL: La Visite, 6000-7000 ft.
Myrmelachista (Myrmelachista) ramulorum Wheeler, 1908. Wheeler, 1936: 206; Menozzi & Russo, 1930: 165; Lubertazzi,
2019: 134
Myrmelachista sp.†—Baroni Urbani (1995: 20) [Fossil in Dominican amber]
Nylanderia Emery, 1906
Nylanderia bibadia LaPolla & Kallal, 2019: 408 *DR > TL: Parque Nacional Sierra de Bahoruco, elev. 1423 m
Nylanderia caerula LaPolla & Kallal, 2019: 411 *DR > TL: Parque Nacional Sierra de Bahoruco, elev. 407 m
Nylanderia disatra LaPolla & Kallal, 2019: 414 *DR > TL: Near Reserva Cientifica Ebano Verde, elev. 1139 m
Nylanderia esperanza LaPolla & Kallal, 2019: 417 *DR > TL: Parque Nacional del Este, nr. Boca de Yuma, elev. 8 m
Nylanderia fulva (Mayr, 1862). Wheeler & Mann, 1914: 45; Wheeler, 1936: 210
Nylanderia fuscaspecula LaPolla & Kallal, 2019: 418 *DR > TL: Salto de Jimenoa, elev. 620 m
Nylanderia guatemalensis (Forel, 1885). [By synonymy of species below]
=Paratrechina vividula antillana (Forel, 1893). Menozzi & Russo, 1930: 168
Nylanderia guatemalensis itinerans (Forel, 1901). Wheeler & Mann, 1914: 44
Nylanderia metacista LaPolla & Kallal, 2019: 422 *DR > TL: Cachote forest, elev. 1054 m
Nylanderia pini LaPolla & Kallal, 2019: 427 *DR > TL: Parque Nacional Armando Bermúdez, elev. 1037 m
Nylanderia steinheili (Forel, 1893). Wheeler & Mann, 1914: 45; Menozzi & Russo, 1930: 168; Wheeler, 1936: 210; LaPolla &
Kallal, 2019: 437
Nylanderia vetula† LaPolla & Dlussky, 2010: 263 [Fossil in Dominican amber]
=Paratrechina (Nylanderia) sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Paratrechina Motschulsky, 1863
Paratrechina longicornis (Latreille, 1802). Lubertazzi, 2019: 141 [Introduced]
=Paratrechina longicornis hagemanni (Forel, 1901). Wheeler & Mann, 1914: 45; Wheeler, 1936: 210
Zatania LaPolla, Kallal & Brady, 2012
Zatania darlingtoni (Wheeler, 1936: 210). Lubertazzi, 2019: 199 *H > TL: Between La Visite and Kenscoff, at an altitude of
5000-7000 ft.
Zatania electra† LaPolla, Kallal & Brady, 2012: 206 [Fossil in Dominican amber]
=Prenolepis sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Zatania gibberosa (Roger, 1863). Fontenla, 2000: 82; LaPolla, Kallal & Brady, 2012: 209; Lubertazzi, 2019: 199
Subfamily LEPTANILLOIDINAE
Tribe LEPTANILLOIDINI
Leptanilloides Mann, 1923
Leptanilloides sp.†—Wu (1997: 211) [Fossil in Dominican amber]
Subfamily MYRMICINAE
Tribe ATTINI
Apterostigma Mayr, 1865
Apterostigma sp.†—Baroni Urbani (1995: 20) [Fossil in Dominican amber]
Apterostigma electropilosum† Schultz, 2007: 426 [Fossil in Dominican amber]
Apterostigma eowilsoni† Schultz, 2007: 427 [Fossil in Dominican amber]
Cyphomyrmex Mayr, 1862

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Cyphomyrmex minutus Mayr, 1862. Wheeler & Mann, 1914: 42; Wheeler, 1936: 204; Lubertazzi, 2019: 108
=Cyphomyrmex rimosus flavescens Weber, 1940: 411. Kempf, 1965: 162; Snelling & Longino, 1992: 490
Cyphomyrmex rimosus Spinola, 1851. Lubertazzi, 2019: 109
Cyphomyrmex taino† De Andrade, 2003: 135 [Fossil in Dominican amber]
Mycocepurus Forel, 1893
Mycocepurus smithii (Forel, 1893). Wheeler & Mann, 1914: 42; Lubertazzi, 2019: 132
Mycetomoellerius Solomon et al., 2019
Mycetomoellerius jamaicensis (André, 1893). Mayhé-Nunes & Brandão 2007: 13; Rabeling et al., 2007: 11; Wetterer, 2015:
1176; Lubertazzi, 2019: 196; Solomon et al., 2019 : 948
=Trachymyrmex jamaicensis frontalis Santschi, 1925: 238
Mycetomoellerius haytianus (Wheeler & Mann, 1914: 41). Lubertazzi, 2019: 196; Solomon, 2019: 948
Mycetomoellerius primaevus (Baroni Urbani, 1980a: 4). Solomon et al., 2019 : 948 [Fossil in Dominican amber]
Tribe BASICEROTINI
Eurhopalothrix Brown & Kempf, 1961
Eurhopalothrix floridana Brown & Kempf, 1961. Longino, 2013: 118; Lubertazzi, 2019: 112
Eurhopalothrix sp.†—Baroni Urbani (1995: 20) [Fossil in Dominican amber]
Octostruma Forel, 1912
Octostruma sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Tribe BLEPHARIDATTINI
Wasmannia Forel, 1893
Wasmannia auropunctata (Roger, 1863). Wheeler & Mann, 1914: 38; Menozzi & Russo, 1930: 162; Wheeler, 1936: 200;
Lubertazzi, 2019: 198
Wasmannia sp.†—Baroni Urbani (1995: 20); Wu (1997: 212) [Fossil in Dominican amber]
Tribe CEPHALOTINI
Cephalotes Latreille, 1802
Cephalotes alveolatus† (Vierbergen & Scheven, 1995: 166) [Fossil in Dominican amber]
Cephalotes argentiventris De Andrade in De Andrade & Baroni Urbani, 1999: 104. Lubertazzi, 2019: 99 *DR > TL:
Constanza, 3–4,000 ft.
Cephalotes auricomus (Wheeler, 1936: 202). Lubertazzi, 2019: 99
Cephalotes bloosi† Baroni Urbani & De Andrade in De Andrade & Baroni Urbani, 1999: 549 [Fossil in Dominican amber]
Cephalotes brevispineus† De Andrade in De Andrade & Baroni Urbani, 1999: 240 [Fossil in Dominican amber]
Cephalotes caribicus† De Andrade in De Andrade & Baroni Urbani, 1999: 418 [Fossil in Dominican amber]
Cephalotes decoloratus De Andrade in De Andrade & Baroni Urbani, 1999: 508. Lubertazzi, 2019: 100 *H, *DR > TL:
Diquini
Cephalotes dieteri† Baroni Urbani in De Andrade & Baroni Urbani, 1999: 547 [Fossil in Dominican amber]
Cephalotes flavigaster De Andrade in De Andrade & Baroni Urbani, 1999: 108. Lubertazzi, 2019: 101 *DR > TL: Constanza,
3–4,000 ft.
Cephalotes hamulus (Roger, 1863: 269). Dalla Torre, 1893: 142; Wheeler, 1936: 201; Lubertazzi, 2019: 101 *H, *DR > TL: St.
Domingo
=Cryptocerus haemorrhoidalis (Latreille, 1802). F. Smith, 1854: 223; Dalla Torre, 1893: 142
Cephalotes hispaniolicus† De Andrade in De Andrade & Baroni Urbani, 1999: 298 [Fossil in Dominican amber]
Cephalotes integerrimus† (Vierbergen & Scheven, 1995: 162) [Fossil in Dominican amber]
Cephalotes jansei† (Vierbergen & Scheven, 1995: 161) [Fossil in Dominican amber]
Cephalotes obscurus† (Vierbergen & Scheven, 1995: 164) [Fossil in Dominican amber]
Cephalotes resinae† De Andrade in De Andrade & Baroni Urbani, 1999: 96 [Fossil in Dominican amber]
Cephalotes serratus† (Vierbergen & Scheven, 1995). De Andrade & Baroni Urbani, 1999: 522 [Fossil in Dominican amber]
Cephalotes squamosus† (Vierbergen & Scheven, 1995: 163) [Fossil in Dominican amber]
=Zacryptocerus pseudoaztecus (Vierbergen & Scheven, 1995: 163)
Cephalotes sucinus† De Andrade in De Andrade & Baroni Urbani, 1999: 540 [Fossil in Dominican amber]
Cephalotes taino† De Andrade in De Andrade & Baroni Urbani, 1999: 106 [Fossil in Dominican amber]
Cephalotes unimaculatus (F. Smith, 1854: 221). De Andrade & Baroni Urbani, 1999: 100; Lubertazzi, 2019: 103 *H, *DR >
TL: Erroneously “Brazil”, species is endemic to Hispaniola
Cephalotes vinosus (Wheeler, 1936: 202); Lubertazzi, 2019: 104 *H > TL: Mt. Rochelois
Tribe CREMATOGASTRINI

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Crematogaster Lund, 1831
Crematogaster (Orthocrema) russoi Menozzi in Menozzi & Russo, 1930: 155. Lubertazzi, 2019: 106
Crematogaster (Acrocoelia) sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Crematogaster (Orthocrema) sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Crematogaster (Orthocrema) steinheili Forel, 1881. Wheeler, 1936: 203; Lubertazzi, 2019: 106
Trichomyrmex Mayr, 1865
Trichomyrmex destructor (Jerdon, 1851). Wolcott, 1927: 349; Menozzi & Russo, 1930: 158; Wetterer, 2009: 102 [Introduced]
Tribe DACETINI
Acanthognathus Mayr, 1887
Acanthognathus poinari† Baroni Urbani in Baroni Urbani & De Andrade, 1994: 41 [Fossil in Dominican amber]
Strumigenys F. Smith, 1860
Strumigenys economoi Booher, Prebus & Lubertazzi, 2019: 353 *DR > TL: Jardín Botánico Nacional, 45 m
Strumigenys eggersi (Emery, 1890). Brown, 1960: 46: Bolton, 2000: 184; Wetterer, 2018: 138; Lubertazzi, 2019: 176
Strumigenys electrina† (De Andrade in Baroni Urbani & De Andrade, 1994: 38) [Fossil in Dominican amber]
Strumigenys emmae (Emery, 1890). Wetterer, 2012c: 72; Lubertazzi, 2019: 176 [Introduced]
Strumigenys gundlachi (Roger, 1862). Bolton, 2000: 186; Lubertazzi & Alpert, 2014: 3; Lubertazzi, 2019: 177
Strumigenys lanuginosa Wheeler, 1905. Wetterer, 2017: 731; Lubertazzi, 2019: 177
Strumigenys louisianae Roger, 1863. Brown, 1953: 28; Lubertazzi, 2019: 178
Strumigenys membranifera Emery, 1869. Wetterer, 2011b: 133; Lubertazzi, 2019: 179 [Introduced]
Strumigenys nigrescens (Wheeler, 1911). Brown, 1953: 96; Lubertazzi, 2019: 180
Strumigenys pilosula† Baroni Urbani & de Andrade, 2008: 168 [Fossil in Dominican amber]
Strumigenys poinari† Baroni Urbani & de Andrade, 2008: 173 [Fossil in Dominican amber]
Strumigenys rogeri (Emery, 1890). Wheeler & Mann, 1914: 40; Menozzi & Russo, 1930: 163; Wetterer, 2012a: 4; Lubertazzi,
2019: 181 [Introduced]
Strumigenys schleeorum† (Baroni Urbani & De Andrade, 1994: 35) [Fossil in Dominican amber]
Strumigenys silvestrii Emery, 1906. MacGown, Wetterer & Hill, 2012: 318; Lubertazzi, 2019: 182 [Introduced]
Strumigenys zemi Booher, Prebus & Lubertazzi, 2019: 348 *DR > TL: Loma Guaconejo, 150 m
Tribe FORMICOXENINI
Cardiocondyla Emery, 1869
Cardiocondyla emeryi Forel, 1881. Menozzi & Russo, 1930: 153; Wheeler, 1936: 199; Lubertazzi, 2019: 96 [Introduced]
Cardiocondyla minutior Forel, 1899. Wetterer, 2014: 572; Lubertazzi, 2019: 97 [Introduced]
Cardiocondyla venustula Wheeler, 1908. Wheeler & Mann, 1914: 19; Wheeler, 1936: 199; Lubertazzi, 2019: 98 [Introduced]
Temnothorax Mayr, 1861
Temnothorax caritatis† (De Andrade in De Andrade et al., 1999: 134) [Fossil in Dominican amber]
Temnothorax ciferrii (Menozzi in Menozzi & Russo, 1930: 160). Baroni Urbani, 1978: 433; Lubertazzi, 2019: 187 *DR > TL:
Los Siete Hermanos
Temnothorax creolus (Baroni Urbani, 1978: 437). Lubertazzi, 2019: 187 *DR > TL: Baní, 65 m
Temnothorax dominicanus† (De Andrade in De Andrade et al., 1999: 137) [Fossil in Dominican amber]
Temnothorax flavidulus (Wheeler & Mann, 1914: 37). Baroni Urbani, 1978: 441; Lubertazzi, 2019: 187 *H > TL: Manneville
Temnothorax hispaniolae (Baroni Urbani, 1978: 448). Lubertazzi, 2019: 188 *DR > TL: Cazabita [Loma Casabito], 1250 m
Temnothorax praecreolus† (De Andrade, 1992: 344) [Fossil in Dominican amber]
Temnothorax pastinifer (Emery, 1894). Baroni Urbani, 1978: 474
Temnothorax sallei (Guérin-Méneville, 1852). Wheeler & Mann, 1914: 31; Wheeler, 1936: 200; Baroni Urbani, 1978: 489;
Lubertazzi, 2019: 189
=Macromischa sallei opacinoda Wheeler, 1931. Bolton, 1995: 242
Tribe LASIINI
Acropyga Roger, 1862
Acropyga dubitata (Wheeler & Mann, 1914: 47). Lubertazzi, 2019: 67 *DR > TL: San Francisco Mts. of San Domingo
Acropyga glaesaria† LaPolla, 2005: 22 [Fossil in Dominican amber]
Acropyga parvidens (Wheeler & Mann, 1914: 46). Lubertazzi, 2019: 70 *H > TL: Pétionville
=Acropyga mesonotalis (Weber, 1944: 111)
Tribe MYRMICINI
Pogonomyrmex Mayr, 1868
Pogonomyrmex aterrimus Wheeler, 1936: 197. Johnson & Cover, 2015: 232; Lubertazzi, 2019: 157 *H,*DR > TL: Between La

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Visite and Kenscoff
=Pogonomyrmex schmitti aterrimus Wheeler, 1936.
=Pogonomyrmex schmitti darlingtoni Wheeler, 1936: 197
Pogonomyrmex saucius Wheeler & Mann, 1914: 29. Wheeler, 1936: 196; Johnson & Cover, 2015: 239; Lubertazzi, 2019: 158
*H, *DR > TL: About one-half mile east of Manneville
Pogonomyrmex schmitti Forel, 1901: 339. Johnson & Cover, 2015: 244; Lubertazzi, 2019: 158 *H, *DR > TL: Haïti
=Pogonomyrmex schmitti sublaevigatus Wheeler & Mann, 1914: 29. Wheeler, 1936: 197
Pogonomyrmex sp.†—Baroni Urbani (1995: 20) [Fossil in Dominican amber]
Tribe PHEIDOLINI
Aphaenogaster Mayr, 1853
Aphaenogaster amphioceanica† De Andrade, 1995: 4 [Fossil in Dominican amber]
Aphaenogaster relicta Wheeler & Mann, 1914. Lubertazzi, 2019: 75
Aphaenogaster relicta epinotalis Wheeler & Mann, 1914: 27. Wheeler, 1936: 198; Lubertazzi, 2019: 75 *H > TL: Manneville
Pheidole Westwood, 1841
Pheidole anticua† Casadei-Ferreira, Chaul & Feitosa, 2019: 118 [Fossil in Dominican amber]
Pheidole darlingtoni Wheeler, 1936: 198. Lubertazzi, 2019: 142 *H > TL: Northeastern foothills of the Massif de la Hotte,
about 3000 ft.
Pheidole drepanon Wilson, 2003: 409. Lubertazzi, 2019: 143 *DR > TL: Sierra de Bahoruco, N. Cabo Rojo, km 23, 500 m
Pheidole flavens Roger, 1863. Wheeler & Mann, 1914: 24; Lubertazzi, 2019: 143
Pheidole harlequina Wilson, 2003: 432. Lubertazzi, 2019: 144 *DR > TL: Hotel Nueva Suiza, Constanza, La Vega Prov., 1300
m
Pheidole hispaniolae Wilson, 2003: 707. Lubertazzi, 2019: 145 *DR > TL: 14 km northwest of Bonao, 1100 m
Pheidole indica Mayr, 1879. Lubertazzi, 2019: 146 [Introduced]
=Pheidole teneriffana Forel, 1893. Wetterer, 2011c: 846
Pheidole jamaicensis Wheeler, 1908. Longino & Cox, 2009: 37; Lubertazzi, 2019: 147
=Pheidole bilimeki Mayr, 1870. Wilson, 2003: 378 [Misidentification]
Pheidole jelskii Mayr, 1884. Wheeler & Mann, 1914: 21; Menozzi & Russo, 1930: 153; Wheeler, 1936: 198; Lubertazzi &
Alpert, 2014: 3; Lubertazzi, 2019: 147
Pheidole megacephala (Fabricius, 1793). Wheeler & Mann, 1914: 20; Menozzi & Russo, 1930: 152; Wheeler, 1936: 198;
Wetterer, 2012d: 56; Lubertazzi, 2019: 149 [Introduced]
Pheidole moerens Wheeler, 1908. Wheeler, 1936: 198; Wilson, 2003: 461; Lubertazzi, 2019: 150
Pheidole noar Wilson, 2003: 210. Lubertazzi, 2019: 150 *DR > TL: Cabo Rojo, 5 m
Pheidole primigenia† Baroni Urbani, 1995: 9 [Fossil in Dominican amber]
Pheidole subarmata Mayr, 1884. Lubertazzi & Alpert, 2014: 3. Lubertazzi, 2019: 151
Pheidole susannae Forel, 1886. Lubertazzi, 2019: 153 [Introduced]
Pheidole terresi Wheeler & Mann, 1914: 22. Wilson, 2003: 523; Lubertazzi, 2019: 154 *H > TL: Diquini
=Pheidole terresi illota Wheeler & Mann, 1914: 23
Pheidole sp.? (prope primigenia)†—Baroni Urbani (1995: 12) [Fossil in Dominican amber]
Pheidole tethepa† Wilson, 1985a: 5 [Fossil in Dominican amber]
Tribe SOLENOPSIDINI
Monomorium Mayr, 1855
Monomorium bicolor Emery, 1877. Lubertazzi, 2019: 128 [Introduced]
Monomorium ebeninum Forel, 1891. Wheeler & Mann, 1914: 19; Menozzi & Russo, 1930: 157; Wheeler, 1936: 199;
Lubertazzi, 2019: 129
Monomorium floricola (Jerdon, 1851). Wheeler & Mann, 1914: 19; Menozzi & Russo, 1930: 156; Wetterer, 2010a: 19;
Lubertazzi, 2019: 130 [Introduced]
Monomorium pharaonis (Linnaeus, 1758). Wetterer, 2010c: 115; Lubertazzi & Alpert, 2014: 3; Lubertazzi, 2019: 131
[Introduced]
Oligomyrmex Mayr, 1867
Oligomyrmex sp.†—Baroni Urbani (1995: 20) [Fossil in Dominican amber]
Oxyidris† Wilson, 1985
Oxyidris antillana† Wilson, 1985a: 5 [Fossil in Dominican amber]
Solenopsis Westwood, 1841
Solenopsis geminata (Fabricius, 1804). Wheeler & Mann, 1914: 20; Menozzi & Russo, 1930: 158; Wheeler, 1936: 200;

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Wetterer, 2011a: 27; Lubertazzi, 2019: 172
Solenopsis globularia (F. Smith, 1858). Pacheco & Mackay, 2013: 164; Lubertazzi, 2019: 173
=Solenopsis globularia borinquensis F. Smith, 1858. Wheeler & Mann, 1914: 20; Menozzi & Russo, 1930: 160
=Solenopsis globularia steinheili Forel, 1881. Menozzi & Russo, 1930: 154; Wheeler, 1936: 199
Solenopsis pollux Forel, 1893. Wheeler & Mann, 1914: 20; Lubertazzi, 2019: 173
Solenopsis pygmaea Forel, 1901. Pacheco & Mackay, 2013: 282; Lubertazzi, 2019: 174
Solenopsis (Euophthalma) sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Solenopsis (Solenopsis) sp.†—Wilson (1985e: 266); De Andrade & Baroni Urbani (1999: 893) [As Diplorhoptrum] [Fossil in
Dominican amber]
Solenopsis succinea Emery, 1890. Pacheco & Mackay, 2013: 313; Lubertazzi, 2019: 175
Solenopsis terricola Menozzi, 1931. Lubertazzi & Alpert, 2014: 3; Lubertazzi, 2019: 175
Note: The records of Monomorium salomonis (Linnaeus, 1758) by Wheeler & Mann, 1914: 19 and Menozzi & Russo, 1930:
157 may be misidentifications of M. ebeninum. Monomorium salomonis is not included in the Hispaniola species list
[Lubertazzi, 2019: 132].
Tribe STENAMMINI
Rogeria Emery, 1894
Rogeria alzatei Kluger, 1994: 52. Lubertazzi, 2019: 170
Rogeria brunnea Santschi, 1930. Kluger, 1994: 48; Lubertazzi, 2019: 170
Rogeria curvipubens Emery, 1894. Lubertazzi & Alpert, 2014: 3; Lubertazzi, 2019: 170
Rogeria leptonana Kluger, 1994: 58. Lubertazzi, 2019: 170
Tribe TETRAMORIINI
Tetramorium Mayr, 1855
Tetramorium bicarinatum (Nylander, 1846). Lubertazzi, 2019: 190; Guérin-Méneville, 1852: 79 [as Myrmica cariniceps]
[Introduced]
Tetramorium caldarium (Roger, 1857). Bolton, 1979: 169; Wetterer & García, 2015: 97; Lubertazzi, 2019: 191 [Introduced]
Tetramorium lanuginosum Mayr, 1870. Wetterer, 2010b: 85; Lubertazzi & Alpert, 2014: 3; Lubertazzi, 2019: 192 [Introduced]
Tetramorium lucayanum Wheeler, 1905. Smith, 1937: 852; Lubertazzi, 2019: 193 [Introduced]
Tetramorium simillimum (F. Smith, 1851). Wheeler & Mann, 1914: 38; Menozzi & Russo, 1930: 161; Lubertazzi, 2019: 194
[Introduced]
Tribe INCERTAE SEDIS
Ilemomyrmex† Wilson, 1985
Ilemomyrmex caecus† Wilson, 1985a: 3 [Fossil in Dominican amber]
Subfamily PARAPONERINAE
Tribe PARAPONERINI
Paraponera F. Smith, 1858
Paraponera dieteri† Baroni Urbani, 1994: 5 [Fossil in Dominican amber]
Subfamily PONERINAE
Tribe PONERINI
Anochetus Mayr, 1861
Anochetus ambiguus† De Andrade, 1994: 17 [Fossil in Dominican amber]
Anochetus brevidentatus† MacKay, 1991: 138. De Andrade, 1994: 19 [Fossil in Dominican amber]
Anochetus conisquamis† De Andrade, 1994: 22 [Fossil in Dominican amber]
Anochetus corayi† Baroni Urbani, 1980b: 2. De Andrade, 1994: 24 [Fossil in Dominican amber]
Anochetus exstinctus† De Andrade, 1994: 11 [Fossil in Dominican amber]
Anochetus haytianus Wheeler & Mann, 1914: 15. Lubertazzi, 2019: 72 *H > TL: Manneville
Anochetus intermedius† De Andrade, 1994: 13 [Fossil in Dominican amber]
Anochetus longispinus Wheeler, 1936: 196. Lubertazzi, 2019: 73 *H > TL: Northeastern foothills of the Massif de la Hotte,
3000–4000 ft.
Anochetus lucidus† De Andrade, 1994: 19 [Fossil in Dominican amber]
Anochetus mayri Emery, 1884. Wheeler & Mann, 1914: 15; Lubertazzi, 2019: 73
Hypoponera Santschi, 1938
Hypoponera opaciceps (Mayr, 1887). Wheeler & Mann, 1914: 13; Wheeler, 1936: 196; Lubertazzi, 2019: 120
Hypoponera opacior (Forel, 1893). Menozzi & Russo, 1930: 150; Lubertazzi, 2019: 121
Hypoponera punctatissima (Roger, 1859). Lubertazzi, 2019: 63; Lubertazzi, 2019: 121

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Hypoponera sp.†—Wilson (1985e: 266) [Fossil in Dominican amber]
Note: The records of Hypoponera ergatandria (Forel, 1893) by Wheeler & Mann, 1914: 13 and Hypoponera foeda (Forel,
1893) by Wheeler, 1936: 195 are considered problematic records. These seem to be misidentifications of H. punctatissima (see
Lubertazzi, 2019: 122).
Leptogenys Roger, 1861
Leptogenys antillana Wheeler & Mann, 1914: 14. Wheeler, 1936: 196; Lattke, 2011: 146; Lubertazzi, 2019: 123 *H, *DR >
TL: Pétionville
Leptogenys pubiceps Emery, 1890. Lattke, 2011: 219; Lubertazzi, 2019: 123
=Leptogenys punctaticeps Emery, 1890. Wheeler & Mann, 1914: 14 [Misidentification]
Odontomachus Latreille, 1804
Odontomachus bauri Emery, 1892. Brown, 1976: 130; Lubertazzi, 2019: 137
Odontomachus insularis Guérin-Méneville, 1844. Ashmead, 1900: 321; Wheeler & Mann, 1914: 17; Menozzi & Russo, 1930:
151; Wheeler, 1936: 196; Lubertazzi, 2019: 138
Odontomachus pseudobari† De Andrade, 1994: 9 [Fossil in Dominican amber]
Odontomachus ruginodis Smith, 1937. Lubertazzi, 2019: 139
Odontomachus spinifer† De Andrade, 1994: 7 [Fossil in Dominican amber]
Notes: Following Lubertazzi (2019: 140) Odontomachus bradleyi Brown, 1976: 133 is deleted from the list of Hispaniolan
ants. This species is from South America. Odontomachus haematodus (Linnaeus, 1758) also reported from Hispaniola (see
Lubertazzi, 2019: 140) does not appear to occur on the island.
Pseudoponera Emery, 1900
Pseudoponera sp.†—Wilson (1985e: 266 as Pachycondyla) [Fossil in Dominican amber]
Pseudoponera stigma (Wheeler & Mann, 1914: 13). Menozzi & Russo, 1930: 150; Wetterer, 2012b: 41; MacKay, W. & E.
MacKay, 2010: 518; Lubertazzi, 2019: 167
=Pachycondyla rufescens Wheeler & Mann, 1914: 13
Pseudoponera succedanea (Roger, 1863). MacKay, W. & E. MacKay, 2010: 538; Lubertazzi, 2019: 167
Thaumatomyrmex Mayr, 1887
Thaumatomyrmex sp. 1—Lubertazzi (2019: 136)
Thaumatomyrmex sp. 2—Lubertazzi (2019: 136)
Tribe PLATYTHYREINI
Platythyrea Roger, 1863
Platythyrea dentata† Lattke, 2003: 108 [Fossil in Dominican amber]
Platythyrea procera† Lattke, 2003: 108 [Fossil in Dominican amber]
Platythyrea pumilio† De Andrade, 2004: 644 [Fossil in Dominican amber]
Platythyrea punctata (F. Smith, 1858). Ashmead, 1900: 317; Wheeler & Mann, 1914: 7; Menozzi & Russo, 1930: 150; Kellner
et al., 2010: 737; Lubertazzi, 2019: 155
Platythyrea scalprum† Lattke, 2003: 110 [Fossil in Dominican amber]
Platythyrea strenua Wheeler & Mann, 1914: 6; Lubertazzi, 2019: 156 *H > TL: Diquini
Subfamily PROCERATIINAE
Tribe PROCERATIINI
Discothyrea Roger, 1863
Discothyrea gigas† De Andrade, 1998c: 205 [Fossil in Dominican amber]
Discothyrea sp.†—Baroni Urbani (1995: 20); Wu (1997: 213) [Fossil in Dominican amber]
Discothyrea testacea Roger, 1863. Lubertazzi, 2019: 110
Proceratium Roger, 1863
Proceratium denticulatum† Lattke, 1990: 201; Baroni Urbani & De Andrade, 2003: 66 [Fossil in Dominican amber]
Proceratium dominicanum† De Andrade in Baroni Urbani & De Andrade, 2003: 182 [Fossil in Dominican amber]
Proceratium gibberum† De Andrade in Baroni Urbani & De Andrade, 2003: 68 [Fossil in Dominican amber]
Proceratium longiscapus De Andrade in Baroni Urbani & De Andrade, 2003: 190. Lubertazzi, 2019: 161 *DR > TL: Bonao
Proceratium poinari† Baroni Urbani & De Andrade, 2003: 178 [Fossil in Dominican amber]
Proceratium taino De Andrade in Baroni Urbani & De Andrade, 2003: 186. Lubertazzi, 2019: 161 *DR > TL: La Vega Prov.,
PN. A. Bermúdez, 4 km SW Ciénaga, 1200 m
Subfamily PSEUDOMYRMECINAE
Tribe PSEUDOMYRMECINI
Pseudomyrmex Lund, 1831

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Pseudomyrmex antiquus† Ward, 1992: 57 [Fossil in Dominican amber]
Pseudomyrmex avitus† Ward, 1992: 59 [Fossil in Dominican amber]
Pseudomyrmex baros† Ward, 1992: 63 [Fossil in Dominican amber]
Pseudomyrmex coruscus† Ward, 1992: 65 [Fossil in Dominican amber]
Pseudomyrmex cubaensis (Forel, 1910). Ward, 1992: 80; Lubertazzi, 2019: 163
Pseudomyrmex flavidulus (F. Smith, 1858). Wheeler & Mann, 1914: 17
Pseudomyrmex haytianus (Forel, 1901: 342). Wheeler & Mann, 1914: 17; Wheeler, 1936: 196; Ward, 1992: 80; Lubertazzi,
2019: 163 *H > TL: Haïti
Pseudomyrmex macrops† Ward, 1992: 66 [Fossil in Dominican amber]
Pseudomyrmex nexilis† Ward, 1992: 68 [Fossil in Dominican amber]
Pseudomyrmex oryctus† Ward, 1992: 69 [Fossil in Dominican amber]
Pseudomyrmex prioris† Ward, 1992: 70 [Fossil in Dominican amber]
Pseudomyrmex simplex (F. Smith, 1877). Ward, 1992: 75; Lubertazzi, 2019: 164
Pseudomyrmex subater (Wheeler & Mann, 1914). Ward, 1992: 80; Lubertazzi, 2019: 165
Pseudomyrmex succinus† Ward, 1992: 71 [Fossil in Dominican amber]
Pseudomyrmex thecolor† Ward, 1992: 73 [Fossil in Dominican amber]
Pseudomyrmex vicinus† Ward, 1992: 74 [Fossil in Dominican amber]
Note: The record of Pseudomyrmex gracilis (Fabricius, 1804) for Hispaniola by Ward, 1992: 80 was later found to be erroneous
and the species record excluded (Wetterer, 2010d: 538).

Family MUTILIIDAE
Subfamily MUTILIINAE
Ephuta Say, 1835
Ephuta clavigera† Brothers, 2003: 103 [Fossil in Dominican amber]
Ephuta prima Genaro, 1997: 264 *DR > TL: Prov. Barahona, cerca de Filipinas, mina Larimar
Subfamily SPHAEROPTHALMINAE
Dasymutilla Ashmead, 1899
Dasymutilla albifasciatus† Manley & Poinar, 1999: 49 [Fossil in Dominican amber]
Dasymutilla bouvieri (André, 1898: 51). Mickel, 1928: 27; Nonveiller, 1990: 56; Genaro, 1997: 264; Manley & Pitts, 2007: 33;
Williams & Pitts, 2013: 432 *H, *DR > TL: Saint-Domingue
Dasymutilla dominica† Manley & Poinar, 1991: 201 [Fossil in Dominican amber]
Dasymutilla melancholica (F. Smith, 1879: 223). Ashmead, 1900: 316; Mickel, 1928: 27; Nonveiller, 1990: 58; Manley &
Pitts, 2007: 67 *H, *DR > TL: St. Domingo
Dasymutilla militaris nigriceps (Cresson, 1865). Manley & Pitts, 2007: 68
Dasymutilla spiniscapula Manley & Pitts, 2007: 92 *DR > TL: Pedernales Prov., 21 km N Cabo Rojo
Pseudomethoca Ashmead, 1896
Pseudomethoca flaviceps (André, 1906: 72). Nonveiller, 1990: 46; Mickel, 1928: 27 *H > TL: Haïti
Pseudomethoca merengue Genaro, 1997: 266 *DR > TL: Prov. Independencia, Los Ríos, Lago Enriquillo
Sphaeropthalma Blake, 1871
Sphaeropthalma (Photopsis) retifera (Dow, 1931: 2). Nonveiller, 1990: 72

Family POMPILIDAE
Subfamily CEROPALINAE
Irenangelus Schulz, 1906
Irenangelus hispaniolae Evans, 1969: 431. Kimsey & Wasbauer, 2004: 662; Waichert et al., 2012: 5 *DR > TL: Confluence of
Río Duey and Río Haina, Sec. El Duey, Prov. San Cristóbal
Subfamily NOTOCYPHINAE
Notocyphus F. Smith, 1855
Notocyphus anacaona Rodriguez & Pitts in Waichert et al., 2012: 32 *DR > TL: Hato Mayor, Parque Los Haitises, 3 km W
Cueva de Arena, 20 m
Subfamily PEPSINAE
Ageniella Banks, 1911
Ageniella (Ageniella) bruesi (Banks, 1928). Waichert et al., 2012: 7
Ageniella (Ageniella) domingensis (Banks, 1944: 186). Waichert et al., 2012: 8 *DR > TL: Near and southeast of Constanza,

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Valle Nuevo, 3–7000 ft.
Ageniella (Ageniella) violaceipes (Cresson, 1865). Waichert et al., 2012: 10
Ageniella (Ameragenia) ursula (Banks, 1944: 184). Waichert et al., 2012: 10 *DR > TL: Villa Altagracia
Ageniella (Priophanes) dowii (Banks, 1938). Waichert et al., 2012: 9
=Priocnemis arioles Banks, 1944: 185
Auplopus Spinola, 1841
Auplopus bellus (Cresson, 1865). Banks, 1928: 7; Waichert et al., 2012: 12
Auplopus charlesi Waichert & Pitts in Waichert et al., 2012: 11 *DR > TL: Pedernales, 26 km N Cabo Rojo, 730 m
Caliadurgus Pate, 1946
Caliadurgus maestris Alayo, 1969. Waichert et al., 2012: 13
Dipogon Fox, 1897
Dipogon (Deuteragenia) marlowei Waichert & Pitts in Waichert et al., 2012: 14 *DR > TL: Pedernales, La Abeja, 38 km NNW
Cabo Rojo, 1250 m
Entypus Dahlbom, 1844
Entypus caeruleus (Linnaeus, 1758). Waichert et al., 2012: 15
Entypus sulphureicornis (Palisot de Beauvois). Waichert et al., 2012: 15
Entypus manni (Banks, 1928: 6). Waichert et al., 2012: 15 *H > TL: Port-au-Prince
Entypus ochrocerus Dahlbom, 1843. Day, 1974: 93; Waichert et al., 2012: 16
=Pompilus flammipennis F. Smith, 1855: 155. Day, 1974: 93
Epipompilus Kohl, 1884
Epipompilus pulcherrimus (Evans, 1955). Waichert et al., 2012: 6
Hemipepsis Dahlbom, 1844
Hemipepsis toussainti (Banks, 1928: 5). Waichert et al., 2012: 17
Pepsis Fabricius, 1805
Pepsis marginata Palisot de Beauvois, 1809. F. Smith, 1855: 199; Vardy, 2002: 67; Snelling & Torres, 2004: 366; Waichert et
al., 2012: 17
Pepsis rubra (Drury, 1773). Wolcott, 1927: 358; Vardy, 2000: 63; Snelling & Torres, 2004: 367; Waichert et al., 2012: 18
=Pepsis quadrata Lepeletier, 1845. F. Smith, 1855: 198; Vardy, 2000: 64
Pepsis ruficornis (Fabricius, 1781). Banks, 1928: 5; Vardy, 2005: 232; Waichert et al., 2012: 19
Priocnemis Schiödte, 1837
Priocnemis cornica (Say, 1836). Waichert et al., 2012: 20
Priocnessus Banks, 1925
Priocnessus vancei Waichert & Pitts in Waichert et al., 2012: 21 *DR > TL: Pedernales, Sierra de Bahoruco, Aceitillar, 23.6 km
NE Pedernales, 1560 m
Subfamily POMPILINAE
Anoplius Dufour, 1834
Anoplius (Anoplius) fulgidus (Cresson, 1865). Evans, 1951: 339; Waichert et al., 2012: 23
Anoplius (Arachnophroctonus) americanus ambiguus (Dahlbom, 1845). Snelling & Torres, 2004: 369; Waichert et al., 2012: 24
Anoplius (Arachnophroctonus) hispaniolae Evans, 1966: 324. Snelling & Torres, 2004: 371; Waichert et al., 2012: 26
Anoplius (Notiochares) amethystinus amethystinus (Fabricius, 1793). F. Smith, 1855: 199; Cresson, 1867: 149; Evans, 1950:
229; Waichert et al., 2012: 25
Anoplius planeta† Rodriguez & Pitts in Rodriguez et al., 2016: 90 [Fossil in Dominican amber]
Aporinellus Banks, 1912
Aporinellus medianus Banks, 1917. Waichert et al., 2012: 27
Drepanaporus Bradley, 1944
Drepanaporus antillarum (Bradley, 1944: 107). Waichert et al., 2012: 28; Rodriguez et al., 2014: 715
Drepanaporus bachata Rodriguez et al., 2014: 717 *DR > TL: South of Bayahibe, La Altagracia
Drepanaporus collaris (Cresson, 1865). Waichert et al., 2012: 29; Rodriguez et al., 2014: 716
Episyron Schiödte, 1837
Episyron conterminus cressoni (Dewitz, 1881). Genaro, 1995: 12; Waichert et al., 2012: 30
=Episyron conterminus posterus (Fox, 1893). Genaro, 1995: 12.
Poecilopompilus Ashmead, 1902
Poecilopompilus mixtus (Fabricius, 1798). Evans, 1972: 5; Genaro, 1996: 431; Waichert et al., 2012: 33
Psorthaspis Banks, 1911

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Psorthaspis hispaniolae Bradley, 1944: 50. Waichert et al., 2012: 34 *DR, *H > TL: Constanza
Psorthaspis naomi (F. Smith, 1855: 177). Cresson, 1867: 135; Bradley, 1944: 73; Waichert et al., 2012: 34 *H, *DR > TL: St.
Domingo
Tachypompilus Ashmead, 1902
Tachypompilus ferrugineus bicolor (Banks, 1938). Genaro, 1995: 12; Waichert et al., 2012: 35
Tainopompilus† Rodriguez & Pitts, 2015
Tainopompilus argentum† Rodriguez & Pitts in Rodriguez et al., 2016: 92 [Fossil in Dominican amber]

Family RHOPALOSOMATIDAE
Rhopalosoma Cresson, 1865
Rhopalosoma haitiense Townes, 1977: 23 *H > TL: Manneville
Rhopalosoma hispaniola† Lohrmann in Lohrmann et al., 2019: 35 [Fossil in Dominican amber]
Rhopalosoma poeyi Cresson, 1865. Townes, 1977: 23
=Sibyllina aenigmatica Westwood, 1868: 329. Brues, 1943: 315

Family SCOLIIDAE
Subfamily SCOLIINAE
Campsomeris Guérin-Méneville, 1838
Campsomeris atrata (Fabricius, 1793). Smith, 1855: 106
Campsomeris dorsata (Fabricius, 1787). Bradley, 1928: 333
Campsomeris tolteca (Saussure, 1857). Bradley, 1928: 331
Campsomeris tricincta (Fabricius, 1775). Rohwer, 1927: 152
Campsomeris trifasciata (Fabricius, 1793). Wolcott, 1927: 231; Bradley, 1928: 321

Family TIPHIIDAE
Myzine Latreille, 1803
Myzine apicale sulphurea Krombein, 1942: 347 *H, *DR > TL: Port-au-Prince
Myzine ephippium ephippium (Fabricius, 1775). Krombein, 1942: 316
Myzine haemorrhoidalis haemorrhoidalis (Fabricius, 1775). Krombein, 1942: 335
Myzine striata boxi Krombein, 1942: 326 *H, *DR > TL: Port-au-Prince
Myzine wolcotti Krombein, 1942: 328 *H, *DR > TL: Kenscoff
Tiphia Fabricius, 1775
Tiphia (Tiphia) haiti Allen & Krombein, 1961: 62
Tiphia (Tiphia) hispaniolae Wolcott, 1938. Allen & Krombein, 1961: 58
Tiphia (Tiphia) kenscoffi Allen & Krombein, 1961: 60 *H > TL: Kenscoff
Tiphia (Tiphia) paupi Allen & Krombein, 1961: 62
Tiphia (Tiphia) sandae Allen & Krombein, 1961: 60 *DR > TL: San Domingo
Tiphia (Tiphia) wolcotti Allen & Krombein, 1961: 61 *H > TL: Kenscoff

Family VESPIDAE
Subfamily EUMENINAE
Eumenes Latreille, 1802
Eumenes (Eumenes) pictus F. Smith, 1857: 32. Ashmead, 1900: 312; Grandinete et al., 2018: 25 *H, *DR > TL: St. Domingo
Euodynerus Dalla Torre, 1904
Euodynerus jeititia Carpenter & Genaro, 2011: 7
Euodynerus haitiensis (Bequaert & Salt, 1931: 783) *H > TL: Grande Anse
Monobia Saussure, 1852
Monobia batesi Bequaert, 1940c: 830 *H > TL: Swamps north of Dessalines
Omicron Saussure, 1855
Omicron lacerum Giordani Soika, 1978: 166 *H, *DR > TL: Santo Domingo
Pachodynerus Saussure, 1875
Pachodynerus nasidens (Latreille, 1812). Wolcott, 1927: 358
Pachodynerus tibialis (Saussure, 1853). Saussure, 1875: 242
Parancistrocerus Bequaert, 1925

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Parancistrocerus domingensis Bequaert & Salt, 1931: 772
Parancistrocerus lutzi Bequaert & Salt, 1931: 778 *H > TL: Fond Parisien
Zeta Saussure, 1855
Zeta abdominale (Drury, 1770). Carpenter & Genaro, 2011: 14 *H, *DR > TL: Port-au-Prince
=Zethus abdominalis hispaniolae (Bequaert & Salt, 1931: 772)
=Eumenes abdominalis colona Saussure, 1875: 106. Ashmead, 1900: 312
Zethus Fabricius, 1805
Zethus (Zethus) albopictus F. Smith, 1857: 15. Saussure, 1875: 43; Ashmead, 1900: 312; Bohart & Stange, 1965: 102 *H, *DR
> TL: St. Domingo
Zethus (Zethusculus) jurinei Saussure, 1852. F. Smith, 1857: 15; Bohart & Stange, 1965: 142; Stange, 2004: 215 *DR > TL:
Apparently by error, type locality in original publication says “Carac.” [Caracas, Venezuela]. Type material would indicate
mislabeling of lectotype male (Stange, 2004b).
Subfamily POLISTINAE
Agelaia Lepeletier de St. Fargeau, 1835
Agelaia electra† Carpenter & Grimaldi, 1997: 2 [Fossil in Dominican amber]
Mischocyttarus Saussure, 1853
Mischocyttarus (Monocyttarus) phthisicus (Fabricius, 1793). Zikán, 1949: 171
Polistes Latreille, 1802
Polistes (Aphanilopterus) crinitus crinitus (Felton, 1765). Wolcott, 1927: 359; Carpenter, 1996: 27
Polistes (Aphanilopterus) incertus Cresson, 1865. Carpenter, 1996: 29
Polistes (Aphanilopterus) minor Palisot de Beauvois, 1818. Carpenter, 1996: 31
Polistes (Aphanilopterus) poeyi haitiensis Bequaert & Salt, 1931: 792. Carpenter, 1996: 32 *H, *DR > TL: Bizeton
Polistes (Palisotius) major major Palisot de Beauvois, 1818. Wolcott, 1927: 359; Carpenter, 1996: 30

Superfamily CHRYSIDOIDEA
Family BETHYLIDAE
Subfamily BETHYLINAE
Archaeopristocera† Terayama, 2004
Archaeopristocera miki† Terayama, 2004: 44 [Fossil in Dominican amber]
Parasierola Cameron, 1883
Parasierola sp.†—Poinar (1992: 201) [Fossil in Dominican amber]
Prosierola Kieffer, 1905
Prosierola cubana Evans, 1964. Azevedo, 2008: 4
=Prosierola insularis Evans, 1964: 188. Perez-Gelabert, 2008: 229
Subfamily EPYRINAE
Anisepyris Kieffer, 1906
Anisepyris darlingtoni Evans, 1959: 73. Barbosa & Azevedo, 2018: 51 *H, *DR > TL: Port-au-Prince & vicinity
Anysepyris bogotensis (Kieffer, 1910). Barbosa & Azevedo, 2018: 74
Anysepyris dominicanus Evans, 1966. Barbosa & Azevedo, 2018: 52
Anisepyris excisus Evans, 1959: 72. Barbosa & Azevedo, 2018: 52 *H, *DR > TL: Port-au-Prince
Anysepyris finduilasi Barbosa & Azevedo, 2018: 161
Anysepyris gradatus† Sorg, 1988. Barbosa & Azevedo, 2018: 132 [Fossil in Dominican amber]
Anisepyris grisselli Evans, 1979: 456. Barbosa & Azevedo, 2018: 235 *DR > TL: 30 km N Cabo Rojo, Prov. Pedernales
Anysepyris jocundus Evans, 1966. Barbosa & Azevedo, 2018: 54
Anysepyris venustus Evans, 1964. Barbosa & Azevedo, 2018: 240
Anisepyris wolcotti Evans, 1959: 72 *H > TL: Port-au-Prince
Bakeriella Kieffer, 1910
Bakeriella floridana Evans, 1964. Kawada & Azevedo, 2003: 468
Cephalonomia Westwood, 1833
Cephalonomia stephanoderis Betrem, 1960. Contreras & Camilo, 2007: 45 [Purposely introduced]
Epyris Westwood, 1832
Epyris hispaniolae Evans, 1964: 108. Evans, 1969: 242 *H > TL: Etang Lachaux, SW Peninsula, under 1000 ft.
Epyris manni Evans, 1964: 110. Evans, 1969: 195
Holepyris Kieffer, 1904

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Holepyris gracilis Kieffer, 1908. Evans, 1977: 544
Holepyris hispaniolae Evans, 1977: 565
Holepyris longulus Evans, 1977: 549 *DR > TL: Loma Rucilla and mts. N, 5–8000 ft.
Laelius Ashmead, 1893
Laelius muesebecki (Evans, 1965). Evans, 1979: 456; Waichert & Azevedo, 2009: 23
Nesepyris Bridwell, 1920
Nesepyris antilleanus Evans, 1979: 458 *DR > TL: Dominican Republic (on Verawood log) intercepted at quarantine
Subfamily PRISTOCERINAE
Dissomphalus Ashmead, 1893
Dissomphalus bicavatus Evans, 1979. Azevedo, 2001: 176
Dissomphalus singularis Evans, 1962: 77 *H > TL: Camp Perrin
Dissomphalus thysanus Azevedo, 2003: 50 *DR > TL: Pedernales, 24 km N Cabo Rojo, 610 m
Dissomphalus triangularis Azevedo, 1999: 323
Pseudisobrachium Kieffer, 1904
Pseudisobrachium terresi Mann, 1915: 164

Family CHRYSIDIDAE
Subfamily AMISEGINAE
Adelphe Mocsáry, 1890
Adelphe acuta Kimsey, 2008: 162 *DR > TL: La Vega, La Ciénaga, Parque Nacional A. Bermúdez
Adelphe argentea Kimsey, 1993: 206 *DR > TL: 9.5 km N of Cabo Rojo, Pedernales Prov.
Adelphe dominicana Kimsey, 1993: 206 *DR > TL: 14.5 km N of Cabo Rojo, Pedernales Prov.
Adelphe masneri Kimsey, 1986: 202 *DR > TL: Loma El Peñón, La Cumbre, Puerto Plata
Adelphe minuta Kimsey, 1993: 209 *DR > TL: 9.5 km N of Cabo Rojo, Pedernales Prov.
Adelphe paracubana Kimsey, 2008: 171
Adelphe taino (Krombein, 1957). Kimsey, 2008: 157
Subfamily CHRYSIDINAE
Caenochrysis Kimsey & Bohart, 1981
Caenochrysis bosci (Buysson, 1898: 562) *H, *DR > TL: St.-Domingue
Ceratochrysis Cooper, 1952
Ceratochrysis dominicana† Engel, 2006: 500 [Fossil in Dominican amber]
Chrysis Linnaeus, 1767
Chrysis insularis Guérin-Méneville, 1842. Kimsey & Bohart, 1990: 423
Chrysis purpuriventris Cresson, 1865. Kimsey & Bohart, 1990: 454
Holopyga Dahlbom, 1845
Holopyga maculata (Fabricius, 1798: 258). Kimsey & Bohart, 1990: 233 *DR, *H > TL: St. Domingue
Holopyga ventralis (Say, 1824) [By synonymy of species below]
=Holopyga dohrni Dahlbom, 1854. Buysson, 1898: 519

Family DRYINIDAE
Subfamily ANTEONINAE
Anteon Jurine, 1807
Anteon dominicanum Olmi, 1991: 203. Olmi & Virla, 2014: 59 *DR > TL: 21 km N Cabo Rojo, Pedernales Prov.
Anteon panamense Olmi, 1984. Olmi & Virla, 2014: 82
Subfamily APHELOPINAE
Aphelopus Dalman, 1823
Aphelopus poinari† Olmi, 1998: 153. Olmi & Virla, 2014: 29 [Fossil in Dominican amber]
Aphelopus surinamensis Olmi, 1984. Olmi & Virla, 2014: 23
Aphelopus trinitatis Olmi, 1984. Olmi & Virla, 2014: 24
Aphelopus tschirnhausi Olmi, 2011: 44. Olmi & Virla, 2014: 28
Crovettia Olmi, 1984
Crovettia plaumanniana Olmi, 1984. Olmi & Virla, 2014: 38
Subfamily BOCCHICINAE
Bocchus Ashmead, 1893

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Bocchus vetustus† Olmi, 1991: 243. Olmi & Virla, 2014: 212 [Fossil in Dominican amber]
Subfamily DRYININAE
Dryinus Latreille, 1804
Dryinus alametallus† Olmi & Guglielmino in Olmi et al., 2011: 45. Olmi & Virla, 2014: 248 [Fossil in Dominican amber]
Dryinus antilleanus (Evans, 1969). Olmi et al., 2000: 150. Olmi & Virla, 2014: 222
Dryinus grimaldii† Olmi, 1995: 254. Olmi & Virla, 2014: 305 [Fossil in Dominican amber]
Dryinus hymenaeaphilus† Olmi, 1995: 255 [Fossil in Dominican amber]
Dryinus obrieni (Olmi, 1991: 293). Olmi & Virla, 2014: 293 *DR > TL: 16 km E San José de Ocoa, Peravia Prov.
Dryinus palaeodominicanus† Currado & Olmi, 1983: 330. Olmi, 1995: 266; Olmi et al., 2011: 48; Olmi & Virla, 2014: 250
[Fossil in Dominican amber]
Dryinus paleomexicanus† Olmi, 1995. Olmi & Virla, 2014: 313 [Fossil in Dominican amber]
Dryinus poinari† Olmi, 1997: 161. Olmi & Virla, 2014: 314 [Fossil in Dominican amber]
Dryinus priscus† Olmi, 1997: 77; Olmi et al., 2011: 50; Olmi & Virla, 2014: 253 [Fossil in Dominican amber]
Dryinus pristinus† Olmi, 1997: 79; Olmi et al., 2011: 51; Olmi & Virla, 2014: 254 [Fossil in Dominican amber]
Dryinus rastnitsyni† Olmi & Guglielmo, 2011: 511; Olmi & Virla, 2014: 307 [Fossil in Dominican amber]
Dryinus smetanai Olmi, 2008: 222. Olmi & Virla, 2014: 239 *DR > TL: Puerto Plata Prov., Marapicá, edge of gallery forest
near Iberostar Costa Dorada Resort
Dryinus tainensis Olmi, 2008: 224. Olmi & Virla, 2014: 276 *DR > TL: La Vega Prov., 12 km N junction Hwys 1 and 12
Dryinus valens Olmi & Virla, 2014: 303 *DR > TL: Pedernales Prov., Sierra Bahoruco, 26 km N Cabo Rojo, 730 m
Dryinus vetustus† Olmi, 1995: 258. Olmi & Virla, 2014: 316 [Fossil in Dominican amber]
Harpactosphecion Haupt, 1944
Harpactosphecion scheveni† Olmi, 2004b: 16. Olmi & Virla, 2014: 319 [Fossil in Dominican amber]
Harpactosphecion sucinum† (Olmi, 1987: 220). Olmi & Virla, 2014: 321 [Fossil in Dominican amber]
Thaumatodryinus Perkins, 1905
Thaumatodryinus miocenicus† Olmi, 1995: 263. Olmi & Virla, 2014: 335 [Fossil in Dominican amber]
Subfamily GONATOPODINAE
Gonatopus Ljungh, 1810
Gonatopus breviforceps Kieffer, 1904. Olmi et al., 2000: 155; Olmi & Virla, 2014: 416
Gonatopus cubensis (Richards, 1969). Olmi et al., 2000: 153; Olmi & Virla, 2014: 360
Gonatopus haitianus (Olmi, 1986: 86). Olmi et al., 2000: 154

Family EMBOLEMIDAE
Embolemus Westwood, 1833
Embolemus kheeli Olmi, 2004a: 2 *DR > TL: La Altagracia, Punta Cana Resort
Embolemus subtilis Olmi, 1995. Olmi, 2004a: 2

Family SCLEROGIBBIDAE
Probethylus Ashmead, 1902
Probethylus callani Richards, 1939. Olmi, 2004c: 95
Probethylus poinari† Olmi, 2004c: 118 [Fossil in Dominican amber]
Pterosclerogibba† Olmi, 2004
Pterosclerogibba antiqua† Olmi, 2004c: 186 [Fossil in Dominican amber]

Family SCOLEBYTHIDAE
Clystopsenella Kieffer, 1911
Clystopsenella mirabilis† Engel, 2015: 4 [Fossil in Dominican amber]
Clystopsenella sp.†—Poinar & Poinar (1999: 210) [Fossil in Dominican amber]
Pristapenesia Brues, 1933
Pristapenesia inopinatus† (Prentice et al., 1996: 805) [Fossil in Dominican amber]

Superfamily APOIDEA
Family CRABRONIDAE
Subfamily ASTATINAE
Astata Latreille, 1796

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 311
Astata (Astata) dominica Pate, 1948: 230 *DR > TL: San Domingo
Subfamily BEMBICINAE
Tribe BEMBICINI
Bembix Fabricius, 1775
Bembix americana antilleana Fabricius, 1793. Genaro, 2006: 53
Bicyrtes Lepeletier de St. Fargeau, 1845
Bicyrtes spinosus (Fabricius, 1794). Genaro, 2006: 53
Crabro Geoffroy, 1762
Crabro sp.†—Poinar (1992: 213) [Fossil in Dominican amber]
Crossocerus Lepeletier de St. Fargeau & Brullé, 1835
Crossocerus sp.†—Poinar (1992: 213) [Fossil in Dominican amber]
Hoplisoides Gribodo, 1884
Hoplisoides alaya (Pate, 1947b: 96). Bohart, 2000: 224 *DR > TL: San Domingo
Hoplisoides arispilos Bohart, 2000: 224. Amarante, 2002: 15 *DR > TL: Tortuguero, Azua Prov.
Hoplisoides azuae Bohart, 2000: 226. Amarante, 2002: 15 *DR > TL: Tortuguero, Azua Prov.
Hoplisoides dominicanus Bohart, 2000: 228. Amarante, 2002: 15 *DR > TL: Playa
Tortuguero
Sphecius Dahlbom, 1844
Sphecius hogardii (Latreille, 1809). F. Smith, 1856: 336; Genaro, 2006: 52
Stictia Illiger, 1807
Stictia signata (Linnaeus, 1758). Wolcott, 1927: 358; Genaro, 2006: 53
Tribe NYSSONINI
Epinysson Pate, 1935
Epinysson borinquinensis (Pate, 1937). Genaro, 2007a: 58
Nysson Latreille, 1802
Nysson sp.†—Poinar & Poinar (1999: 210) [Misspelled Nyssonia] [Fossil in Dominican amber]
Zanysson Rohwer, 1921
Zanysson armatus (Cresson, 1865). Genaro, 1995: 12
Zanysson caonao Genaro, 2007a: 53 *DR > TL: Los Olivares, provincia Pedernales
Subfamily CRABRONINAE
Tribe CRABRONINI
Ectemnius Dahlbom, 1845
Ectemnius (Apoctemnius) craesus (Lepeletier & Brullé, 1835). Pate, 1947a: 12; Genaro, 2006: 54
Ectemnius (Hypocrabro) taino Pate, 1947a: 22 *DR > TL: Santo Domingo
Lindocerus Antropov, 2016
Lindocerus paleomystax† (Bennett & Engel, 2006: 2). Antropov, 2016: 614 [Fossil in Dominican amber]
Rhopalum Stephens, 1829
Rhopalum (Corynopus) lomae Leclercq, 2002: 66 *DR > TL: Loma El Peñón, La Cumbre, 676 m, P. Plata
Tribe OXYBELINI
Oxybelus Latreille, 1796
Oxybelus analis Cresson, 1865. Genaro, 2006: 54
Oxybelus marginatus F. Smith, 1856. Bohart, 1993: 160
Oxybelus romingeri Bohart, 1993: 167 *DR > TL: Montecristi
Tribe MISCOPHINI
Lyroda Say, 1837
Lyroda antillana Genaro & Portuondo, 2001: 45. Genaro, 2006: 54
Lyroda sp.—Genaro (1995: 12)
Tribe LARRINI
Nitela Latreille, 1809
Nitela sp.†—Poinar & Poinar (1999: 210) [Misspelled Nitella] [Fossil in Dominican amber]
Tachysphex Kohl, 1883
Tachysphex alayoi Pulawski, 1974. Genaro, 2006: 55
Tachysphex dominicanus Pulawski, 1988: 79. Genaro, 2006: 55
Tachysphex quisqueyus Pulawski, 1988: 156 *DR > TL: Prov. Pedernales, Cabo Rojo

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Tachytes Panzer, 1806
Tachytes chrysopyga (Spinola, 1841). Perez-Gelabert, 2008: 241
Tachytes distinctus F. Smith, 1856. Genaro, 2006: 55
Liris Fabricius, 1805
Liris labiatus (Fabricius, 1793). Genaro, 2006: 56
=Larrada ignipennis F. Smith, 1856: 288
Liris luctuosus (F. Smith, 1856: 289)
Liris trifasciatus (F. Smith, 1856). Genaro, 2006: 56
Tribe TRYPOXYLINI
Pison Spinola, 1808
Pison antiquum† Antropov & Pulaski, 1996: 16 [Fossil in Dominican amber]
Pison menkei† Bennett & Engel, 2008: 114 [Fossil in Dominican amber]
Trypoxylon Latreille, 1796
Trypoxylon dominicanum† Prentice & Poinar, 1993: 288 [Fossil in Dominican amber]
Trypoxylon electrum† Antropov, 1995: 125 [Fossil in Dominican amber]
Trypoxylon eucharis† Prentice & Poinar, 1993: 286 [Fossil in Dominican amber]
Trypoxylon pallidiventre† Prentice & Poinar, 1993: 288 [Fossil in Dominican amber]
Trypoxylon rohweri Richards, 1934: 251 *H > TL: Port-au-Prince
Trypoxylon subimpressum F. Smith, 1856: 380. Genaro, 2006: 56
=Trypoxylon excavatum F. Smith, 1856. Genaro, 2006: 56
Trypoxylon (Trypoxylon) orientinum Richards, 1969. Genaro, 2006: 56
Trypoxylon (Trypoxylon) n. sp. 1—Genaro (2006: 57)
Trypoxylon (Trypargilum) mexicanum (Saussure, 1867). Genaro, 2007b: 3
Trypoxylon (Trypargilum) saussurei Rohwer, 1912. Genaro, 2008: 48 [Introduced]
=Trypoxylon texense Saussure, 1867. Genaro, 1995: 12; Perez-Gelabert, 2008: 241
Subfamily PEMPHREDONINAE
Tribe PSENINI
Pluto Pate, 1937
Pluto atricornis (Malloch, 1933). Van Lith, 1979: 194
Pseneo Malloch, 1933
Pseneo alayoni Genaro, 2017: 13 *DR > TL: Parque Nacional A. Bermúdez, Antón Sape Bueno, Trillo de la Hispaniola
Tribe PEMPHREDONINI
Stigmus Panzer, 1804
Stigmus sp.—Finnamore (1995: 265)
Subfamily PHILANTINAE
Tribe CERCERINI
Cerceris Latreille, 1802
Cerceris ayti Genaro, 2017: 5 *H > TL: Port-au-Prince
Cerceris howardevansi Genaro, 2004: 761 *DR > TL: Prov. Monte Cristi, Paroli [beach]
Cerceris laevigata F. Smith, 1856: 465. Ashmead, 1900: 307; Ferguson, 1983: 487 *DR > TL: St. Domingo
Cerceris jeiti Genaro, 2009: 101 *DR > TL: Parque Nacional del Este, Boca de Yuma, Prov. La Altagracia
Cerceris surieli Genaro, 2017: 6 *DR > TL: Las Abejas, Sierra de Bahoruco
Tribe PHILANTHINI
Trachypus Klug, 1810
Trachypus gerstaeckeri Dewitz, 1881. Rubio Espina, 1975: 39

Family SPHECIDAE
Tribe SCELIPHRINI
Podium Fabricius, 1805
Podium fulvipes Cresson, 1865. Genaro, 2006: 50
Podium tau (Dalla Torre, 1897). Ashmead, 1900: 309; Bohart & Menke, 1976: 97; Genaro, 2006: 50
=Pepsis T Palisot de Beauvois, 1805. Fernald, 1907: 418
Sceliphron Klug, 1801
Sceliphron (Sceliphron) assimile (Dahlbom, 1843). Wolcott, 1927: 357

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 313
Sceliphron (Sceliphron) caementarium (Drury, 1773). Pulawski, 2003: 3
=Pelopaeus lunatus Latreille, 1805. Palisot de Beauvois, 1806: 50
Sceliphron (Sceliphron) jamaicense (Fabricius, 1775). Genaro, 2006: 50
Sceliphron fasciatum (Lepeletier, 1845). Bohart & Menke, 1976: 105
Note: The record of Sceliphron madraspatanum (Fabricius, 1781). Pulawski, 2003: 3 with the synonym Sceliphron lugubre
Christ, 1791. Ashmead, 1900: 309 seems to be an erroneous identification.
Tribe SPHECINI
Isodontia Patton, 1880
Isodontia poeyi antillana Genaro, 2017: 11 *DR > TL: Prov. La Altagracia, Cueva de Berna, Boca de Yuma
Prionyx Van der Linden, 1827
Prionyx thomae (Fabricius, 1775). Genaro, 2006: 51
Sphex Linnaeus, 1758
Sphex dorsalis Lepeletier, 1845. Evans, 1972: 8; Genaro, 2006: 51
=Chlorion (Proterosphex) spiniger (Kohn, 1890). Fernald, 1907: 392
Sphex ichneumoneus (Linnaeus, 1758). Fernald, 1907: 399
Sphex jamaicensis (Drury, 1773) [By synonymy of species below]
=Chlorion (Isodontia) costipennis (Spinola, 1851). Fernald, 1907: 351
Sphex mandibularis Cresson, 1869. Bohart & Menke, 1976: 115; Genaro, 2006: 51
Sphex resinipes (Fernald, 1906: 386). Bohart & Menke, 1976: 116
=Sphex rufipes Lepeletier, 1845. F. Smith, 1856: 260; Ashmead, 1900: 308
Sphex servillei Lepeletier de St. Fargeau, 1845 [By synonymy of species below]
=Chlorion (Proterosphex) chichimecum (Saussure, 1867). Fernald, 1907: 407
Note: The record of Sphex caliginosus Erichson, 1849 by Fernald, 1907: 403 for Santo Domingo is apparently an error as this
species is considered in more recent catalogues to be distributed in the Lesser Antilles, Mexico and Brazil.

Family COLLETIDAE
Subfamily DIPHAGLOSSINAE
Tribe CAUPOLICANINI
Caupolicana Spinola, 1851
Caupolicana (Alayoapis) notabilis (F. Smith, 1861). Ashmead, 1900: 305; Friese, 1902: 197; Michener, 1966: 749; Genaro,
2007c: 248
Subfamily HYLAEINAE
Hylaeus Fabricius, 1793
Hylaeus (Prosopis) hispaniolensis Genaro, 2016: 41 *DR > TL: Antón Sape Bueno, Trillo de la Hispaniola, Parque Bermúdez
Hylaeus (Prosopis?) n. sp.—Genaro (2007c: 248)
Hylaeus (Hylaeana) n. sp.—Genaro (2007c: 248)
Subfamily XEROMELISSINAE
Tribe CHILICOLINI
Chilicola Spinola, 1851
Chilicola (Hylaesoma) electrodominica† Engel, 2000: 455 [Fossil in Dominican amber]
Chilicola (Hylaeosoma) gracilis† Michener & Poinar, 1996: 358 [Fossil in Dominican amber]

Family ANDRENIDAE
Subfamily PANURGINAE
Tribe PROTANDRENINI
Heterosarus Robertson, 1918
Heterosarus sp. n.—Genaro (2007: 248)
Perdita F. Smith, 1853
Perdita islacabritosensis Ascher & Engel, 2012: 596 *DR > TL: Isla Cabrito, Descubierta, Prov. Independencia
Protandrena Cockerell, 1896
Protandrena eickworti† (Rozen, 1996: 347) [Fossil in Dominican amber]

Family HALICTIDAE
Subfamily HALICTINAE

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Tribe HALICTINI
Agapostemon Guérin-Méneville, 1844
Agapostemon (Agapostemon) alayoi Roberts, 1972: 461. Moure & Hurd, 1987: 204; Genaro, 2007c: 249 *H > TL: Port-au-
Prince
Agapostemon (Agapostemon) centratus (Vachal, 1903). Moure & Hurd, 1987: 205; Genaro, 2007c: 249
Agapostemon hispaniolicus Roberts, 1972: 484. Moure & Hurd, 1987: 204 *DR, *H > TL: Port-au-Prince & vic.
Agapostemon (Agapostemon) insularis Roberts, 1972: 485. Moure & Hurd, 1987: 203; Genaro, 2007c: 249
Agapostemon (Agapostemon) kohliellus (Vachal, 1903). Moure & Hurd, 1987: 204; Genaro, 2007c: 249
Agapostemon (Agapostemon) poeyi (Lucas, 1856). Moure & Hurd, 1987: 203; Genaro, 2007c: 249
Agapostemon (Notagapostemon) luzzii† Engel & Breitkreuz, 2013: 2 [Fossil in Dominican amber]
Agapostemon viridulus sp. group—Eickwort (1988: 236)
Halictus Latreille, 1804
Halictus (Halictus) poeyi Lepeletier, 1841. Genaro, 2007c: 249
Lasioglossum Curtis, 1833
Lasioglossum (Dialictus) busckiellum (Cockerell, 1915). Genaro, 2007c: 249; Gibbs, 2018: 46
Lasioglossum (Dialictus) ferreri (Baker, 1906). Genaro, 2007c: 249; Gibbs, 2018: 27
Lasioglossum (Dialictus) gundlachii (Baker, 1906). Genaro, 2007c: 249; Gibbs, 2018: 35
Lasioglossum (Dialictus) havanense (Baker, 1906). Genaro, 2007c: 249
Lasioglossum (Dialictus) mestrei (Baker, 1906). Genaro, 2007c: 249
Lasioglossum (Dialictus) parvum (Cresson, 1865). Genaro, 2007c: 249
Lasioglossum (Dialictus) sp. n. 1—Genaro (2007c: 249)
Nesosphecodes Engel, 2006
Nesosphecodes halictophagus Engel, 2006: 6. Genaro, 2007c: 249 *DR > TL: Pedernales, 60 km NW Cabo Rojo, Las Abejas
cloud forest, 1200 m
Sphecodes Latreille, 1804
Sphecodes sp.—Eickwort (1988: 236)
Tribe AUGOCHLORINI
Augochlora F. Smith, 1853
Augochlora (Augochlora) alcyone F. Smith, 1879: 45. Ashmead, 1900: 304; Cockerell, 1910: 489; Moure & Hurd, 1987: 256;
Genaro, 2007c: 249 *DR > TL: St. Domingo
Augochlora (Augochlora) cubana Cockerell, 1910. Moure & Hurd, 1987: 258
Augochlora (Augochlora) elegans Cresson, 1865. Friese, 1902: 197. Moure & Hurd, 1987: 260; Genaro, 2007c: 249
Augochlora (Augochlora) magnifica Cresson, 1865. Moure & Hurd, 1987: 263; Genaro, 2007c: 249
Augochlora (Augochlora) regina F. Smith, 1853. Moure & Hurd, 1987: 268; Genaro, 2007c: 249
Augochlora (Oxystoglossella) haitiensis (Vachal, 1911). Moure & Hurd, 1987: 273 Haiti?
Augochlora sp.—Genaro (2007: 249)
Neocorynura Schrottky, 1910
Neocorynura (Neocorynura) electra† Engel, 1995: 318 [Fossil in Dominican amber]
Oligochlora† Engel, 1996
Oligochlora eickworti† Engel, 1996: 338 [Fossil in Dominican amber]
Oligochlora grimaldii† Engel, 1997: 98 [Fossil in Dominican amber]
Oligochlora marquettorum† Engel & Rightmyer, 2000: 432 [Fossil in Dominican amber]
Oligochlora micheneri† Engel, 1996: 340 [Fossil in Dominican amber]
Oligochlora semirugosa† Engel, 2009: 4 [Fossil in Dominican amber]
Pereirapis Moure, 1943
Pereirapis semiaurata (Spinola, 1853) [By synonymy of species below]
=Pereirapis cerasis (Vachal, 1911). Moure & Hurd, 1987: 247
Tribe CAENOHALICTINI
Eickwortapis† Michener & Poinar, 1997
Eickwortapis dominicana† Michener & Poinar, 1997: 356 [Fossil in Dominican amber]
Nesagapostemon† Engel, 2009
Nesagapostemon moronei† Engel, 2009: 7 [Fossil in Dominican amber]

Family MEGACHILIDAE

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Subfamily MEGACHILINAE
Tribe LITHURGINI
Lithurgus Latreille, 1825
Lithurgus (Lithurgopsis) antilleorum Michener, 1989: 161. Genaro, 2007c: 249
Tribe MEGACHILINI
Coelioxys Latreille, 1809
Coelioxys (Cyrtocoelioxys) danielperezi Genaro, 2009: 13 *DR > TL: Cerro Santo Tomás, Parque Botánico de Jánico, Santiago
Prov., 493 m
Coelioxys (Neocoelioxys) vigilans F. Smith, 1879: 105. Ashmead, 1900: 303; Friese, 1902: 200; Genaro, 2007c: 250
Megachile Latreille, 1802
Megachile (Callomegachile) rufipennis (Fabricius, 1793). Genaro, 2007c: 249 [Introduced]
Megachile (Callomegachile) solitaria F. Smith, 1879: 80. Raw, 2007: 25 [Introduced]
Megachile (Callomegachile) torrida F. Smith, 1853. Genaro, 2007c: 249 [Introduced]
Megachile (Eutricharaea) concinna F. Smith, 1879: 79. Ashmead, 1900: 302; Friese, 1902: 199; Genaro, 2007c: 250
[Introduced]
Megachile (Leptorachis) curta Cresson, 1865. Genaro, 2007c: 249
Megachile (Melanosarus) sedula F. Smith, 1879: 79. Ashmead, 1900: 302; Friese, 1902: 200; Genaro, 2007c: 250; Raw, 2007:
62 *DR, *H > TL: St. Domingo
Megachile (Pseudocentron) deceptrix F. Smith, 1879: 81. Ashmead, 1900: 302; Friese, 1902: 199; Genaro, 2007c: 250; Raw,
2007: 76 *DR, *H > TL: St. Domingo
Megachile (Pseudocentron) elongata F. Smith, 1879: 80. Ashmead, 1900: 302; Friese, 1902: 199; Genaro, 2007c: 250; Raw,
2007: 77
Megachile (Pseudocentron) luctifera Spinola, 1841. Raw, 2007: 80
Megachile (Pseudomegachile) lanata (Fabricius, 1775). Genaro, 2007c: 249; Raw, 2007: 86 [Introduced]
Megachile (Sayapis) dentipes Vachal, 1909. Genaro, 2007c: 250; Raw, 2007: 92
Megachile (Sayapis) zaptlana Cresson, 1878. Genaro, 2007c: 250
Thaumatosoma F. Smith, 1865
Thaumatosoma (Chalicodomopsis) glaesaria† (Engel, 1999: 4); Engel et al., 2012: 2 [Fossil in Dominican amber]
Tribe OSMIINI
Heriades Spinola, 1808
Heriades (Neotrypetes) currani Michener, 1943. Genaro, 2007c: 249
Osmia Panzer, 1806
Osmia (Diceratosmia) stangei Genaro, 2001b: 1029 *DR > TL: Prov. Monte Cristi, playa Paroli

Family APIDAE
Subfamily APINAE
Tribe ANTHOPHORINI
Anthophora Latreille, 1803
Anthophora (Mistacanthophora) hilaris F. Smith, 1879: 123. Ashmead, 1900: 300; Friese, 1902: 198; Brooks, 1999: 115.
Genaro, 2007c: 250
Anthophora (Mistacanthophora) hispaniolae Brooks, 1999: 118. Genaro, 2007c: 250 *H, *DR > TL: 20 km S Constanza
Anthophora (Mistacanthophora) tricolor (Fabricius, 1775). Brooks, 1999: 120. Genaro, 2007c: 250
Tribe APINI
Apis Linnaeus, 1758
Apis mellifera Linnaeus, 1758. Wolcott, 1927: 362; Hedstrom, 1988: 553; Genaro, 2007c: 252 [Introduced]
Tribe CENTRIDINI
Centris Fabricius, 1805
Centris (Centris) poecila Lepeletier, 1841. Genaro, 2007c: 251
=Centris apicalis F. Smith, 1874. Friese, 1902: 199
Centris (Centris) decolorata Lepeletier, 1841. F. Smith, 1874: 364; Friese, 1902: 199; Genaro, 2007c: 251
Centris (Centris) insularis F. Smith, 1874: 367. Ashmead, 1900: 300; Friese, 1902: 199; Genaro, 2007c: 251 *H, *DR > TL:
San Domingo
Centris (Centris) testacea Lepeletier, 1841: 165. F. Smith, 1874: 365; Friese, 1902: 199; Wolcott, 1927: 66; Genaro, 2015: 1
*H, *DR > TL: Original: “Patrie inconnue.” Endemic to Hispaniola.

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Centris (Hemisiella) lanipes (Fabricius, 1775). Genaro, 2007c: 251
Centris (Heterocentris) analis (Fabricius, 1804). Genaro, 2007c: 251
=Centris fulviventris Cresson, 1865. Friese, 1902: 199
Centris (Trachina) simillima F. Smith, 1874: 370. Friese, 1902: 199; Genaro, 2007c: 251
Centris (Xanthemisia) domingensis Dalla Torre, 1986: 304. Friese, 1902: 199; Vivallo, 2014: 64 *DR > TL: St. Domingo
=Centris thoracica F. Smith, 1874: 370 [Preoccupied name]
Tribe ERICROCIDINI
Mesocheira Lepeletier & Serville, 1825
Mesocheira bicolor (Fabricius, 1804). Genaro, 2007c: 251
Mesoplia Lepeletier, 1841
Mesoplia (Mesoplia) pilicrus (Friese, 1902: 200). Genaro, 2007c: 251 *H > TL: Port-au-Prince
Mesoplia (Mesoplia) rufipes (Perty, 1833). Genaro, 2007c: 251
Tribe EUCERINI
Melissodes Berthold, 1827
Melissodes (Ecplectica) haitiensis LaBerge, 1961. Genaro, 2007c: 251
Melissodes mimica Cresson, 1869. Genaro, 2007c: 251
Melissodes sp.—Genaro (2007c: 251)
Melissoptila Holmberg, 1884
Melissoptila (Ptilomelissa) micheneri Genaro, 2001a. Genaro, 2007c: 251
Tribe EUGLOSSINI
Euglossa Latreille, 1802
Euglossa moronei† Engel, 1999: 4 [Fossil in Dominican amber]
Paleoeuglossa† Poinar, 1999
Paleoeuglossa melissiflora† Poinar, 1998: 29 [Fossil in Dominican amber]
Tribe EXOMALOPSINI
Anthophorula Cockerell, 1898
Anthophorula (Anthophorula) persephone† Engel in Engel et al., 2012: 5 [Fossil in Dominican amber]
Exomalopsis Spinola, 1853
Exomalopsis (Exomalopsis) analis Spinola, 1853. Genaro, 2007c: 250
Exomalopsis (Exomalopsis) bahamica Timberlake, 1980. Genaro, 2007c: 250
Exomalopsis (Exomalopsis) pulchella Cresson, 1865. Genaro, 2007c: 251
Exomalopsis (Exomalopsis) similis Cresson, 1865. Genaro, 2007c: 251
Exomalopsis sp.—Genaro (2007: 251)
Tribe MELECTINI
Xeromelecta Linsley, 1939
Xeromelecta (Nesomelecta) haitensis Michener, 1948. Genaro, 2007c: 251
Tribe MELIPONINI
Proplebeia† Michener, 1999
Proplebeia abdita† Greco & Engel in Greco et al., 2011: 491 [Fossil in Dominican amber]
Proplebeia dominicana† (Wille & Chandler, 1964: 188); Michener, 1982: 39; Michener & Poinar, 1997: 360; Camargo et al.,
2000: 3 [Fossil in Dominican amber]
Proplebeia vetusta† Camargo et al., 2000: 11 [Fossil in Dominican amber]
Proplebeia tantilla† Camargo et al., 2000: 13 [Fossil in Dominican amber]
Subfamily NOMADINAE
Tribe EPEOLINI
Triepeolus Robertson, 1901
Triepeolus nisibonensis Genaro, 2001b: 1033. Genaro, 2007c: 251 *DR > TL: Prov. La Altagracia, Nisibón
Triepeolus danieli Genaro, 2014: 23 *DR > TL: Cueva del Puente, Guaraguao, Parque Nacional del Este.
Triepeolus victori Genaro, 1998a: 92. Genaro, 2007c: 251 *DR > TL: Prov. Barahona, near Filipinas, Larimar mine
Tribe NOMADINI
Nomada Scopoli, 1770
Nomada krugii Cresson, 1878. Genaro, 2007c: 250
Nomada pilipes (Cresson, 1865). Genaro, 2007c: 250
Nomada sp. n.—Genaro (2007c: 250)

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Subfamily XYLOCOPINAE
Tribe CERATININI
Ceratina Latreille, 1802
Ceratina (Ceratinula) guarnacciana Genaro, 1998b: 97. Genaro, 2007c: 252 *DR > TL: Los Llanos, San Pedro de Macorís
Ceratina (Ceratinula) polita Friese, 1902: 198. Genaro, 1998b: 96; Genaro, 2007c: 251 *H > TL: Haïti
Ceratina n. sp.—Genaro (2007c: 251)
Tribe XYLOCOPINI
Xylocopa Latreille, 1802
Xylocopa (Neoxylocopa) mordax F. Smith, 1874. Ashmead, 1900: 302; Friese, 1902: 198; Genaro, 2007c: 252

Order LEPIDOPTERA
General classification follows Heppner (1995).

Taxonomic summary:
Suborder Glossata
Superfamily Nepticuloidea
Family Opostegidae
Superfamily Incurvarioidea
Family Heliozelidae
Superfamily Carposinoidea
Family Carposinidae
Superfamily Tineoidea
Family Acrolophidae
Family Tineidae
Family Psychidae
Family Gracillaridae
Superfamily Urodoidea
Family Urodidae
Superfamily Gelechioidea
Family Oecophoridae
Family Gelechiidae
Family Choreutidae
Family Batrachedridae
Family Agonoxenidae
Family Cosmopterigidae
Family Stenomidae
Superfamily Yponomeutoidea
Family Heliodinidae
Family Plutellidae
Family Yponomeutidae
Family Lyonetiidae
Superfamily Hyblaeoidea
Family Hyblaeidae
Superfamily Pyraloidea
Family Thyrididae
Family Crambidae
Family Pyralidae
Superfamily Pterophoroidea
Family Pterophoridae
Superfamily Cossoidea
Family Cossidae
Family Epipyropidae
Family Limacodidae

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Superfamily Castnioidea
Family Castniidae
Superfamily Tortricoidea
Family Tortricidae
Superfamily Geometroidea
Family Uraniidae
Family Geometridae
Family Sematuridae
Superfamily Hesperioidea
Family Hesperiidae
Superfamily Papilionoidea
Family Papilionidae
Family Pieridae
Family Lycaenidae
Family Riodinidae
Family Nymphalidae
Superfamily Bombycoidea
Family Saturniidae
Superfamily Sphingoidea
Family Sphingidae
Superfamily Noctuoidea
Family Notodontidae
Family Erebidae
Family Noctuidae
Superfamily Lasiocampoidea
Family Lasiocampidae

Suborder GLOSSATA
Superfamily NEPTICULOIDEA
Family OPOSTEGIDAE
Pseudopostega Kozlov, 1985
Pseudopostega venticola (Walsingham, 1897: 140). Davis, 1989: 77

Superfamily INCURVARIOIDEA
Family HELIOZELIDAE
Heliozela Herrich-Schäffer, 1853
Heliozela ahenea Walsingham, 1897: 110. Forbes, 1930: 136; Heppner, 1984: 19

Superfamily CARPOSINOIDEA
Family CARPOSINIDAE
Carposina Herrich-Schäffer, 1853
Carposina (Hypopremna) bullata Meyrick, 1913. Davis, 1969: 37

Superfamily TINEOIDEA
Family ACROLOPHIDAE
Acrolophus Poey, 1832
Acrolophus arcasalis (Walker, 1858: 153). Walsingham, 1891: 515; Walsingham, 1897: 173; Heppner, 1984: 19
Acrolophus australis (Walsingham, 1897: 166). Forbes, 1930: 153; Heppner, 1984: 19
Acrolophus indecora (Walker, 1863: 5). Heppner, 1984: 20
=Acrolophus mimasalis (Walker, 1858: 154). Walsingham, 1891: 515; Walsingham, 1897: 173; Forbes, 1930: 163; Heppner,
1984: 20
=Tachasara languidalis (Walker, 1865). Heppner, 1984: 20
Acrolophus occultum (Walsingham, 1897: 169). Heppner, 1984: 20
Acrolophus sp.†—Grimaldi & Engel (2005: 575); Sohn et al., 2012: 30 [Fossil in Dominican amber]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 319
Acrolophus umbratipalpis (Walsingham, 1891: 511). Walsingham, 1897: 170; Heppner, 1984: 20

Family TINEIDAE
Lepyrotica Meyrick, 1921
Lepyrotica fragilella (Walsingham, 1897: 161). Heppner, 1984: 21 *H > TL: Port-au-Prince
Opogona Zeller, 1853
Opogona simplex (Walsingham, 1897: 141). Heppner, 1984: 22
Phereoeca Hinton & Bradley, 1956
Phereoeca uterella (Walsingham, 1897). Flores Sasso & Prieto Vicioso, 2016: 77
Protodarcia Forbes, 1931
Protodarcia tischeriella (Walsingham, 1897: 161). Forbes, 1930: 156; Heppner, 1984: 23
Tinea Linnaeus, 1758
Tinea cretella Walsingham, 1897: 160. Heppner, 1984: 23
Tinea familiaris Zeller, 1877. Walsingham, 1897: 164; Forbes, 1930: 159

Family PSYCHIDAE
Subfamily OIKETICINAE
Oiketicus Guilding, 1827
Oiketicus kirbyi Guilding, 1827. Santoro, 1960: 33; Marcano, 1964: 122
Subfamily PENESTOGLOSSINAE
Pterogyne Davis, 1975
Pterogyne insularis Davis, 1975: 11. Heppner, 1984: 24
Subfamily PSYCHINAE
Cryptothelea Duncan & Westwood, 1841
Cryptothelea watsoni (Jones, 1923). Davis, 1964: 52; Davis, 1975: 25; Heppner, 1984: 24
Lumacra Davis, 1964
Lumacra haitiensis Davis, 1964: 67. Davis, 1975: 27; Heppner, 1984: 24
Paucivena Davis, 1975
Paucivena hispaniolae Davis, 1975: 15. Heppner, 1984: 24
Thyridopteryx Stephens, 1835
Thyridopteryx ephemeraeformis (Haworth, 1803). Davis, 1975: 29

Family GRACILLARIDAE
Key reference: De Prins and De Prins (2005).
Subfamily GRACILLARIINAE
Acrocercops Wallengren, 1881
Acrocercops undifraga Meyrick, 1931: 47. Busck, 1933: 174; Heppner, 1984: 26
Dialectica Walsingham, 1897
Dialectica permixtella Walsingham, 1897: 152. Heppner, 1984: 26; De Prins & De Prins, 2005: 174
Dialectica sanctaecrucis Walsingham, 1897. Busck, 1933: 173; De Prins & De Prins, 2005: 174
Eucosmophora Walsingham, 1897
Eucosmophora dives Walsingham, 1897. Davis & Wagner, 2005: 24
Parectopa Clemens, 1860
Parectopa pulverella (Walsingham, 1897: 153). Forbes, 1930: 143; Heppner, 1984: 26
Parectopa undosa (Walsingham, 1897: 153). Forbes, 1930: 143; Heppner, 1984: 26
Subfamily PHYLLOCNISTINAE
Phyllocnistis Zeller, 1848
Phyllocnistis citrella Stainton, 1856. Zuluaga & Mesa, 2000: 197; De Prins & De Prins, 2005: 250 [Introduced]

Superfamily URODOIDEA
Family URODIDAE
Urodus Herrich-Schäffer, 1858
Urodus mirella (Möschler, 1890). Davis, 1986: 90

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Superfamily GELECHIOIDEA
Family OECOPHORIDAE
Subfamily DEPRESSARIINAE
Gonionota Zeller, 1877
Gonionota rosacea (Forbes, 1931: 379). Clarke, 1971: 31; Heppner, 1984: 28
Subfamily ETHMIINAE
Ethmia Hübner, 1819
Ethmia abraxasella abraxasella (Walker, 1864: 1016). Walsingham, 1891: 528; Walsingham, 1897: 90; Forbes, 1930: 134;
Forbes, 1931: 379; Powell, 1973: 125; Heppner, 1984: 29
Ethmia confusella (Walker, 1863: 531). Walsingham, 1891: 527; Walsingham, 1897: 89; Busck, 1933: 163; Powell, 1973: 162;
Heppner, 1984: 30
=Cryptolechia strigosella Walker, 1864: 710. Heppner, 1984: 30
Ethmia kirbyi (Möschler, 1890). Powell, 1973: 118
Ethmia nivosella (Walker, 1864: 817). Walsingham, 1897: 114; Powell, 1973: 129
Ethmia notatella (Walker, 1863: 536). Walsingham, 1891: 528; Walsingham, 1897: 90; Forbes, 1930: 133; Forbes, 1931: 379;
Powell, 1973: 154; Heppner, 1984: 30
Ethmia paucella (Walker, 1863: 530). Walsingham, 1891: 527; Walsingham, 1897: 90; Powell, 1973: 156; Heppner, 1984: 30
*H, *DR > TL: St. Domingo
Subfamily STENOMATINAE
Cerconota Meyrick, 1915
Cerconota anonella (Sepp, 1852). SEA, 1999: 11

Family GELECHIIDAE
Subfamily ANACAMPSINAE
Pectinophora Busck, 1917
Pectinophora gossypiella (Saunders, 1844). Wolcott, 1927: 299; Smith & Audant, 1930: 974; Forbes, 1931: 378 [Introduced]
Sitotroga Heinemann, 1870
Sitotroga cerealella (Olivier, 1789). Wolcott, 1927: 297 [Introduced]
Subfamily DICHOMERINAE
Cymotricha Meyrick, 1923
Cymotricha melissia (Walsingham, 1911). Abud, 1981: 34; Schmutterer, 1990: 505
Dichomeris Hübner, 1818
Dichomeris sp.—Marcano (1981: 34) [As Trichotaphe]
Subfamily GELECHIINAE
Aristotelia Hübner, 1825
Aristotelia penicillata (Walsingham, 1897: 70). Heppner, 1984: 45; Forbes, 1930: 117
Aristotelia pudibundella (Zeller, 1873). Walsingham, 1897: 66; Forbes, 1930: 116
Aristotelia trossulella Walsingham, 1897: 67. Heppner, 1984: 45
Keiferia Busck, 1939
Keiferia lycopersicella (Walsingham, 1897). Smith & Audant, 1930: 977; Schmutterer, 1990: 561 [Introduced]
Phthorimaea Meyrick, 1902
Phthorimaea operculella (Zeller, 1873). Wolcott, 1927: 297; Smith & Audant, 1930: 975; Schmutterer, 1990: 549
Recurvaria Haworth, 1828
Recurvaria kittella (Walsingham, 1897: 65). Heppner, 1984: 47 *H > TL: Port-au-Prince
Stegasta Meyrick, 1904
Stegasta capitella (Fabricius, 1794). Forbes, 1930: 129; Heppner, 1984: 47
Telphusa Chambers, 1872
Telphusa perspicua (Walsingham, 1897: 72). Forbes, 1931: 372; Heppner, 1984: 47 *H > TL: Cap Haitien
Thiotricha Meyrick, 1886
Thiotricha godmani (Walsingham, 1892). Walsingham, 1897: 79
Tildenia Povolny, 1967
Tildenia gudmannella (Walsingham, 1897: 77). Forbes, 1930: 126
Tuta Kieffer & Jörgensen, 1910
Tuta absoluta (Meyrick, 1917). Verheggen & Fontus, 2019: 349 [Introduced]

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 321
Family CHOREUTIDAE
Subfamily CHOREUTINAE
Hemerophila Hübner, 1817
Hemerophila biferana Walker, 1863: 418. Walsingham, 1891: 529; Walsingham, 1897: 120; Walsingham, 1909–1915: 316;
Heppner, 1995: 115
Tortyra Walker, 1863
Tortyra auriferalis Walker, 1863. Walsingham, 1891: 528; Walsingham, 1897: 121; Heppner, 1995: 115

Family BRATRACHEDRIDAE
Subfamily BRATRACHEDINAE
Homaledra Busck, 1900
Homaledra sabalella (Chambers, 1880). Wolcott, 1927: 304; Smith & Audant, 1930: 977; Schmutterer, 1990: 134

Family AGONOXENIDAE
Pammeces Zeller, 1863
Pammeces lithochroma Walsingham, 1897: 103. Heppner, 1984: 43 *DR > TL: Puerto Plata
Pammeces picticornis (Walsingham, 1897: 102). Forbes, 1930: 138; Vives Moreno, 1988: 104; Becker, 1999: 160; Baldizzone
et al., 2006: 136

Family COSMOPTERIGIDAE
Subfamily COSMOPTERIGINAE
Anoncia Clarke, 1941
Anoncia?sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]
Cosmopterix Hübner, 1825
Cosmopterix abnormalis Walsingham, 1897: 106. Heppner, 1984: 43; Koster, 2010: 282
Cosmopterix attenuatella (Walker, 1864). Walsingham, 1897: 105; Koster, 2010: 566
Cosmopterix interfracta Meyrick, 1922. Koster, 2010: 349
Cosmopterix magophila Meyrick, 1919. Koster, 2010: 365
Pyroderces Herrich-Schäffer, 1853
Pyroderces rileyi (Walsingham, 1882). Wolcott, 1927: 303; Forbes, 1930: 10; Schmutterer, 1990: 62 [Introduced]
=Pyroderces stigmatophora (Walsingham, 1897: 104)
Pyroderces? sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]

Family STENOMIDAE
Mothonica Walsingham, 1912
Mothonica ocellea Forbes, 1930. Kisliuk & Coole, 1933: 3; Duckworth, 1969: 7

Superfamily YPONOMEUTOIDEA
Family HELIODINIDAE
Embola Walsingham, 1909
Embola melanotela Hsu in Hsu & Powell, 2005: 67

Family PLUTELLIDAE
Plutella Schrank, 1802
Plutella xylostella (Linnaeus, 1758). Schmutterer, 1990: 418

Family YPONOMEUTIDAE
Subfamily ATTEVINAE
Atteva Walker, 1854
Atteva fulviguttata (Zeller, 1873). Becker, 2009: 252
Atteva rawlinsi Becker, 2009: 351 *DR > TL: Barahona, 9.2 km NW Paraiso, confluence of Río Nizao and Río Cortico, 230 m
Atteva siderea (Walsingham, 1892: 533). Walsingham, 1897: 113; Becker, 2009: 352

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Family LYONETIIDAE
Subfamily BUCCULATRIGINAE
Perileucoptera Silvestri, 1943
Perileucoptera coffeella (Guérin-Méneville, 1842). Wolcott, 1927: 305; Smith & Audant, 1930: 974
Subfamily CEMIOSTOMINAE
Bucculatrix Zeller, 1839
Bucculatrix thurberiella (Busck, 1914). Schmutterer, 1990: 318

Superfamily HYBLAEOIDEA
Family HYBLAEIDAE
Hyblaea Fabricius, 1793
Hyblaea puera (Cramer, 1777). Walker, 1857: 979; Marcano, 1986: 2/86; SEA, 1999: 26

Superfamily PYRALOIDEA
Family THYRIDIDAE
Subfamily STRIGLININAE
Banisia Walker, 1863
Banisia furva illicta Whalley, 1976: 165

Family CRAMBIDAE
Subfamily ACENTROPINAE
Chrysendeton Grote, 1881
Chrysendeton claudialis (Walker, 1859). Hampson, 1897: 151; Heppner, 1995: 49
Petrophila Guilding, 1830
Petrophila gratalis (Walker, 1865d: 1335). Hampson, 1897: 149; Heppner, 1995: 50
Petrophila insulalis (Walker, 1862). Walker, 1865d: 1333; Hampson, 1897: 150; Heppner, 1995: 50
Petrophila malcusalis (Schaus, 1924). Heppner, 1995: 50
Subfamily CRAMBINAE
Diatraea Guilding, 1828
Diatraea crambidoides (Grote, 1880). Marcano, 1964: 93
Diatraea saccharalis (Fabricius, 1794). Druce, 1891: 296; Wolcott, 1927: 292; Santoro, 1960: 138; Schmutterer, 1990: 46;
Heppner, 1995: 37
=Crambus leucaniella Walker, 1863. Heppner, 1995: 37
Fissicrambus Bleszynski, 1963
Fissicrambus fissiradiellus (Walker, 1863: 160). Druce, 1891: 291; Heppner, 1995: 40
=Crambus curtellus Walker, 1863. Heppner, 1995: 40
Fissicrambus haytiellus (Zincken, 1821). Bleszinski, 1967: 106; Heppner, 1995: 40
Mesolia Ragonot, 1889
Mesolia plurimella (Walker, 1863: 162). Druce, 1891: 297; Heppner, 1995: 34
Microcrambus Bleszynski, 1963
Microcrambus discludellus (Möschler, 1890). Bleszinski, 1967: 107; Heppner, 1995: 41
=Crambus domingellus (Schaus, 1922). Schaus, 1940: 398; Heppner, 1995: 41
Microcrambus francescellus (Schaus, 1922). Bleszinski, 1967: 107; Heppner, 1995: 41
Microcrambus podalirius Bleszynski, 1967: 87. Heppner, 1995: 41 *DR > TL: San Francesco Mts.
Prionapteryx Stephens, 1834
Prionapteryx eugraphis (Walker, 1863: 188). Heppner, 1995: 34
Subfamily EVERGESTINAE
Trischistognatha Warren, 1892
Trischistognatha pyrenealis (Walker, 1859b: 580). Heppner, 1995: 53
Subfamily GLAPHYRIINAE
Alatuncusia Amsel, 1956
Alatuncusia canalis (Walker, 1866: 1717). Heppner, 1995: 45
Dichogama Lederer, 1863
Dichogama decoralis (Walker, 1865d: 1380). Heppner, 1995: 45

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 323
Hellula Guenée, 1854
Hellula phidilealis (Walker, 1859). SEA, 1999: 27
Hellula rogatalis (Hulst, 1886). Domínguez, 1997: 71
Schacontia Dyar, 1914
Schacontia rasa Solis & Goldstein in Goldstein et al., 2013: 61
Schacontia themis Solis & Goldstein in Goldstein et al., 2013: 54
Subfamily MIDILINAE
Odilla Schaus, 1940
Odilla noralis Schaus, 1940. Hayden, 2012: 313
Subfamily ODONTIINAE
Cliniodes Guenée, 1854
Cliniodes muralis Hayden, 2012: 262
Cliniodes nacrealis Munroe, 1964: 532. Heppner, 1995: 52; Hayden, 2012: 290
Mimoschinia Warren, 1892
Mimoschinia rufofascialis (Stephens, 1834). Heppner, 1995: 52
=Botys perviana Walker, 1865c: 804. Heppner, 1995: 52
=Botys thalialis Walker, 1859b: 582. Heppner, 1995: 52
=Pyralis gelidalis Walker, 1865d: 1229. Heppner, 1995: 52
Subfamily PYRAUSTINAE S. L.
Achyra Guenée, 1849
Achyra bifidalis (Fabricius, 1794). Heppner, 1995: 55
=Phlyctaenodes inornatalis (Walker, 1865d: 1456). Heppner, 1995: 55
Achyra rantalis (Guenée, 1854). Capps, 1967: 47; Heppner, 1995: 55
=Ebulea murcialis Walker, 1859b: 746. Heppner, 1995: 55
=Scopula diotimealis Walker, 1859. Heppner, 1995: 55
Agathodes Guenée, 1854
Agathodes designalis Guenée, 1854. Domínguez, 1997: 71
Apogeshna Munroe, 1956
Apogeshna acestealis (Walker, 1859a: 352). Heppner, 1995: 64
Arthromastix Warren, 1890
Arthromastix lauralis Walker, 1859a: 358. Druce, 1891: 224; Heppner, 1995: 74
Asciodes Guenée, 1854
Asciodes gordialis Guenée, 1854. Heppner, 1995: 74
=Scoparia quietalis Walker, 1859c: 825. Heppner, 1995: 74
Azochis Walker, 1859
Azochis rufidiscalis Hampson, 1904. Domínguez, 1997: 71
Bicilia Amsel, 1956
Bicilia iarchasalis (Walker, 1859c: 983). Heppner, 1995: 76
=Pyralis differalis Walker, 1865d: 1228. Heppner, 1995: 76
Bocchoropsis Amsel, 1956
Bocchoropsis pharaxalis Walker, 1859. Santoro, 1960: 100. SEA, 1999: 126
Ceratoclasis Lederer, 1863
Ceratoclasis delimitalis (Guenée, 1854). Walker, 1859b: 571; Druce, 1891: 244; Heppner, 1995: 74
Conchylodes Guenée, 1854
Conchylodes diphteralis (Geyer, 1832). Walker, 1859a: 494; Schaus, 1940: 345
Conchylodes hebraealis Guenée, 1854. Walker, 1859a: 494; Heppner, 1995: 75
Conchylodes hedonialis (Walker, 1859a: 470). Heppner, 1995: 75
Cyclocena Möschler, 1890
Cyclocena lelex (Cramer, 1777). Perez-Gelabert, 2008: 252
Desmia Westwood, 1832
Desmia funeralis (Hübner, 1796). Domínguez, 1997: 71
Desmia niveiciliata E. Hering, 1906. Heppner, 1995: 66
Desmia ploralis (Guenée, 1854). Domínguez, 1997: 71
Desmia ufeus (Cramer, 1777). Heppner, 1995: 66

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=Desmia divisalis Walker, 1865d: 1292. Heppner, 1995: 66
Diacme Warren, 1892
Diacme adipaloides (Grote & Robinson, 1867). Domínguez, 1997: 71
Diaphania Hübner, 1818
Diaphania antillia Munroe, 1960: 165. Heppner, 1995: 68
Diaphania busccalis (Schaus, 1920: 216). Heppner, 1995: 68
Diaphania costata (Fabricius, 1794) [By synonymy of species below]
=Margaronia imitalis (Guenée, 1854). Walker, 1859b: 520; Schaus, 1940: 357
Diaphania elegans (Möschler, 1890). Domínguez, 1997: 70
Diaphania glauculalis (Guenée, 1854). Heppner, 1995: 68
=Margarodes lustralis Guenée, 1854. Walker, 1859b: 520; Heppner, 1995: 69
Diaphania hyalinata (Linnaeus, 1767). Wolcott, 1927: 291; Schmutterer, 1990: 436; Domínguez, 1997: 71
Diaphania indica (Saunders, 1851). Domínguez, 1997: 71
Diaphania lualis (Herrich-Schäffer, 1871). Domínguez, 1997: 71
Diaphania nitidalis (Cramer, 1781). Walker, 1859b: 511; Schmutterer, 1990: 442
Diaphania quadristigmalis (Guenée, 1854). Walker, 1859b: 519; Abud Antún, 1981: 26/81
Diaphantania Möschler, 1890
Diaphantania candacalis (Felder & Rogenhofer, 1875). Heppner, 1995: 76
Diaphantania ceresalis (Walker, 1859b: 581). Heppner, 1995: 76
Epicorsia Hübner, 1818
Epicorsia oedipodalis (Guenée, 1854). Walker, 1859b: 569; Heppner, 1995: 54
Ercta Walker, 1859
Ercta vittata (Fabricius, 1794). Heppner, 1995: 66
=Ercta tipulalis Walker, 1859a: 426. Heppner, 1995: 66
Eulepte Hübner, 1825
Eulepte gastralis (Guenée, 1854). Walker, 1859b: 570; Heppner, 1995: 72
Glyphodes Guenée, 1854
Glyphodes sibillalis sibillalis Walker, 1859a: 495. Heppner, 1995: 69
Helvibotys Munroe, 1976
Helvibotys carnifex (Felder & Rogenhofer, 1875). Schaus, 1940: 376; Heppner, 1995: 57
Helvibotys panopealis (Walker, 1859). Heppner, 1995: 57
=Botys coecilialis Walker, 1859b: 581. Heppner, 1995: 57
=Rhodaria catenalis Walker, 1865d: 1282. Heppner, 1995: 57
Herpetogramma Lederer, 1863
Herpetogramma agavealis (Walker, 1859b: 574). Heppner, 1995: 61
Herpetogramma antillalis (Schaus, 1920: 218). Heppner, 1995: 61
Herpetogramma bipunctalis (Fabricius, 1794). Wolcott, 1927: 290; Heppner, 1995: 61; SEA, 1999: 57
=Botys lycialis Walker, 1859b: 572. Heppner, 1995: 61
Herpetogramma phaeopteralis (Guenée, 1854). Heppner, 1995: 61
=Botys vecordalis Guenée, 1854. Walker, 1859b: 571; Heppner, 1995: 61
Hileithia Snellen, 1875
Hileithia magualis (Guenée, 1854). Heppner, 1995: 62
=Asopia curtalis Walker, 1865d: 1305. Heppner, 1995: 62
=Botys belusalis Walker, 1859b: 582. Heppner, 1995: 62
=Samea medealis Walker, 1859a: 353. Heppner, 1995: 62
Hymenia Hübner, 1825
Hymenia perspectalis (Hübner, 1796). Walker, 1859a: 395; Domínguez, 1997: 71
Lamprosema Hübner, 1823
Lamprosema pelealis (Walker, 1859a: 339). Heppner, 1995: 63
Lygropia Lederer, 1863
Lygropia imparalis (Walker, 1865d: 1300). Heppner, 1995: 59
=Physematia rotundalis Felder & Rogenhofer, 1875. Heppner, 1995: 59
Lygropia tripunctata (Fabricius, 1794). Wolcott, 1927: 291; Heppner, 1995: 59
=Botys campalis Guenée, 1854. Walker, 1859b: 570; Heppner, 1995: 59

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 325
Mabra Moore, 1885
Mabra russoi Schaus in Russo, 1940: 139. Heppner, 1995: 78
Marasmia Lederer, 1863
Marasmia cochrusalis (Walker, 1859c: 959). Heppner, 1995: 75
Marasmia trapezalis (Guenée, 1854). Wolcott, 1927: 290; Schmutterer, 1990: 100; Heppner, 1995: 75
=Botys creonalis Walker, 1859b: 579. Heppner, 1995: 75
Maruca Walker, 1859
Maruca testulalis (Geyer, 1832). Santoro, 1960: 222; Schmutterer, 1990: 355
Maruca vitrata (Fabricius, 1787). Serra, 2005: 124
Microthyris Lederer, 1863
Microthyris anormalis (Guenée, 1854). Heppner, 1995: 59
=Botys helcitalis Walker, 1859b: 574. Heppner, 1995: 59
Neoleucinodes Capps, 1948
Neoleucinodes imperialis (Guenée, 1854). Walker, 1859a: 393; Heppner, 1995: 63
Nomophila Hübner, 1825
Nomophila nearctica Munroe, 1973: 190
Omiodes Guenée, 1854
Omiodes humeralis Guenée, 1854. Walker, 1859b: 740; Heppner, 1995: 70
=Botys peleusalis Walker, 1859b: 575. Heppner, 1995: 70
=Botys gnomalis Walker, 1859b: 580. Heppner, 1995: 70
Omiodes indicata (Fabricius, 1775). Heppner, 1995: 69
=Botys connexalis Walker, 1865d: 1394. Heppner, 1995: 69
Omiodes simialis Guenée, 1854. Druce, 1891: 253
Ommatospila Lederer, 1863
Ommatospila descriptalis (Walker, 1865d: 1222). Heppner, 1995: 73
Ommatospila marcaeusalis (Walker, 1859). Heppner, 1995: 73
=Leucinodes venustalis (Walker, 1865d: 1312). Heppner, 1995: 73
Palpita Hübner, 1808
Palpita flegia (Cramer, 1777). Druce, 1891: 224; Domínguez, 1997: 71
Palpita quadistrigmalis (Guenée, 1854). Domínguez, 1997: 71
Palpita kimballi Munroe, 1959. Domínguez, 1997: 71
Palpita viettei Munroe, 1959: 642
Pantographa Lederer, 1863
Pantographa prorogata (Hampson, 1912). Schmutterer, 1990: 169
Phaedropsis Warren, 1890
Phaedropsis domingalis (Schaus, 1920). Heppner, 1995: 58
Phaedropsis hecalialis (Walker, 1859b: 573). Heppner, 1995: 59
Phaedropsis stictigramma (Hampson, 1912). Domínguez, 1997: 71
Pilemia Möschler, 1881
Pilemia periusalis (Walker, 1859). Wolcott, 1927: 291
Pleuroptya Meyrick, 1890
Pleuroptya silicalis (Guenée, 1854). Perez-Gelabert, 2008: 254
Polygrammodes Guenée, 1854
Polygrammodes elevata (Fabricius, 1794). SEA, 1999: 32
Proleucinodes Capps, 1948
Proleucinodes impuralis (Felder & Rogenhofer, 1875). Heppner, 1995: 63
Psara Snellen, 1875
Psara dryalis (Walker, 1859b: 573). Heppner, 1995: 74
Pyrausta Schrank, 1802
Pyrausta carnifex (Felder, 1860). Schaus, 1940: 376
Pyrausta tyralis (Guenée, 1854). Domínguez, 1997: 71
Salbia Guenée, 1854
Salbia haemorrhoidalis Guenée, 1854 [By synonymy of species below]
=Asopia dircealis Walker, 1859a: 365

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Samea Guenée, 1854
Samea multiplicalis (Guenée, 1854). Domínguez, 1997: 71
Sathria Lederer, 1863
Sathria internitalis (Guenée, 1854). Walker, 1859b: 761; Druce, 1891: 241; Heppner, 1995: 75
=Megaphysa serenalis Walker, 1865d: 1309. Heppner, 1995: 75
Spoladea Guenée, 1854
Spoladea recurvalis (Fabricius, 1775). Walker, 1859a: 396; Druce, 1891: 257
Syllepte Hübner, 1823
Syllepte opalisans (Felder & Rogenhofer, 1875). Heppner, 1995: 61
Syngamia Guenée, 1854
Syngamia florella (Stoll, 1781). Heppner, 2010: 113
Syngamia sp.—Schmutterer (1990: 48)
Synclera Lederer, 1863
Synclera jarbusalis (Walker, 1859a: 352). Heppner, 1995: 70
=Zebronia cottalis Walker, 1859b: 964. Heppner, 1995: 70
Terastia Guenée, 1854
Terastia meticulosalis Guenée, 1854. Walker, 1859a: 379; Druce, 1891: 215; Heppner, 1995: 72
Tomopteryx Philippi, 1873
Tomopteryx pterophoralis (Walker, 1865d: 1340). Walsingham, 1909–1915: 453; Heppner, 1995: 63
Udea Guenée, 1845
Udea secernalis (Möschler, 1890). Schaus, 1940: 343
Subfamily SCHOENOBIINAE
Carectocultus Blanchard, 1975
Carectocultus perstrialis (Hübner, 1825). Heppner, 1995: 41
=Crambus serriradiellus (Walker, 1863: 161). Heppner, 1995: 41
Subfamily SPILOMELINAE
Sufetula Walker, 1859
Sufetula diminutalis (Walker, 1866). Hayden, 2013: 10

Family PYRALIDAE
Subfamily CHRYSAUGINAE
Bonchis Walker, 1862
Bonchis munitalis (Lederer, 1863). Druce, 1891: 196; Heppner, 1995: 82
=Vurna instructalis Walker, 1865d: 1189. Heppner, 1995: 82
Carcha Walker, 1859
Carcha hersilialis Walker, 1859a: 282. Druce, 1891: 195; Heppner, 1995: 82
=Pyralis dispansalis Walker, 1865d: 1228. Heppner, 1995: 82
Murgisca Walker, 1863
Murgisca cervinalis Walker, 1863: 12. Heppner, 1995: 85
Parachmidia Hampson, 1896
Parachmidia fervidalis (Walker, 1865d: 1461). Heppner, 1995: 85
=Scopula prodigalis Walker, 1865d: 1461. Heppner, 1995: 85
Subfamily EPIPASCHIINAE
Deuterolita Lederer, 1863
Deuterolita ragonoti (Möschler, 1890). Alvarez, 1992: 12/92
Jocara Walker, 1863
Jocara fragilis Walker, 1863. Heppner, 1995: 91
Macalla Walker, 1859
Macalla thyrsisalis Walker, 1859. Howard & Solis, 1989: 473
Subfamily GALLERIINAE
Corcyra Ragonot, 1885
Corcyra cephalonica (Stainton, 1865). Smith & Audant, 1930: 978
Galleria Fabricius, 1798
Galleria mellonella (Linnaeus, 1758). Smith & Audant, 1930: 978; Heppner, 1995: 88

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 327
=Vindana obliquella Walker, 1866: 1706. Heppner, 1995: 88
Subfamily PHYCITINAE
Amegarthria Neunzig & Dow, 1993
Amegarthria cervicalis (Dyar, 1919). Neunzig, 2003: 7
Amyelois Amsel, 1956
Amyelois transitella (Walker, 1863). Heppner, 1995: 94
=Nephopteryx notatalis Walker, 1863. Heppner, 1995: 94
Anabasis Heinrich, 1956
Anabasis ochrodesma (Zeller, 1881). Neunzig, 2003: 7
Anadelosemia Dyar, 1919
Anadelosemia texanella (Hulst, 1892). Neunzig, 2003: 7
Ancylostomia Ragonot, 1893
Ancylostomia stercorea (Zeller, 1848). Heinrich, 1956: 95
Anegcephalesis Dyar, 1917
Anegcephalesis arctella (Ragonot, 1893). Neunzig, 2003: 7
Anypsipyla Dyar, 1914
Anypsipyla univitella Dyar, 1914. Neunzig, 2003: 7
Aptunga Heinrich, 1956
Aptunga culmenicola Neunzig, 1996: 783 *DR > TL: Independencia, 15 km NE Los Arroyos (Pedernales), summit of Sierra de
Baoruco, 2260 m
Aptunga setadebilia Neunzig, 1996: 777 *DR > TL: La Vega, 18 km SE Constanza, 2310 m
Aptunga vega Neunzig, 1996: 781 *DR > TL: La Vega, 23 km SE Constanza, 2225 m
Australephestiodes Neunzig, 1990
Australephestiodes stictella Hampson, 1901. Neunzig, 2003: 7
Bema Dyar, 1914
Bema neuricella (Zeller, 1848). Neunzig, 2003: 8
Cactoblastis Ragonot, 1901
Cactoblastis cactorum (Berg, 1885). Heppner, 2000: 20 [Introduced]
Caristanius Heinrich, 1956
Caristanius pellucidellus (Ragonot, 1888). Neunzig, 2003: 8
Caristanius tripartitus Neunzig, 1996: 799 *DR > TL: La Vega Prov., Hotel Montaña, ca. 520 m, 10 km NE Jarabacoa
Caudellia Dyar, 1904
Caudellia pilosa Neunzig, 1996: 785 *DR > TL: La Vega Prov., Hotel Montaña, ca. 520 m, 10 km NE Jarabacoa
Chararica Heinrich, 1956
Chararica circiimperfecta Neunzig, 1996: 775 *DR > TL: Independencia, 4 km S Los Pinos, Loma de Vientos, 475 m
Coptarthria Ragonot, 1893
Coptarthria dasypyga (Zeller, 1881). Neunzig, 2003: 8
Crocidomera Zeller, 1848
Crocidomera fissuralis (Walker, 1863: 58). Heinrich, 1956: 33; Heppner, 1995: 93
=Myelois adorea Felder & Rogenhofer, 1875. Heppner, 1995: 93
Crocidomera imitata Neunzig, 1990. Neunzig, 2003: 8
Cryptobables Zeller, 1848
Cryptobables sp.—Schmutterer (1990: 275)
Dasypyga Ragonot, 1887
Dasypyga independencia Neunzig, 1996: 793 *DR > TL: Independencia, 3 km ESE El Aguacate, north slope Sierra de
Bahoruco, 1980 m
Davara Walker, 1859
Davara caricae (Dyar, 1913). Heinrich, 1956: 74
Davara interjecta Heinrich, 1956: 75
Davara rufulella (Ragonot, 1888). Neunzig, 2003: 8
Dioryctria Zeller, 1846
Dioryctria amatella (Hulst, 1887). Domínguez, 1997: 71
Dioryctria dominguensis Neunzig, 1996: 791 *DR > TL: Independencia, 3 km ESE El Aguacate, north slope Sierra de
Bahoruco, 1980 m

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Dioryctria postmajorella Neunzig, 1996: 789 *DR > TL: Independencia, 3 km ESE El Aguacate, north slope Sierra de
Bahoruco, 1980 m
Ectomyelois Heinrich, 1956
Ectomyelois decolor (Zeller, 1881). Neunzig, 2003: 8
Ectomyelois muriscis (Dyar, 1914). Neunzig, 2003: 8
Elasmopalpus Blanchard, 1852
Elasmopalpus lignosellus (Zeller, 1848). Schmutterer, 1990: 99
Ephestia Guenée, 1845
Ephestia cautella (Walker, 1883). SEA, 1999: 128
Ephestia elutella (Hübner, 1796). SEA, 1999: 128
Ephestia kuhniella Zeller, 1879. SEA, 1999: 128
Erelieva Heinrich, 1956
Erelieva quantulella (Hulst, 1887). Heinrich, 1956: 308
Etiella Zeller, 1839
Etiella zinckenella (Treitschke, 1832). Santoro, 1960: 220; Neunzig, 2003: 8
Fundella Zeller, 1848
Fundella argentina Dyar, 1919. Heinrich, 1956: 61; Neunzig, 2003: 8
Fundella ignobilis Heinrich, 1945. Heinrich, 1956: 61; Neunzig, 2003: 8
Fundella pelluscens Zeller, 1848. Heinrich, 1956: 60; Neunzig, 2003: 9
Hemiptiloceroides Neunzig & Dow, 1993
Hemiptiloceroides deltus Neunzig & Dow, 1993. Neunzig, 2003: 9
Homeosoma Curtis, 1833
Homeosoma electella (Hulst, 1887). Abud, 1993: 3/93; Neunzig, 2003: 9
Hypargyria Ragonot, 1888
Hypargyria slosonella (Hulst, 1900). Neunzig, 2003: 9
Hypochalcia Hübner, 1825
“Hypochalcia” cervinistrigalis Walker, 1863: 45. Heppner, 1995: 104
Hypsipyla Ragonot, 1888
Hypsipyla grandella (Zeller, 1848). Neunzig, 2003: 9 [Introduced]
Lascelina Heinrich, 1956
Lascelina pedernalensis Neunzig, 1996: 796 *DR > TL: Pedernales, along Rio Mulito, 13 km N Pedernales, 230 m
Lipographis Ragonot, 1887
Lipographis subosseella Hulst, 1892. Neunzig, 2003: 9
Moodna Hulst, 1890
Moodna antilleana Neunzig, 1996: 783 *DR > TL: Dajabón Prov., 13 km S Loma de Cabrera, ca. 400 m
Moodnopsis Dyar, 1914
Moodnopsis portoricensis Heinrich, 1956. Neunzig, 2003: 9
Nefundella Neunzig, 1986
Nefundella munroei Neunzig, 2003: 10 *DR > TL: San Juan, 8 km NE Vallejuelo, 690 m
Olyca Walker, 1857
Olyca phryganoides Walker, 1857: 726. Heinrich, 1956: 243; Heppner, 1995: 101
Oncolabis Zeller, 1848
Oncolabis anticella Zeller, 1848. Neunzig, 2003: 9
Oryctometopia Ragonot, 1887
Oryctometopia fossulatella Ragonot, 1888. Neunzig, 2003: 9
Ozamia Hampson, 1901
Ozamia lucidalis (Walker, 1863: 39). Heppner, 1995: 101
Ozamia plagata Neunzig, 1996: 787 *DR > TL: Pedernales, 14.5 km N Cabo Rojo, 165 m
Peadus Heinrich, 1956
Peadus burdettellus (Schaus, 1913). Neunzig, 2003: 9
Phycitodes Hampson, 1917
Phycitodes olivaceella (Ragonot, 1888). Neunzig, 2003: 9
Plodia Guenée, 1845
Plodia interpunctella (Hübner, 1813). Marcano, 1964: 100

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 329
Salebria Zeller, 1846
Salebria infusella (Zeller, 1848). Walker, 1866: 1714; Heppner, 1995: 104
Stylopalpia Hampson, 1901
Stylopalpia lunigerella Hampson, 1901. Neunzig, 2003: 9
Ufa Walker, 1863
Ufa rubedinella (Zeller, 1848). Druce, 1891: 283; Heinrich, 1956: 171; Heppner, 1995: 97
=Acrobasis translucida Walker, 1863. Heppner, 1995: 97
=Nephopteryx rufescentalis Walker, 1863. Heppner, 1995: 97
Varneria Dyar, 1904
Varneria albiornatella Neunzig, 1996: 787 *DR > TL: Peravia, 3 km SW La Nuez, tributary to Río Las Cuevas, 1870 m
Zamagiria Dyar, 1914
Zamagiria laidion (Zeller, 1881). Neunzig, 2003: 9
Zamagiria rawlinsi Neunzig, 1996: 797 *DR > TL: Azua, east side of crest Sierra Martin Garcia, 7 km WNW Barrero, 860 m
Subfamily PYRALINAE
Pseudasopia Grote, 1873
Pseudasopia intermedialis (Walker, 1862). Heppner, 1995: 80
Pyralis Linnaeus, 1758
Pyralis manihotalis Guenée, 1854. Druce, 1891: 550; Heppner, 1995: 81
=Pyralis vetusalis Walker, 1859c: 891. Heppner, 1995: 81

Superfamily PTEROPHOROIDEA
Family PTEROPHORIDAE
Adaina Tutt, 1905
Adaina ipomoeae Bigot & Etienne, 2009. Matthews & Gielis, 2011: 1
Adaina thomae (Zeller, 1877). Walsingham, 1897: 60
Exelastis Meyrick, 1907
Exelastis montischristi (Walsingham, 1897: 59). Forbes, 1930: 77; Heppner, 1995: 107
Leioptilus Wallengren, 1860
Leioptilus agraphodactylus (Walker, 1864: 941). Walsingham, 1897: 61; Heppner, 1995: 108
Leioptilus inquinatus (Zeller, 1893). Walsingham, 1897: 59; Walsingham, 1909–1915: 446; Forbes, 1930: 77
Oidaematophorus Wallengren, 1862
Oidaematophorus ossipellis (Walsingham, 1897: 60). Heppner, 1995: 108 *DR > TL: Puerto Plata

Superfamily COSSOIDEA
Family COSSIDAE
Subfamily ZEUZERINAE
Psychonoctua Grote, 1866
Psychonoctua personalis Grote, 1866. Marcano, 1964: 123

Family EPIPYROPIDAE
Fulgoraecia Newman, 1851
Fulgoraecia cucullata (Heinrich, 1931: 2). Heppner, 1995: 127 *H > TL: Port-au-Prince

Family LIMACODIDAE
Heuretes Grote & Robinson, 1868
Heuretes daidaleos Epstein & Miller, 1990: 711. Heppner, 1995: 130 *DR > TL: Dajabón Prov., Rio Massacre, Balneario Don
Miguel, 7 km SW Dajabón, 40 m elevation
Heuretes divisus Epstein & Miller, 1990: 712. Heppner, 1995: 130 *DR > TL: Pedernales Prov., La Abeja, 38 km NNW Cabo
Rojo, 1250 m

Superfamily CASTNIOIDEA
Family CASTNIIDAE
Subfamily CASTNIINAE
Ircila Houlbert, 1918

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Ircila hecate (Herrich-Schäffer, 1854). Westwood, 1877: 174 *H, *DR > TL: Insula Haiti

Superfamily TORTRICOIDEA
Family TORTRICIDAE
Key reference: Brown (2005).
Subfamily CHLIDANOTINAE
Ardeutica Meyrick, 1913
Ardeutica melidora Razowski, 1984. Razowski, 1999a: 312
Ardeutica sphenobathra (Meyrick, 1917). Razowski, 1999a: 312
Polyortha Dognin, 1905
Polyortha naevifera Razowski, 1984. Razowski, 1999a: 312
Polyortha nigriguttata Walsingham, 1914. Razowski, 1999a: 312
Polyvena† Poinar & Brown, 1993
Polyvena horatis† Poinar & Brown, 1993: 28 [Fossil in Dominican amber]
Subfamily OLETHREUTINAE
Bactra Stephens, 1834
Bactra philocherda Diakonoff, 1964. Perez-Gelabert, 2008: 258
Cacocharis Walsingham, 1892
Cacocharis cymotoma (Meyrick, 1917). Perez-Gelabert, 2008: 258 [Misspelled Caccocharis]; Brown, 2008: 536
Crocidosema Zeller, 1847
Crocidosema longipalpana (Möschler, 1891) [By synonymy of species below]
=Crocidosema litchivora Baixeras in Brown et al., 2002: 319
Crocidosema plebejana Zeller, 1847. Perez-Gelabert, 2008: 258 (Based on specimen in USNM collection)
Cryptaspasma Walsingham, 1900
Cryptaspasma (Anaphorodes) bipenicilla J. Brown & R. Brown, 2004: 289
Cryptaspasma lugubris (Felder, 1875). Razowski, 1999a: 313
Cydia Hübner, 1825
Cydia ingens (Heinrich, 1926). Perez-Gelabert, 2008: 258
Cydia membrosa (Heinrich, 1926). Perez-Gelabert, 2008: 258
Epiblema Hübner, 1825
Epiblema strenuana (Walker, 1863). Walsingham, 1897: 126; Forbes, 1930: 90
Episimus Walsingham, 1892
Episimus guiana (Busck, 1913). Razowski, 1999a: 313
Episimus transferrana (Walker, 1863). Perez-Gelabert, 2008: 258
Gymnandrosoma Dyar, 1904
Gymnandrosoma aurantianum Costa Lima, 1927. SEA, 1999: 112
=Ecdytolopha torticornis (Meyrick, 1931). Razowski, 1999a: 314
Gymnandrosoma leucothorax Adamski & Brown, 2001: 77
Gymnandrosoma trachycerus Forbes, 1931. Adamski & Brown, 2001: 72
Streblopotamia Diakonoff, 1973
Streblopotamia streblopa (Meyrick, 1936). Razowski & Becker, 2018: 36; Razowski & Brown, 2018: 63
Olethreutes Hübner, 1822
“Olethreutes” subapicana Walker, 1863: 387. Walsingham, 1891: 541. Heppner, 1995: 156
Rhopobota Lederer, 1859
Rhopobota macroceria Razowski, 1999b: 350. Brown, 2005: 535 *DR > TL: Pedernales, 5 km NE Los Arroyos, 1680 m
Rhopobota microceria Razowski, 1999b: 351. Brown, 2005: 535 *DR > TL: Peravia, 3 km SW La Nuez, tributary to Río Las
Cuevas, 1870 m
Rhopobota unidens Razowski, 1999b: 351. Brown, 2005: 536 *DR > TL: La Vega, 9 km SE Constanza, near Valle Nuevo,
1930 m
Rhyacionia Hübner, 1825
Rhyacionia frustrana (Comstock, 1880). Powell & Miller, 1978: 22; Razowski, 1999a: 314
Rhyacionia pallidicosta Razowski, 1999a: 313. Brown, 2005: 539 *DR > TL: Pedernales, 9.7 km NE Los Arroyos, 2070 m
Strepsicrates Meyrick, 1888
Strepsicrates smithiana Walsingham, 1891. Perez-Gelabert, 2008: 258

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 331
Subfamily TORTRICINAE
Amorbia Clemens, 1860
Amorbia sp.—SEA (1999: 66)
Apotoforma Busck, 1933
Apotoforma monochroma (Walsingham, 1897: 132). Heppner, 1995: 147 *H > TL: Port-au-Prince
Apotoforma negans (Walsingham, 1897: 131); Heppner, 1995: 147
Argyrotaenia Stephens, 1852
Argyrotaenia bisignata Razowski, 1999a: 310. Razowski & Becker, 2000b: 313; Brown, 2005: 133 *DR > TL: Pedernales, 5
km NE Los Arroyos, 1680 m
Argyrotaenia ceramica Razowski, 1999a: 309. Brown, 2005: 134
Argyrotaenia felisana Razowski, 1999a: 309. Razowski & Becker, 2000b: 312; Brown, 2005: 134 *DR > TL: Independencia,
Sierra de Neiba just south of crest, 5 km WNW Angel Feliz, 1780 m
Argyrotaenia mesosignaria Razowski, 1999a: 311. Razowski & Becker, 2000b: 314; Brown, 2005: 136 *DR > TL: La Vega, 9
km SE Constanza, near Valle Nuevo, 1930 m
Argyrotaenia minisignaria Razowski, 1999a: 311. Razowski & Becker, 2000b: 314; Brown, 2005: 136 *DR > TL: Pedernales,
8 km NE Los Arroyos, 1940 m
Argyrotaenia neibana Razowski, 1999a: 310. Razowski & Becker, 2000b: 312; Brown, 2005: 136 *DR > TL: Baoruco, Sierra
de Neiba, Los Guineos on upper Rio Colorado, 630 m
Argyrotaenia nuezana Razowski, 1999a: 309. Razowski & Becker, 2000b: 312; Brown, 2005: 136 *DR > TL: La Vega, 24 km
SE Constanza, 2220 m
Argyrotaenia thamaluncus Razowski, 1999a: 311. Razowski & Becker, 2000b: 314; Brown, 2005: 138 *DR > TL: Peravia, 3
km SW La Nuez, tributary to Río Las Cuevas, 1870 m
Bonagota Razowski, 1986
Bonagota dominicana Razowski, 1999a: 308. Brown, 2005: 159 *DR > TL: Pedernales, 8 km NE Los Arroyos, 1940 m
Claduncaria Razowski, 2000
Claduncaria ochrochlaena (Razowski, 1999a: 312). Brown, 2005: 178 *DR > TL: Pedernales, 5 km NE Los Arroyos, 1680 m
Cochylis Treitschke, 1829
“Cochylis” pimana (Busck, 1907). Razowski, 1999a: 308
Coelostathma Clemens, 1860
Coelostathma parallelana Walsingham, 1897. Forbes, 1930: 84
Eugnosta Hübner, 1825
Eugnosta chalicophora Razowski, 1999a: 308. Brown, 2005: 337 *DR > TL: Pedernales, 30 km N Cabo Rojo, 1070 m
Orthocomotis Dognin, 1905
Orthocomotis independentia Razowski, 1999a: 313. Brown, 2005: 455 *DR > TL: Independencia, Sierra de Neiba just south of
crest, 5 km WNW Angel Feliz, 1780 m
Platynota Clemens, 1860
Platynota flavedana Clemens, 1860. Walsingham, 1897: 134; Forbes, 1930: 85 [Misidentification?]
Platynota restitutana (Walker, 1863: 292). Heppner, 1995: 149
Platynota rostrana (Walker, 1863). Walsingham, 1897: 134; Walsingham, 1909–1915: 215; Heppner, 1995: 149
=Platynota repandana (Walker, 1863: 292). Walsingham, 1891: 499; Walsingham, 1897: 134; Heppner, 1995: 149
Saphenista Walsingham, 1914
Saphenista peraviae Razowski, 1994: 202. Razowski, 1999a: 308; Brown, 2005: 546 *DR > TL: Peravia, 3 km SW La Nuez,
tributary to Rio Las Cuevas, 1870 m
Note: Argyrotaenia ochrochroa Razowski, 1999: 310 originally described from Turks and Caicos was erroneously cited as from
the Dominican Republic as Argyrotaenia ochrotona by Razowski & Becker (2000b: 312) [Misspelling]. The mistaken
Dominican Republic distribution was later repeated by Brown, 2005: 136 and Perez-Gelabert, 2008: 259.

Superfamily GEOMETROIDEA
Family URANIIDAE
Manidia Westwood, 1879
Manidia excavata (Walker, 1854: 9). Westwood, 1879: 528

Family GEOMETRIDAE
Subfamily ENNOMINAE

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Arilophia Rindge, 1990
Arilophia rawlinsi Rindge, 1990: 26. Scoble, 1999: 750 *DR > TL: La Abeja, 1250 m, 38 km NNW Cabo Rojo, Pedernales
Cyclomia Guenée, 1857
Cyclomia mopsaria mopsaria Guenée, 1858. Walker, 1861b: 981; Schaus, 1940: 313; Scoble, 1999: 203
=Alydda? calidata Walker, 1862a: 1081. Scoble, 1999: 203
=Alydda? ebuleata Walker, 1862a: 1082. Scoble, 1999: 203
=Alydda? scopulata Walker, 1862a: 1082. Scoble, 1999: 203
=Audia mixtalis Walker, 1863: 18. Scoble, 1999: 203
=Coremia divisata Walker, 1863: 1714. Scoble, 1999: 203
=Cyclomia iodaria Guenée, 1858. Walker, 1861b: 981; Scoble, 1999: 203
=Fidonia eugraphiata Walker, 1863: 1669. Scoble, 1999: 203
=Macaria? turturaria Walker, 1861b: 896. Scoble, 1999: 203
=Omyza? acontiaria Walker, 1862. Scoble, 1999: 203
Digrammia Gumppenberg, 1887
Digrammia heliothidata (Guenée, 1858). Walker, 1861b: 891; Scoble, 1999: 230
=Acidalia restorata Walker, 1861b: 726. Scoble, 1999: 230
=Aspilates subcolumbata Walker, 1863: 1676. Scoble, 1999: 230
Epimecis Hübner, 1825
Epimecis detexta Walker, 1860b: 452. Druce, 1891: 82; Scoble, 1999: 286
Epimecis hortaria (Fabricius, 1794). Domínguez, 1997: 72
Epimecis matronaria (Guenée, 1857). Domínguez, 1997: 71
Epimecis scolopaiae scolopaiae Drury, 1773. Scoble, 1999: 287
=Epimecis nyctalemonaria Walker, 1860b: 453. Scoble, 1999: 287
Epimecis scolopaiae transitaria Walker, 1860b: 454. Scoble, 1999: 287
Erastria Hübner, 1813
Erastria decrepitaria decrepitaria (Hübner, 1823). Druce, 1891: 44; Scoble, 1999: 294
=Hyperetes combinataria Walker, 1863: 1501. Scoble, 1999: 294
=Hyperythra versatiliaria Guenée, 1858. Walker, 1860a: 125; Scoble, 1999: 294
Euchlaena Hübner, 1823
Euchlaena amoenaria astylusaria Walker, 1860. Domínguez, 1997: 72
Iridopsis Warren, 1894
Iridopsis idonearia idonearia (Walker, 1860b: 352). Schaus, 1940: 323; Scoble, 1999: 517
Iridopsis monticola (Rindge, 1966: 210). Scoble, 1999: 517 *DR > TL: Hotel Montaña, Jarabacoa, La Vega
Macaria Curtis, 1826
Macaria abydata Guenée, 1858. Scoble, 1999: 564
=Macaria acidaliata Walker, 1861b: 893. Scoble, 1999: 564
Macaria everiata Guenée, 1858. Walker, 1861b: 892; Schaus, 1940: 311; Scoble, 1999: 567
=Alicia? rithrusaria Walker, 1860a: 102. Scoble, 1999: 567
=Pyrinia tegularia Walker, 1860a: 228. Scoble, 1999: 567
Macaria inoptata Walker, 1861b: 892. Scoble, 1999: 568
=Semiothisa dominicata Folder & Rogenhofer, 1875. Scoble, 1999: 568
Macaria paleolata Guenée, 1858. Walker, 1861b: 892; Schaus, 1940: 311; Scoble, 1999: 570
=Macaria spilogastrata Walker, 1861b: 894. Scoble, 1999: 570
Microsema Hübner, 1823
Microsema immaculata (Warren, 1897: 493). Scoble, 1999: 603 *DR > TL: San Domingo
Nepheloleuca Butler, 1883
Nepheloleuca complicata (Guenée, 1858). Walker, 1860a: 7; Schaus, 1940: 315; Scoble, 1999: 637
Nepheloleuca floridata (Grote, 1883). Domínguez, 1997: 72
Numia Guenée, 1857
Numia terebintharia Guenée, 1858. Walker, 1861a: 623; Scoble, 1999: 648
=Geometra diffissa Walker, 1861a: 510. Scoble, 1999: 648
=Geometra factaria Walker, 1861a: 511. Scoble, 1999: 648
=Geometra subcelata Walker, 1861a: 510. Scoble, 1999: 648
=Geometra subvectaria Walker, 1861a: 509. Scoble, 1999: 648

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 333
Oxydia Guenée, 1857
Oxydia vesulia transponens (Walker, 1860a: 20). Druce, 1891: 26; Domínguez, 1997: 72; Scoble, 1999: 685
Patalene Herrich-Schäffer, 1854
Patalene epionata (Guenée, 1858). Scoble, 1999: 706
=Drepanodes oemearia Walker, 1860a: 73. Scoble, 1999: 706
=Drepanodes pappiaria Walker, 1860a: 72. Scoble, 1999: 706
=Drepanodes pionaria Walker, 1860a: 71. Druce, 1891: 29; Scoble, 1999: 706
=Drepanodes tellesaria Walker, 1860a: 71. Scoble, 1999: 707
Patalene nicoaria (Walker, 1860a: 74). Scoble, 1999: 707
Patalene nutriaria (Walker, 1860a: 72). Scoble, 1999: 707
Pero Herrich-Schäffer, 1855
Pero nerisaria (Walker, 1860a: 188). Poole, 1987: 44; Scoble, 1999: 732
=Azelina? vetustaria Walker, 1866: 1550. Schaus, 1940: 317; Poole, 1987: 44; Scoble, 1999: 732
Pero rica Poole, 1987: 63. Scoble, 1999: 734 *DR > TL: Pinar Bonito, 2 mi. S. Constanza
Pero sella Poole, 1987: 45. Scoble, 1999: 734 *H > TL: La Visite and vic., La Selle Range, 5–7000 ft.
Phrygionis Hübner, 1825
Phrygionis bicornis Scoble, 1994: 126. Scoble, 1999: 749 *DR > TL: East side of crest, Sierra Martín García, 7 km WNW
Barrero, 860 m
Phrygionis ferreus Scoble, 1994: 130. Scoble, 1999: 749 *DR > TL: Elias Piña: Sierra de Neiba at crest, 5.5 km NNW Angel
Feliz, 1800 m
Phrygionis rawlinsi Scoble, 1994: 136. Scoble, 1999: 750 *DR > TL: Pedernales: 5 km NE Los Arroyos, 1680 m
Pyrinia Hübner, 1818
Pyrinia fusilineata Walker, 1863: 1489. Scoble, 1999: 812
Renia Guenée, 1854
Renia fraternalis (J. B. Smith, 1895). Domínguez, 1997: 72
Sabulodes Guenée, 1857
Sabulodes subopalaria (Walker, 1860a: 46). Scoble, 1999: 829
=Choerodes subclararia Walker, 1860a: 221. Scoble, 1999: 829
Semiothisa Hübner, 1818
“Semiothisa” cosmiata Walker, 1861b: 894. Scoble, 1999: 883
“Semiothisa” increta (Walker, 1861b: 895). Schaus, 1940: 311; Scoble, 1999: 884
Sphacelodes Guenée, 1857
Sphacelodes haitiaria Oberthür, 1923: 254. Scoble, 1999: 899
Thysanopyga Herrich-Schäffer, 1855
Thysanopyga abdominaria (Guenée, 1858). Walker, 1861b: 1018; Druce, 1891: 135; Scoble, 1999: 937
=Caberodes bilbisaria Walker, 1860a: 252. Scoble, 1999: 938
=Hyperythra? agasusaria Walker, 1860a: 242. Scoble, 1999: 937
“Thysanopyga” nicetaria (Guenée, 1858: 971). Schaus, 1940: 320; Scoble, 1999: 938
=Epione liboraria Walker, 1863: 1495. Scoble, 1999: 938
=Ephyra lucidiferata Walker, 1863: 1574. Scoble, 1999: 938
=Pachydia cyclata Walker, 1861b: 1018. Scoble, 1999: 938
=Tephrina confiniaria Walker, 1861b: 959. Scoble, 1999: 938
“Thysanopyga” proditata (Walker, 1861a: 633). Scoble, 1999: 939
Subfamily GEOMETRINAE
Synchlora Guenée, 1857
Synchlora herbaria herbaria (Fabricius, 1794). Schaus, 1940: 305; Scoble, 1999: 913
=Geometra congruata Walker, 1861a: 511. Druce, 1891: 84; Scoble, 1999: 913
=Iodis indeclararia Walker, 1861a: 541. Scoble, 1999: 913
=Racheospila sitellaria Guenée, 1858. Walker, 1861a: 581; Scoble, 1999: 913
Subfamily LARENTIINAE
Disclisioprocta Wallengren, 1861
Disclisioprocta stellata (Guenée, 1858). Walker, 1862b: 1366; Scoble, 1999: 235
Euphyia Hübner, 1825
Euphyia perturbata Walker, 1862b: 1367. Druce, 1891: 160; Scoble, 1999: 324

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 =Cidaria absitaria Felder & Rogenhofer, 1875. Scoble, 1999: 324
=Pterocypha deturbata Walker, 1862b: 1368. Scoble, 1999: 324
Obila Walker, 1869
Obila pannosata (Guenée, 1858). Walker, 1862b: 1366; Scoble, 1999: 652
=Anaitis dominataria Walker, 1862b: 1446. Scoble, 1999: 652
=Pterocypha delineata Walker, 1862b: 1367. Druce, 1891: 156; Scoble, 1999: 652
Pterocypha Herrich-Schäffer, 1855
Pterocypha floridata (Walker, 1863). Scoble, 1999: 806
=Pterocypha defensata Walker, 1862b: 1366. Scoble, 1999: 807
Triphosa Stephens, 1829
Triphosa affirmata (Guenée, 1858). Scoble, 1999: 949
=Scotosia progressata Walker, 1862b: 1348. Scoble, 1999: 949
Xanthorhoe Hübner, 1825
Xanthorhoe divisata (Walker, 1863: 1714). Scoble, 1999: 964
Subfamily OENOCHROMINAE
Almodes Guenée, 1857
Almodes terraria Guenée, 1858. Walker, 1861a: 608; Schaus, 1940: 292; Scoble, 1999: 35
=Polysemia stellidaria Guenée, 1858. Walker, 1862b: 1372; Scoble, 1999: 35
Subfamily STERRHINAE
Acratodes Guenée, 1857
Acratodes phakellurata Guenée, 1858. Walker, 1861b: 871; Scoble, 1999: 16
Cyclophora Hübner, 1822
Cyclophora nanaria (Walker, 1861a: 633). Scoble, 1999: 206
Euacidalia Packard, 1873
Euacidalia externata (Walker, 1863: 1598). Scoble, 1999: 301
Eumacrodes Warren, 1905
Eumacrodes yponomeutaria (Guenée, 1858). Scoble, 1999: 317
=Acidalia semidecora Walker, 1863: 1597. Scoble, 1999: 317
Leptostales Möschler, 1890
Leptostales noctuata (Guenée, 1858). Walker, 1861b: 806; Scoble, 1999: 537
Leptostales phorcaria (Guenée, 1858). Walker, 1861a: 726; Scoble, 1999: 538
Lobocleta Warren, 1906
Lobocleta dativaria Schaus, 1940. Schaus, 1940: 303
Lobocleta inermaria (Guenée, 1858). Walker, 1861a: 725; Scoble, 1999: 548
Lobocleta monogrammata (Guenée, 1858). Walker, 1861a: 725; Schaus, 1940: 303; Scoble, 1999: 548
Lobocleta nymphidiata (Guenée, 1858). Walker, 1861a: 726; Scoble, 1999: 548
Lobocleta perditaria (Walker, 1866: 1626). Scoble, 1999: 549
Pleuroprucha Möschler, 1890
Pleuroprucha rudimentaria (Guenée, 1858). Scoble, 1999: 767
Ptychamalia Prout, 1932
Ptychamalia botydata (Walker, 1861a: 727). Druce, 1891: 116; Scoble, 1999: 808
Scopula Schrank, 1802
Scopula micrata (Guenée, 1858). Walker, 1861a: 727; Scoble, 1999: 855
Semaeopus Herrich-Schäffer, 1855
Semaeopus castaria (Guenée, 1858). Walker, 1861b: 802; Scoble, 1999: 877
=Acidalia concinnata Felder & Rogenhofer, 1875. Scoble, 1999: 877
=Acidalia solitaria Walker, 1861a: 728. Scoble, 1999: 877
Semaeopus indignaria indignaria (Guenée, 1858). Scoble, 1999: 879
=Drepanodes absconditaria Walker, 1863: 1488. Scoble, 1999: 879
Sterrha Hübner, 1825
Sterrha curvicauda Schaus, 1940: 304

Family SEMATURIDAE
Mania Hübner, 1821

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Mania aegisthus (Fabricius, 1781). Hampson, 1918: 368

Superfamily HESPERIOIDEA
Key references: Schwartz (1989), Smith, Miller and Miller (1994). Updated and revised according to catalog of Neotropical
butterflies edited by Lamas (2004).
Family HESPERIIDAE
Subfamily HESPERIINAE
Argon Evans, 1955
Argon sp.—Schwartz (1989: 67)
Atalopedes Scudder, 1872
Atalopedes mesogramma apa Comstock, 1944: 561. Schwartz, 1989: 87
Calpodes Hübner, 1819
Calpodes ethlius (Stoll, 1782). Schwartz, 1989: 102
Choranthus Scudder, 1872
Choranthus haitensis Skinner, 1920. Schwartz, 1989: 92 *H, *DR > TL: Port de Paix
Choranthus maria Minno, 1990: 55 *H, *DR > TL: Puerto Plata Prov.: 1.5 km E. Sosua
Choranthus melissa Gali, 1983. Schwartz, 1989: 95 *DR > TL: Prov. Pedernales: Las Abejas, 11 km NW Aceitillar, 1220 m
Choranthus schwartzi Gali, 1983. Schwartz, 1989: 94 *DR *H > TL: Prov. La Vega: 10 km W Jayaco, 915 m
Copaeodes Speyer, 1877
Copaeodes stillmani Bell & Comstock, 1948: 18. Schwartz, 1989: 77
Cymaenes Scudder, 1872
Cymaenes tripunctus tripunctus (Herrich-Schäffer, 1865). Schwartz, 1989: 72
Euphyes Scudder, 1872
Euphyes singularis insolata (Butler, 1878). Schwartz, 1989: 99
Hesperia Fabricius, 1793
Hesperia nabokovi (Bell & Comstock, 1948: 19). Schwartz, 1989: 89 *H, *DR > TL: Thomazeau
Hylephila Billberg, 1820
Hylephila phyleus phyleus (Drury, 1773). Schwartz, 1989: 85
Nyctelius Hayward, 1948
Nyctelius nyctelius nyctelius (Latreille, 1824). Schwartz, 1989: 112
Ochlodes Scudder, 1872
Ochlodes batesi (Bell, 1935: 63). Schwartz, 1989: 97 *DR, *H > TL: La Selle Mts.
Panoquina Hemming, 1934
Panoquina ocola distipuncta Johnson & Matusik, 1988: 245. Schwartz, 1989: 110
Panoquina lucas (Fabricius, 1793). (By synonymy of species below)
=Panoquina sylvicola woodruffi Watson, 1937. Schwartz, 1989: 103
Perichares Scudder, 1872
Perichares philetes philetes (Gmelin, 1790). Takizawa et al., 2003: 16
Polites Scudder, 1872
Polites baracoa loma Evans, 1955. Schwartz, 1989: 81 *H, *DR > TL: Haiti
Pyrrhocalles Mabille, 1904
Pyrrhocalles antiqua antiqua (Herrich-Schäffer, 1863). Schwartz, 1989: 65
Rhinton Godman, 1900
Rhinton bushi Watson, 1937. Schwartz, 1989: 75
Synapte Mabille, 1904
Synapte malitiosa adoceta Schwartz & Sommer, 1986. Schwartz, 1989: 71
Wallengrenia Berg, 1897
Wallengrenia otho druryi (Latreille, 1824). Schwartz, 1989: 82
Subfamily PYRGINAE
Tribe EUDAMINI
Aguna Williams, 1927
Aguna asander haitensis (Mabille & Boullet, 1912). Schwartz, 1989: 28
Astraptes Hübner, 1819
Astraptes anaphus anausis (Goodman & Salvin, 1896). Schwartz, 1989: 38

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Astraptes alardus heriul (Mabille & Boullet, 1912). Schwartz, 1989: 40
Astraptes christyi (Sharpe, 1898). Schwartz, 1989: 39. Steinhauser et al., 2017: 87
Astraptes talus (Cramer, 1777). Schwartz, 1989: 35
Cabares Godman & Salvin, 1894
Cabares potrillo (Lucas, 1857). Schwartz, 1989: 43
Chioides Lindsey, 1921
Chioides ixion (Plötz, 1880). Schwartz, 1989: 25 *H, *DR > TL: Erroneously Centr.[al] Am. [erica]
=Eudamus myrto Mabille, 1903: 21
Epargyreus Hübner, 1819
Epargyreus spanna Evans, 1952. Schwartz, 1989: 21 *H, *DR > TL: Santo Domingo
Phocides Hübner, 1819
Phocides pigmalion bicolora (Boddaert, 1783). Schwartz, 1989: 17
Polygonus Hübner, 1825
Polygonus leo leo (Gmelin, 1790). Schwartz, 1989: 23
Polythrix Watson, 1893
Polythrix octomaculata (Ménétriés, 1855). Schwartz, 1989: 35
Proteides Hübner, 1825
Proteides mercurius sanchesi Bell & Comstock, 1948: 4. Schwartz, 1989: 19
Urbanus Hübner, 1807
Urbanus dorantes cramptoni Comstock, 1944. Schwartz, 1989: 32
Urbanus proteus domingo (Scudder, 1872: 69). Schwartz, 1989: 29
Tribe PYRGINI
Eantis Boisduval, 1836
Eantis papinianus (Poey, 1832). Sourakov, 2010: 115
=Achlyodes mithridates sagra Evans, 1953. Schwartz, 1989: 48
=Achlyodes thraso (Hübner, 1807). SEA, 1999: 111
Anastrus Hübner, 1824
Anastrus sempiternus dilloni (Bell & Comstock, 1948). Schwartz, 1989: 50
Burca Bell & Comstock, 1948
Burca hispaniolae Bell & Comstock, 1948: 14. Schwartz, 1989: 43 *H, *DR > TL: Freres
Burca stillmani Bell & Comstock, 1948: 12. Schwartz, 1989: 43 *H, *DR > TL: Río Yaque, 10 miles south of Monte Cristi
Chiomara Godman & Salvin, 1899
Chiomara mithrax (Möschler, 1879). Henríquez, 2008: 39
Ephyriades Hübner, 1819
Ephyriades zephodes (Hübner, 1825). Schwartz, 1989: 57
Erynnis Schrank, 1801
Erynnis zarucco (Lucas, 1857). Schwartz, 1989: 59
Gesta Evans, 1953
Gesta gesta (Herrich-Schäffer, 1863). Schwartz, 1989: 52
Pyrgus Hübner, 1819
Pyrgus crisia Herrich-Schäffer, 1865. Schwartz, 1989: 63
Pyrgus oileus (Linnaeus, 1767). Schwartz, 1989: 60

Superfamily PAPILIONOIDEA
Family PAPILIONIDAE
Subfamily PAPILIONINAE
Tribe LEPTOCIRCINI
Neographium Möhn in Bauer & Frankenbach, 2002
Neographium zonaria (Butler, 1869: 271). Schwartz, 1989: 123 *H, *DR > TL: St. Domingo
Tribe PAPILIONINI
Heraclides Hübner, 1819
Heraclides androgeus epidaurus (Godman & Salvin, 1890). Schwartz, 1989: 115; Sourakov, 2009: 117
Heraclides aristodemus aristodemus (Esper, 1794). Schwartz, 1989: 127
Heraclides aristor (Godart, 1819). Schwartz, 1989: 138 *H, *DR > TL: Port-au-Prince

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Heraclides machaonides (Esper, 1796). Coutsis, 1983: 114; Schwartz, 1989: 131 *H, *DR > TL: Port-au-Prince
Heraclides pelaus imerius (Godart, 1819). Schwartz, 1989: 142; Sourakov, 2009: 117
Papilio Linnaeus, 1758
Papilio demoleus demoleus Linnaeus, 1758. Guerrero et al., 2004: 227 [Introduced]
Tribe TROIDINI
Battus Scopoli, 1777
Battus polydamas polycrates (Höpffer, 1865). Schwartz, 1989: 119
Battus zetides (Munroe, 1971: 142). Coutsis, 1983: 114; Schwartz, 1989: 115; Johnson & Matusik, 1993: 78 *DR > TL: St.
Domingo
=Papilio zetes Westwood, 1847: 36

Family PIERIDAE
Subfamily COLIADINAE
Abaeis Hübner, 1819
Abaeis nicippe (Cramer, 1782). Schwartz, 1989: 184
Abaeis nicippiformis (Munroe, 1947). Schwartz, 1989: 186 *H, *DR > TL: Ravine of Pétionville, ca. 1600 ft.
Anteos Hübner, 1819
Anteos clorinde (Godart, 1824). Coutsis, 1983: 113; Schwartz, 1989: 200
Anteos maerula (Fabricius, 1775). Schwartz, 1989: 197
Aphrissa Butler, 1873
Aphrissa godartiana godartiana (Swainson, 1821). Schwartz, 1989: 222
Aphrissa orbis browni (Munroe, 1947: 1). Schwartz, 1989: 220
Aphrissa statira hispaniolae (Munroe, 1947: 1). Schwartz, 1989: 225
Eurema Hübner, 1819
Eurema daira palmira (Poey, 1852). Schwartz, 1989: 173
Eurema elathea elathea (Cramer, 1777). Schwartz, 1989: 176
Eurema priddyi (Lathy, 1898: 223). Schwartz, 1989: 180; Pérez-Asso et al., 2017: 50
Kricogonia Reakirt, 1864
Kricogonia lyside (Godart, 1819). Schwartz, 1989: 190
Nathalis Boisduval, 1836
Nathalis iole Boisduval, 1836. Schwartz, 1989: 188
Phoebis Hübner, 1819
Phoebis agarithe antillia Brown, 1929. Schwartz, 1989: 211
Phoebis argante rorata (Butler, 1869). Schwartz, 1989: 208
Phoebis editha (Butler, 1870). Coutsis, 1983: 113; Schwartz, 1989: 216
Phoebis philea thalestris (Illiger, 1801). Schwartz, 1989: 204
Phoebis sennae sennae (Linnaeus, 1758). Coutsis, 1983: 113; Schwartz, 1989: 213
Pyrisitia Butler, 1870
Pyrisitia dina mayobanex (Bates, 1939). Schwartz, 1989: 165
Pyrisitia euterpiformis (Munroe, 1947: 3). Schwartz, 1989: 161
Pyrisitia larae (Herrich-Schaffer, 1862). Pérez-Asso et al., 2017: 50
Pyrisitia leuce memulus (Butler, 1871). Schwartz, 1989: 165
Pyrisitia lisa euterpe (Ménétriés, 1832). Schwartz, 1989: 158
Pyrisitia proterpia (Fabricius, 1775). Schwartz, 1989: 170
Pyrisitia pyro (Godart, 1819). Schwartz, 1989: 181 *H, *DR > TL: Not stated
Rhabdodryas Godman & Salvin, 1889
Rhabdodryas trite watsoni (Brown, 1929). Schwartz, 1989: 218
Zerene Hübner, 1819
Zerene cesonia cynops (Butler, 1873). Schwartz, 1989: 195
Subfamily DISMORPHIINAE
Dismorphia Hübner, 1819
Dismorphia spio (Godart, 1819). Schwartz, 1989: 222
Subfamily PIERINAE
Tribe PIERINI

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Ascia Scopoli, 1777
Ascia monuste eubotea (Godart, 1819). Schwartz, 1989: 149
Ganyra Billberg, 1820
Ganyra josephina josephina (Godart, 1819). Schwartz, 1989: 146
Glutophrissa Butler, 1887
Glutophrissa drusilla boydi (Comstock, 1943). Schwartz, 1989: 152
Glutophrissa punctifera (d’Almeida, 1939). Takizawa et al., 2003: 33
Melete Swainson, 1832
Melete salacia salacia (Godart, 1819). Schwartz, 1989: 155

Family LYCAENIDAE
Subfamily POLYOMMATINAE
Brephidium Scudder, 1876
Brephidium exilis isophthalma (Herrich-Schäffer, 1862). Schwartz, 1989: 257
Cyclargus Nabokov, 1945
Cyclargus ammon noeli (Comstock & Huntington, 1943). Schwartz, 1989: 266
Cyclargus kathleena Johnson & Matusik, 1992: 3 *DR > TL: Circa 4 km from upper Las Abejas, in xeric pine forest, pine
forest bottom, @1000 m
Cyclargus sorpresus Johnson & Matusik, 1992: 3 *DR > TL: 21 km marker near “Las Mercedes” on Ideal Dominicana S. A.
company road from Cabo Rojo to Aceitillar, Pedernales Prov., alt. @500 m
Hemiargus Hübner, 1818
Hemiargus hanno ceraunus (Fabricius, 1793). Schwartz, 1989: 264
=Hemiargus hanno watsoni Comstock & Huntington, 1943. Schwartz, 1989: 262
Leptotes Scudder, 1876
Leptotes cassius theonus (Lucas, 1857). Schwartz, 1989: 258
Leptotes perkinsae idealus Johnson & Matusik, 1988: 243. Schwartz, 1989: 261 *DR > TL: Bottom of Las Abejas
Pseudochrysops Nabovkov, 1945
Pseudochrysops bornoi bornoi (Comstock & Huntington, 1943). Schwartz, 1989: 268
Subfamily THECLINAE
Tribe EUMAEINI
Allosmaitia Clench, 1964
Allosmaitia coelebs (Herrich-Schäffer, 1862). Takizawa et al., 2003: 49
Chlorostrymon Clench, 1961
Chlorostrymon maesites (Herrich-Schäffer, 1865). Schwartz, 1989: 236
Chlorostrymon simaethis (Drury, 1773). Schwartz, 1989: 235
Electrostrymon Clench, 1961
Electrostrymon angelia (Hewitson, 1874). Schwartz, 1989: 233
Electrostrymon minikyanos Johnson & Matusik, 1988: 223. Schwartz, 1989: 234; Robbins & Lamas, 2002: 206 *DR > TL:
“Middle Abejas”, 1500 m, Prov. Pedernales [Nomen dubium]
Ministrymon Clench, 1961
Ministrymon azia (Hewitson, 1873). Schwartz, 1989: 238
Nesiostrymon Clench, 1964
Nesiostrymon celida (Lucas, 1857). Robbins in Robbins & Lamas, 2005: 136
=Nesiostrymon celida baorucoensis Johnson, 1991: 9. Robbins in Robbins & Lamas, 2005: 136
Rekoa Kaye, 1904
Rekoa bourkei (Kaye, 1924). Robbins, 1991: 45
=Heterosmaitia abeja (Johnson & Matusik, 1988: 232). Schwartz, 1989: 231
Strymon Hübner, 1818
Strymon acis petioni (Comstock & Huntington, 1943). Schwartz, 1989: 242
Strymon bazochii gundlachianus (Bates, 1935). Schwartz, 1989: 255
Strymon christophei (Comstock & Huntington, 1943: 85). Schwartz, 1989: 250 *H > TL: Port-au-Prince
Strymon columella (Fabricius, 1793). Schwartz, 1989: 244
Strymon limenia (Hewitson, 1868). Schwartz, 1989: 251
Strymon monopeteinus Schwartz & J. Y. Miller, 1985. Schwartz, 1989: 253 *DR > TL: Prov. María Trinidad Sánchez: 1 km S

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Cruce de Rincón, s. l. [sea level]
Strymon toussainti (Comstock & Huntington, 1943). Schwartz, 1989: 247; Robbins & Lamas, 2002: 202
=Strymon andrewi Johnson & Matusik, 1988: 228. Schwartz, 1989: 249; Robbins & Lamas, 2002: 202
Note: The species Terra hispaniola Johnson & Matusik, 1988: 241 and Tmolus victoria Johnson & Matusik in Schwartz, 1989:
525, were shown by Robbins & Lamas (2002) not to occur in Hispaniola.

Family RIODINIDAE
Theope Doubleday, 1847
Theope sp.†—DeVries & Poinar (1997: 1138) [Fossil in Dominican amber]
Voltinia Stichel, 1910
Voltinia dramba† Robbins & Harvey in Hall et al., 2004: 797. Peñalver & Grimaldi (2006: 3) [Fossil in Dominican amber]

Family NYMPHALIDAE
Subfamily APATURINAE
Asterocampa Röber, 1916
Asterocampa idyja idyja (Geyer, 1828). Schwartz, 1989a: 379
Doxocopa Hübner, 1819
Doxocopa thoe (Godart, 1824). Schwartz, 1989a: 381 *DR, *H > TL: Erroneously “Brasil”, this species is endemic to
Hispaniola
Subfamily BIBLIDINAE
Tribe CYRESTINI
Marpesia Hübner, 1818
Marpesia chiron chiron (Fabricius, 1775). Schwartz, 1989a: 358
Marpesia eleuchea dospassosi Munroe, 1971: 190. Schwartz, 1989a: 360
Tribe BIBLIDINI
Archimestra Munroe, 1949
Archimestra teleboas (Ménétriés, 1832). Schwartz, 1989a: 336 *H > TL: Haiti
Biblis Fabricius, 1807
Biblis hyperia (Cramer, 1779). Schwartz, 1989a: 343
Eunica Hübner, 1819
Eunica monima (Cramer, 1782). Schwartz, 1989a: 332
Eunica tatila tatilista Kaye, 1926. Schwartz, 1989a: 329
Hamadryas Hübner, 1806
Hamadryas februa ferox (Staudinger, 1886). Garzón-Orduña et al., 2018: 61
=Hamadryas amphichloe diasia (Fruhstorfer, 1916). Schwartz, 1989a: 345
Lucinia Hübner, 1823
Lucinia cadma torrebia (Ménétriés, 1832). Lathy, 1903: 203; Schwartz, 1989a: 340
Myscelia Doubleday, 1844
Myscelia aracynthia (Dalman, 1823). Schwartz, 1989a: 324 *DR > TL: Unknown.
Tribe DYNAMININI
Dynamine Hübner, 1819
Dynamine alexae† Peñalver & Grimaldi, 2006: 7 [Fossil in Dominican amber]
Dynamine serina zetes (Ménétriés, 1832). Schwartz, 1989a: 334
Subfamily CHARAXINAE
Tribe ANAEINI
Anaea Hübner, 1819
Anaea troglodyta (Fabricius, 1775). Schwartz, 1989: 364 *DR, *H > TL: “America”. Endemic to Hispaniola
Fountainea Rydon, 1971
Fountainea johnsoni (Avinoff & Shoumatoff, 1941). Coutsis, 1983: 113; Schwartz, 1989a: 368
Memphis Hübner, 1819
Memphis verticordia (Hübner, 1831). Schwartz, 1989a: 370 *DR, *H > TL: Not stated. Endemic to Hispaniola
Siderone Hübner, 1823
Siderone nemesis nemesis Illiger, 1801. Dottax & Pierre, 2009: 404
Tribe PREPONINI

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Archaeoprepona Fruhstorfer, 1915
Archaeoprepona demophoon insulicola (Godart, 1823). Schwartz, 1989a: 376
Subfamily DANAINAE
Tribe EUPLOEINI
Anetia Robineau-Desvoidy, 1863
Anetia briarea briarea (Godart, 1819). Schwartz, 1989a: 474
Anetia jaegeri (Ménétriés, 1832). Schwartz, 1989a: 480
Anetia pantheratus pantheratus (Martyn, 1797). Schwartz, 1989a: 478
Lycorea Doubleday, 1847
Lycorea halia cleobaea (Godart, 1819). Schwartz, 1989a: 471
Lycorea halia halia (Hübner, 1816) [By synonymy of species below]
=Lycorea pieteri Lamas, 1978. Wetherbee, 1986: 20
Tribe DANAINI
Danaus Linnaeus, 1758
Danaus cleophile (Godart, 1819). Schwartz, 1989a: 467
Danaus eresimus tethys Forbes, 1944. Schwartz, 1989a: 460
Danaus gilippus cleothera (Godart, 1819). Schwartz, 1989a: 458
Danaus plexippus megalippe (Hübner, 1826). Schwartz, 1989a: 463
Subfamily HELICONIINAE
Tribe ARGYNNINI
Euptoieta Doubleday, 1848
Euptoieta claudia claudia (Cramer, 1779). Schwartz, 1989a: 287
Euptoieta hegesia hegesia (Cramer, 1779). Schwartz, 1989a: 289
Tribe HELICONIINI
Agraulis Boisduval & LeConte, 1836
Agraulis vanillae insularis Maynard, 1889. Schwartz, 1989a: 274
Dryas Hübner, 1807
Dryas iulia fucatus (Boddaert, 1783). Schwartz, 1989a: 277
Eueides Hübner, 1816
Eueides isabella melphis (Godart, 1819). Schwartz, 1989a: 281
Heliconius Kluk, 1802
Heliconius charitonia churchi Comstock & Brown, 1950. Schwartz, 1989a: 283
Subfamily ITHOMIINAE
Tribe GODYRIDINI
Greta Hemming, 1934
Greta diaphanus quisqueya (Fox, 1963: 177). Schwartz, 1989a: 453 *DR > TL: Mt. Diego de Ocampo, 3–4000 ft.
=Greta diaphana calimete Schwartz, 1982. Schwartz, 1989a: 455
=Greta diaphana charadra Schwartz, 1982. Schwartz, 1989a: 454
=Greta diaphana galii Schwartz, 1982. Schwartz, 1989a: 456
Subfamily LIBYTHEINAE
Libytheana Michener, 1943
Libytheana terena (Godart, 1819). Schwartz, 1989a: 270; Kawahara, 2013: 57
Subfamily LIMENITIDINAE
Adelpha Hübner, 1819
Adelpha fessonia lapitha Hall, 1929. Schwartz, 1989a: 323
Adelpha gelania gelania (Godart, 1824). Schwartz, 1989a: 320
Subfamily NYMPHALINAE
Tribe COEINI
Colobura Billberg, 1820
Colobura dirce wolcotti (Comstock, 1942). Schwartz, 1989a: 354
Historis Hübner, 1819
Historis acheronta semele (Bates, 1939). Schwartz, 1989a: 349
Historis odius odius (Fabricius, 1775). Schwartz, 1989a: 350
Tribe KALLIMINI

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Anartia Hübner, 1819
Anartia jatrophae saturata Staudinger, 1884. Schwartz, 1989a: 311
Anartia lytrea (Godart, 1819). Schwartz, 1989a: 315 *DR, *H > TL: San Domingo
=Anartia dominica Skinner, 1889: 86
Hypolimnas Hübner, 1819
Hypolimnas misippus (Linnaeus, 1764). Schwartz, 1989a: 306
Junonia Hübner, 1819
Junonia evarete (Cramer, 1779). Takizawa et al., 2003: 81
Junonia genoveva (Cramer, 1780). Schwartz, 1989a: 307
Siproeta Hübner, 1823
Siproeta stelenes stelenes (Linnaeus, 1758). Schwartz, 1989a: 318
Tribe NYMPHALINI
Hypanartia Hübner, 1821
Hypanartia paullus (Fabricius, 1793). Schwartz, 1989a: 304
Vanessa Fabricius, 1807
Vanessa atalanta rubria (Fruhstorfer, 1909). Schwartz, 1989a: 302
Vanessa cardui (Linnaeus, 1758). Schwartz, 1989a: 300
Vanessa virginiensis (Drury, 1773). Schwartz, 1989a: 298
Tribe MELITAEINI
Antillea Higgins, 1959
Antillea pelops pelops (Drury, 1773). Schwartz, 1989a: 296
Anthanassa Scudder, 1875
Anthanassa frisia (Poey, 1832). Schwartz, 1989a: 293
Atlantea Higgins, 1959
Atlantea cryptadia Sommer & Schwartz, 1980. Schwartz, 1989a: 293 *H, *DR > TL: Departement de L’Ouest: Boutilliers
Road, 734–857 m
Subfamily SATYRINAE
Tribe SATYRINI
Calisto Hübner, 1823
Calisto ainigma Johnson, Quinter & Matusik, 1987. Schwartz, 1989a: 445 *DR > TL: La Vega Prov., 930 m in central portion
of Cordillera Central, along small stream between Hotel Pinar Dorado cabañas and highway from hotel grounds about 4 km
northwest to central Jarabacoa
Calisto arcas Bates, 1939: 48. Schwartz, 1989a: 395; Sourakov & Zakharov, 2011: 202 *DR > TL: Valle Nuevo, S. E.
Constanza, c. 7000 ft.
Calisto archebates (Ménétriés, 1832). Schwartz, 1989a: 384; Sourakov & Zakharov, 2011: 202 *DR, *H > TL: Haiti
Calisto azua Pérez-Asso et al., 2016: 422 *DR > TL: Hatillo, El Número
Calisto bahoruco Pérez-Asso, Núñez Aguila & Genaro in Núñez Aguila et al., 2017: 3 *DR > TL: Villa Nizao, Paraíso
Calisto batesi Clench, 1943: 23. Schwartz, 1989a: 410; Sourakov & Zakharov, 2011: 202 *DR > TL: Loma del Toro, foothills
of the Cordillera Central, south of Santiago, ±5000 ft.
Calisto chrysaoros Bates, 1935. Bates, 1939: 48; Michener, 1943: 3; Schwartz, 1989a: 389; Sourakov & Zakharov, 2011: 202;
Núñez Aguila et al., 2017: 751 *H, *DR > TL: Roche Croix, La Hotte, 5000 ft.
Calisto chrysaoros chrysaoros Bates, 1935. Nuñez Aguila et al., 2017: 751
=Calisto clenchi Schwartz & Gali, 1984: 10. Schwartz, 1989a: 441; Nuñez Aguila et al., 2017: 751
Calisto chrysaoros galii Schwartz, 1983: 3. Schwartz, 1989a: 391*DR > TL: 18 km SE Constanza, 1586 m, Prov. La Vega
=Calisto galii choneupsilon Schwartz, 1985. Schwartz, 1989a: 393; Nuñez Aguila et al., 2017: 751
=Calisto galii galii Schwartz, 1983: 4. Schwartz, 1989a: 391; Nuñez Aguila et al., 2017: 751
Calisto clydoniata Schwartz & Gali, 1984: 10. Schwartz, 1989a: 439; Núñez Aguila et al., 2017: 748 *DR > TL: La Estrelleta:
1 km SW Puesto Pirámide 204, 6200 ft.
Calisto confusa Lathy, 1899: 227. Michener, 1943: 5; Schwartz, 1989a: 414; Sourakov & Zakharov, 2011: 202 *DR, *H > TL:
Haiti
Calisto crypta Gali, 1985: 3. Schwartz, 1989a: 427; Sourakov & Zakharov, 2011: 202; Pérez-Asso et al., 2016: 428 *DR > TL:
Prov. Monte Cristi, near Monte Cristi
Calisto debarriera Clench, 1943. Sourakov & Zakharov, 2011: 202 *H > TL: Debarriere [Desbarriere], La Hotte Mt., 4000 ft.
=Calisto confusa debarriera Clench, 1943: 25. Schwartz, 1989a: 422

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 =Calisto gonzalezi Schwartz, 1988. Schwartz, 1989a: 421
=Calisto montana Clench, 1943. Schwartz, 1989a: 438
=Calisto neiba Schwartz & Gali, 1984: 2. Schwartz, 1989a: 424
Calisto eleleus Bates, 1935. Schwartz, 1989a: 442; Sourakov & Zakharov, 2011: 202 *H, *DR > TL: Mt. Bourette (5000 ft.)
Calisto franciscoi Gali, 1985: 4. Schwartz, 1989a: 429; Sourakov & Zakharov, 2011: 202; Pérez-Asso et al., 2016: 425 *DR >
TL: Prov. Azua, Tábara Abajo
=Calisto hendersoni Gali, 1985: 6. Schwartz, 1989a: 433
Calisto grannus amazona Gonzalez, 1987: 6. Schwartz, 1989a: 402; Sourakov & Zakharov, 2011: 202 *DR > TL: Prov.
Santiago, entrance to Valle de Tetero, 6300 ft. (1922 m)
Calisto grannus dilemma Gonzalez, 1987. Schwartz, 1989a: 400; Sourakov & Zakharov, 2011: 202 *DR > TL: Prov. La Vega,
6 km SSE Constanza, 4600 ft. (1483 m)
Calisto grannus dystacta Gonzalez, 1987: 8. Schwartz, 1989a: 403; Sourakov & Zakharov, 2011: 202 *DR > TL: Prov.
Santiago, Loma Leonor, 19 km SW Monción, 2000 ft. (645 m)
Calisto grannus grannus Bates, 1939: 49. Schwartz, 1989a: 399; Sourakov & Zakharov, 2011: 202 *DR > TL: Valle Nuevo, S.
E. Constanza, c. 7000 ft.
Calisto grannus micheneri Clench, 1943: 115. Schwartz, 1989a: 407; Sourakov & Zakharov, 2011: 202 *DR > TL: Loma del
Toro, foothills of Cordillera Central, about 5000 ft., S of Santiago
Calisto grannus micrommata Schwartz & Gali, 1984: 6. Schwartz, 1989a: 404; Sourakov & Zakharov, 2011: 202 *DR > TL: 2
km NE Puesto Pirámide 2004, 1586 m
Calisto grannus phoinix Gonzalez, 1987: 5. Schwartz, 1989a: 401; Sourakov & Zakharov, 2011: 202; Nuñez Aguila et al.,
2017: 21 *DR > TL: La Palma, 19 km W Jayaco, 1007 m, Prov. La Vega
Calisto grannus sommeri Schwartz & Gali, 1984. Schwartz, 1989a: 405; Sourakov & Zakharov, 2011: 202 *H, *DR > TL:
Forêt des Pins, 1586 m
Calisto hysius (Godart, 1819). Michener, 1943: 4; Schwartz, 1989a: 408; Sourakov & Zakharov, 2011: 202 *H, *DR > TL:
Sud: 10.7 km WNW Les Platons (citadel), Caye Paul 1120 m
Calisto hysius aleucosticha Correa & Schwartz, 1986. Schwartz, 1989a: 413; Sourakov & Zakharov, 2011: 202 *DR > TL:
Prov. La Vega, Buena Vista, 11 km NW Jarabacoa, 640 m
Calisto loxias Bates, 1935. Schwartz, 1989a: 387 *H > TL: Roche Croix, La Hotte Mountains, at an elevation of about 5000 ft.
Calisto lyceius Bates, 1935. Michener, 1943: 6; Schwartz, 1989a: 426; Sourakov & Zakharov, 2011: 202; Pérez-Asso et al.,
2016: 424 *DR > TL: Saona Island
Calisto mariposa Pérez-Asso et al., 2016: 418 *DR > TL: Sierra Prieta, Santo Domingo
Calisto neochma Schwartz, 1991: 1 *DR > TL: Prov. Santiago: Monte El Rubio, 900 m
Calisto obscura Michener, 1943: 5. Schwartz, 1989a: 417; Sourakov & Zakharov, 2011: 202 *H, *DR > TL: Pétionville, 1600
ft. altitude
Calisto pauli Johnson & Hedges, 1998: 47 *H > TL: Sud: 10.7 km WNW Les Platons (citadel), Caye Paul 1120 m
Calisto pulchella Lathy, 1899. Bates, 1939: 50; Wolcott, 1927: 267; Schwartz, 1989a: 446; Sourakov & Zakharov, 2011: 202
*DR, *H > TL: Haiti
=Calisto pulchella darlingtoni Clench, 1943: 28. Schwartz, 1989a: 450; Sourakov & Zakharov, 2011: 202; Pérez-Asso et al.,
2016: 413
Calisto raburni Gali, 1985: 8. Schwartz, 1989a: 434; Sourakov & Zakharov, 2011: 202; Pérez-Asso et al., 2016: 415 *DR >
TL: Prov. Independencia, 7 km NE El Aguacate, 519 m
Calisto samana Pérez-Asso et al., 2016: 428 *DR > TL: Cueva del Agua, Samaná
Calisto schwartzi Gali, 1985: 7. Schwartz, 1989a: 436; Pérez-Asso et al., 2016: 421 *DR > TL: Prov. Pedernales, 1 km N
Aceitillar, 1281 m
Calisto tasajera Gonzalez et al., 1991: 1; Sourakov & Zakharov, 2011: 202; Pérez-Asso et al., 2016: 419 *DR > TL: Prov. San
Juan, Loma de Tasajera, 2142 m
Calisto thomasi Johnson & Hedges, 1998: 50 *H > TL: Sud: 10.7 km WNW Les Platons (citadel), Caye Paul 1120 m
Calisto tragius Bates, 1935. Schwartz, 1989a: 396 *H > TL: La Visite, in the La Selle range, 5000 to 7000 ft.
Calisto victori Pérez-Asso et al., 2016: 422 *DR > TL: Camino entre Caseta 2 y Caseta 1, Parque Nacional Sierra de Bahoruco
Calisto wetherbeei Schwartz & Gonzalez, 1988: 2. Schwartz, 1989a: 388 *DR > TL: Summit, Loma Nalga de Maco, ca. 1900
m, Prov. Elías Piña
Calisto woodsi Johnson & Hedges, 1998: 49 *H > TL: Sud: 10.7 km WNW Les Platons (citadel), Caye Paul 1120 m

Superfamily BOMBYCOIDEA

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 343
Family SATURNIIDAE
Subfamily HEMILEUCINAE
Hispaniodirphia Lemaire, 1999
Hispaniodirphia lemaireiana Rougerie & Herbin, 2006: 54 *DR > TL: Prov. Pedernales, km 40 of the road from Puerto
Escondido to Pedernales, Sierra de Bahoruco National Park, alt. 1740 m asl
Hispaniodirphia plana (Walker, 1855: 1436). Lemaire, 1999: 177 *DR > TL: St. Domingo

Superfamily SPHINGOIDEA
Family SPHINGIDAE
Key reference: Haxaire (2016).
Subfamily SPHINGINAE
Tribe ACHERONTIINI
Agrius Hübner, 1819
Agrius cingulatus (Fabricius, 1775). Walker, 1856: 215; Butler, 1877: 608; Druce, 1891: 21; Schreiber, 1978: 39; Domínguez,
1997: 67; Miyata, 1997: 19; Haxaire, 2016: 62
Tribe SPHINGINI
Cocytius Hübner, 1819
Cocytius antaeus antaeus (Drury, 1773). Walker, 1856: 200; Butler, 1877: 599; Forbes, 1930: 52; Schreiber, 1978: 39; Miyata,
1997: 19; Eitschberger, 2006: 191
Cocytius duponchel (Poey, 1832). Butler, 1877: 599; Druce, 1891: 17; Schreiber, 1978: 39; Miyata, 1997: 19; Haxaire, 2016: 62
=Amphonyx kofleri Eitschberger, 2006: 177. Haxaire, 2016: 62
=Amphonyx rivularis Butler, 1875. Eitschberger, 2006: 180
Lintneria Butler, 1876
Lintneria tricolor (Clark, 1923). Haxaire, 2016: 62
Manduca Hübner, 1807
Manduca afflicta hispaniolana Brechlin & Haxaire, 2014: 69 *DR > TL: Puerto Plata, 800 m
Manduca brontes haitiensis (Clark, 1916: 41). Cary, 1951: 120; Schreiber, 1978: 42; Miyata, 1997: 19; Kitching & Cadiou,
2000: 138; Haxaire, 2016: 62
Manduca caribbeus (Cary, 1952: 198). Schreiber, 1978: 39; Haxaire, 2016: 62
Manduca johanni (Cary, 1958). Haxaire, 2016: 62
Manduca rustica cubana (Gehlen, 1928). Haxaire, 2016: 62
=Protoparce dominicana Gehlen, 1928: 391
Manduca sexta paphus (Linnaeus, 1764). Haxaire, 2016: 62
Nannoparce Rothschild & Jordan, 1903
Nannoparce poeyi poeyi Grote, 1867. Cary, 1951: 120; Schreiber, 1978: 44; Haxaire, 2016: 62
Neococytius Hodges, 1971
Neococytius cluentius (Cramer, 1775). Walker, 1856: 200; Butler, 1877: 600; Schreiber, 1978: 39; Eitschberger, 2006: 197;
Haxaire, 2016: 62
Sphinx Linnaeus, 1758
Sphinx tricolor Clark, 1923. Schreiber, 1978: 45; Haxaire, 2016: 62
Subfamily SMERINTHINAE
Tribe AMBULYCINI
Protambulyx Rothschild & Jordan, 1903
Protambulyx strigilis strigilis (Linnaeus, 1771). Walker, 1856: 121; Butler, 1877: 579; Druce, 1891: 16; Schreiber, 1978: 46;
Domínguez, 1997: 67; Miyata, 1997: 20; Haxaire, 2016: 62
=Ambulyx rubripennis Butler, 1877: 579
Subfamily MACROGLOSSINAE
Tribe DILOPHONOTINI
Aellopos Hübner, 1819
Aellopos blaini Herrich-Schaffer, 1869. Haxaire, 2016: 68
Aellopos clavipes (Cramer, 1776). Haxaire, 2016: 68
Aellopos tantalus (Linnaeus, 1758). Walker, 1856: 88; Butler, 1877: 530; Druce, 1891: 1; Schreiber, 1978: 53 [needs
confirmation]
Callionima Lucas, 1857

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Callionima calliomenae (Schaufuss, 1870). Schreiber, 1978: 50; Miyata, 1997: 20; Haxaire, 2016: 63
=Calliomma ellacombei Rothschild, 1894: 74
=Calliomma lutescens (Butler, 1875a: 5). Butler, 1877: 540
Callionima falcifera falcifera (Gehlen, 1943). Haxaire, 2016: 63
Enyo Hübner, 1819
Enyo lugubris lugubris (Linnaeus, 1771). Walker, 1856: 113; Butler, 1877: 540; Druce, 1891: 6; Schreiber, 1978: 54;
Domínguez, 1997: 68; Haxaire, 2016: 63
Enyo ocypete (Linnaeus, 1758). Schreiber, 1978: 54; Haxaire, 2016: 63
=Enyo danum (Cramer, 1779). Butler, 1877: 541; Druce, 1891: 7
Erinnyis Hübner, 1819
Erinnyis alope alope (Drury, 1770). Walker, 1856: 225; Schreiber, 1978: 49; Domínguez, 1997: 67; Miyata, 1997: 20; Haxaire,
2016: 63
Erinnyis crameri (Schaus, 1898). Miyata, 1997: 20; Haxaire, 2016: 63
Erinnyis ello ello (Linnaeus, 1758). Walker, 1856: 224; Butler, 1877: 603; Druce, 1891: 19; Forbes, 1931: 344; Schreiber, 1978:
49; Schmutterer, 1990: 489; Domínguez, 1997: 67; Miyata, 1997: 20; Haxaire, 2016: 63
Erinnyis guttularis (Walker, 1856: 227). Butler, 1877: 604; Forbes, 1931: 344; Cary, 1951: 120; Schreiber, 1978: 49;
Domínguez, 1997: 67; Haxaire, 2016: 63
Erinnyis lassauxii (Boisduval, 1859). Schreiber, 1978: 49; Domínguez, 1997: 67; Miyata, 1997: 20; Haxaire, 2016: 63
=Dilophonota omphaleae (Boisduval, 1870). Butler, 1877: 603; Druce, 1891: 19
Erinnyis obscura (Fabricius, 1775). Druce, 1891: 19; Walker, 1856: 226; Butler, 1877: 604; Forbes, 1931: 344; Schreiber,
1978: 49; Haxaire, 2016: 63
=Dilophonota rhaebus (Boisduval, 1870). Druce, 1891: 19
=Erinnyis domingonis (Butler, 1875b: 258). Butler, 1877: 604; Schreiber, 1978: 49; Domínguez, 1997: 67
Erinnyis oenotrus (Cramer, 1782). Walker, 1856: 227; Butler, 1877: 603; Druce, 1891: 19; Schreiber, 1978: 49; Miyata, 1997:
20; Haxaire, 2016: 63
Eupyrrhoglossum Grote, 1865
Eupyrrhoglossum sagra (Poey, 1832). Haxaire, 2016: 68
Isognathus Felder & Felder, 1862
Isognathus rimosus molitor Rothschild & Jordan, 1915. Cary, 1951: 120; Schreiber, 1978: 48; Domínguez, 1997: 67; Haxaire,
2016: 63
Pachylia Walker, 1856
Pachylia ficus (Linnaeus, 1758). Walker, 1856: 189; Butler, 1877: 578; Druce, 1891: 15; Schreiber, 1978: 49; Miyata, 1997: 19;
Haxaire, 2016: 63
=Pachylia undatifascia Butler, 1877: 578
Pachylia syces insularis (Hübner, 1822). Butler, 1877: 578; Schreiber, 1978: 50; Miyata, 1997: 19 [needs confirmation]
Pachylioides Hodges, 1971
Pachylioides resumens (Walker, 1856: 190). Butler, 1877: 579; Druce, 1891: 16; Schreiber, 1978: 50; Haxaire, 2016: 63
Perigonia Herrich-Schäffer, 1854
Perigonia caryae Cadiou & Rawlins, 1998: 613. Haxaire, 2016: 66 *DR > TL: La Vega, near mouth Arroyo Los Dajaos, 5 km
E Manabao, 740 m
Perigonia desotoi Haxaire, 2016: 66 *DR > TL: Piste de El Escucharo a Constanza, environ 9 km au nord de Constanza, 1820
m
Perigonia glaucescens Walker, 1856: 103. Butler, 1877: 532; Schreiber, 1978: 53
Perigonia lefebraei (Lucas, 1846). Schreiber, 1978: 53
Perigonia lusca (Fabricius, 1777). Walker, 1856: 101; Butler, 1877: 532; Druce, 1891: 3; Schreiber, 1978: 53; Domínguez,
1997: 68; Miyata, 1997: 20; Haxaire, 2016: 65
Perigonia manni Clark, 1935: 28. Cary, 1951: 120; Schreiber, 1978: 53; Miyata, 1997: 21; Haxaire, 2016: 65
Phrixus Hübner, 1819
Phrixus caicus (Cramer, 1777). Walker, 1856: 228; Butler, 1877: 605; Druce, 1891: 20; Schreiber, 1978: 49; Haxaire, 2016: 63
Pseudosphinx Burmeister, 1855
Pseudosphinx tetrio (Linnaeus, 1771). Butler, 1877: 610; Druce, 1891: 22; Wolcott, 1927: 273; Schreiber, 1978: 48;
Domínguez, 1997: 67; Miyata, 1997: 20; Haxaire, 2016: 63
Tribe PHILAMPELINI
Eumorpha Hübner, 1807

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Eumorpha fasciatus fasciatus (Sulzer, 1776). Butler, 1877: 600; Schreiber, 1978: 55; Domínguez, 1997: 68; Haxaire, 2016: 68
Eumorpha labruscae labruscae (Linnaeus, 1758). Walker, 1856: 178; Butler, 1877: 578; Druce, 1891: 14; Wolcott, 1927: 274;
Schreiber, 1978: 56; Domínguez, 1997: 68; Miyata, 1997: 21; Haxaire, 2016: 68
Eumorpha strenua (Ménétriès, 1857). Walker, 1865: 30; Butler, 1877: 574; Schreiber, 1978: 55; Haxaire, 2016: 68
=Dupo domingonis Rothschild, 1894: 83
Eumorpha vitis vitis (Linnaeus, 1758). Walker, 1856: 176; Butler, 1877: 574; Druce, 1891: 13; Wolcott, 1927: 274; Schreiber,
1978: 55; Domínguez, 1997: 68; Miyata, 1997: 21; Haxaire, 2016: 68
=Philampelus linnei (Grote & Robinson, 1865). Butler, 1877: 574; Druce, 1891: 13
Tribe MACROGLOSSINI
Cautethia Grote, 1865
Cautethia carsusi Haxaire & Schmitt, 2001: 606
Cautethia insolita Haxaire, 2016: 63 *DR > TL: Piste de El Escucharo a Constanza, environ 9 km au nord de Constanza, 1820
m
Cautethia noctuiformis noctuiformis (Walker, 1856: 232). Butler, 1877: 602; Cary, 1951: 120; Schreiber, 1978: 56; Kitching &
Cadiou, 2000: 87
Hyles Hübner, 1819
Hyles lineata lineata (Fabricius, 1775). Druce, 1891: 13; Schreiber, 1978: 61; Domínguez, 1997: 68; Miyata, 1997: 21
Xylophanes Hübner, 1819
Xylophanes chiron necchus (Cramer, 1779). Walker, 1856: 132; Domínguez, 1997: 68
=Chaerocampa haitensis Butler, 1875a: 9. Butler, 1877: 565
Xylophanes clarki Ramsden, 1921. Haxaire, 2016: 69
Xylophanes pluto (Fabricius, 1777). Schreiber, 1978: 57; Miyata, 1997: 21; Haxaire, 2016: 69
=Calliomma thorates (Hübner, 1831). Walker, 1856: 151; Butler, 1877: 540; Druce, 1891: 5
Xylophanes rhodocera (Walker, 1856: 184). Butler, 1877: 567; Cary, 1951: 121; Schreiber, 1978: 57; Haxaire, 2016: 68
Xylophanes tersa tersa (Linnaeus, 1771). Walker, 1856: 131; Butler, 1877: 563; Druce, 1891: 10; Schreiber, 1978: 60;
Domínguez, 1997: 68; Miyata, 1997: 21; Haxaire, 2016: 69
Notes: The species names Cauthethia grotei grotei H. Edwards, 1882, Xylophanes falco (Walker, 1856), Xylophanes irrorata
(Grote, 1865), Xylophanes porcus (Hubner, 1823) and Manduca quinquemaculata (Haworth, 1803), previously included in the
checklist of Perez-Gelabert (2008) are believed to be misidentifications. They are here deleted from the Hispaniolan faunal list.

Superfamily NOCTUOIDEA
Family NOTODONTIDAE
Subfamily DIOPTINAE
Caribojosia Rawlins & Miller, 2008
Caribojosia youngi Rawlins & Miller, 2008: 211. Miller, 2010: 783 *DR > TL: Independencia, Sierra de Neiba just south of
crest, 5 km NNW Angel Feliz, 1780 m
Eremonidia Rawlins & Miller, 2008
Eremonidia mirifica Rawlins & Miller, 2008: 207. Miller, 2010: 102 *DR > TL: Independencia, Sierra de Neiba just south of
crest, 5 km NNW Angel Feliz, 1780 m
Subfamily DUDUSINAE
Antillisa Thiaucourt, 2006
Antillisa barbuti Thiaucourt, 2006: 2 *DR > TL: La Vega, route Constanza á Valle Nuevo, km 19, 1,940 m
Antillisa vincenti Thiaucourt, 2006: 3 *DR > TL: Independencia, route de Puerto Escondido á Pedernales, km 41.5, 1,620 m
Subfamily HEMICERATINAE
Hemiceras Guenée, 1852
Hemiceras domingonis Dyar, 1908: 53
Subfamily HETEROCAMPINAE
Schizuroides Thiacourt, 2016
Schizuroides desotoi Thiacourt, 2016: 14 *DR > TL: La Palma de Constanza, 600 m
Tainocampa Thiacourt, 2016
Tainocampa nigra Thiacourt, 2016: 2 *DR > TL: Duarte, environs de San Francisco de Macoris, Loma Quita Espuela Nat.
Res., 780 m
Subfamily NYSTALEINAE
Hippia Möschler, 1878

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Hippia vittipalpis (Walker, 1857: 1106)
Nystalea Guenée, 1852
Nystalea ebalea (Cramer, 1779). Marcano, 1981: 103
Nystalea aequipars Walker, 1858. Weller, 1990: 41
Nystalea indiana Grote, 1884. Domínguez, 1997: 72
Pentobesa Schaus, 1901
Pentobesa smithsoni Weller, 1991: 805 *DR > TL: La Estrelleta Prov., 4 km SE Rio Limpio, ca. 760 m
Symmerista Hübner, 1821
Symmerista albifrons (Smith, 1797). Domínguez, 1997: 72

Family EREBIDAE
Key references: Watson & Goodger (1986); Vincent & Laguerre (2014).
Subfamily ARCTIINAE
Tribe ARCTIINI
Subtribe ARCTIINA
Caribarctia Ferguson, 1985
Caribarctia bertrandae Vincent, 2006: 223 *DR > TL: Réserve Scientifique Valle Nuevo, route de Constanza a Valle Nuevo,
km 19, 1940 m
Caribarctia cardinalis Ferguson, 1985: 247 *DR > TL: Hotel Nueva Suiza, Constanza, 1164 m, La Vega Prov.
Virbia Herrich-Schaefer, 1835
Virbia insularis (Gaede, 1923: 12) *H > TL: Port-au-Prince
Hypercompe Hübner, 1819
Hypercompe decora (Walker, 1855: 693). Watson & Goodger, 1986: 30; Kirby, 1892: 215; Hampson, 1901: 372
=Ecpantheria haitiensis Oberthür, 1881: 109
Subtribe CALLIMORPHINA
Utetheisa Hübner, 1819
Utetheisa bella (Linnaeus, 1758). Wolcott, 1927: 275; Domínguez, 1997: 72
Utetheisa ornatrix ornatrix (Linnaeus, 1758). Forbes, 1917: 341; Forbes, 1930: 39
=Utetheisa ornatrix stretchii (Butler, 1872). Forbes, 1917: 341; Forbes, 1930: 39
Subtribe CTENUCHINA
Clystea Watson, 1980
Clystea rubipectus (Schaus, 1898: 139). Forbes, 1917: 339
Ctenucha Kirby, 1837
Ctenucha editha (Walker, 1856: 1661)
Ctenucha quadricolor (Walker, 1856: 1867)
Empyreuma Hübner, 1818
Empyreuma affinis Rothschild, 1912. Forbes, 1917: 339
=Empyreuma haitiensis Rothschild, 1912. Forbes, 1917: 339
Empyreuma lichas (Fabricius, 1781). Walker, 1854: 211; Walker, 1856: 1622
Empyreuma pugione (Linnaeus, 1767). Forbes, 1917: 339; Forbes, 1930: 26; Wolcott, 1927: 274
Episcepsis Butler, 1877
Episcepsis dominicensis Rothschild, 1911: 44
Eucereon Hübner, [1819]
Eucereon ochrota Hampson, 1905: 431
Eunomia Hübner, 1818
Eunomia columbina (Fabricius, 1793). Forbes, 1930: 22
Galethalea Butler, 1876
Galethalea portoricensis Pinheiro, 2016: 360
Horama Hübner, 1819
Horama panthalon panthalon (Grote, 1860). Domínguez, 1997: 73
=Mastigocera tibialis Butler, 1876b: 373
Horama rawlinsi McCabe, 1992: 245 *DR > TL: 26 km N Cabo Rojo, 730 m
Lymire Harris, 1839
Lymire candida Forbes, 1917: 344

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Lymire edwardsi (Grote, 1881). Domínguez, 1997: 73
Napata Walker, 1854
Napata munda (Walker, 1856: 1680)
Nelphe Herrich-Schäffer, 1858
Nelphe trimaculata Pinheiro, 2016: 488 *H > TL: Haiti
Subtribe CTENUCHINA
Cosmosoma Hübner, [1823]
Cosmosoma achemon (Fabricius, 1781: 162)
Cosmosoma auge (Linnaeus, 1767: 807)
Cosmosoma odilae Laguerre, 2014: 5 *DR > TL: Road El Arroyo - El Aguacate, PK 5 (Pedernales), alt. 1995 m
Nyridela Lucas, 1857
Nyridela chalciope (Hübner, 1827). Forbes, 1917: 339
Phoenicoprocta Bruce, 1898
Phoenicoprocta parthenii (Fabricius, 1793). Forbes, 1930: 20
=Glaucopis multicincta Walker, 1854: 163. Walker, 1856: 1606; Forbes, 1930: 20
Subtribe PERICOPINA
Composia Hübner, 1820
Composia credula (Fabricius, 1775). Bates, 1933: 121; Butler, 1872: 53; Bryk, 1931: 8
=Composia sybaris Moschler, 1890: 116; Walker, 1856: 1655; Forbes, 1930: 40; Wolcott, 1948: 580
Ctenuchidia Grote, 1865
Ctenuchidia subcyanea (Walker, 1845). Watson & Goodger, 1986: 19
Hyalurga Hübner, 1819
Hyalurga vinosa (Drury, 1773). Watson & Goodger, 1986: 36
=Lauron domingonis Butler, 1876a: 120
Stenognatha Felder, 1874
Stenognatha flinti Todd, 1982: 316. Watson & Goodger, 1986: 36 *DR > TL: 12 km N. of Constanza
Subtribe PHAEGOPTERINA
Ammalo Walker, 1855
Ammalo helops (Cramer, [1776]: 113). Strand, 1919: 35; Kirby, 1892: 212
Bituryx Hampson, 1901
Bituryx lanceolata (Walker, 1856: 1702). Watson & Goodger, 1986: 26 *DR > TL: St. Domingo
Calidota Dyar, 1901
Calidota strigosa (Walker, 1855: 736). Moschler, 1890: 115; Clemens, 1861: 535; Watson & Goodger, 1986: 19; Vincent &
Laguerre, 2010: 179 *DR > TL: St. Domingo
Halysidota Hübner, 1819
Halysidota ata Watson, 1980: 50. Watson & Goodger, 1986: 22
=Halysidota tesselaris (J. E. Smith, 1797). Domínguez, 1997: 72 [Misidentification]
=Halysidota cinctipes Grote, 1865, Strand, 1919: 73 [Misidentification]
Lophocampa Harris, 1841
Lophocampa albitegula Vincent, 2011: 73 *DR > TL: Track from Los Arroyos to El Aguacate pK 5.2, 1990 m
Lophocampa baorucoensis Vincent, 2005a: 4 *DR > TL: Independencia, Sierra Baoruco national Park, route de Puerto
Escondido à Pedernales, kilomètre 41.5, 1620 m
Lophocampa duarteiensis Vincent, 2005b: 11 *DR > TL: Duarte, environs de San Francisco de Macorís, Réserve Scientifique
Loma Quita Espuela, 750 m alt. (650 m du sommet)
Lophocampa hispaniola Vincent, 2009: 47 *DR > TL: Réserve scientifique Loma Quita Espuela, 650 mètres du sommet, alt.
750 m
Lophocampa latepunctata Vincent, 2005b: 13 *DR > TL: Independencia, piste la Descubierta à Aniceto Martínez, kilomètre
32, 1750 m
Lophocampa lesieuri Vincent, 2005a: 1 *DR > TL: Independencia, piste la Descubierta à Aniceto Martínez, kilomètre 32, 1750
m
Lophocampa lineata Vincent, 2011: 71 *DR > TL: Road El Blanco to Constanza, pK [kilometer post] 10, Ebano Verde
Scientific Reserve, 1360 m
=Halisidota caryae (Harris, 1841). Domínguez, 1997: 72 [Misidentification]
Lophocampa neibaensis Vincent, 2005b: 10 *DR > TL: Independencia, piste la Descubierta à Aniceto Martinez, km 32, 1750

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m
Opharus Walker, 1855
Opharus bipunctata Vincent & Laguerre in Vincent et al., 2009: 70 *DR > TL: route Puerto Escondido à Pedernales, km 41,5,
1620 m
Pareuchaetes Grote, 1900
Pareuchaetes insulata (Walker, 1855: 734). Clemens, 1861: 534; Druce, 1891: 392; Strand, 1919: 36; Butler, 1875: 38; Forbes,
1931: 342
Robinsonia Grote, [1866]
Robinsonia formula Grote, [1866]; Forbes, 1931: 342
Westindia Vincent in Vincent et al., 2014
Westindia haxairei Vincent in Vincent et al., 2014: 293 *DR > TL: Road El Blanco to Constanza pK 10, Reserva Scientifica
Ebano Verde, alt. 1360 m
Tribe LITHOSIINI
Aethosia Hampson, 1900
Aethosia ectrocta Hampson, 1900: 385 *H > TL: Haiti
Areva Walker, 1854
Areva trigemmis (Hübner, 1824–1831). Walker, 1854: 538
Boenasa Walker, 1865
Boenasa angelica Schaus, 1924: 30
Boenasa nigrorosea Walker, 1865e: 266 *DR, *H > TL: Haiti
Mulona Walker, 1866
Mulona lapidaria Walker, 1866: 1896 *H, *DR > TL: St. Domingo
Torycus Herrich-Schäffer, 1866
Torycus domingonis Schaus, 1924: 29

Family NOCTUIDAE
Key reference: Poole (1989).
Subfamily ACONTIINAE
Acontia Ochsenheimer, 1816
Acontia tetragona Walker, 1857b: 786. Druce, 1891: 305; Poole, 1989: 18
Chobata Walker, 1857
Chobata discalis Walker, 1857b: 838. Schaus, 1940: 213; Poole, 1989: 248
=Chobata concludens Walker, 1865c: 791. Poole, 1989: 248
Cobubatha Walker, 1863
Cobubatha metaspilaris Walker, 1863: 126. Poole, 1989: 258
=Agrophila signiferana Walker, 1865c: 775. Poole, 1989: 258
Cydosia Duncan & Westwood, 1841
Cydosia nobilitella Cramer, 1779. Walker, 1856a: 1683; Domínguez, 1997: 69; Poole, 1989: 302
=Cydosia chrysorrhaeella Guenée, 1879. Poole, 1989: 302
=Cydosia cyanella Guenée, 1879. Poole, 1989: 302
=Cydosia graciella Guenée, 1879. Poole, 1989: 302
=Eggyna submutata Walker, 1866: 1895. Walsingham, 1891: 541; Schaus, 1940: 206; Poole, 1989: 302
Eublemma Hübner, 1821
Eublemma recta (Guenée, 1852). Poole, 1989: 387
=Anthophila flammicincta Walker, 1865c: 801. Poole, 1989: 387
Eublemma rosescens Hampson, 1898. Poole, 1989: 392
Marimatha Walker, 1865
Marimatha trajectalis Walker, 1865d: 1519. Poole, 1989: 624
Ponometia Herrich-Schäffer, 1868
Ponometia exigua (Fabricius, 1793). Poole, 1989: 832
=Ponometia costalis (Walker, 1857b: 787). Druce, 1891: 308; Poole, 1989: 832
=Ponometia dimidiata (Walker, 1865c: 783). Poole, 1989: 832
Spragueia Grote, 1875
Spragueia dama (Guenée, 1852). Poole, 1989: 927

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 349
 =Agrophila transmutata Walker, 1865c: 776. Druce, 1891: 301; Poole, 1989: 927
Spragueia margana (Fabricius, 1794). Poole, 1989: 927
=Agrophila rudisana Walker, 1865c: 776. Druce, 1891: 302; Poole, 1989: 927
=Spragueia canofusa Hampson, 1898. Poole, 1989: 926
Tarachidia Hampson, 1898
Tarachidia venustula (Walker, 1865c: 783). Poole, 1989: 950
=Acontia discoidalis Walker, 1866: 1962. Poole, 1989: 950
Subfamily AGARISTINAE
Caularis Walker, 1858
Caularis undulans Walker, 1857. Schaus, 1940: 205; Todd, 1966: 2; Poole, 1989: 227
Seirocastnia Grote, 1866
Seirocastnia tribuna (Hübner, 1825). Druce, 1891: 36; Poole, 1989: 904
=Seirocastnia continua (Walker, 1856a: 1646). Poole, 1989: 904
Subfamily AMPHIPYRINAE
Antachara Walker, 1858
Antachara rotundata Walker, 1858b: 1741. Poole, 1989: 94
=Laphygma lignigera Walker, 1865b: 650. Poole, 1989: 94
Callopistria Hübner, 1821
Callopistria floridensis Guenée, 1852. Domínguez, 1997: 69
Catabena Walker, 1865
Catabena vitrina (Walker, 1857). Poole, 1989: 210
Condica Walker, 1856
Condica albigera (Guenée, 1852). Poole, 1989: 265
=Apamea intermittens Walker, 1858b: 1686. Druce, 1891: 273; Poole, 1989: 265
Condica circuita (Guenée, 1852). Walker, 1858b: 1687; Poole, 1989: 266
=Celaena prolifera Walker, 1857a: 732. Poole, 1989: 266
Condica concisa (Walker, 1856). Poole, 1989: 265
=Perigea imbella Walker, 1858b: 1692. Poole, 1989: 265
Condica mobilis (Walker, 1856b: 277). Poole, 1989: 265
=Celaena inclinata Walker, 1857a: 732. Poole, 1989: 265
Condica punctifera (Walker, 1857). Poole, 1989: 266
=Celaena semifurca Walker, 1857a: 732. Poole, 1989: 266
Condica sufficiens (Walker, 1858b: 1692). Poole, 1989: 267
Condica sutor (Guenée, 1852). Poole, 1989: 266
=Perigea otiosa Walker, 1858b: 1693. Poole, 1989: 266
Cropia Walker, 1858
Cropia indigna (Walker, 1857c: 1115). Poole, 1989: 283
Cropia subapicalis (Walker, 1857c: 1117). Poole, 1989: 283
Dypterygia Stephens, 1829
Dypterygia punctirena (Walker, 1857a: 586). Schaus, 1940: 190; Frank & McCoy, 1992: 12
Hampsonodes Nye, 1975
Hampsonodes ampliplaga (Walker, 1858b: 1687). Poole, 1989: 485
Micrathetis Hampson, 1908
Micrathetis triplex (Walker, 1857). Poole, 1989: 660; Domínguez, 1997: 69
=Micratethis spilomela (Walker, 1865b: 648). Poole, 1989: 660
Speocropia Hampson, 1908
Speocropia scriptura (Walker, 1858b: 1723). Schaus, 1940: 188; Poole, 1989: 919
=Xylina transversalis Walker, 1858b: 1734. Poole, 1989: 919
Subfamily BAGISARINAE
Bagisara Walker, 1858
Bagisara repanda (Fabricius, 1793). Poole, 1989: 154
=Anthophila erecta Walker, 1857b: 828. Druce, 1891: 499; Poole, 1989: 154
=Atethmia subusta Hübner, 1818. Druce, 1891: 290; Christy, 1898: 368; Domínguez, 1997: 69
=Laphygma trilinea Walker, 1865b: 649. Poole, 1989: 154

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Bagisara tristicta Hampson, 1898. Poole, 1989: 154
Subfamily CATOCALINAE
Achaea Hübner, 1823
Achaea ablunaris (Guenée, 1852). Poole, 1989: 8
=Geria restituta Walker, 1858a: 1366. Poole, 1989: 11
Alabama Grote, 1865
Alabama argillacea (Hübner, 1823). Druce, 1891: 336; Wolcott, 1927b: 286; Smith & Audant, 1930: 974; Domínguez, 1997:
69
Anomis Hübner, 1821
Anomis editrix (Guenée, 1852). Druce, 1891: 339; Poole, 1989: 88
=Gonitis albitacta Walker, 1857c: 999. Poole, 1989: 88
=Gonitis cosmioides Walker, 1857c: 1000. Poole, 1989: 89
=Gonitis fractifera Walker, 1857c: 998. Poole, 1989: 90
Anomis erosa (Hübner, 1821). Walker, 1857: 986; Druce, 1891: 336; Butler, 1894: 298; Domínguez, 1997: 69
Anomis exacta Hübner, 1822. Walker, 1857: 989; Druce, 1891: 337; Poole, 1989: 89
=Anomis conducta Walker, 1857c: 990. Poole, 1989: 89
Anomis flava (Fabricius, 1775). Domínguez, 1997: 69
Anomis impasta Guenée, 1852. Domínguez, 1997: 69
Anomis luridula Guenée, 1852. Poole, 1989: 91
=Anomis derogata Walker, 1857c: 990. Poole, 1989: 89
Anomis properans (Walker, 1857c: 999). Poole, 1989: 92
Antiblemma Hübner, 1821
Antiblemma brevipennis (Walker, 1865c: 1039). Poole, 1989: 96
Antiblemma leucospila (Walker, 1865c: 998). Poole, 1989: 96
Antiblemma rufinans (Guenée, 1852). Poole, 1989: 98
=Antiblemma subcinerascens (Walker, 1858b: 1614). Druce, 1891: 415; Poole, 1989: 98
Antiblemma sterope (Walker, 1858). Druce, 1891: 415; Poole, 1989: 98
=Antiblemma obliterata (Walker, 1858b: 1600). Poole, 1989: 97
Anticarsia Hübner, 1818
Anticarsia gemmatalis Hübner, 1818. Walker, 1858b: 1560; Druce, 1891: 400; Poole, 1989: 99; Schmutterer, 1990: 364;
Domínguez, 1997: 68
=Plaxia spiloleuca Walker, 1858b: 1624. Poole, 1989: 99
=Plaxia subducta Walker, 1858b: 1624. Poole, 1989: 99
=Thermesia monstratura (Walker, 1858b: 1564). Poole, 1989: 99
Ascalapha Hübner, 1809
Ascalapha odorata (Linnaeus, 1758). Walker, 1858a: 1290; Druce, 1891: 376; Wolcott, 1927b: 285
Azeta Guenée, 1852
Azeta quassa Walker, 1858b: 1581. Druce, 1891: 413; Poole, 1989: 153
Azeta repugnalis Hübner, 1831. Druce, 1891: 402; Poole, 1989: 153
=Azeta mirzah Guenée, 1852. Walker, 1858b: 1579; Poole, 1989: 153
=Chabora undulifera Walker, 1865c: 1114. Poole, 1989: 153
=Thermesia fusilinea Walker, 1858b: 1564. Poole, 1989: 153
Baniana Walker, 1858
Baniana relapsa (Walker, 1858b: 1836). Schaus, 1940: 249; Poole, 1989: 156
=Baniana significans Walker, 1858b: 1843. Poole, 1989: 156
Batina Walker, 1865
Batina marginalis Walker, 1865c: 865. Poole, 1989: 158
Bulia Walker, 1858
Bulia confirmans (Walker, 1857c: 1157). Poole, 1989: 180; Pogue & Laughlin, 2002: 136
=Bulia propria Walker, 1857c: 1170. Poole, 1989: 180
Caenurgia Walker, 1858
Caenurgia adusta (Walker, 1865c: 988). Poole, 1989: 182
Casandria Walker, 1857
Casandria ferrocana Walker, 1857. Poole, 1989: 210

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 351
=Laphygma nigriscripta Walker, 1865b: 649. Poole, 1989: 210
Antapistis Hübner, 1825
Antapistis leucospila (Walker, 1865). Poole, 1989: 94
=Nazuda digestalis Walker, 1865c: 1118. Poole, 1989: 94
Celiptera Guenée, 1852
Celiptera levina Stoll, 1782. Druce, 1891: 387; Poole, 1989: 229
=Ophiusa variolosa Walker, 1858a: 1421. Poole, 1989: 229
Coenipeta Hübner, 1818
Coenipeta mollis Walker, 1865c: 892. Poole, 1989: 260
Concana Walker, 1858
Concana mundissima Walker, 1857b: 940. Poole, 1989: 263
Diphthera Hübner, 1809
Diphthera festiva (Fabricius, 1775). Thöny, 1997: 25
=Diphthera hieroglyphica (Cramer, 1777). Walker, 1856b: 145; Druce, 1891: 263; Wolcott, 1927b: 285
Elousa Walker, 1857
Elousa albicans Walker, 1857c: 1118. Poole, 1989: 354; Frank & McCoy, 1992: 12
=Masebia famelica Walker, 1858b: 1772. Poole, 1989: 354
Ephyrodes Guenée, 1852
Ephyrodes cacata Guenée, 1852. Walker, 1858b: 1587; Druce, 1891: 409; Poole, 1989: 361
=Ephyrodes exprimens Walker, 1858b: 1589. Poole, 1989: 361
=Ephyrodes implens Walker, 1858b: 1588. Druce, 1891: 409; Poole, 1989: 361
Ephyrodes omicron Guenée, 1852. Walker, 1858b: 1588; Poole, 1989: 361
Epidromia Guenée, 1852
Epidromia lienaris (Hübner, 1823). Becker, 2001: 962
=Thermesia suffusa Walker, 1858b: 1561. Druce, 1891: 391; Poole, 1989: 362; Becker, 2001: 962
=Thermesia valida Walker, 1865c: 953; Becker, 2001: 962
Epidromia pyraliformis (Walker, 1858b: 1562). Poole, 1989: 362; Frank & McCoy, 1992: 12
=Epidromia concisa Walker, 1865a: 107. Poole, 1989: 362
=Poaphila profecta Walker, 1858b: 1836. Poole, 1989: 362
=Phurys arenosa Walker, 1865c: 997. Poole, 1989: 362
=Phurys pedestris Walker, 1858b: 1844. Poole, 1989: 362
=Thermesia credibilis Walker, 1858b: 1565. Poole, 1989: 362
=Thermesia sigillata Walker, 1858b: 1562. Poole, 1989: 362
Epitausa Walker, 1857
Epitausa laetabilis Walker, 1857. Poole, 1989: 367
Eudocima Billberg, 1820
Eudocima serpentifera (Walker, 1857c: 1218). Druce, 1891: 361; Poole, 1989: 400
Eulepidotis Hübner, 1823
Eulepidotis addens (Walker, 1858b: 1768). Poole, 1989: 404
Eulepidotis modestula (Herrich-Schäffer, 1869). Schaus, 1940: 248
Glympis Walker, 1859
Glympis arenalis (Walker, 1865d: 1196). Poole, 1989: 460
Glympis eubolialis (Walker, 1865d: 1153). Schaus, 1940: 251; Poole, 1989: 460
=Aluaca anaitisalis (Walker, 1865d: 1153). Poole, 1989: 460
Gonodonta Hübner, 1818
Gonodonta bidens bidens Geyer, 1832. Todd, 1959: 38
Gonodonta incurva (Sepp, 1840). Todd, 1959: 45; Domínguez, 1997: 68
Gonodonta nitidimacula Guenée, 1852. Wolcott, 1927b: 289; Todd, 1959: 47
Gonodonta nutrix (Stoll, 1780). Walker, 1857b: 949; Druce, 1891: 254; Domínguez, 1997: 68
Gonodonta unica Neumoegen, 1891. Domínguez, 1997: 68
Gonodonta uxoria (Cramer, 1780). Todd, 1959: 26
Hemeroblemma Hübner, 1818
Hemeroblemma numeria (Drury, 1773). Druce, 1891: 363; Schaus, 1940a: 243
=Peosina mexicana Guenée, 1852. Walker, 1857c: 1243

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Hemeroblemma rengus (Poey, 1832). Walker, 1858a: 1252; Druce, 1891: 366
Hemeroplanis Hübner, 1818
Hemeroplanis aurora (Walker, 1865c: 1039). Todd, 1960: 510; Poole, 1989: 496
Hemeroplanis scopulepes Haworth, 1809. Todd, 1960: 507
=Hemeroplanis pyraloides Hübner, 1823. Druce, 1891: 412
Hypocala Guenée, 1852
Hypocala andremona (Stoll, 1781). Druce, 1891: 359; Poole, 1989: 540
=Hypocala pierreti Guenée, 1852. Walker, 1857c: 1173; Poole, 1989: 540
Hypogrammodes Hampson, 1913
Hypogrammodes balma (Guenée, 1852). Walker, 1857c: 1096; Poole, 1989: 541
Isogona Guenée, 1852
Isogona scindens (Walker, 1858a: 1829). Poole, 1989: 556
=Isogona ordinans (Walker, 1858b: 1837). Druce, 1891: 383; Poole, 1989: 556
Kakopoda Smith, 1900
Kakopoda progenies (Guenée, 1852). Poole, 1989: 558
=Kakopoda directa (Walker, 1857c: 1061). Poole, 1989: 558
=Kakopoda infausta (Walker, 1857c: 1059). Poole, 1989: 558
Lesmone Hübner, 1818
Lesmone formularis (Geyer, 1837). Walker, 1858a: 1339; Druce, 1891: 377; Poole, 1989: 574
=Bendis fuscifascia Walker, 1858b: 1823. Poole, 1989: 574
=Bendis impar Guenée, 1852. Poole, 1989: 574
Letis Hübner, 1821
Letis intracta Walker, 1858a: 1266. Poole, 1989: 575
Letis mycerina Cramer, 1777. Walker, 1858a: 1265; Druce, 1891: 369; Poole, 1989: 574
=Letis atricolor (Guenée, 1852). Walker, 1858a: 1266; Poole, 1989: 574
Litoprosopus Grote, 1869
Litoprosopus haitiensis Hampson, 1926. Poole, 1989: 599; Frank & McCoy, 1992: 12
Litoprosopus hatuey (Poey, 1832). Walker, 1857b: 855
Litoprosopus puncticosta Hampson, 1926. Poole, 1989: 599; Barbut & Lalanne-Cassou, 2009: 413
Massala Walker, 1865
Massala obvertens (Walker, 1858b: 1580). Poole, 1989: 628
Melipotis Hübner, 1818
Melipotis acontioides (Guenée, 1852). Abud, 1981: 83
Melipotis contorta (Guenée, 1852). Schaus, 1940: 236; Poole, 1989: 642
=Bolina bistriga (Walker, 1857c: 1155). Druce, 1891: 356; Poole, 1989: 642
Melipotis famelica (Guenée, 1852). Domínguez, 1997: 69
=Melipotis bivittata (Walker, 1857c: 1156). Druce, 1891: 356
Melipotis fasciolaris (Hübner, 1825). Walker, 1857c: 1147; Druce, 1891: 354; Domínguez, 1997: 68
Melipotis indomita (Walker, 1857). Domínguez, 1997: 68
Melipotis januaris (Guenée, 1852). Walker, 1857c: 1149; Poole, 1989: 643; Domínguez, 1997: 69
=Bolina excavans Walker, 1857c: 1154. Poole, 1989: 643
=Bolina subtilis Walker, 1857c: 1156. Druce, 1891: 357; Poole, 1989: 644
Melipotis jucunda Hübner, 1818. Domínguez, 1997: 69
Melipotis lucigera (Walker, 1857c: 1152). Poole, 1989: 643
Melipotis ochrodes (Guenée, 1852). Walker, 1857c: 1151; Druce, 1891: 355; Poole, 1989: 644
=Bolina parens Walker, 1857c: 1154. Poole, 1989: 644
=Bolina terminifera Walker, 1857c: 1151. Poole, 1989: 644
Melipotis perpendicularis (Guenée, 1852). Poole, 1989: 643; Domínguez, 1997: 68
=Bolina glaucipennis Walker, 1857c: 1153. Poole, 1989: 643
Melipotis strigifera (Walker, 1857c: 1153). Poole, 1989: 644
=Coronta surrepens Walker, 1857c: 1171. Poole, 1989: 644
Metallata Möschler, 1890
Metallata absumens (Walker, 1862). Poole, 1989: 653
=Thermesia alacris Walker, 1865c: 1041. Poole, 1989: 652

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 353
=Thermesia conficita Walker, 1865c: 1042. Poole, 1989: 653
=Thermesia funesta Walker, 1865c: 1040. Poole, 1989: 653
=Thermesia saturata Walker, 1865c: 1043. Poole, 1989: 653
Metria Hübner, 1823
Metria irresoluta (Walker, 1858). Poole, 1989: 657
Mocis Hübner, 1823
Mocis diffluens (Guenée, 1852). Druce, 1891: 388; Poole, 1989: 669
=Remigia mensuralis Walker, 1858a: 1499. Poole, 1989: 669
Mocis disseverans (Walker, 1858a: 1495). Druce, 1891: 389; Poole, 1989: 669; Domínguez, 1997: 70
=Remigia persubtilis Walker, 1858a: 1497. Poole, 1989: 669
Mocis exscindens (Walker, 1858a: 1500). Poole, 1989: 668
Mocis latipes (Guenée, 1852). Walker, 1858a: 1494; Druce, 1891: 388; Domínguez, 1997: 70
Mocis repanda (Fabricius, 1794). Walker, 1858a: 1497; Wolcott, 1927b: 283; Poole, 1989: 669
=Remigia remanens Walker, 1858a: 1498. Poole, 1989: 669
Ophisma Guenée, 1852
Ophisma tropicalis Guenée, 1852. Poole, 1989: 731
=Ophisma detrahens Walker, 1858a: 1368. Druce, 1891: 379; Poole, 1989: 731
Ophiusa Ochsenheimer, 1816
Ophiusa salmus Guenée, 1852: 267. Walker, 1858a: 1421
Oraesia Guenée, 1852
Oraesia basiplaga (Walker, 1865c: 847). Poole, 1989: 737
Oraesia excitans Walker, 1857b: 942. Druce, 1891: 252; Schaus, 1940: 259; Poole, 1989: 737
Oraesia honesta Walker, 1857b: 943. Poole, 1989: 737
Panula Guenée, 1852
Panula inconstans (Guenée, 1852). Walker, 1858b: 1813; Druce, 1891: 353; Poole, 1989: 766; Domínguez, 1997: 69
=Alamis olivacea Walker, 1865c: 876. Poole, 1989: 766
=Athyrma tetragona Walker, 1865c: 965. Poole, 1989: 766
=Ophiusa narrans Walker, 1858b: 1828. Poole, 1989: 766
=Poaphila consequens Walker, 1858b: 1835. Poole, 1989: 766
=Poaphila contacta Walker, 1858a: 1475. Poole, 1989: 766
=Poaphila subordinata Walker, 1865c: 989. Poole, 1989: 766
Pararcte Hampson, 1926
Pararcte immanis (Walker, 1858b: 1810). Poole, 1989: 776
Perasia Hübner, 1823
Perasia flexa (Guenée, 1852). Walker, 1858a: 1482; Poole, 1989: 783
Perasia garnoti (Guenée, 1852). Walker, 1858a: 1483; Druce, 1891: 386
Perasia inficita (Walker, 1865c: 994). Poole, 1989: 783
Plusiodonta Guenée, 1852
Plusiodonta stimulans (Walker, 1857b: 963. Poole, 1989: 817
Ptichodis Hübner, 1818
Ptichodis immunis (Guenée, 1852). Walker, 1858a: 1482; Druce, 1891: 385; Poole, 1989: 857
=Ptichodis tussia (Guenée, 1852). Poole, 1989: 857
Ptichodis infecta (Walker, 1858a: 1487). Poole, 1989: 856
=Ptichodis refracta Walker, 1858a: 1488. Poole, 1989: 857
Radara Walker, 1862
Radara nealcesalis (Walker, 1859: 883). Poole, 1989: 860
Radara tauralis (Walker, 1865d: 1185). Poole, 1989: 860
Renodes Guenée, 1852
Renodes aequalis (Walker, 1865d: 1121). Poole, 1989: 866
Renodes eupithecioides (Walker, 1858b: 1593). Poole, 1989: 866
Syllectra Hübner, 1819
Syllectra ericata (Cramer, 1780). Walker, 1858b: 1542; Druce, 1891: 397; Domínguez, 1997: 69
Syllectra congemmalis Hübner, 1823. Walker, 1858b: 1542; Druce, 1891: 397
Thysania Dalman, 1824

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Thysania zenobia (Cramer, 1777). Walker, 1858a: 1287; Druce, 1891: 374; Domínguez, 1997: 70
Toxonprucha Möschler, 1890
Toxonprucha diffundens (Walker, 1858). Poole, 1989: 966
=Homoptera minuscula Walker, 1865c: 880. Poole, 1989: 966
Toxonprucha excavata (Walker, 1865c: 879). Poole, 1989: 966
Trigonodes Guenée, 1852
Trigonodes lucassi Guenée, 1852. Poole, 1989: 975
=Trigonodes problematica Walker, 1858a: 1448. Poole, 1989: 975
Tyrissa Walker, 1866
Tyrissa recurva Walker, 1866: 1976. Poole, 1989: 982
Zale Hübner, 1818
Zale albidula (Walker, 1865c: 880). Poole, 1989: 1007
Zale fictilis Guenée, 1852. Poole, 1989: 1008
=Homoptera gradata Walker, 1857. Druce, 1891: 343; Poole, 1989: 1008
Zale fuliginosa (Walker, 1857c: 1059). Poole, 1989: 1008
Zale peruncta (Guenée, 1852). Poole, 1989: 1008
=Letis incipiens Walker, 1858a: 1266. Poole, 1989: 1008
Subfamily CHLOEPHORINAE
Collomena Möschler, 1890
Collomena filifera (Walker, 1857a: 719). Schaus, 1940: 223; Poole, 1989: 261
Subfamily EUSTROTIINAE
Tripudia Grote, 1877
Tripudia grapholithoides (Möschler, 1890). Pogue, 2009: 80
Tripudia flavibrunnea Pogue, 2009: 83
Tripudia lamina Pogue, 2009: 85
Subfamily EUTELIINAE
Eutelia Hübner, 1823
Eutelia furcata (Walker, 1865c: 815). Poole, 1989: 421
=Eutelia distracta Walker, 1865c: 819. Poole, 1989: 421
Paectes Hübner, 1818
Paectes asper Pogue, 2013: 140
Paectes nana Walker, 1865b: 425. Pogue, 2013: 134
Subfamily GLOTTULINAE
Xanthopastis Hübner, 1821
Xanthopastis timais (Cramer, 1782). Walker, 1856b: 142; Druce, 1891: 263; Reynoso Salcedo, 1985: 23
Subfamily HADENINAE
Elaphria Hübner, 1818
Elaphria cuprescens (Hampson, 1909). Poole, 1989: 350
Elaphria deliriosa (Walker, 1857a: 731). Poole, 1989: 350
Elaphria nucicolora (Guenée, 1852). Domínguez, 1997: 69
Lacinipolia McDunnough, 1937
Lacinipolia distributa (Möschler, 1886). Todd & Poole, 1981: 184
Leucania Ochsenheimer, 1816
Leucania chejela (Schaus, 1921). Adams, 2001: 194
Leucania clarescens Möschler, 1856. Adams, 2001: 209
Leucania humidicola Guenée, 1852. Adams, 2001: 202
Leucania dorsalis Walker, 1856. Frank & McCoy, 1992: 12; Adams, 2001: 197
Leucania incognita (Barnes & McDunnough, 1918). Adams, 2001: 211
Leucania inconspicua Herrich-Schäffer, 1868. Adams, 2001: 192
Leucania lamisma Adams, 2001: 204 *DR > TL: Pedernales, 37 km N Cabo Rojo, 1500 m
Leucania latiuscula Herrich-Schäffer, 1868. Adams, 2001: 200
Leucania lobrega Adams, 2001: 189
Leucania neiba Adams, 2001: 217 *DR > TL: Elías Piña, North slope Sierra de Neiba, 2 km SW Canada, 7 km WSW Hondo
Valle, 980 m

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 355
Leucania rawlinsi Adams, 2001: 186
Leucania sonroja Adams, 2001: 191 *DR > TL: Pedernales, 26 km N Cabo Rojo, 730 m
Leucania senescens Möschler, 1890. Adams, 2001: 195
Mythimna Ochsenheimer, 1816
Mythimna unipuncta (Haworth, 1809). Schmutterer, 1990: 412; Domínguez, 1997: 69; SEA, 1999: 45
Spodoptera Guenée, 1852
Spodoptera albula (Walker, 1857). Druce, 1891: 296; Poole, 1989: 922; Pogue, 2002: 59
=Spodoptera orbicularis Walker, 1857a: 719. Druce, 1891: 267; Poole, 1989: 925; Pogue, 2002: 59
Spodoptera dolichos (Fabricius, 1794). Pogue, 2002: 71
=Prodenia commelinae Guenée, 1852. Walker, 1857a: 722; Druce, 1891: 258; Pogue, 2002: 71
Spodoptera eridania (Cramer, 1782). Domínguez, 1997: 69; Pogue, 2002: 57
Spodoptera exigua (Hübner, 1808). Todd, 1975: 14; Schmutterer, 1990: 427; Domínguez, 1997: 69; Pogue, 2002: 40
Spodoptera frugiperda (J. E. Smith, 1797). Wolcott, 1927b: 280; Santoro, 1960: 386; Schmutterer, 1990: 21; Domínguez, 1997:
69; Pogue, 2002: 69
=Prodenia signifera Walker, 1856: 722
Spodoptera latifascia (Walker, 1856). Poole, 1989: 926; Domínguez, 1997: 69; Pogue, 2002: 83
=Spodoptera variolosa Walker, 1857a: 722. Druce, 1891: 269; Poole, 1989: 926
Spodoptera ornithogalli (Guenée, 1852). Wolcott, 1927b: 283; Schmutterer, 1990: 571; Domínguez, 1997: 69; Pogue, 2002: 92
Spodoptera pulchella (Herrich-Schäffer, 1868). Pogue, 2002: 94
Subfamily HELIOTHINAE
Helicoverpa Hardwick, 1965
Helicoverpa zea (Boddie, 1850). Schmutterer, 1990: 325
=Helicoverpa obsoleta (Fabricius, 1793). Wolcott, 1927b: 277
Heliothis Ochsenheimer, 1816
Heliothis lucilinea (Walker, 1858b: 1749). Poole, 1989: 492
Heliothis subflexa (Guenée, 1852). Domínguez, 1997: 69
Heliothis virescens (Fabricius, 1777). Poole, 1989: 494; Poole et al., 1993: 6; Domínguez, 1997: 69
=Xanthia viridescens (Walker, 1856: 465). Poole, 1989: 494
Subfamily HERMINIINAE
Bleptina Guenée, 1854
Bleptina acastusalis Walker, 1858c: 122. Schaus, 1940: 275; Poole, 1989: 168
Bleptina atymnusalis (Walker, 1858c: 247). Poole, 1989: 168
Bleptina caradrinalis Guenée, 1854. Poole, 1989: 169
=Bleptina priassalis Walker, 1858c: 123. Poole, 1989: 170
Bleptina hydrillalis Guenée, 1854. Walker, 1858c: 120; Poole, 1989: 169
=Bleptina phanasgalis Walker, 1859. Poole, 1989: 170
=Bleptina ulricusalis Walker, 1858c: 121. Poole, 1989: 170
Bleptina menalcasalis Walker, 1858c: 123. Poole, 1989: 169
Bleptina pithosalis Walker, 1858c: 122. Poole, 1989: 170
=Megatomis ferrilinealis Walker, 1865d: 1165. Poole, 1989: 169
Heterogramma Guenée, 1854
Heterogramma micculalis Guenée, 1854. Walker, 1858c: 148; Poole, 1989: 505
Hypenula Grote, 1876
Hypenula complectalis (Guenée, 1854). Walker, 1858c: 120; Poole, 1989: 536
=Hypenula flesalis Walker, 1858c: 121. Poole, 1989: 536
Lophophora Möschler, 1890
Lophophora thaumasalis (Walker, 1858c: 109). Poole, 1989: 601
Mursa Walker, 1859
Mursa phthisialis (Guenée, 1854). Walker, 1858c: 141; Poole, 1989: 673
Mursa sotiusalis (Walker, 1859: 840). Poole, 1989: 673
=Megatomis judicatalis Walker, 1859: 882. Poole, 1989: 672
=Scopula comptalis Walker, 1865d: 1462. Poole, 1989: 672
Salia Hübner, 1818
Salia ferrigeralis (Walker, 1865d: 1165). Poole, 1989: 886

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Subfamily HYPENINAE
Hypena Schrank, 1802
Hypena concinnulalis Walker, 1865d: 1134. Poole, 1989: 522
Hypena (Ophiuche) lividalis (Hübner, 1796). Druce, 1891: 427; Lödl, 1994: 495
=Hypena scissalis Walker, 1865d: 1133. Poole, 1989: 531; Lödl, 1994: 496
Hypena pacatalis Walker, 1858c: 80; Druce, 1891: 432; Poole, 1989: 529
Hypena subidalis Guenée, 1854. Poole, 1989: 530
=Hypena pilosalis Guenée, 1852. Walker, 1858c: 40; Druce, 1891: 428; Poole, 1989: 530
Hypena vetustalis Guenée, 1852. Walker, 1858c: 40; Druce, 1891: 430
Subfamily NOCTUINAE
Agrotis Ochsenheimer, 1816
Agrotis apicalis (Herrich-Schaffer, 1868). Schmutterer, 1990: 454
Agrotis subterranea (Fabricius, 1794). Poole, 1989: 47; Schmutterer, 1990: 409; Domínguez, 1997: 69
=Agrotis annexa Treitschke, 1825. Walker, 1856: 328
=Agrotis decernens Walker, 1856: 333. Poole, 1989: 47
Agrotis ipsilon (Hufnagel, 1766). Schmutterer, 1990: 454
Anicla Grote, 1874
Anicla infecta (Ochsenheimer, 1816). Domínguez, 1997: 69
=Anicla incivis (Guenée, 1852). Walker, 1856: 330; Druce, 1891: 284
Feltia Walker, 1856
Feltia jaculifera (Guenée, 1852) [By synonymy of species below]
=Feltia ducens Walker, 1856. Domínguez, 1997: 70
Subfamily NOLINAE
Nola Leach, 1815
Nola cereella (Bosc, 1800) [By synonymy of species below]
=Nola sorghiella Riley, 1882. SEA, 1999: 99
Subfamily ONCOCNEMIDINAE
Neogalea Hampson, 1906
Neogalea sunia (Guenée, 1852). Valdéz et al., 2016: 66
Subfamily PANTHEINAE
Panthea Hubner, 1820
Panthea reducta Anweiler, 2009: 104 *DR > TL: Pedernales, P. N. Sierra de Bahoruco, caseta 2, ca. 1790 m
Subfamily PLUSIINAE
Argyrogramma Hübner, 1823
Argyrogramma verruca (Fabricius, 1794). Domínguez, 1997: 70
Autoplusia McDunnough, 1944
Autoplusia egena (Guenée, 1852). Walker, 1857b: 914; Druce, 1891: 332
Mouralia Walker, 1859
Mouralia tinctoides (Guenée, 1852). Poole, 1989: 672
=Mouralia annulifera Walker, 1858b: 1803. Poole, 1989: 672
Plusia Ochsenheimer, 1816
Plusia calceolaris Walker, 1857b: 915. Schaus, 1940: 232; Poole, 1989: 814
Pseudoplusia McDunnough, 1944
Pseudoplusia includens (Walker, 1857b: 914). Poole, 1989: 852; Domínguez, 1997: 70
=Pseudoplusia oo Stoll in Cramer, 1872. SEA, 1999: 121
Trichoplusia McDunnough, 1944
Trichoplusia ni (Hübner, 1803). Schaus, 1940: 232; Poole, 1989: 971; Schmutterer, 1990: 460; Domínguez, 1997: 69
=Plusia humilis Walker, 1857b: 915. Poole, 1989: 971
Subfamily SARROTHRIPINAE
Characoma Walker, 1863
Characoma sp.—Domínguez (1997: 69)
Iscadia Walker, 1857
Iscadia aperta Walker, 1857a: 725. Druce, 1891: 474; Poole, 1989: 554
Subfamily UNKNOWN

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 357
Homoptera Boisduval, 1852
Homoptera terminalis Walker, 1857c: 1061. Poole, 1989: 511
Magusa Walker, 1857
Magusa orbifera (Walker, 1857a: 761). Poole, 1989: 616; Domínguez, 1997: 69
=Magusa strigifera Walker, 1857a: 763. Druce, 1891: 297; Poole, 1989: 616
Notioplusia Lafontaine & Poole, 1991
Notioplusia illustrata (Guenée, 1852). Walker, 1857b: 913; Schaus, 1940: 231
Poaphila Guenée, 1852
Poaphila scita Walker, 1865. Poole, 1989: 818
Phurys Guenée, 1852
Phurys contenta Walker, 1865c: 995. Poole, 1989: 801
Phurys pura Walker, 1865c: 995. Poole, 1989: 801
Selenisa Hayward, 1967
Selenisa humeralis (Walker, 1857c: 1062). Poole, 1989: 904
Selenisa projiciens (Hampson, 1826). Poole, 1989: 904

Superfamily LASIOCAMPOIDEA
Family LASIOCAMPIDAE
Subfamily MACROMPHALINAE
Nesara Walker, 1855
Nesara albida (Walker, 1855: 1493). Becker & Heppner in Heppner, 1996: 23

Order MEGALOPTERA

Family SIALIDAE
Sialis Latreille, 1802
Sialis (Protosialis) casca† Engel & Grimaldi, 2007: 7 [Fossil in Dominican amber]

Order NEUROPTERA
Key reference: Perez-Gelabert & Flint (2001); Engel et al. (2017).

Family ASCALAPHIDAE
Subfamily ASCALAPHINAE
Amoea Lefebvre, 1842
Amoea electrodominicana† Engel & Grimaldi, 2007: 41 [Fossil in Dominican amber]
Ululodes Smith, 1900
Ululodes banksi van der Weele, 1909. Smith, 1931: 819; Perez-Gelabert & Flint, 2001: 10
Ululodes macleayanus (Guilding, 1823). By synonymy of species below
=Ululodes macleayanus venezolensis van der Weele, 1909. Smith, 1931: 818
Ululodes paleonesia† Engel & Grimaldi, 2007: 41 [Fossil in Dominican amber]
Ululodes sanctidomingi van der Weele, 1909: 115. Smith, 1931: 819; Perez-Gelabert & Flint, 2001: 11 *H, *DR > TL: Haiti or
San-Domingo
Ululodes smithi Banks, 1938: 179. Perez-Gelabert & Flint, 2001: 11 *H > TL: Port-au-Prince
Ululodes sp. 1†—Grimaldi & Engel (2007: 44) [Fossil in Dominican amber]
Ululodes sp. 2†—Grimaldi & Engel (2007: 48) [Fossil in Dominican amber]
Ululodes subvertens (Walker, 1853). Smith, 1931: 818
Ululodes villosus (Palisot de Beauvois, 1805: 86). Perez-Gelabert & Flint, 2001: 11 *DR, *H > TL: Saint-Domingue

Family CHRYSOPIDAE
Subfamily APOCHRYSINAE
Joguina Navás, 1912
Joguina constellata (Navás, 1913). Perez-Gelabert & Flint, 2001: 11 (as Lainius). Winterton & Brooks, 2002: 24
Subfamily CHRYSOPINAE
Ceraeochrysa Adams, 1982

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Ceraeochrysa cincta (Schneider, 1851). Perez-Gelabert & Flint, 2001: 12
=Ceraeochrysa rochina (Navás, 1915). Tauber & Flint, 2010: 65
Ceraeochrysa claveri (Navás, 1911). Perez-Gelabert & Flint, 2001: 11; Freitas et al., 2009: 532
=Chrysopa haitiensis Smith, 1931: 806
Ceraeochrysa cubana (Hagen, 1861). Perez-Gelabert & Flint, 2001: 12; Freitas et al., 2009: 536
=Chrysopa freemani Smith, 1931: 811
Ceraeochrysa valida (Banks, 1895). Perez-Gelabert & Flint, 2001: 12; Freitas et al., 2009: 587
=Chrysopa lioni Navás, 1927: 54. Smith, 1931: 812
=Chrysopa damiensis Smith, 1931: 803
=Chrysopa wolcotti Smith, 1931: 810
Chrysopa Leach, 1815
“Chrysopa” domingensis Banks, 1941a: 394. Perez-Gelabert & Flint, 2001: 12 *DR > TL: Constanza
“Chrysopa” sp.†—Spahr (1992: 79) [Fossil in Dominican amber]
Chrysopa? glaesaria† Engel & Grimaldi, 2007: 25 [Fossil in Dominican amber]
Chrysopa? vetula† Engel & Grimaldi, 2007: 27 [Fossil in Dominican amber]
Chrysoperla Steinmann, 1964
Chrysoperla dozieri (Smith, 1931: 808) (as Chrysopa). Perez-Gelabert & Flint, 2001: 12 *H > TL: Damien
Chrysoperla exotera (Navás, 1913). Perez-Gelabert & Flint, 2001: 13
=Chrysopa exterior Navás, 1922. Smith, 1931: 804
=Chrysopa estradai Navás, 1924. Smith, 1931: 809
Chrysoperla externa (Hagen, 1861). Perez-Gelabert & Flint, 2001: 13
Chrysoperla harrisii (Fitch, 1855). Perez-Gelabert & Flint, 2001: 13
Chrysoperla rufilabris (Burmeister, 1839). Perez-Gelabert & Flint, 2001: 13
Chrysopodes Navás, 1913
Chrysopodes (Neosuarius) collaris (Schneider, 1851). Perez-Gelabert & Flint, 2001: 13
=Chrysopa thoracica Walker, 1853: 243. Smith, 1931: 807
Eremochrysa Banks, 1903
Eremochrysa sp.—Perez-Gelabert & Flint (2001: 14)
Leucochrysa MacLachlan, 1868
Leucochrysa (Leucochrysa) christophei Banks, 1938: 292. Perez-Gelabert & Flint, 2001: 14 *H > TL: Desbarriere, Mt. La
Hotte, 4000 ft.
Leucochrysa floridana Banks, 1897. (By synonymy of species below)
=Leucochrysa (Leucochrysa) haitiensis Smith, 1931: 812. Perez-Gelabert & Flint, 2001: 14
Leucochrysa (Leucochrysa) insularis (Walker, 1853). Perez-Gelabert & Flint, 2001: 14
Leucochrysa (Nodita) ocampina Banks, 1941a: 395. Perez-Gelabert & Flint, 2001: 14 *DR > TL: Mt. Diego de Ocampo, 4,000
ft.
Leucochrysa (Nodita) prisca† Engel & Grimaldi, 2007: 27 [Fossil in Dominican amber]
Leucochrysa sp.†—Poinar & Poinar (1999: 203). Perez-Gelabert & Flint (2001: 14) [Fossil in Dominican amber]
Meleoma Fitch, 1855
Meleoma sp.—Perez-Gelabert & Flint (2001: 15)
Nacarina Navás, 1916
Nacarina sp.—Perez-Gelabert (2008: 283)
Plesiochrysa Adams, 1982
Plesiochrysa brasiliensis (Schneider, 1851). Perez-Gelabert & Flint, 2001: 15
=Chrysopa antillana Navás, 1922. Smith, 1931: 810
Plesiochrysa elongata (Navás, 1913). Perez-Gelabert & Flint, 2001: 15

Family CONIOPTERYGIDAE
Subfamily ALEUROPTERYGINAE
Neoconis Enderlein, 1929
Neoconis cubana (Banks, 1938). Perez-Gelabert & Flint, 2001: 15
Neoconis paleocaribis† Grimaldi & Engel in Grimaldi et al., 2013: 12 [Fossil in Dominican amber]
Spiloconis Enderlein, 1907
Spiloconis glaesaria† Meinander, 1998: 33. Perez-Gelabert & Flint, 2001: 15; Grimaldi et al., 2013: 4 [Fossil in Dominican

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 359
amber]
Spiloconis oediloma† Engel & Grimaldi, 2007: 19; Grimaldi et al., 2013: 4 [Fossil in Dominican amber]
Subfamily CONIOPTERYGINAE
Coniopteryx Curtis, 1834
Coniopteryx antiquua† Engel & Grimaldi, 2007: 17 [Fossil in Dominican amber]
Coniopteryx haitiensis Smith, 1931: 819
Coniopteryx (Scotoconiopteryx) fumata Enderlein, 1907. Perez-Gelabert & Flint, 2001: 14
Coniopteryx sp.†—Meinander (1998: 35). Perez-Gelabert & Flint, 2001: 16 [Fossil in Dominican amber]
Semidalis Enderlein, 1905
Semidalis marginalis (Banks, 1930). Perez-Gelabert & Flint, 2001: 16

Family DILARIDAE
Subfamily NALLACHINAE
Nallachius Navás, 1909
Nallachius pulchellus (Banks, 1938). Perez-Gelabert & Flint, 2001: 16

Family HEMEROBIIDAE
Subfamily HEMEROBIINAE
Hemerobius Linnaeus, 1758
Hemerobius antigonus Banks, 1941a: 396. Perez-Gelabert & Flint, 2001: 17 *DR > TL: Valle Nuevo to Constanza, 7,000 ft.
Hemerobius darlingtoni Banks, 1938. Perez-Gelabert & Flint, 2001: 17
Hemerobius domingensis Banks, 1941a: 395. Perez-Gelabert & Flint, 2001: 17
Hemerobius gaitoi Monserrat, 1996. Perez-Gelabert & Flint, 2001: 17
Hemerobius sp.†—Poinar & Poinar (1999: 203). Perez-Gelabert & Flint, 2001: 17 [Fossil in Dominican amber]
Subfamily MEGALOMINAE
Megalomus Rambur, 1842
Megalomus minor Banks, 1905. Perez-Gelabert & Flint, 2001: 17
=Megalomus lioni Navás, 1927. Smith, 1931: 800
Subfamily MICROMINAE
Micromus Rambur, 1842
Micromus posticus (Walker, 1853). Perez-Gelabert & Flint, 2001: 18
Micromus subanticus (Walker, 1853). Perez-Gelabert & Flint, 2001: 18
=Micromus haitiensis Smith, 1931: 800
Micromus sp.—Perez-Gelabert & Flint (2001: 18).
Nusalala Navás, 1913
Nusalala cubana (Hagen, 1886). Monserrat, 2000: 112; Perez-Gelabert & Flint, 2001: 18
=Nusalala damiensis Smith, 1931: 800
Nusalala payasi Monserrat, 2000: 128 *DR > TL: Jarabacoa, 530 m
Subfamily NOTOBIELLINAE
Notiobiella Banks, 1909
Notiobiella israeli (Alayo, 1968). Perez-Gelabert & Flint, 2001: 19
Notiobiella thaumasta† Oswald, 2000: 298. Perez-Gelabert & Flint, 2001: 19 [Fossil in Dominican amber]
Subfamily SYMPHEROBINAE
Sympherobius Banks, 1904
Sympherobius amiculus (Fitch, 1856). Perez-Gelabert & Flint, 2001: 19
Sympherobius sp.†—Engel & Grimaldi (2007: 22) [Fossil in Dominican amber]

Family MANTISPIDAE
Subfamily MANTISPINAE
Climaciella Enderlein, 1910
Climaciella cubana Enderlein, 1910. Perez-Gelabert, 2008: 284; Hoffman et al., 2017: 3
Dicromantispa Hoffman in Penny, 2002
Dicromantispa moronei† Engel & Grimaldi, 2007: 14 [Fossil in Dominican amber]
Dicromantispa sayi (Banks, 1897). Perez-Gelabert & Flint, 2001: 19; Ohl, 2004: 169; Hoffman et al., 2017: 4

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Feroseta† Poinar, 2006
Feroseta priscus† Poinar, 2006: 412 [Fossil in Dominican amber]
Leptomantispa Hoffman in Penny, 2002
Leptomantispa hispaniolaensis Hoffman in Hoffman et al., 2017: 8 *DR > TL: 14 km W Puerto Plata
=Species of Mantispa pulchella group—Hoffman (1992: 186); Perez-Gelabert & Flint (2001: 20)
Zeugomantispa Hoffman, 2002
Zeugomantispa minuta (Fabricius, 1775). Ohl, 2004: 206; Hoffman et al., 2017: 10

Family MYRMELEONTIDAE
Tribe ACANTHACLISINI
Vella Navás, 1913
Vella eggerti Esben-Petersen, 1928. Miller & Stange, 2011: 5
=Vella haitiensis Smith, 1931: 815. Perez-Gelabert & Flint, 2001: 20; Stange, 2004: 362
Tribe BRACHYNEMURINI
Peruveleon Miller & Stange, 2011
Peruveleon indiges (Walker, 1860: 189). Smith, 1931: 816; Perez-Gelabert & Flint, 2001: 20; Stange, 2004: 226; Miller &
Stange, 2011: 7
Tribe MYRMELEONTINI
Myrmeleon Linnaeus, 1767
Myrmeleon insertus Hagen, 1861. Smith, 1931: 817; Perez-Gelabert & Flint, 2001: 21; Stange, 2004: 311; Miller & Stange,
2011: 9
Porrerus Navás, 1913
Porrerus dominicanus† Poinar & Stange, 1996: 384; Perez-Gelabert & Flint, 2001: 21; Stange, 2004: 339; Miller & Stange,
2011: 10 [Fossil in Dominican amber]
Tribe NEMOLEONTINI
Eremoleon Banks, 1901
Eremoleon cerverai (Navás, 1921). Smith, 1931: 814; Stange, 2004: 169; Perez-Gelabert & Flint, 2001: 20; Stange, 2004: 169;
Miller & Stange, 2011: 12
Eremoleon cerverinus (Navás, 1921). Stange, 2004: 169; Perez-Gelabert & Flint, 2001: 20; Stange, 2004: 169; Miller &
Stange, 2011: 13
Eremoleon petrophila Miller & Stange, 2011: 14 *DR > TL: San Juan Prov., El Capá, 17 km NE Vallejuelo
Eremoleon phasma Miller & Stange, 2011: 15 *DR > TL: Independencia Prov., Los Ríos, Lago Enriquillo
Purenleon Stange, 2002
Purenleon bistictus (Hagen, 1861). Smith, 1931: 815; Perez-Gelabert & Flint, 2001: 21; Stange, 2004: 215; Miller & Stange,
2011: 17
Purenleon imbellis (Banks, 1941b: 102). Perez-Gelabert & Flint, 2001: 21; Stange, 2004: 216; Miller & Stange, 2011: 19 *H >
TL: Port-au-Prince
Purenleon minor (Banks, 1927). Perez-Gelabert & Flint, 2001: 21; Stange, 2004: 216; Miller & Stange, 2011: 119
Purenleon nunezi Miller & Stange, 2011: 20 *DR > TL: San Juan Prov., El Capá, 17 km NE Vallejuelo
Purenleon woodruffi Miller & Stange, 2011: 25 *DR > TL: Montecristi Prov., 9 km north of Villa Elisa

Order ODONATA
Key references: Daigle (1996), Flint et al. (2006).

Taxonomic summary:
Suborder Zygoptera
Family Coenagrionidae
Family Lestidae
Family Megapodagrionidae
Family Protoneuridae
Family Synlestidae
Suborder Anisoptera
Family Aeshnidae
Family Gomphidae

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 361
 Family Libellulidae

Suborder ZYGOPTERA
Family COENAGRIONIDAE
Enallagma Charpentier, 1840
Enallagma civile (Hagen, 1861). Flint et al., 2006: 69
Enallagma coecum (Hagen, 1861). Flint et al., 2006: 69
Diceratobasis Kennedy, 1920
Diceratobasis melanogaster Garrison, 1986: 62. Flint et al., 2006: 69 *DR > TL: La Vega Prov., roadside cuts along Constanza
Rd., 15 km E of Autopista Duarte, elev. 1000 m
Diceratobasis worki† Poinar, 1996: 382 [Fossil in Dominican amber]
Ischnura Charpentier, 1840
Ischnura capreolus (Hagen, 1861). Flint et al., 2006: 70
Ischnura hastata (Say, 1839). Flint et al., 2006: 70
Ischnura ramburii (Selys, 1842). Flint et al., 2006: 70
Ischnura velteni† Bechly, 2000: 2 [Fossil in Dominican amber]
Leptobasis Hagen, 1868
Leptobasis vacillans Hagen, 1877. Flint et al., 2006: 71
Nehalennia Selys, 1850
Nehalennia minuta (Selys, 1857). Flint et al., 2006: 71
Neoerythromma Kennedy, 1920
Neoerythromma cultellatum (Hagen, 1876). Flint et al., 2006: 71
Telebasis Selys, 1865
Telebasis dominicana (Selys, 1857). Flint et al., 2006: 71
Telebasis vulnerata (Hagen, 1861). Flint et al., 2006: 71

Family LESTIDAE
Lestes Leach, 1815
Lestes forficula Rambur, 1842. Flint et al., 2006: 72
Lestes scalaris Gundlach, 1888. Flint et al., 2006: 72
Lestes spumarius Hagen, 1862. Flint et al., 2006: 72

Family MEGAPODAGRIONIDAE
Hypolestes Hagen, 1868
Hypolestes hatuey Torres-Cambas in Torres-Cambas et al., 2015: 218 *DR > TL: Hatillo, Arroyo Bermejo
=Hypolestes trinitatis (Gundlach, 1888). Flint et al., 2006: 72 [Misidentification]
=Ortholestes abbotti Calvert, 1893: 382

Family PROTONEURIDAE
Protoneura Selys, 1857
Protoneura dunklei Daigle, 1990: 81. Flint et al., 2006: 73; von Ellenrieder & Garrison, 2017: 25 *DR > TL: La Vega Prov.,
Arroyo Ana Marie, 10.9 km SW of Autopista Duarte on road to Jarabacoa
Protoneura sanguinipes Westfall, 1987: 94. Flint et al., 2006: 73; von Ellenrieder & Garrison, 2017: 35 *DR > TL: Arroyo
Bermejo, 4 km NNE of Hatillo and Autopista Duarte, Distrito Nacional
Protoneura viridis Westfall, 1964. Flint et al., 2006: 73; von Ellenrieder & Garrison, 2017: 41

Family SYNLESTIDAE
Phylolestes Christiansen, 1948
Phylolestes ethelae Christiansen, 1948: 258. Westfall, 1976: 67; Flint et al., 2006: 73 *H, *DR > TL: La Visite, La Selle Mts.,
4,000 ft.

Suborder ANISOPTERA
Family AESHNIDAE
Anax Leach, 1815

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Anax amazili (Burmeister, 1839). Flint et al., 2006: 73
Anax concolor Brauer, 1865. Flint et al., 2006: 73
Anax junius (Drury, 1861). Flint et al., 2006: 73
Coryphaeschna Williamson, 1903
Coryphaeschna adnexa (Hagen, 1861). Flint et al., 2006: 74
Coryphaeschna viriditas Calvert, 1952. Flint et al., 2006: 74
Gynacantha Rambur, 1842
Gynacantha nervosa Rambur, 1842. Flint et al., 2006: 74
Rhionaeshna Förster, 1909
Rhionaeshna psilus (Calvert, 1947). Flint et al., 2006: 74
Triacanthagyna Selys, 1883
Triacanthagyna septima (Selys, 1857). Fernández-Martínez, 2004: 27; Flint et al., 2006: 74
Triacanthagyna trifida Rambur, 1842. Flint et al., 2006: 74; Marquez-Rodríguez, 2019: 359

Family GOMPHIDAE
Aphylla Selys, 1854
Aphylla caraiba (Selys, 1854). Flint et al., 2006: 74
Progomphus Selys, 1854
Progomphus integer Hagen, 1878. Flint et al., 2006: 75 [Record needs verification]
Progomphus serenus Hagen, 1878. Flint et al., 2006: 75
Progomphus tennesseni Daigle, 1996: 367. Flint et al., 2006: 75 *DR > TL: La Vega Prov., Arroyo Guasara, just below Salto
Guasara
Progomphus zephyrus Needham, 1941. Daigle, 1993c: 198; Flint et al., 2006: 75 *DR > TL: Distrito Nacional, Arroyo
Bermejo, 4 km NNE of Hatillo

Family LIBELLULIDAE
Brachymesia Kirby, 1889
Brachymesia furcata (Hagen, 1861). Flint et al., 2006: 75
Brachymesia herbida Gundlach, 1888. Flint et al., 2006: 75
Cannaphila Kirby, 1889
Cannaphila insularis Kirby, 1889. Flint et al., 2006: 75
Dythemis Hagen, 1861
Dythemis rufinervis (Burmeister, 1839). Flint et al., 2006: 75
Erythemis Hagen, 1861
Erythemis attala (Selys, 1857). Flint et al., 2006: 76 [Record needs verification]
Erythemis plebeja (Burmeister, 1839). Flint et al., 2006: 76
Erythemis simplicicollis (Say, 1839). Flint et al., 2006: 76
Erythemis vesciculosa (Fabricius, 1775). Flint et al., 2006: 76
Erythrodiplax Brauer, 1868
Erythrodiplax berenice (Drury, 1770). Flint et al., 2006: 76
Erythrodiplax fervida (Erichson, 1848). Flint et al., 2006: 76
Erythrodiplax justiniana (Selys, 1857). Flint et al., 2006: 77
Erythrodiplax umbrata (Linnaeus, 1758). Flint et al., 2006: 77
Idiataphe Cowley, 1934
Idiataphe cubensis (Scudder, 1866). Flint et al., 2006: 77
Macrodiplax Brauer, 1868
Macrodiplax balteata (Hagen, 1861). Flint et al., 2006: 77
Macrothemis Hagen, 1868
Macrothemis celeno (Selys, 1857). Flint et al., 2006: 78
Miathyria Kirby, 1889
Miathyria marcella (Selys, 1857). Flint et al., 2006: 78
Miathyria simplex (Rambur, 1842). Flint et al., 2006: 78
Micrathyria Kirby, 1889
Micrathyria aequalis (Hagen, 1861). Flint et al., 2006: 78

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Micrathyria didyma (Selys, 1857). Flint et al., 2006: 78
Micrathyria dissocians Calvert, 1906. Flint et al., 2006: 78
Micrathyria hagenii Kirby, 1890. Flint et al., 2006: 78
Orthemis Hagen, 1861
Orthemis species ferruginea group. Flint et al., 2006: 79
Pantala Hagen, 1861
Pantala flavescens Fabricius, 1798. Flint et al., 2006: 79
Pantala hymenaea (Say, 1839). Flint et al., 2006: 79
Perithemis Hagen, 1861
Perithemis domitia (Drury, 1773). Flint et al., 2006: 79
Scapanea Kirby, 1889
Scapanea frontalis (Burmeister, 1839). Flint et al., 2006: 80
Sympetrum Newman, 1833
Sympetrum gilvum (Hagen, 1861). Flint et al., 2006: 80
Tauriphila Kirby, 1889
Tauriphila australis (Hagen, 1867). Flint et al., 2006: 80
Tholymis Hagen, 1867
Tholymis citrina Hagen, 1867. Flint et al., 2006: 80
Tramea Hagen, 1861
Tramea abdominalis (Rambur, 1842). Flint et al., 2006: 81
Tramea binotata (Rambur, 1842). Flint et al., 2006: 81
Tramea calverti Muttkowski, 1910. Flint et al., 2006: 81
Tramea insularis Hagen, 1861. Flint et al., 2006: 81
Tramea onusta Hagen, 1861. Flint et al., 2006: 81

Order ORTHOPTERA
Key reference: Perez-Gelabert (2001).

Taxonomic summary:
Suborder Caelifera
Family Ripipterygidae
Family Tridactylidae
Family Tetrigidae
Family Episactidae
Family Pyrgomorphidae
Family Acrididae
Family Romaleidae
Suborder Ensifera
Family Tettigoniidae
Family Gryllidae
Family Phalangopsidae
Family Trigonidiidae
Family Gryllotalpidae
Family Mogoplistidae
Family Myrmecophilidae
Family Anostostomatidae
Family Gryllacrididae

Suborder CAELIFERA
Family RIPIPTERYGIDAE
Mirhipipteryx Günther, 1969
Mirhipipteryx antillarum† Heads, 2010a: 208 [Fossil in Dominican amber]

Family TRIDACTYLIDAE

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Subfamily TRIDACTYLINAE
Archaeoellipes† Heads, 2010
Archaeoellipes engeli† Heads, 2010a: 207. Gorochov, 2010: 669 [Fossil in Dominican amber]
Ellipes Scudder, 1902
Ellipes dominicana† Poinar, 2018: 1 [Fossil in Dominican amber]
Ellipes minuta histrionica (Saussure, 1896). Wolcott, 1927: 152; Perez-Gelabert, 2001: 64

Family TETRIGIDAE
Subfamily CLADONOTINAE
Tribe MUCROTETTIGINI
Antillotettix Perez-Gelabert, 2004
Antillotettix electrum† Heads, 2009: 70 [Fossil in Dominican amber]
Antillotettix nanus Perez-Gelabert, 2004: 111 *DR > TL: Pedernales Prov., 30 km N Cabo Rojo, 1070 m
Baeotettix† Heads, 2009
Baeotettix lottiae† Heads, 2009: 72 [Fossil in Dominican amber]
Bahorucotettix Perez-Gelabert et al., 1998
Bahorucotettix larimar Perez-Gelabert et al., 1998: 196. Perez-Gelabert, 2001: 65 *DR > TL: Barahona Prov., Larimar mine,
near Filipinas, Eastern Sierra de Bahoruco
Electrotettix† Heads & Thomas in Heads et al., 2014
Electrotettix attenboroughi† Heads & Thomas in Heads et al., 2014: 90 [Fossil in Dominican amber]
Haitianotettix Perez-Gelabert et al., 1998
Haitianotettix tuberculatus Perez-Gelabert et al., 1998: 198. Perez-Gelabert, 2001: 65 *H > TL: Pedernales Prov., 5 km NE of
Los Arroyos, western Sierra de Bahoruco, 1,680 m
=Haitianotettix monstruosus Perez-Gelabert et al., 1998: 199. Perez-Gelabert, 2001: 65; Perez-Gelabert, 2009: 900
Hottettix Perez-Gelabert et al., 1998
Hottettix haitianus Perez-Gelabert et al., 1998: 195. Perez-Gelabert, 2001: 65 *H > TL: Dept. du Sud, northeastern foothills of
Massif de la Hotte, 2–4,000 ft.
Mucrotettix Perez-Gelabert et al., 1998
Mucrotettix gibbosus Perez-Gelabert et al., 1998: 192. Perez-Gelabert, 2001: 65 *DR > TL: Santiago Prov., Mte. Diego de
Ocampo, 1,000
Mucrotettix spinifer Perez-Gelabert et al., 1998: 193. Perez-Gelabert, 2001: 65 *DR > TL: La Vega Prov., Los Tablones, Parque
Nacional A. Bermúdez, 1,245 m
Sierratettix Perez-Gelabert et al., 1998
Sierratettix carinatus Perez-Gelabert et al., 1998: 194. Perez-Gelabert, 2001: 65 *DR > TL: Barahona Prov., Sierra de
Bahoruco, on way to Cortico, 1,350 m
Tiburonotus Perez-Gelabert et al., 1998
Tiburonotus peninsularis Perez-Gelabert et al., 1998: 197. Perez-Gelabert, 2001: 65 *H > TL: Northeastern foothills of La
Hotte, Dept. du Sud, 3–4,000 ft.
Truncotettix Perez-Gelabert et al., 1998
Truncotettix fronterizus Perez-Gelabert et al., 1998: 191. Perez-Gelabert, 2001: 65 *DR, *H > TL: Independencia Prov., Los
Pinos del Edén, 530 m
Truncotettix interruptus Perez-Gelabert et al., 1998: 190. Perez-Gelabert, 2001: 65 *DR > TL: Monseñor Nouel Prov., Arroyo
Toro Arriba, near Bonao, 250 m
Subfamily TETRIGINAE
Micronotus Hancock, 1902
Micronotus quadriundulatus (Redtenbacher, 1892). Hancock, 1900: 25; Perez-Gelabert, 2000: 17; Perez-Gelabert, 2001: 65
Paratettix Bolívar, 1887
Paratettix freygessneri Bolívar, 1878. Perez-Gelabert, 2000: 16; Perez-Gelabert, 2001: 65
=Merotettix pristinus Morse, 1899: 199

Family EPISACTIDAE
Subfamily ESPAGNOLINAE
Antillacris J.A.G. Rehn & J.W. Rehn, 1939
Antillacris eumenes Perez-Gelabert, Hierro, Dominici & Otte, 1997: 143. Perez-Gelabert, 2001: 65 *DR > TL: Elías Piña Prov.,

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Intramontane valley on Loma de Las Tayotas, Loma Nalga de Maco Massif, 1,200 m
Antillacris explicatrix J.A.G. Rehn & J.W. Rehn, 1939: 202. Perez-Gelabert, 2001: 65 *DR > TL: Loma Rucilla, elevation
5000–8000 ft.
Antillacris inflaticercus Perez-Gelabert, Hierro, Dominici & Otte, 1997: 143. Perez-Gelabert, 2001: 65 *DR > TL:
Independencia Prov., Sierra de Neiba, just South of crest, 5 m NNW of Angel Feliz, 1,780 m
Espagnola J.A.G. Rehn & J.W. Rehn, 1939
Espagnola darlingtoni J.A.G. Rehn & J.W. Rehn, 1939: 192. Perez-Gelabert, 2001: 65 *DR > TL: Jarabacoa, 1500–4000 ft.
Espagnoleta Perez-Gelabert, 2000
=Espagnolina Perez-Gelabert, Hierro & Otte, 1997
Espagnoleta microptera (Perez-Gelabert, Hierro & Otte, 1997: 154). Perez-Gelabert, 2001: 66 *DR > TL: Peravia Prov., about
3 km from town of Los Anones, S of San José de Ocoa
Espagnolopsis Perez-Gelabert, Hierro, Dominici & Otte, 1997
Espagnolopsis breviptera Perez-Gelabert, Hierro & Otte, 1997: 159. Perez-Gelabert, 2001: 66 *DR > TL: La Vega Prov.,
Parque Nacional A. Bermúdez, from 1 km after La Ciénaga de Manabao to Los Tablones, 1,250 m
Espagnolopsis exaltata Perez-Gelabert & Rowell, 2006: 242 *DR > TL: Ocoa Prov., Hilltop on way to Palos Grandes, NE San
José de Ocoa, 1437 m
Espagnolopsis ornatipennis Perez-Gelabert, Hierro, & Otte, 1997: 157. Perez-Gelabert, 2001: 66 *DR > TL: San Cristóbal
Prov., 2 km after La Colonia
Neibamastax Rowell & Perez-Gelabert, 2006
Neibamastax divergentis (Perez-Gelabert, Hierro, Dominici & Otte, 1997: 142). Perez-Gelabert, 2001: 66; Rowell & Perez-
Gelabert, 2006: 224 *DR > TL: Independencia Prov., Los Pinos del Edén, 520 m
Paleomastacris† Perez-Gelabert, Hierro, Dominici & Otte, 1997
Paleomastacris ambarinus† Perez-Gelabert, Hierro, Dominici & Otte, 1997: 150. Perez-Gelabert, 2001: 66 [Fossil in
Dominican amber]
Tainacris Perez-Gelabert, Hierro, Dominici & Otte, 1997
Tainacris nitaina Perez-Gelabert, Hierro, Dominici & Otte, 1997: 140. Perez-Gelabert, 2001: 66 *DR > TL: Azua Prov., Las
Yayitas, 225 m
Tainacris quisqueiana Perez-Gelabert, Hierro, Dominici & Otte, 1997: 142. Perez-Gelabert, 2001: 66 *DR > TL: Peravia Prov.,
km 2 road Los Martínez—Los Ranchos, near Ocoa

Family PYRGOMORPHIDAE
Jaragua Perez-Gelabert, Dominici & Hierro, 1995
Jaragua oviedensis Perez-Gelabert, Dominici & Hierro, 1995: 34. Perez-Gelabert, 2000: 17; Perez-Gelabert, 2001: 66 *DR >
TL: Pedernales Prov., 10 km SW of Oviedo, sea level
Jaragua serranus Perez-Gelabert, Dominici & Hierro, 1995: 35. Perez-Gelabert, 2001: 66 *DR > TL: 1 km S of Los Pinos del
Edén, on La Descubierta - Los Pinos road, ~520 m

Family ACRIDIDAE
Subfamily COPIOCERINAE
Dellia Stål, 1878
Dellia bayahibe Perez-Gelabert, 2003: 32 *DR > TL: La Altagracia Prov., 2 km N Bayahibe
Dellia ciceroana Perez-Gelabert & Otte, 2012: 9 *DR > TL: Independencia Prov., ~2 km E Puerto Escondido, 450 m
Dellia dominicensis Perez-Gelabert, Dominici, Hierro & Otte, 1995: 158. Perez-Gelabert, 2001: 66 *DR > TL: Azua Prov., Las
Yayitas, 225 m
Dellia monticola Perez-Gelabert & Otte, 1999: 6. Perez-Gelabert, 2001: 66 *DR > TL: Prov. Pedernales, El Aceitillar, km 25
carr. de la Alcoa, 600 m
Dellia roseomaculata Perez-Gelabert & Otte, 1999: 5. Perez-Gelabert, 2001: 66 *DR > TL: Prov. Elías Piña, Loma Las
Tayotas, macizo de Loma Nalga de Maco, 760 m
Dellia viridissima Perez-Gelabert & Otte, 2012: 8 *DR > TL: Barahona Prov., forest near La Mina, Cortico, Monteada Nueva,
1,250 m
Subfamily CYRTACANTHACRIDINAE
Schistocerca Stål, 1873
Schistocerca pallens (Thunberg, 1870). Harvey, 1981: 70; Perez-Gelabert, 2000: 19; Perez-Gelabert, 2001: 66
Schistocerca quisqueya J.A.G. Rehn & Hebard, 1938: 221. Dirsh, 1974: 162; Perez-Gelabert, 2000: 19; Perez-Gelabert, 2001:

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66 *DR > TL: Moca, Prov. Espaillat
Schistocerca serialis serialis (Thunberg, 1815). Harvey, 1981: 67; Perez-Gelabert, 2001: 66
Subfamily GOMPHOCERINAE
Amblytropidia Stål, 1873
Amblytropidia hispaniolana Perez-Gelabert, Dominici, Hierro & Otte, 1995: 155. Perez-Gelabert, 2000: 17; Perez-Gelabert,
2001: 67 *DR > TL: Azua Prov., hills above Playa Monte Río, 15–20 m
Orphulella Giglio-Tos, 1894
Orphulella decisa (Walker, 1870: 757). J.A.G. Rehn & Hebard, 1938: 205; Otte, 1979: 66; Perez-Gelabert, 2000: 17; Perez-
Gelabert, 2001: 67 *H, *DR > TL: San Domingo
Orphulella nesicos Otte, 1979: 66. Perez-Gelabert, 2000: 17; Perez-Gelabert, 2001: 67 *H, *DR > TL: La Vega Prov., 5 km
east of Constanza, 500—1500 m
Orphulella punctata (De Geer, 1773). Perez-Gelabert, 2001: 67
Orphulella scudderi (Bolívar, 1888). Perez-Gelabert, 2001: 67
Rhammatocerus Saussure, 1861
Rhammatocerus cyanipes (Fabricius, 1775). Perez-Gelabert, 2001: 67
Subfamily LEPTYSMINAE
Stenacris F. Walker, 1870
Stenacris caribea (J.A.G. Rehn & Hebard, 1938: 217). Perez-Gelabert, 2000: 19; Perez-Gelabert, 2001: 67
Subfamily MELANOPLINAE
Duartettix Perez-Gelabert & Otte, 2000
Duartettix montanus Perez-Gelabert & Otte, 2000: 130. Perez-Gelabert, 2001: 67 *DR > TL: La Vega Prov., Valle Nuevo,
between La Pirámide and La Neverita, 2,500–2400 m
Melanoplus Stål, 1873
Melanoplus perezi Otte, 2013: 75 *DR > TL: La Vega Prov., between Bonao and Constanza, 500–1500 m
=Melanoplus sanguinipes (Fabricius, 1798). Perez-Gelabert, 2003: 543 [Misidentification]
Subfamily OEDIPODINAE
Sphingonotus Fieber, 1852
Sphingonotus haitensis haitensis (Saussure, 1861). Wolcott, 1927: 146; J.A.G. Rehn & Hebard, 1938: 211; Perez-Gelabert,
2001: 67
Subfamily OMMATOLAMPIDINAE
Acridurus Perez-Gelabert, Dominici, Hierro & Otte, 1995
Acridurus neibanus Perez-Gelabert, Dominici, Hierro & Otte, 1995: 165. Perez-Gelabert, 2001: 67 *DR > TL: Independencia
Prov., beginning of road from La Descubierta to Los Pinos del Edén
Acridurus robustus Perez-Gelabert, Dominici, Hierro & Otte, 1995: 163. Perez-Gelabert, 2000: 18; Perez-Gelabert, 2001: 67
*DR > TL: Pedernales Prov., rocky cliffs of Playa Inglesa, Parque Nacional Jaragua, S of southern end of Laguna de Oviedo
Acridurus yayitas Perez-Gelabert, Dominici, Hierro & Otte, 1995: 164. Perez-Gelabert, 2000: 18; Perez-Gelabert, 2001: 67
*DR > TL: Azua Prov., Las Yayitas, 225 m
Hispanacris Perez-Gelabert, Dominici, Hierro & Otte, 1995
Hispanacris oreades Perez-Gelabert, Dominici, Hierro & Otte, 1995: 165. Perez-Gelabert, 2000: 18; Perez-Gelabert, 2001: 67
*DR > TL: Pirámide 204, Sierra de Neiba
Hispanotettix Perez-Gelabert, Dominici, Hierro & Otte, 1995
Hispanotettix nitidus Perez-Gelabert, Dominici, Hierro & Otte, 1995: 161. Perez-Gelabert, 2000: 18; Perez-Gelabert, 2001: 68
*DR > TL: Azua Prov., Las Yayitas, 225 m
Tergoceracris Perez-Gelabert & Otte, 2003
Tergoceracris ebanoverde Perez-Gelabert & Otte, 2003: 5 *DR > TL: La Vega Prov., La Sal, Reserva Científica Ebano Verde
Tergoceracris ocampensis Perez-Gelabert & Otte, 2003: 5 *DR > TL: Santiago Prov., Monte Diego de Ocampo, summit, 1,300
m

Family ROMALEIDAE
Subfamily ROMALEINAE
Xyleus Gistel, 1848
Xyleus discoideus rosulentus (Stål, 1878). Perez-Gelabert, Bastardo & Hierro, 2010: 77 [Introduced]

Suborder ENSIFERA

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Family TETTIGONIIDAE
Subfamily CONOCEPHALINAE
Conocephalus Thunberg, 1815
Conocephalus (Conocephalus) cinereus Thunberg, 1815. Perez-Gelabert, 2001: 68
Conocephalus (Conocephalus) insularis (Morse, 1905). Perez-Gelabert, 2001: 68
Erechthis Bolívar, 1888
Erechthis gundlachi Bolívar, 1888. Perez-Gelabert, 2014: 38; Perez-Gelabert & Yong, 2017: 286
Erechthis ayiti De Luca & Morris, 2016: 53 *H, *DR > TL: Coffee Exp. Sta., Fond des Negres
Neoconocephalus Karny, 1907
Neoconocephalus affinis (Palisot de Beauvois, 1805: 219). Perez-Gelabert, 2001: 68
Neoconocephalus maxillosus (Fabricius, 1775). Perez-Gelabert, 2001: 68
Neoconocephalus occidentalis (Saussure, 1859: 208). Perez-Gelabert, 2001: 68 *H, *DR > TL: Haiti
Neoconocephalus pinicola T. Walker & Greenfield, 1983: 375. Perez-Gelabert, 2001: 68 *DR > TL: Km 25–33 on Alcoa road,
Baoruco Mts., 700-1250 m
Neoconocephalus triops (Linnaeus, 1758: 430). Wolcott, 1927: 148; Perez-Gelabert, 2001: 68
Pyrgocorypha Stål, 1873
Pyrgocorypha uncinata (Harris, 1841). Perez-Gelabert, 2014: 41
Unicorniella Yong, 2019
Unicorniella hatueyi Yong, 2019: 235
Subfamily MECONEMATINAE
Miophlugis† Gorochov, 2010
Miophlugis rostratus† Gorochov, 2010: 662 [Fossil in Dominican amber]
Phlugis Stål, 1860
Phlugis sp.—Perez-Gelabert (2001: 68)
Subfamily PHANEROPTERINAE
Anaulacomera Stål, 1873
Anaulacomera sp.—Perez-Gelabert (2001: 68)
Microcentrum Scudder, 1862
Microcentrum decoratum (F. Walker, 1869: 373). Perez-Gelabert, 2001: 68
Phoebolampta Brunner von Wattenwyl, 1878
Phoebolampta excellens (F. Walker, 1869: 371). Wolcott, 1927: 148; Perez-Gelabert, 2001: 68 *DR > TL: St. Domingo
Phoebolampta subaequale (F. Walker, 1869: 372). Perez-Gelabert, 2001: 69 *DR > TL: St. Domingo
Turpilia Stål, 1874
Turpilia appendiculata Brunner, 1878: 326. Wolcott, 1927: 148; Perez-Gelabert, 2001: 69 *H > TL: Port-au-Prince
Subfamily PSEUDOPHYLLINAE
Anacaona Yong, 2019
Anacaona bellatrix Yong, 2019: 247 *DR > TL: Barahona province: Cabral Municipality, km 7–9 of road from Cabral to Polo,
222 m
Caribophyllum Rehn, 1947
Caribophyllum necopinum Rehn, 1947: 4. Perez-Gelabert, 2001: 69 *H, *DR > TL: Pétionville
Polyancistrus Serville, 1831
Polyancistrus abbotti Rehn, 1936: 308. Perez-Gelabert, 2001: 69 *H > TL: Vicinity of La Visite, Massif de la Selle, 5000–7000
ft.
Polyancistrus atlas atlas Rehn, 1936: 286. Perez-Gelabert, 2001: 69 *H > TL: Coffee Experiment Station, Fond des Negres
Polyancistrus atlas pilanus Rehn, 1936: 291. Perez-Gelabert, 2001: 69 *H > TL: Furcy, west side of Massif de la Selle
Polyancistrus darlingtoni Rehn, 1936: 302. Perez-Gelabert, 2001: 69 *H > TL: Mount La Hotte, 5000—7800 ft.
Polyancistrus gerulus Rehn, 1936: 295. Perez-Gelabert, 2001: 69 *H > TL: Miragoâne
Polyancistrus loripes Rehn, 1936: 299. Perez-Gelabert, 2001: 69 *DR > TL: Constanza, La Vega Prov., 3600 ft.
Polyancistrus serrulatus oreiotes Rehn, 1936: 281. Perez-Gelabert, 2001: 69 *DR > TL: Manabao, 900 m
Polyancistrus serrulatus serrulatus (Palisot de Beauvois, 1805: 218). Perez-Gelabert, 2001: 69 *DR > TL: St. Domingue
Spelaeala Rehn, 1943
Spelaeala bondi Rehn, 1943: 5. Perez-Gelabert, 2001: 69 *DR > TL: Loma Rucilla and mountains to the north, elevation 5000–
8000 ft.
Spelaeala scobina Rehn, 1943: 11. Perez-Gelabert, 2001: 69 *DR > TL: Loma Rucilla and mountains to the north, elevation

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8000 ft.

Family GRYLLIDAE
Classification follows Orthoptera Species File Online (Cigliano et al., 2017).
Subfamily HAPITHINAE
Tribe HAPITHINI
Carylla Otte & Perez-Gelabert, 2009
Carylla cadabra Otte & Perez-Gelabert, 2009: 266 *DR > TL: Jaiquí Picado, Santiago Prov., 531 m
Carylla vivida Otte & Perez-Gelabert, 2009: 264 *DR > TL: Juan Dolio, San Pedro de Macorís Prov.
Knyella Otte & Perez-Gelabert, 2009
Knyella vivax Otte & Perez-Gelabert, 2009: 237 *DR > TL: Arroyazo, Reserva Científica Ebano Verde, 3500 ft.
Margarettia Otte & Perez-Gelabert, 2009
Margarettia epaxia Otte & Perez-Gelabert, 2009: 236 *DR > TL: ~100 m up from El Sitio del Agua, N Los Bolos, Sierra de
Neiba, 1520 m
Margarettia fotuto Otte & Perez-Gelabert, 2009: 236 *DR > TL: Los Tablones, Parque Nacional Armando Bermúdez
Margarettia guajey Otte & Perez-Gelabert, 2009: 235 *DR > TL: La Sal, Reserva Científica Ebano Verde, 1043 m
Margarettia jaiba Otte & Perez-Gelabert, 2009: 236 *DR > TL: La Compartición, Parque Nacional Armando Bermúdez
Margarettia naniki Otte & Perez-Gelabert, 2009: 236 *DR > TL: Down from Cuevita, Valle Nuevo, 2096 m
Margarettia yarari Otte & Perez-Gelabert, 2009: 236 *DR > TL: La Jarda (monte abajo), Padre Las Casas, Azua Prov.
Sabelo Otte & Perez-Gelabert, 2009
Sabelo tutrix Otte & Perez-Gelabert, 2009: 231 *DR > TL: 11.5 km S of Constanza
Sipho Otte & Perez-Gelabert, 2009
Sipho atabey Otte & Perez-Gelabert, 2009: 235 *DR > TL: 2 km on Road to Los Martínez, Ocoa Prov., 510 m
Walkerana Otte & Perez-Gelabert, 2009 [Considered a Nomen nudum by Dubois, 2017: 77]
=Walkerella Otte & Perez-Gelabert, 2009: 257 [Preoccupied name]
Walkerana clandestina (Otte & Perez-Gelabert, 2009: 257). Otte, 2009: 488 *DR > TL: Hato Mayor, Parque Los Haitises, 3 km
N of Cueva de Arena
Walkerana enstates (Otte & Perez-Gelabert, 2009: 257). Otte, 2009: 488 *DR > TL: Down from Monteada Nueva, Barahona
Prov., 970 m
Walkerana encollos (Otte & Perez-Gelabert, 2009: 258). Otte, 2009: 488 *DR > TL: Arroyazo, Reserva Científica Ebano
Verde, 3500 ft.
Walkerana mira (Otte & Perez-Gelabert, 2009: 257). Otte, 2009: 488 *DR > TL: Río Limpio, Elías Piña Prov., 781 m
Walkerana ochleros (Otte & Perez-Gelabert, 2009: 258). Otte, 2009: 489 *DR > TL: La Sal, Reserva Científica Ebano Verde,
1010 m
Walkerana pedetes (Otte & Perez-Gelabert, 2009: 258). Otte, 2009: 489 *DR > TL: Cortico, Monteada Nueva, Barahona Prov.,
1433 m
Walkerana rhopheticos (Otte & Perez-Gelabert, 2009: 259). Otte, 2009: 489 *DR > TL: 5 km NE dirt rd off Km 41 Carretera
Duarte, San Cristóbal Prov., 285 m
Walkerana timens (Otte & Perez-Gelabert, 2009: 258). Otte, 2009: 489 *DR > TL: Arroyazo, Reserva Científica Ebano Verde,
990 m
Walkerana tinniens (Otte & Perez-Gelabert, 2009: 259). Otte, 2009: 489 *DR > TL: Loma del Toro, Sierra de Bahoruco, 2310
m
Walkerana vexans (Otte & Perez-Gelabert, 2009: 258). Otte, 2009: 489 *DR > TL: Diferencia, Parque Nacional Armando
Bermúdez, 750 m
Subtribe HAPITHINA
Hapithus Uhler, 1864
Hapithus (Antillicharis) Otte & Perez-Gelabert, 2009. Gorochov, 2017: 16 (Antillicharis downgraded to subgenus of Hapithus)
=Alexandrina Otte & Perez-Gelabert, 2009
Hapithus (Antillicharis) abejas (Otte & Perez-Gelabert, 2009: 248) *DR > TL: Upper Las Abejas, Parque Nacional Sierra de
Bahoruco, 1310 m
Hapithus (Antillicharis) adaptos (Otte & Perez-Gelabert, 2009: 252) *DR > TL: 3 km S Montecristi
Hapithus (Antillicharis) adorabilis (Otte & Perez-Gelabert, 2009: 253) *DR > TL: Cortico, Barahona Prov., in bromeliads
Hapithus (Antillicharis) agastos (Otte & Perez-Gelabert, 2009: 242) *DR > TL: Trudillé, Parque Nacional Jaragua
Hapithus (Antillicharis) aphenges (Otte & Perez-Gelabert, 2009: 242) *DR > TL: ~2 km S Los Pinos del Edén, Sierra de Neiba

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Hapithus (Antillicharis) celerans (Otte & Perez-Gelabert, 2009: 252) *DR > TL: Monte Río, about 1 km S of beach
Hapithus (Antillicharis) cresbios (Otte & Perez-Gelabert, 2009: 254) *DR > TL: 21 km N of Cabo Rojo, 1200 ft.
Hapithus (Antillicharis) cuphos (Otte & Perez-Gelabert, 2009: 242) *DR > TL: ~5 km W of Las Americas Airport, nr.
LADOM
Hapithus (Antillicharis) demissus (Otte & Perez-Gelabert, 2009: 249) *DR > TL: ~1 km SE caseta no. 1, Parque Nacional
Sierra de Bahoruco, 1153 m
Hapithus (Antillicharis) domingensis (Saussure, 1878: 613). Perez-Gelabert, 2001: 71; Otte & Perez-Gelabert, 2009: 246 *H,
*DR > TL: St.-Domingue
Hapithus (Antillicharis) empsychos (Otte & Perez-Gelabert, 2009: 255) *DR > TL: Road down from La Hoz, Sierra de
Bahoruco, 1170 m
Hapithus (Antillicharis) energos (Otte & Perez-Gelabert, 2009: 253) *DR > TL: ~2 km on road to Los Martínez, Ocoa Prov.,
468 m
Hapithus (Antillicharis) epholcos (Otte & Perez-Gelabert, 2009: 247) *DR > TL: Km 26 Rd. Cabo Rojo-Aceitillar, 680 m
Hapithus (Antillicharis) eribombos (Otte & Perez-Gelabert, 2009: 249) *DR > TL: ~1 km SE caseta no. 1, Parque Nacional
Sierra de Bahoruco, 1153 m
Hapithus (Antillicharis) errabundus (Otte & Perez-Gelabert, 2009: 251) *DR > 3 km ESE El Aguacate, Independencia Prov.
Hapithus (Antillicharis) habros (Otte & Perez-Gelabert, 2009: 250) *H > TL: Kenscoff, 1500 m
Hapithus (Antillicharis) illectans (Otte & Perez-Gelabert, 2009: 254) *DR > TL: Río Limpio, Elías Piña Prov., 720 m
Hapithus (Antillicharis) illex (Otte & Perez-Gelabert, 2009: 254) *DR > TL: La Jarda (Monte Abajo), Padre Las Casas, Azua
Prov.
Hapithus (Antillicharis) nanion (Otte & Perez-Gelabert, 2009: 251) *DR > TL: Baoruco Mountains, 250 m
Hapithus (Antillicharis) pannychios (Otte & Perez-Gelabert, 2009: 248) *DR > TL: 4 km W of Las Americas Airport, Santo
Domingo
Hapithus (Antillicharis) pelliciens (Otte & Perez-Gelabert, 2009: 249) *DR > TL: Guizo, Monteada Nueva
Hapithus (Antillicharis) properatos (Otte & Perez-Gelabert, 2009: 253) *DR > TL: East of Polo, Barahona Prov.
Hapithus (Antillicharis) regificus (Otte & Perez-Gelabert, 2009: 247) *DR > TL: 30 km N Cabo Rojo, Pedernales Prov.
Hapithus (Antillicharis) regillus (Otte & Perez-Gelabert, 2009: 247) *DR > TL: 30 km N Cabo Rojo, Pedernales Prov.
Hapithus (Antillicharis) saukros (Otte & Perez-Gelabert, 2009: 250) *H > TL: Kenscoff, 1500 m
Hapithus (Antillicharis) similis (F. Walker, 1869). Wolcott, 1927: 149; Perez-Gelabert, 2001: 71; Otte & Perez-Gelabert, 2009:
248
Hapithus (Antillicharis) tychaeos (Otte & Perez-Gelabert, 2009: 253) *DR > TL: Km 54 Rd. Azua - Barahona, Barahona Prov.,
70 m
Hapithus (Antillicharis) tyrannicos (Otte & Perez-Gelabert, 2009: 252) *DR > TL: 37 km N Cabo Rojo, 1500 m
Hapithus (Antillicharis) vigil (Otte & Perez-Gelabert, 2009: 247) *DR > Arroyazo, Reserva Científica Ebano Verde, 990 m
Hapithus (Antillicharis) vigilax (Otte & Perez-Gelabert, 2009: 255) *DR > TL: Dry forest of Monte Río, Azua Prov., 40 ft.
Hapithus (Antillicharis) vivus (Otte & Perez-Gelabert, 2009: 255) *DR > TL: Playa Blanca, Parque Nacional Jaragua
Hapithus (Antillicharis) zatheos (Otte & Perez-Gelabert, 2009: 252) *DR > TL: Loma del Toro, Sierra de Bahoruco
Hapithus (Hapithus) Uhler, 1864. Gorochov, 2017: 15
Hapithus (Hapithus) bellus Otte & Perez-Gelabert, 2009: 230 *H > TL: Port-au-Prince
Hapithus (Hapithus) cabralense Otte & Perez-Gelabert, 2009: 229 *DR > TL: ~4 km S of Cabral, Barahona Prov., 105 m
Hapithus (Hapithus) cerbatana Otte & Perez-Gelabert, 2009: 228 *DR > TL: Jardin Botánico Nacional, Reserva Forestal,
Santo Domingo
Hapithus (Hapithus) cucurucho Otte & Perez-Gelabert, 2009: 229 *DR > TL: Cuevita, Valle Nuevo, 2076 m
Hapithus (Hapithus) elisae Otte & Perez-Gelabert, 2009: 228 *DR > TL: Around Caseta No. 1, Parque Nacional Sierra de
Bahoruco, 1239 m
Hapithus (Hapithus) epidendrios (Otte & Perez-Gelabert, 2009: 263) *DR > TL: La Malena de Boca Chica
Hapithus (Hapithus) fraudans (Otte & Perez-Gelabert, 2009: 263) *DR > TL: ~2 km N Bayahibe, La Altagracia Prov.
Hapithus (Hapithus) lucreciae Otte & Perez-Gelabert, 2009: 229 *DR > TL: Sierra Prieta, Villa Mella
Hapithus (Hapithus) mabuya Otte & Perez-Gelabert, 2009: 228 *DR > TL: La Travesía, Barahona Prov., 900 m
Hapithus (Hapithus) mythicos Otte & Perez-Gelabert, 2009: 228 *DR > TL: Arroyazo, Reserva Científica Ebano Verde, 3500
ft.
Hapithus (Hapithus) polymechanus Otte & Perez-Gelabert, 2009: 229 *DR > TL: 2 km SW of Canada, N slope of Sierra de
Neiba, 980 m
Hapithus (Hapithus) pudens (Otte & Perez-Gelabert, 2009: 261) *DR > TL: Sierra Prieta, Villa Mella, 500 ft.

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Hapithus (Hapithus) sajoma (Otte & Perez-Gelabert, 2009: 263) *DR > TL: San José de las Matas
Hapithus (Hapithus) samanense Otte & Perez-Gelabert, 2009: 228 *DR > TL: La Laguna, N. El Valle, Samaná Prov., 54 m
Hapithus (Hapithus) tenuicornis (F. Walker, 1869: 75). Perez-Gelabert, 2001: 71 *H, *DR > TL: St. Domingo
Hapithus (Laurepa) Walker, 1869. Gorochov, 2017: 16 (Laurepa downgraded to subgenus of Hapithus)
=Orochirus Bolívar, 1888
Hapithus (Laurepa) antiphonos (Otte & Perez-Gelabert, 2009: 270) *DR > TL: Las Yayitas, Azua Prov., 240 m
Hapithus (Laurepa) catacrotos (Otte & Perez-Gelabert, 2009: 274) *DR > TL: ~4 km SE Montecristi, 27 m
Hapithus (Laurepa) ecplecticos (Otte & Perez-Gelabert, 2009: 271) *DR > TL: Down from Majagual, nr. Galván, Sierra de
Neiba, 141 m
Hapithus (Laurepa) epagogos (Otte & Perez-Gelabert, 2009: 272) *DR > TL: Juan Dolio, San Pedro de Macorís Prov.
Hapithus (Laurepa) epimeles (Otte & Perez-Gelabert, 2009: 271) *DR > TL: Near Angel Feliz, Sierra de Neiba,
Hapithus (Laurepa) gavisa (Otte & Perez-Gelabert, 2009: 271) *DR > TL: Gran Chorra, Parque Nacional del Este
Hapithus (Laurepa) gymnopta (Otte & Perez-Gelabert, 2009: 272) *DR > TL: Km 26 Rd. Cabo Rojo-Aceitillar, Pedernales
Prov., 680 m
Hapithus (Laurepa) krugi (Saussure, 1878). Perez-Gelabert, 2001: 71; Yong & Perez-Gelabert, 2014: 419
Hapithus (Laurepa) paraxynticos (Otte & Perez-Gelabert, 2009: 275) *DR > TL: El Cajuil, Oviedo, 52 m
Hapithus (Laurepa) periphantos (Otte & Perez-Gelabert, 2009: 274) *DR > TL: Caseta no. 3, Parque Nacional Sierra de
Bahoruco, 1941 m
Hapithus (Laurepa) phasma (Otte & Perez-Gelabert, 2009: 270) *DR > TL: Pueblo Nuevo, Baní, 97 m
Hapithus (Laurepa) prosplatos (Otte & Perez-Gelabert, 2009: 270) *DR > TL: La Malena de Boca Chica, Santo Domingo
Prov.
Hapithus (Laurepa) semnos (Otte & Perez-Gelabert, 2009: 274) *DR > TL: 23.5 km N Cabo Rojo
Hapithus (Laurepa) spectrum (Otte & Perez-Gelabert, 2009: 270) *DR > TL: Km 8 Cabo Rojo - Aceitillar, 27 m
Hapithus (Laurepa) trepida (Otte & Perez-Gelabert, 2009: 271) *DR > TL: Jardín Botánico Nacional, reserva forestal, Santo
Domingo
Hapithus (Laurepa) turbulenta (Otte & Perez-Gelabert, 2009: 276) *DR > TL: Road down from La Hoz, Sierra de Bahoruco,
1110 m
Hapithus (Mashiyana) Otte & Perez-Gelabert, 2009. Gorochov, 2017: 16 (Mashiyana downgraded to subgenus of Hapithus)
Hapithus (Mashiyana) evanidus (Otte & Perez-Gelabert, 2009: 235) *DR > TL: Cortico, Monteada Nueva, Barahona Prov.,
1433 m
Jabulania Otte & Perez-Gelabert, 2009
Jabulania clancularia Otte & Perez-Gelabert, 2009: 490 *DR > TL: 5 km NE Los Arroyos, Pedernales Prov., 1680 m
Stenogryllus Saussure, 1878
Stenogryllus bromia Otte & Perez-Gelabert, 2009: 480 *DR > TL: Caseta no. 3, Parque Nacional Sierra de Bahoruco, 1941 m
Stenogryllus diadelos Otte & Perez-Gelabert, 2009: 477 *DR > TL: Las Yayitas, Azua Prov., 240 m
Stenogryllus disjectus Otte & Perez-Gelabert, 2009: 477 *DR > TL: La Mina de Cortico, Monteada Nueva, 1250 m
Stenogryllus epimeces Otte & Perez-Gelabert, 2009: 482 *DR > TL: Around Caseta No. 1, Parque Nacional Sierra de
Bahoruco, 1239 m
Stenogryllus exallos Otte & Perez-Gelabert, 2009: 480 *DR > TL: 0.5 Km E Federación Campesinos de Blanco, Bonao,
Monseñor Nouel Prov., 577 m
Stenogryllus fremens Otte & Perez-Gelabert, 2009: 480 *DR > TL: ~100 m up from El Sitio del Agua, N Los Bolos, Sierra de
Neiba, 1520 m
Stenogryllus phthisicus Saussure, 1878: 695. Perez-Gelabert, 2001: 70
Stenogryllus psophodes Otte & Perez-Gelabert, 2009: 477 *DR > TL: ~1 km SE caseta no. 1, Parque Nacional Sierra de
Bahoruco, 1153 m
Subfamily PODOSCIRTINAE
Tribe PAROECANTHINI
Ectotrypa Saussure, 1874
=Chremon Rehn, 1930. Gorochov, 2011: 251
Ectotrypa anembatos (Otte & Perez-Gelabert, 2009: 485). Gorochov, 2011: 253 *DR > TL: ~ 2.5 km N La Colonia, San
Cristóbal Prov., 600 m
Ectotrypa apithanon (Otte & Perez-Gelabert, 2009: 485). Gorochov, 2011: 253 *DR > TL: Río Limpio, Elías Pina Prov., 769 m
Ectotrypa eribombos (Otte & Perez-Gelabert, 2009: 485). Gorochov, 2011: 253 *DR > TL: East of Polo, Barahona Prov.
Ectotrypa jaiqui (Otte & Perez-Gelabert, 2009: 485). Gorochov, 2011: 253 *DR > TL: Jaiquí Picado, Santiago Prov., 531 m

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Ectotrypa procax (Otte & Perez-Gelabert, 2009: 487). Gorochov, 2011: 253 *DR > TL: Mano Matuey, San Cristóbal Prov., 460
m
Ectotrypa repentinus (Rehn, 1930: 87). Perez-Gelabert, 2001: 71; Gorochov, 2011: 253 *H > TL: Fond des Negres
Paroecanthus Saussure, 1874
Paroecanthus scinax Otte & Perez-Gelabert, 2009: 491 *DR > TL: La Poza de Agua Nueva, El Curro, Sierra Martín García,
~800 m
Paroecanthus sporadicos Otte & Perez-Gelabert, 2009: 490 *DR > TL: Sierra Martín García, 7 km WNW of Barrero
Paroecanthus valens Otte & Perez-Gelabert, 2009: 491 *DR > TL: La Travesía, nr. Larimar Mine, Eastern Sierra de Bahoruco,
850 m
Tafalisca F. Walker, 1869
Tafalisca lurida F. Walker, 1869: 53. Perez-Gelabert, 2001: 69
Subfamily GRYLLINAE
Hispanogryllus Otte & Perez-Gelabert, 2009
Hispanogryllus adectos Otte & Perez-Gelabert, 2009: 45 *DR > TL: La Laguna, N. El Valle, Samaná Prov., 54 m
Tribe GRYLLINI
Anurogryllus Saussure, 1877
Anurogryllus ellops Otte & Perez-Gelabert, 2009: 44 *DR > TL: Matadero, Peravia Prov., 1600 ft.
Anurogryllus hierroi Otte & Perez-Gelabert, 2009: 45 *DR > TL: Near Jura River, Peralta, Azua Prov., 510 m
Anurogryllus muticus caraibeus Saussure, 1774. Wolcott, 1927: 150; Perez-Gelabert, 2001: 69
Anurogryllus nyctinomos Otte & Perez-Gelabert, 2009: 44 *DR > TL: ~2 km S Los Pinos del Eden, Sierra de Neiba
Anurogryllus vibrans Otte & Perez-Gelabert, 2009: 44 *DR > TL: La Poza de Agua Nueva, El Curro, Sierra Martín García,
~800 m
Gryllus Linnaeus, 1758
Gryllus assimilis assimilis (Fabricius, 1775). Wolcott, 1927: 150; Perez-Gelabert, 2001: 70
Scapsipedus Saussure, 1877
Scapsipedus bastardoae Otte & Perez-Gelabert, 2009: 38 *DR > TL: Río Los Dajaos [Correction of original spelling, species
dedicated to Dominican biologist Ruth H. Bastardo]
=Scapsipedus bastardoi Otte & Perez-Gelabert, 2009: 38
Tribe MODICOGRYLLINI
Geogryllus Otte & Perez-Gelabert, 2009
Geogryllus arndti (Randell, 1964: 1561). Perez-Gelabert, 2001: 70 *H > TL: Fond des Negres
Geogryllus bondi (Randell, 1964: 1560). Perez-Gelabert, 2001: 70 *H > TL: Morne Tranchant, 5000 ft.
Geogryllus dosenos Otte & Perez-Gelabert, 2009: 47 *DR > TL: S slope of Morne Formon, Massif de la Hotte, 1405 m
Geogryllus dyscheres Otte & Perez-Gelabert, 2009: 48 *DR > TL: Aserradero, ~26 km S Puerto Escondido, Parque Nacional
Sierra de Bahoruco, 1635 m
Geogryllus excavator Otte & Perez-Gelabert, 2009: 48 *DR > TL: ~100 m up from El Sitio del Agua, N Los Bolos, Sierra de
Neiba, 1520 m
Geogryllus oryctes Otte & Perez-Gelabert, 2009: 48 *DR > TL: Oviedo
Geogryllus uhleri (Randell, 1964: 1562). Perez-Gelabert, 2001: 70 *H > TL: Haiti
Gryllodes Saussure, 1874
Gryllodes sigillatus (F. Walker, 1869). Otte, 2006: 227 [Introduced]
Miogryllus Saussure, 1877
Miogryllus bellator Otte & Perez-Gelabert, 2009: 41 *DR > TL: Boca de la Cañada, Parque Nacional Jaragua, 18 Km S Oviedo
Miogryllus catacustes Otte & Perez-Gelabert, 2009: 41 *DR > TL: Los Jobos, Azua Prov., 469 m
Miogryllus ergaticos Otte & Perez-Gelabert, 2009: 41 *DR > TL: Around Caseta No. 1, Parque Nacional Sierra de Bahoruco,
1239 m
Miogryllus pugnans Otte & Perez-Gelabert, 2009: 40 *DR > TL: Road down from La Hoz, Sierra de Bahoruco, 1110 m
Miogryllus rehni (Randell, 1964: 1563). Perez-Gelabert, 2001: 70; Perez-Gelabert & Otte, 2009: 40 *H > TL: Port-au-Prince
Subfamily NEMOBIINAE
Tribe PTERONEMOBIINI
Hygronemobius Hebard, 1928
Hygronemobius epia Otte & Perez-Gelabert, 2009: 105 *DR > TL: Arroyazo, Reserva Cientifica Ebano Verde, 990 m
Pteronemobius Jacobson in Jacobson & Bianchi, 1904
Pteronemobius cantor Otte & Perez-Gelabert, 2009: 108 *DR > TL: 3 km West of Cueva de Arena, Parque Los Haitises

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Pteronemobius caryochron Otte & Perez-Gelabert, 2009: 107 *DR > TL: La Mina de Cortico, Monteada Nueva, Barahona
Prov., 1250 m
Pteronemobius setiger Otte & Perez-Gelabert, 2009: 108 *DR > TL: Arroyazo, Reserva Científica Ebano Verde
Subfamily OECANTHINAE
Oecanthus Serville, 1831
Oecanthus allardi T. Walker & Gurney, 1960. Perez-Gelabert, 2001: 70
Subfamily PENTACENTRINAE
Tribe NEMOBIOPSINI
Grossoxipha† Vickery & Poinar, 1994
Grossoxipha yaque† Vickery & Poinar, 1994: 18. Perez-Gelabert, 2001: 72 [Fossil in Dominican amber]
Grossoxipha feminea† Gorochov, 2010: 444 [Fossil in Dominican amber]
Nemobiopsis Bolivar, I., 1890
Nemobiopsis cortico Otte & Perez-Gelabert, 2009: 545 *DR > TL: 1 km ESE Cortico Barahona Prov., 1347 m
Nemobiopsis diadromos Otte & Perez-Gelabert, 2009: 547 *DR > TL: Rodeo, 0.5 km E Presa de Blanco, Monseñor Nouel
Prov.
Nemobiopsis eratos Otte & Perez-Gelabert, 2009: 545 *DR > TL: 25 km N of Cabo Rojo, 700 m
Nemobiopsis indocilis Otte & Perez-Gelabert, 2009: 545 *DR > TL: La Travesía, nr. Larimar Mine, Eastern Sierra de
Bahoruco, 850 m
Nemobiopsis jabase Otte & Perez-Gelabert, 2009: 548 *DR > TL: Las Abejas, Parque Nacional Sierra de Bahoruco, 1310 m
Nemobiopsis metanasticos Otte & Perez-Gelabert, 2009: 548 *DR > TL: 26 km N of Cabo Rojo, Pedernales Prov.
Nemobiopsis planeticos Otte & Perez-Gelabert, 2009: 547 *DR > TL: Km 26 Rd. Cabo Rojo - Aceitillar, 2500 ft.
Nemobiopsis vagabundus Otte & Perez-Gelabert, 2009: 548 *DR > TL: Along Río Mulito, N of Pedernales
Proanaxipha† Vickery & Poinar, 1994
Proanaxipha bicolorata† Vickery & Poinar, 1994: 16. Perez-Gelabert, 2001: 72 [Fossil in Dominican amber]
Proanaxipha latoca† Vickery & Poinar, 1994: 15. Perez-Gelabert, 2001: 72 [Fossil in Dominican amber]
Proanaxipha madgesuttonae† Heads & Penney in Heads, Penney & Green, 2012: 113 [Fossil in Dominican amber]
Velapia Otte & Perez-Gelabert, 2009
Velapia epaxia Otte & Perez-Gelabert, 2009: 548 *DR > TL: Los Tablones - Agüita Fría, Parque Nacional A. Bermúdez
Velapia myrmex Otte & Perez-Gelabert, 2009: 548 *DR > TL: After Pyramid 204, Sierra de Neiba, 1450 m

Family PHALANGOPSIDAE
Subfamily LUZARINAE
Amphiacusta Saussure, 1874
Amphiacusta abatos Otte & Perez-Gelabert, 2009: 587 *DR > TL: Larimar mine, Filipinas, Barahona Prov.
Amphiacusta aetheria Otte & Perez-Gelabert, 2009: 582 *DR > TL: Mano Matuey, 2 km N of La Colonia, San Cristóbal Prov.
Amphiacusta annulipes (Serville, 1831). Wolcott, 1927: 150; Perez-Gelabert, 2001: 70
Amphiacusta apoplanos Otte & Perez-Gelabert, 2009: 584 *DR > TL: Precise locality unknown
Amphiacusta dapanera Otte & Perez-Gelabert, 2009: 587 *DR > TL: ~4 km on rd. to Honduras, nr. second bridge, Peravia
Prov.
Amphiacusta dimalea Otte & Perez-Gelabert, 2009: 583 *DR > TL: Bucán de Tuí, Parque Nacional Jaragua, sea level
Amphiacusta dissimilis Desutter-Grandcolas, 1997: 110. Perez-Gelabert, 2001: 70 *DR > TL: San Francisco mountains, San
Cristóbal Prov.
Amphiacusta domingensis Desutter-Grandcolas, 1997: 108. Perez-Gelabert, 2001: 71 *DR > TL: San Francisco mountains, San
Cristóbal Prov.
Amphiacusta eritheles Otte & Perez-Gelabert, 2009: 582 *DR > TL: La Jarda (monte abajo), Padre Las Casas, Azua Prov.
Amphiacusta exacerbans Otte & Perez-Gelabert, 2009: 584 *DR > TL: Down from Blanco, Bonao, Monseñor Nouel Prov., 696
m
Amphiacusta expansa Otte & Perez-Gelabert, 2009: 583 *DR > TL: Boca de la Cañada, 15 km S Oviedo
Amphiacusta extenta Otte & Perez-Gelabert, 2009: 586 *DR > TL: Los Tablones, Parque Nacional Armando Bermúdez
Amphiacusta fascinans Otte & Perez-Gelabert, 2009: 583 *DR > TL: Bayahibe, La Altagracia Prov.
Amphiacusta ficticia Otte & Perez-Gelabert, 2009: 587 *DR > TL: Near Angel Feliz, Sierra de Neiba
Amphiacusta haitiana Desutter-Grandcolas, 1997: 108. Perez-Gelabert, 2001: 71 *H > TL: Port-au-Prince
Amphiacusta haitianella Desutter-Grandcolas, 1997: 110. Perez-Gelabert, 2001: 71 *H > TL: Pétionville
Amphiacusta haitiensis Desutter-Grandcolas, 1997: 110. Perez-Gelabert, 2001: 71 *H > TL: Fond des Negres

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Amphiacusta hespera Otte & Perez-Gelabert, 2009: 583 *DR > TL: Sierra Prieta, Santo Domingo, 133 m
Amphiacusta hispaniolae Desutter-Grandcolas, 1997: 107. Perez-Gelabert, 2001: 71 *H > TL: Furcy, Dept. de l’Ouest
Amphiacusta mingodo Otte & Perez-Gelabert, 2009: 582 *DR > TL: San Francisco mountains [San Cristóbal Prov.]
Amphiacusta minima Desutter-Grandcolas, 1997: 112. Perez-Gelabert, 2001: 71 *DR > TL: 1 mile W of Ciudad Trujillo
Amphiacusta mythica Otte & Perez-Gelabert, 2009: 582 *DR > TL: Pueblo Nuevo, Baní, Peravia Prov., 97 m
Amphiacusta obstipa Otte & Perez-Gelabert, 2009: 587 *DR > TL: La Furnia, Barreras, Sierra Martín García, Azua Prov.
Amphiacusta ominosa Otte & Perez-Gelabert, 2009: 584 *DR > TL: ~3 km SE Montecristi, very dry forest
Amphiacusta opsia Otte & Perez-Gelabert, 2009: 582 *DR > TL: 2 km N Federación Campesinos de Blanco, Bonao, Monseñor
Nouel Prov., 577 m
Amphiacusta pavida Otte & Perez-Gelabert, 2009: 587 *DR > TL: La Travesía, nr. Larimar mine, Eastern Sierra de Bahoruco
Amphiacusta pomierense Otte & Perez-Gelabert, 2009: 584 *DR > TL: Anthropological reserve, El Pomier cave, San Cristóbal
Prov.
Amphiacusta strena Otte & Perez-Gelabert, 2009: 583 *DR > TL: Charquito Prieto, El Curro, Sierra Martín García, 731 m
Amphiacusta tachydromos Otte & Perez-Gelabert, 2009: 584 *DR > TL: 0.5 km E Federación Campesinos de Blanco, Bonao,
Monseñor Nouel Prov., 577 m
Amphiacusta taracte Otte & Perez-Gelabert, 2009: 582 *DR > TL: Loma Quita Espuela, rangers quarters, Duarte Prov.
Amphiacusta ultima Desutter-Grandcolas, 1997: 112. Perez-Gelabert, 2001: 71 *DR > TL: 5 km N Villa Elisa, in cave,
Montecristi Prov.
Amphiacusta vespera Otte & Perez-Gelabert, 2009: 583 *DR > TL: Jardín Botánico Nacional, Reserva Forestal, Santo
Domingo
Antilliclodes Otte & Perez-Gelabert, 2009
Antilliclodes bermudezi Otte & Perez-Gelabert, 2009: 667 *DR > TL: La Ciénaga - Los Tablones, Parque Nacional Armando
Bermúdez
Antilliclodes fugitans Otte & Perez-Gelabert, 2009: 667 *DR > TL: Aguas Blancas, S. Constanza, La Vega Prov.
Araneagryllus† Heads, 2010
Araneagryllus dylani† Heads, 2010b: 58 [Fossil in Dominican amber]
Cophella Hebard, 1928
Cophella ceroton Otte & Perez-Gelabert, 2009: 676 *DR > TL: La Travesía, Sierra de Bahoruco, Barahona Prov., 900 m
Cophella diva Otte & Perez-Gelabert, 2009: 676 *DR > TL: Arroyo Toro, Monseñor Nouel Prov.
Cophella drasticor Otte & Perez-Gelabert, 2009: 676 *DR > TL: Cortico, Monteada Nueva, cloud forest
Cophella evides Otte & Perez-Gelabert, 2009: 676 *DR > TL: ~100 m up from El Sitio del Agua, Sierra de Neiba, 1520 m
Cophella evoplon Otte & Perez-Gelabert, 2009: 676 *DR > TL: Manaclar near Los Anones, Ocoa Prov.
Cophella nychios Otte & Perez-Gelabert, 2009: 677 *DR > TL: ~1 km Road to Monteada Nueva, Barahona Prov., 1059 m
Cophella ramireza Otte & Perez-Gelabert, 2009: 677 *DR > TL: Mina de Oro Pueblo Viejo, Sanchez Ramírez Prov.
Cophella setosa Otte & Perez-Gelabert, 2009: 677 *DR > TL: Reserva de Agua, El Aceitillar, Sierra de Bahoruco
Cophella soteriodes Otte & Perez-Gelabert, 2009: 677 *DR > TL: Guizo, Monteada Nueva, Barahona Prov., 831 m
Dominicophus Yong, 2017
Dominicophus sheylae (Armas & Hernández-Triana, 2014: 113) *DR > TL: Cueva de Neno, Talanquera, Sección Los Cacaos,
Samaná Prov.
Dominicophus terueli Yong, 2017: 4 *DR > TL: Cueva de Berna, Parque Nacional del Este
Uvaroviella Chopard, 1923
Uvaroviella tabulatum Otte & Perez-Gelabert, 2009: 671 *DR > TL: Boca de la Cañada, 18 km S Oviedo, Parque Nacional
Jaragua
Subfamily PARAGRYLLINAE
Tribe PARAGRYLLINI
Yoyuteris Ruíz & Otte, 1996
Yoyuteris barahona Ruíz & Otte, 1996: 231. Perez-Gelabert, 2001: 71 *DR > TL: 4.5 km S of Barahona, Barahona Prov.
Yoyuteris cerritus Otte & Perez-Gelabert, 2009: 674 *DR > TL: La Sal, Reserva Científica Ebano Verde
Yoyuteris hylobates Otte & Perez-Gelabert, 2009: 674 *DR > TL: La Jarda (monte abajo), Padre Las Casas, Azua Prov.
Yoyuteris hylonomos Otte & Perez-Gelabert, 2009: 674 *DR > TL: East of Polo [top of Loma La Hoz]

Family TRIGONIDIIDAE
Subfamily TRIGONIDIINAE
Tribe TRIGONIDIINI

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Abanaxipha† Vickery & Poinar, 1994
Abanaxipha incongrua† Vickery & Poinar, 1994: 17. Perez-Gelabert, 2001: 71 [Fossil in Dominican amber]
Abanaxipha longispina† Vickery & Poinar, 1994: 17. Perez-Gelabert, 2001: 72; Gorochov, 2010: 446 [Fossil in Dominican
amber]
Anaxipha Saussure, 1874
Anaxipha amica Otte & Perez-Gelabert, 2009: 134 *DR > TL: La Travesia, nr. Larimar Mine, Eastern Sierra de Bahoruco, 850
m
Anaxipha antictypos Otte & Perez-Gelabert, 2009: 132 *DR > TL: 2 km N of Bayahibe, La Altagracia Prov.
Anaxipha antidupos Otte & Perez-Gelabert, 2009: 137 *DR > TL: 3.3 km NE Los Arroyos, Sierra de Bahoruco
Anaxipha aphaura Otte & Perez-Gelabert, 2009: 136 *DR > TL: Way down from Palos Grandes, NE of San José de Ocoa,
Ocoa Prov., 1440 m
Anaxipha apseudes Otte & Perez-Gelabert, 2009: 132 *DR > TL: La Sal, Reserva Científica Ebano Verde, 1043 m
Anaxipha arena Otte & Perez-Gelabert, 2009: 134 *DR > TL: 3 km W of Cueva de Arena, Parque Los Haitises
Anaxipha athanes Otte & Perez-Gelabert, 2009: 136 *DR > TL: Cortico, Monteada Nueva, Barahona Prov., 1433 m
Anaxipha charitopis Otte & Perez-Gelabert, 2009: 136 *DR > TL: La Mina de Cortico, Monteada Nueva, Barahona Prov.,
1250 m
Anaxipha cregyos Otte & Perez-Gelabert, 2009: 137 *DR > TL: Km 17 Rd. Cabo Rojo - Aceitillar, Pedernales Prov., 1400 ft.
Anaxipha dominica† Vickery & Poinar, 1994: 14. Perez-Gelabert, 2001: 72 [Fossil in Dominican amber]
Anaxipha eminens Otte & Perez-Gelabert, 2009: 134 *DR > TL: ~100 m up from El Sitio del Agua, N Los Bolos, Sierra de
Neiba, 1520 m
Anaxipha eperatos Otte & Perez-Gelabert, 2009: 137 *DR > TL: ~4 km E of Montecristi
Anaxipha euclastos Otte & Perez-Gelabert, 2009: 137 *DR > TL: Manaclar, near Los Anones, Ocoa Prov.
Anaxipha festina Otte & Perez-Gelabert, 2009: 136 *DR > TL: Río Limpio, Elias Piña Prov., 720 m
Anaxipha furtiva Otte & Perez-Gelabert, 2009: 137 *DR > TL: Margen Río Mulito, 19 km N Pedernales
Anaxipha hypsipetes Otte & Perez-Gelabert, 2009: 134 *DR > TL: Río Limpio, Elias Piña Prov., 769 m
Anaxipha lathrios Otte & Perez-Gelabert, 2009: 137 *DR > TL: Río de los Negros, Parque Nacional Armando Bermúdez
Anaxipha phoxi Otte & Perez-Gelabert, 2009: 136 *DR > TL: Guizo, Monteada Nueva, Barahona Prov., 831 m
Anaxipha sp.†—Spahr (1992: 81) [Fossil in Dominican amber]
Anaxipha vivax Otte & Perez-Gelabert, 2009: 129 *DR > TL: Hilltop on way to Palos Grandes, NE of San José de Ocoa, Ocoa
Prov., 1437 m
Cyrtoxipha Brunner von Wattenwyl, 1873
Cyrtoxipha abares Otte & Perez-Gelabert, 2009: 143 *DR > TL: 5 km NNE Botoncillo, Montecristi Prov.
Cyrtoxipha aguanueva Otte & Perez-Gelabert, 2009: 145 *DR > TL: La Poza de Agua Nueva, El Curro, Sierra Martín García,
~800 m
Cyrtoxipha aleptos Otte & Perez-Gelabert, 2009: 145 *DR > TL: Paraiso, Barahona Prov.
Cyrtoxipha apiata Otte & Perez-Gelabert, 2009: 146 *DR > TL: El Cajuil, Oviedo, Pedernales Prov., 52 m
Cyrtoxipha cryptadia Otte & Perez-Gelabert, 2009: 143 *DR > TL: Upper Las Abejas, Sierra de Bahoruco, 1350 m
Cyrtoxipha ctilos Otte & Perez-Gelabert, 2009: 145 *DR > TL: Trail Los Tablones - La Ciénaga, Parque Nacional Armando
Bermúdez, La Vega Prov.
Cyrtoxipha electrina† Gorochov, 2010: 448 [Fossil in Dominican amber]
Cyrtoxipha errans Otte & Perez-Gelabert, 2009: 145 *DR > TL: 3.3 km NE of Los Arroyos, Sierra de Bahoruco
Cyrtoxipha eucrines Otte & Perez-Gelabert, 2009: 145 *DR > TL: Arroyazo, Reserva Científica Ebano Verde, 990 m
Cyrtoxipha hosios Otte & Perez-Gelabert, 2009: 143 *DR > TL: Slopes of Loma La Torre, 5 km SE Polo
Cyrtoxipha illegibilis† Gorochov, 2010: 449 [Fossil in Dominican amber]
Cyrtoxipha itame Otte & Perez-Gelabert, 2009: 146 *DR > TL: Juan Dolio, San Pedro de Macoris Prov.
Cyrtoxipha laurelae Otte & Perez-Gelabert, 2009: 145 *DR > TL: Loma de las Tayotas, Elías Piña Prov.
Cyrtoxipha tempestiva Otte & Perez-Gelabert, 2009: 146 *DR > TL: Loma de los Vientos, 4 km S of Los Pinos, Sierra de
Neiba
Cyrtoxipha vespertina Otte & Perez-Gelabert, 2009: 144 *DR > TL: Paraiso, Barahona Prov.

Family GRYLLOTALPIDAE
Subfamily GRYLLOTALPINAE
Neoscapteriscus Cadena-Castaňeda, 2015
Neoscapteriscus abbreviatus (Scudder, 1869). Perez-Gelabert, 2001: 72

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Neoscapteriscus didactylus (Latreille, 1804). Perez-Gelabert, 2001: 72

Family MOGOPLISTIDAE
Cycloptilum Scudder, 1868
Cycloptilum absconditum Otte & Perez-Gelabert, 2009: 741 *DR > TL: Paraiso, Barahona Prov.
Cycloptilum abstrusum Otte & Perez-Gelabert, 2009: 742 *DR > TL: 2 km N Bayahibe, La Altagracia Prov.
Cycloptilum adecton Otte & Perez-Gelabert, 2009: 742 *DR > TL: Pueblo Nuevo, Baní, Peravia Prov.
Cycloptilum ambrosion Otte & Perez-Gelabert, 2009: 742 *DR > TL: Nr. km 8 rd. Cabo Rojo - Aceitillar
Cycloptilum animosum Otte & Perez-Gelabert, 2009: 742 *DR > TL: Los Pinos del Edén, Sierra de Neiba
Cycloptilum antimimon Otte & Perez-Gelabert, 2009: 743 *DR > TL: Monte Río, Azua Prov.
Cycloptilum aphanton Otte & Perez-Gelabert, 2009: 743 *DR > TL: Way down from Palos Grandes, NE of San José de Ocoa,
Ocoa Prov., 1440 m
Cycloptilum celatum Otte & Perez-Gelabert, 2009: 743 *DR > TL: La Colonia, San Cristóbal Prov., 773 m
Cycloptilum cineticon Otte & Perez-Gelabert, 2009: 743 *DR > TL: Paraiso, Barahona Prov.
Cycloptilum clandestinum Otte & Perez-Gelabert, 2009: 743 *DR > TL: El Cajuil, Oviedo, Pedernales Prov., 52 m
Cycloptilum contectum (Rehn & Hebard, 1912: 203). Perez-Gelabert, 2001: 70 *H > TL: Haiti
Cycloptilum crypton Otte & Perez-Gelabert, 2009: 744 *DR > TL: Sierra Prieta, Santo Domingo, 133 m
Cycloptilum epimonon Otte & Perez-Gelabert, 2009: 744 *DR > TL: Mano Matuey, San Cristóbal Prov., 460 m
Cycloptilum halticon Otte & Perez-Gelabert, 2009: 744 *DR > TL: Mano Matuey, San Cristóbal Prov., 460 m
Cycloptilum hypoclopon Otte & Perez-Gelabert, 2009: 744 *DR > TL: Dry forest of Monte Río, Azua Prov., 40 ft.
Cycloptilum inops Otte & Perez-Gelabert, 2009: 747 *DR > TL: Sierra Prieta, Santo Domingo, 133 m
Cycloptilum liberum Otte & Perez-Gelabert, 2009: 744 *DR > TL: Juan Dolio, San Pedro de Macorís Prov.
Cycloptilum nesydrion Otte & Perez-Gelabert, 2009: 745 *DR > TL: Pueblo Nuevo, Baní, Peravia Prov., 97 m
Cycloptilum occultum Otte & Perez-Gelabert, 2009: 745 *DR > TL: Monte Río, ~4 km W of beach, Azua Prov., 70 m
Cycloptilum opertanium Otte & Perez-Gelabert, 2009: 745 *DR > TL: Playa Blanca, Parque Nacional Jaragua, nr. sea level
Cycloptilum panurgon Otte & Perez-Gelabert, 2009: 745 *DR > TL: Caseta Parques en Diferencia, Parque Nacional Armando
Bermúdez, Santiago Prov., 740 m
Cycloptilum pusillulum Otte & Perez-Gelabert, 2009: 745 *DR > TL: Pueblo Nuevo, Baní, Peravia Prov.
Cycloptilum sanum Otte & Perez-Gelabert, 2009: 745 *DR > TL: 11 kms N of Entrance to Honduras, W of Baní, Peravia Prov.
Cycloptilum thymicon Otte & Perez-Gelabert, 2009: 746 *DR > TL: Way down from Palos Grandes, NE of San José de Ocoa,
Ocoa Prov., 1440 m
Ornebius Guérin-Méneville, 1844
Ornebius ambericus† Vickery & Poinar, 1994: 18. Perez-Gelabert, 2001: 70 [Fossil in Dominican amber]
Pseudarachnocephalus† Gorochov, 2010
Pseudarachnocephalus dominicanus† Gorochov, 2010: 437 [Fossil in Dominican amber]
Pseudarachnocephalus latiusculus† Gorochov, 2010: 439 [Fossil in Dominican amber]

Family MYRMECOPHILIDAE
Myrmecophilus Berthold, 1827
Myrmecophilus (Myrmophilina) americanus Saussure, 1877. Perez-Gelabert, 2001: 70; Perez-Gelabert, 2008: 291; Wetterer &
Hugel, 2008: 157.
=Myrmecophyla prenolepidis Mann, 1915: 161

Family ANOSTOSTOMATIDAE
Subfamily ANOSTOSTOMATINAE
Licodia F. Walker, 1869
Licodia grandis Rehn, 1930: 365. Perez-Gelabert, 2001: 72 *H > TL: Coffe Station, Fond des Negres, Massif de la Hotte
Licodia pallipes F. Walker, 1869: 210. Perez-Gelabert, 2001: 72 *DR > TL: St. Domingo
Lutosa F. Walker, 1869
Lutosa obliqua F. Walker, 1869: 211. Perez-Gelabert, 2001: 72

Family GRYLLACRIDIDAE
Subfamily GRYLLACRIDINAE
Abelona Karny, 1937

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Abelona parvula (F. Walker, 1870: 473). Perez-Gelabert, 2001: 72 *DR > TL: St. Domingo
=Gryllacris haitensis Brunner von Wattenwyl, 1888: 364
Brachybaenus Karny, 1937
Brachybaenus domingensis Perez-Gelabert, 2019: 66 *DR > TL: Paso de la Perra, nr. La Ciénaga, La Vega Prov.

Order PHASMATODEA
Key reference: Otte and Brock (2005).

Taxonomic summary:
Suborder Anareolatae
Family Heteronemiidae
Family Phasmatidae
Suborder Areolatae
Family Pseudophasmatidae

Suborder ANAREOLATAE
Family HETERONEMIIDAE
Subfamily HETERONEMIINAE
Clonistria Stål, 1875
Clonistria dominicana† Poinar, 2011: 223 [Fossil in Dominican amber]
Clonistria haita (Westwood, 1859). Otte & Brock, 2005: 107 *H, *DR > TL: Insula St. Domingo
=Bacteria dryas (Westwood, 1859). Otte & Brock, 2005: 107
=Bacteria spinosa Burmeister, 1838). Otte & Brock, 2005: 107

Family PHASMATIDAE
Subfamily CLADOMORPHINAE
Tribe HAPLOPODINI
Cephaloplopus Hennemann et al., 2016
Cephaloplopus euchlorus Hennemann et al., 2016: 47 *DR > TL: Sierra Prieta, Santo Domingo
Cephaloplopus laetus Hennemann et al., 2016: 52 *DR > TL: Azua Prov., La Poza de Agua Nueva, El Curro, Sierra Martín
García, ~800 m
Cephaloplopus pulchellus Hennemann et al., 2016: 54 *DR > TL: Pedernales Prov., Boca de la Cañada, 15 km S Oviedo
Haplopus Burmeister, 1838
Haplopus bicuspidatus de Haan, 1842: 128. Hennenmann et al., 2016: 108 *H, *DR > TL: Described without type locality
Haplopus brachypterus Hennemann et al., 2016: 113 *H, *DR > TL: Entrance to Sabana Iglesia, Santiago Prov., 372 m
Haplopus intermedius Hennemann et al., 2016: 117 *DR > TL: Valverde Prov., ~2 km N Maizal
Haplopus micropterus (St. Fargeau & Audinet-Serville, 1828). Hennemann et al., 2016: 121
Haplopus cythereus Westwood, 1859: 86. Otte & Brock, 2005: 151 *H, *DR > TL: Insula St. Domingo, Haiti
Haplopus ligius Westwood, 1859: 89. Otte & Brock, 2005: 151 *H, *DR > TL: St. Domingo
Haplopus spinipes (Palisot de Beauvois, 1805). Westwood, 1859: 87; Otte & Brock, 2005: 152 *H, *DR > TL: St. Domingo
Parhaplopus Hennemann et al., 2016
Parhaplopus evadne (Westwood, 1859: 85). Otte & Brock, 2005: 151; Hennemann et al., 2016: 156 *H, *DR > TL: Insula St.
Domingo
Parhaplopus navarroi Hennemann et al., 2016: 161 *DR > TL: Monseñor Nouel Prov., Rodeo, ~0.5 km E Presa de Blanco,
Bonao
Tribe HESPEROPHASMATINI
Hesperophasma Rehn, 1901
Hesperophasma planulum (Westwood, 1859: 34). Otte & Brock, 2005: 151 *H, *DR > TL: St. Domingo
Lamponius Stål, 1875
Lamponius sp.—Delfosse (2007: 15)
Lamponius nigli Yong, 2017: 97 *DR > TL: South of Boca de Yuma, La Altagracia Prov.
Taraxippus Moxey, 1971
Taraxippus paliurus Moxey, 1971: 70. Perez-Gelabert, 2000: 23; Otte & Brock, 2005: 330 *H, *DR > TL: High mountains near
St. Louis du Nord

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Suborder AREOLATAE
Family PSEUDOPHASMATIDAE
Subfamily PSEUDOPHASMATINAE
Anisomorpha Gray, 1835
Anisomorpha clara Conle et al., 2006: 886 *DR > TL: Down from Cuevita, Valle Nuevo, 2096 m
Malacomorpha Rehn, 1906
Malacomorpha bastardoae Conle et al., 2008: 15 *DR > TL: Upper Las Abejas, Parque Nacional Sierra de Bahoruco, 1310 m
Malacomorpha hispaniola Conle et al., 2008: 22 *DR, *H > TL: Furcy
Malacomorpha macaya Conle et al., 2008: 33 *H > TL: Camino a Pic le Ciele, 1484 m
Malacomorpha minima Conle et al., 2008: 35 *H > TL: Parc Natl. La Visite, depression, 1850 m
Malacomorpha multipunctata Conle et al., 2008: 38 *DR > TL: ~7 km on way to Caseta no. 1, Parque Nacional Sierra de
Bahoruco, 777 m
Malacomorpha obscura Conle et al., 2008: 42 *DR > TL: 9.7 km NE Los Arroyos, Pedernales, 2070 m
Malacomorpha spinicollis (Burmeister, 1838). Otte & Brock, 2005: 287; Conle et al., 2008: 52 *H, *DR > TL: Port-au-Prince
Tainophasma Conle et al., 2014
Tainophasma monticola Conle et al., 2014: 31 *DR > TL: La Vega Prov., Top of La Pelona, Parque Nacional Armando
Bermúdez

Order PLECOPTERA
Family PERLIDAE
Dominiperla† Stark & Lentz, 1992
Dominiperla antiqua† Stark & Lentz, 1992: 95 [Fossil in Dominican amber]

Order PSOCODEA
Key reference: Lienhard and Smithers (2002). Classification follows Yoshizawa et al. (2006).

Taxonomic summary:
Suborder Amblycera
Family Menopodidae
Family Trichodectidae
Suborder Anoplura
Family Pediculidae
Family Phthiridae
Suborder Trogiomorpha
Infraorder Atropetae
Family Lepidopsocidae
Family Psoquillidae
Family Psyllipsocidae
Suborder Troctomorpha
Infraorder Amphientometae
Family Amphientomidae
Family Troctopsocidae
Infraorder Nanopsocetae
Family Liposcelididae
Suborder Psocomorpha
Infraorder Epipsocetae
Family Cladiopsocidae
Family Dolabellopsocidae
Family Epipsocidae
Infraorder Caeciliusetae
Family Amphipsocidae
Family Caeciliusidae
Infraorder Homilopsocidea

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 Family Archipsocidae
Family Elipsocidae
Family Lachesillidae
Family Philotarsidae
Infraorder Psocetae
Family Hemipsocidae
Family Myopsocidae
Family Psocidae

Suborder AMBLYCERA
Family MENOPODIDAE
Menacanthus Neumann, 1912
Menacanthus stramineum (Nitzsch, 1818). Marcano, 1964: 14
Menopon Ewing, 1929
Menopon biseriatum Piaget, 1880. Wolcott, 1927: 171; Russo, 1927: 42
Menopon gallinae (Linnaeus, 1758). Marcano, 1964: 14

Family TRICHODECTIDAE
Bovicola Ewing, 1929
Bovicola bovis (Linnaeus, 1758). Marcano, 1964: 14
Bovicola caprae (Gurlt, 1843). Marcano, 1964: 13
Bovicola ovis (Schrank, 1871). Marcano, 1964: 14
Felicola Ewing, 1929
Felicola subrostratus (Burmeister, 1838). Marcano, 1964: 14
Werneckiella Eichler, 1940
Werneckiella equi (Denny, 1842). Marcano, 1964: 14

Suborder ANOPLURA
Family PEDICULIDAE
Haematopinus Leach, 1815
Haematopinus asini (Linnaeus, 1758). Marcano, 1964: 14
Haematopinus eurysternus Denny, 1842. Marcano, 1964: 14
Haematopinus suis (Linnaeus, 1758). Marcano, 1964: 14
Haematopinus tuberculatus (Burmeister, 1839). Marcano, 1964: 14
Pediculus Linnaeus, 1758
Pediculus humanus Linnaeus, 1758. Russo, 1927: 42
=Pediculus humanus capitis Latreille, 1803. Wolcott, 1927: 173; Russo, 1927: 42
=Pediculus humanus corporis Latreille, 1803. Wolcott, 1927: 173

Family PHTHIRIDAE
Pthirus Leach, 1815
Pthirus pubis (Linnaeus, 1758). Wolcott, 1927: 173; Russo, 1927: 42

Suborder TROGIOMORPHA
Infraorder ATROPETAE
Family LEPIDOPSOCIDAE
Echmepteryx Aaron, 1886
Echmepteryx sp.†—Mockford (1986: 112) [Fossil in Dominican amber]
Echmepteryx (Loxopholia) dominicanus† Hakim, Huang & Azar, 2018: 61 [Fossil in Dominican amber]

Family PSOQUILLIDAE
Rhyopsocus Hagen, 1876
Rhyopsocus sp.†—Mockford (1986: 112) [Fossil in Dominican amber]

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Family PSYLLIPSOCIDAE
Psyllipsocus Selys, 1872
Psyllipsocus sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]

Suborder TROCTOMORPHA
Infraorder AMPHIENTOMETAE
Family AMPHIENTOMIDAE
Lithoseopsis Mockford, 1993
Lithoseopsis sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]

Family TROCTOPSOCIDAE
Troctopsocoides Mockford & García Aldrete, 2014
Troctopsocoides gracilis† Mockford & García Aldrete, 2014: 163 [Fossil in Dominican amber]

Infraorder NANOPSOCETAE
Family LIPOSCELIDIDAE
Belaphopsocus Badonnel, 1955
Belaphopsocus dominicus† Grimaldi & Engel, 2006: 628 [Fossil in Dominican amber]
Liposcelis Motschulsky, 1853
Liposcelis divinatoria (Müller, 1776). Marcano, 1964: 13
Note: Nomen dubium. Species unrecognizable from the original description and type lost. This is a complex of species.
Liposcelis sp.†—Grimaldi & Engel (2006: 631) [Fossil in Dominican amber]

Suborder PSOCOMORPHA
Infraorder EPIPSOCETAE
Family CLADIOPSOCIDAE
Cladiopsocus Roesler, 1940
Cladiopsocus sp.†—Mockford (1986: 112) [Fossil in Dominican amber]
Spurostigma Eertmoed, 1973
Spurostigma hispaniola Eertmoed, 1973: 406; Lienhard & Smithers, 2002: 126 *DR > TL: 68 m from Ciudad Trujillo,
Highway K, Prov. La Vega
Spurostigma sp.†—Mockford (1986: 112) [Fossil in Dominican amber]

Family DOLABELLOPSOCIDAE
Isthmopsocus Eertmoed, 1973
Isthmopsocus sp.†—Mockford (1986: 112) [Fossil in Dominican amber]

Family EPIPSOCIDAE
Epipsocus Hagen, 1866
Epipsocus sp.†—Mockford (1986: 112) [Fossil in Dominican amber]
Mesepipsocus Badonnel, 1969
Mesepipsocus icarus (Banks, 1941a: 392). Mockford, 1998: 86; Lienhard & Smithers, 2002: 120 *DR > TL: Constanza to
Valle Nuevo, 3 to 7,000 ft.
Mesepipsocoides García-Aldrete & Casasola-González, 2008
Mesepipsocoides perezgelaberti García-Aldrete & Casasola-González, 2008: 219 *DR > TL: Barahona prov., 1 km on road to
Monteada Nueva, 1,059 m

Infraorder CAECILIUSETAE
Family AMPHIPSOCIDAE
Polypsocus Hagen, 1866
Polypsocus fasciatus Banks, 1908. Mockford, 1974: 125; Lienhard & Smithers, 2002: 209

Family CAECILIUSIDAE
Enderleinella Badonnel, 1932

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Enderleinella sp.†—Poinar & Poinar (1999: 202) [Misspelled Enderleenella] [Fossil in Dominican amber]
Stenocaecilius Mockford, 1999
Stenocaecilius antillanus (Banks, 1938: 288). Mockford, 1966: 153; Lienhard & Smithers, 2002: 147
Stenocaecilius insularum (Mockford, 1966: 157). Lienhard & Smithers, 2002: 149
Valenzuela Navás, 1924
Valenzuela alcinus (Banks, 1941a: 390). Mockford, 1969: 78; Mockford, 1999: 350; Lienhard & Smithers, 2002: 152 *DR >
TL: Loma Rucilla and mountains north, 5 to 8,000 ft.
Valenzuela caligonoides Mockford, 1969: 95; Mockford, 1999: 350; Lienhard & Smithers, 2002: 157 *H > TL: Kenscoff
Valenzuela caligonus (Banks, 1941a: 590). Mockford, 1969: 94; Mockford, 1999: 350; Lienhard & Smithers, 2002: 157 *DR >
TL: Constanza to Valle Nuevo, 3 to 7,000 ft.
Valenzuela dificilis (Mockford, 1969: 96). Lienhard & Smithers, 2002: 161
Valenzuela elegans (Mockford, 1969: 98); Mockford, 1999: 350; Lienhard & Smithers, 2002: 162 *H > TL: Kenscoff
Valenzuela nebulosus (Mockford, 1969: 79). Mockford, 1999: 350; Lienhard & Smithers, 2002: 174 *DR > TL: Santo
Domingo
Valenzuela subelegans Mockford, 1969: 104; Mockford, 1999: 350; Lienhard & Smithers, 2002: 183 *H > TL: Kenscoff
Xanthocaecilius Mockford, 1989
Xanthocaecilius sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]

Infraorder HOMILOPSOCIDEA
Family ARCHIPSOCIDAE
Archipsocus Hagen, 1882
Archipsocus sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]

Family ELIPSOCIDAE
Elipsocus Hagen, 1866
Elipsocus modestus Banks, 1938: 286. Lienhard & Smithers, 2002: 343 *H > TL: La Visite, La Selle Range, 5–7000 ft.

Family LACHESILLIDAE
Lachesilla Westwood, 1840
Lachesilla acuminiforceps García Aldrete, 1996: 107. Lienhard & Smithers, 2002: 226
Lachesilla aethiopica (Enderlein, 1902). Mockford, 1974: 142; García Aldrete, 1996: 111; Lienhard & Smithers, 2002: 227
Lachesilla dilatiforceps García Aldrete, 1996: 109. Lienhard & Smithers, 2002: 231
Lachesilla rena Sommerman, 1946. García Aldrete, 1996: 114; Lienhard & Smithers, 2002: 241
Lachesilla riegeli Sommerman, 1946. García Aldrete, 1996: 115; Lienhard & Smithers, 2002: 241
Lachesilla sandersoni Mockford, 1974. García Aldrete, 1996: 111; Lienhard & Smithers, 2002: 242
Lachesilla tectorum Badonnel, 1931. García Aldrete, 1996: 114; Lienhard & Smithers, 2002: 243
Nanolachesilla Mockford & Sullivan, 1986
Nanolachesilla haitiana Mockford & Sullivan, 1986: 62. Lienhard & Smithers, 2002: 224 *H > TL: Kenscoff, La Découverte

Family PHILOTARSIDAE
Aaroniella Mockford, 1952
Aaroniella achrysa (Banks, 1941a: 391). Mockford, 1974: 133; Lienhard & Smithers, 2002: 331
Aaroniella sp.†—Mockford (1986: 112) [As Broadheadia] [Fossil in Dominican amber]
Garcialdretia Mockford, 2007
Garcialdretia dardana (Banks, 1941a: 389). Lienhard & Smithers, 2002: 328; Mockford, 2007: 134 *DR > TL: Constanza to
Valle Nuevo, 7,000 ft.

Infraorder PSOCETAE
Family HEMIPSOCIDAE
Hemipsocus Selys, 1872
Hemipsocus sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]

Family MYOPSOCIDAE
Lophopterygella Enderlein, 1907

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Lophopterygella haitiensis Banks, 1938: 286. Lienhard & Smithers, 2002: 480 *H > TL: Desbarriere, Mt. La Hotte, 4000 ft.
Myopsocus Hagen, 1866
Myopsocus antillanus (Mockford, 1974: 157). Lienhard & Smithers, 2002: 482
Myopsocus arthuri† Nel et al., 2006: 64 [Fossil in Dominican amber]
Myopsocus minutus (Mockford, 1974: 155). Lienhard & Smithers, 2002: 488
Myopsocus taurus Banks, 1941a: 388. Lienhard & Smithers, 2002: 491 *DR > TL: Loma Vieja, south of Constanza, 6,000 ft.

Family PSOCIDAE
Blaste Kolbe, 1880
Blaste sp.†—Mockford (1986: 112) [Fossil in Dominican amber]
Blastopsocus Roesler, 1943
Blastopsocus sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]
Indiopsocus Mockford, 1974
Indiopsocus sp.†—Poinar & Poinar (1999: 202) [Fossil in Dominican amber]
Mecampsis Enderlein, 1925
Mecampsis insolita (Banks 1938: 285). Lienhard & Smithers, 2002: 439 *H > TL: La Visite, La Selle Range, 5–7000 ft.
Mecampsis laconia (Banks 1941: 387). Lienhard & Smithers, 2002: 439 *DR > TL: Valle Nuevo, southeast of Constanza,
7,000 ft.
Psocus Fabricius, 1798
Psocus cyllarus Banks 1941: 386. Lienhard & Smithers, 2002: 425 *DR > TL: Loma Rucilla, 5 to 10,000 ft.
Psocus oneitus Banks 1941: 387. Lienhard & Smithers, 2002: 425 *DR > TL: Valle Nuevo, south of Constanza, 7,000 ft.
Ptycta Enderlein, 1925
Ptycta sp.†—Mockford (1986: 112) [Fossil in Dominican amber]
Trichadenotecnum Enderlein, 1909
Trichadenotecnum sp.†—Mockford (1986: 112) [Fossil in Dominican amber]

Order SIPHONAPTERA
Family PULICIDAE
Atopopsyllus† Poinar, 2015
Atopopsyllus cionus† Poinar, 2015: 1235 [Fossil in Dominican amber]
=Rhopalopsyllus sp.†—Poinar (1995: 789) [Fossil in Dominican amber]
Echidnophaga Olliff, 1886
Echidnophaga gallinacea (Westwood, 1875). Wolcott, 1927: 335
Eospilopsyllus Beaucournu & Perrichot, 2012
Eospilopsyllus kobberti† Beaucournu & Perrichot in Perrichot, Beaucournu & Velten, 2012: 57 [Fossil in Dominican amber]
Pulex Linnaeus, 1758
Pulex (Pulex) irritans Linnaeus, 1758. Wolcott, 1927: 333; Santoro, 1960: 452
Pulex (Pulex) larimerius† Lewis & Grimaldi, 1997: 4 [Fossil in Dominican amber]

Family TUNGIDAE
Tunga Jarocki, 1838
Tunga penetrans (Linnaeus, 1758). Wolcott, 1927: 336

Order STREPSIPTERA

Taxonomic summary:
Suborder Stylopidia
Family Bohartillidae
Family Corioxenidae
Family Elenchidae
Family Halictophagidae
Family Myrmecolacidae

Suborder STYLOPIDIA

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Family BOHARTILLIDAE
Bohartilla Kinzelbach, 1969
Bohartilla joachimscheveni† Kinzelbach & Pohl, 1994: 65. Pohl & Kinzelbach, 1995: 203 [Fossil in Dominican amber]
Bohartilla kinzelbachi† Kathiritamby & Grimaldi, 1993: 33 [Fossil in Dominican amber]
Bohartilla megalognatha Kinzelbach, 1969. Cook, 2015: 214
Bohartilla megalognatha† Kinzelbach, 1969. Kathiritamby & Grimaldi, 1993: 32. Pohl & Kinzelbach, 1995: 202 [Fossil in
Dominican amber]

Family CORIOXENIDAE
Corioxenos Blair, 1936
Corioxenos antestiae Blair, 1936. Cook, 2001: 397
Pseudoxenos Saunders, 1872
Pseudoxenos ashmeadi (Pierce, 1909: 146). Blackwelder, 1945: 489
Pseudoxenos fasciatai (Pierce, 1909: 142). Blackwelder, 1945: 489

Family ELENCHIDAE
Protelencholax Kinzelbach, 1979
Protelencholax schleei† Kinzelbach, 1979: 6. Pohl & Kinzelbach, 1994: 65 [Fossil in Dominican amber]

Family HALICTOPHAGIDAE
Halictophagus Curtis, 1831
Halictophagus dominicus Cook, 2013: 574 *DR > TL: Parque del Este, 2.9 km SW Boca de Yuma
Halictophagus trigonodontos Cook, 2013: 570 *DR > TL: Parque del Este, Caseta de Guaraguao

Family MYRMECOLACIDAE
Caenocholax Pierce, 1909
Caenocholax brodzynskyi† Kathiritamby & Grimaldi, 1993: 36 [Fossil in Dominican amber]
Caenocholax dominicensis† Kathiritamby & Grimaldi, 1993: 36 [Fossil in Dominican amber]
Caenocholax sp.†—Poinar & Poinar (1999: 145) [Fossil in Dominican amber]
Myrmecholax Westwood, 1861
Myrmecholax glaesi† Kinzelbach, 1983: 31 [Fossil in Dominican amber]
Stichotrema Hofeneder, 1910
Stichotrema beckeri† Kinzelbach, 1983: 33. Pohl & Kinzelbach, 1994: 63 [Fossil in Dominican amber]
Stichotrema dominicanum† Kinzelbach & Pohl, 1994: 64 [Fossil in Dominican amber]
Stichotrema kathirithambyi Cook, 2014: 67 *DR > TL: 6 km NW Paraiso, Río Nizao, Barahona
Note: Palaeomyrmecolax neotropicallis† originally described by Kogan & Poinar, 2010: 228 as Stylops neotropicallis was later
shown by Kogan et al., 2014 to be from Baltic rather than Dominican amber.

Order THYSANOPTERA

Taxonomic summary:
Suborder Terebrantia
Family Aeolothripidae
Family Thripidae
Suborder Tubulifera
Family Phlaeothripidae

Suborder TEREBRANTIA
Family AEOLOTHRIPIDAE
Franklinothrips Back, 1912
Franklinothrips sp.—Russo (1936: 42)
Franklinothrips vespiformis (Crawford, 1909). Russo, 1936: 42; Sakimura, 1986: 350; Hoddle et al., 2002: 259

Family THRIPIDAE

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Subfamily DENDROTHRIPINAE
Leucothrips Reuter, 1904
Leucothrips piercei (Morgan, 1913). Hoddle et al., 2002: 259
Subfamily PANCHAETOTHRIPINAE
Anisopilothrips Stannard & Mitri, 1962
Anisopilothrips venustulus (Priesner, 1923). Sakimura, 1986: 350; Mound & Marullo, 1996: 79
Caliothrips Daniel, 1904
Caliothrips insularis (Hood, 1928). Nakahara, 1991: 99; Mound & Marullo, 1996: 81
Dinurothrips Hood, 1913
Dinurothrips hookeri Hood, 1913. Russo, 1936: 44
Heliothrips Haliday, 1836
Heliothrips haemorrhoidalis (Bouché, 1833). Russo, 1936: 42; Zuluaga & Mesa, 2000: 126; Hoddle et al., 2002: 259
Retithrips Marchal, 1910
Retithrips syriacus (Mayet, 1890). Abud, 1997: 10/97
Selenothrips Karny, 1911
Selenothrips rubrocinctus (Giard, 1901). Russo, 1936: 42; Schmutterer, 1990: 159; Hoddle et al., 2002: 259
Subfamily SERICOTHRIPINAE
Neohydatothrips John, 1929
Neohydatothrips burungae (Hood, 1935). SEA, 1999: 57
Subfamily THRIPINAE
Bolacothrips Uzel, 1895
Bolacothrips striatopennatus (Schmutz, 1913). Medrano, 2014: 98 [Introduced]
Chaetanaphothrips Priesner, 1926
Chaetanaphothrips leeuweni (Karny, 1914). Hoddle et al., 2002: 259
Chaetanaphothrips orchidii (Moulton, 1907). Abud, 1993: 17/93
Chirothrips Haliday, 1936
Chirothrips mexicanus Crawford, 1909. Sakimura, 1986: 351
“Chirothrips” spinosus (Moulton, 1946: 56). Jacot-Guillarmod, 1971: 485
Corynothrips Williams, 1913
Corynothrips stenopterus Williams, 1913. Abud, 1990: 2/90
Dendrothripoides Bagnall, 1923
Dendrothripoides innoxius (Karny, 1914). Sakimura, 1986: 351
Frankliniella Karny, 1910
Frankliniella bruneri Karny, 1910. Hoddle et al., 2002: 259
Frankliniella cephalica (Crawford, 1910). Abud, 1993: 6/93
Frankliniella cubensis Hood, 1925. Sakimura, 1986: 354; Nakahara, 1997: 363; Hoddle et al., 2002: 259
Frankliniella insularis (Franklin, 1908). Russo, 1936: 42; Hoddle et al., 2002: 259
Frankliniella ipomoeae Moulton, 1946: 95 *H > TL: Damien
Frankliniella kelliae Sakimura, 1981: 488. Sakimura, 1986: 355; Nakahara, 1997: 368
Frankliniella musaeperda Hood, 1952: 137. Nakahara, 1997: 369
Frankliniella occidentalis (Pergande, 1895). Nakahara, 1997: 369; Zuluaga & Mesa, 2000: 197 [Introduced]
Frankliniella parvula Hood, 1925. Harrison, 1963: 64; Sakimura, 1986: 355; Mound & Marullo, 1996: 148; Nakahara, 1997:
372
Frankliniella schultzei Trybom, 1910. Sakimura, 1986: 355; Nakahara, 1997: 373
Frankliniella tenuicornis (Uzel, 1895). Schmutterer, 1990: 410
Frankliniella williamsi Hood, 1915. Nakahara, 1997: 377
Microcephalothrips Bagnall, 1926
Microcephalothrips abdominalis (Crawford, 1910). Medrano, 2014: 98 [Introduced]
Plesiothrips Hood, 1915
Plesiothrips perplexus (Beach, 1896). Sakimura, 1986: 356
Rhamphothrips Karny, 1913
Rhamphothrips pandens Sakimura, 1983. Mound & Marullo, 1996: 183
Scirtothrips Shull, 1909
Scirtothrips dorsalis Hood, 1919. Medrano, 2014: 98 [Introduced]

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Sericothrips Haliday, 1836
Sericothrips sp.—Russo (1936: 44)
Thrips Linnaeus, 1758
Thrips florum Schmutz, 1913. Nakahara, 1994: 55
Thrips palmi Karny, 1925. Schmutterer, 1990: 433; Nakahara, 1994: 95; Zuluaga & Mesa, 2000: 197 [Introduced]
Thrips simplex (Morison, 1930) [By synonymy of species below]
=Taeniothrips gladioli Moulton & Steinweden, 1931. SEA, 1999: 63
Thrips tabaci (Lindemann, 1889). Wolcott, 1927: 212; Russo, 1936: 44

Suborder TUBULIFERA
Family PHLAEOTHRIPIDAE
Subfamily IDOLOTHRIPINAE
Allothrips Hood, 1908
Allothrips watsoni watsoni Hood, 1939. Stannard, 1957: 53
Bolothrips Priesner, 1926
Bolothrips sp.—Russo (1936: 45)
Ethirothrips Karny, 1925
Ethirothrips brevis (Bagnall, 1921). Mound & Palmer, 1992: 329
Neosmerinthothrips Schmutz, 1913
Neosmerinthothrips collaris (Bagnall, 1917). Mound & Marullo, 1996: 228
=Gastrothrips dominicanus Hood, 1935: 170. Mound & Marullo, 1996: 228
Neosmerinthothrips sp.—Sakimura (1986: 360)
Nesothrips Kirkaldy, 1907
Nesothrips lativentris (Karny, 1913). Sakimura, 1986: 360; Sakimura, 1972: 668
Subfamily PHLAEOTHRIPINAE
Aleurodothrips Franklin, 1909
Aleurodothrips fasciapennis (Franklin, 1908). Sakimura, 1986: 357
Gynaikothrips Zimmermann, 1900
Gynaikothrips ficorum (Marchal, 1908). Wolcott, 1927: 210 [Introduced]
Gynaikothrips uzeli (Zimmerman, 1900). SEA, 1999: 78 [Introduced]
Haplothrips Amyot & Serville, 1843
Haplothrips gowdeyi (Franklin, 1908). Russo, 1936: 44; Hoddle et al., 2002: 259
Holopothrips Hood, 1914
Holopothrips seini (Watson, 1927: 59). Stannard, 1957: 53
Holopothrips tabebuia Cabrera & Segarra, 2008: 232
Holopothrips sp.—Medina et al. (1994: 186)
Karnyothrips Watson, 1923
Karnyothrips flavipes (Jones, 1912). Sakimura, 1986: 358
Karnyothrips melaleucus (Bagnall, 1911). Sakimura, 1986: 359
Trachythrips Hood, 1930
Trachythrips brevitubus (Stannard, 1957: 34) *H > TL: Fond Verrettes at Morne Fen
Trybomia Karny, 1911
Trybomia sp.—Russo (1936: 44)

Order TRICHOPTERA
Key reference: Flint and Sykora (2004).

Taxonomic summary:
Suborder Annulipalpia
Superfamily Philopotamoidea
Family Philopotamidae
Superfamily Psychomyioidea
Family Ecnomidae
Family Polycentropodidae

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 Family Xiphocentronidae
Superfamily Hydropsychoidea
Family Hydropsychidae
Suborder Integripalpia
Superfamily Glossosomatoidea
Family Glossosomatidae
Superfamily Hydroptiloidea
Family Hydroptilidae
Superfamily Rhyacophiloidea
Family Hydrobiosidae
Infraorder Brevitentoria
Superfamily Leptoceroidea
Family Calamoceratidae
Family Leptoceridae
Family Odontoceridae
Superfamily Sericomastoidea
Family Helicopsychidae

Suborder ANNULIPALPIA
Superfamily PHILOPOTAMOIDEA
Family PHILOPOTAMIDAE
Subfamily CHIMARRINAE
Chimarra Stephens, 1829
Chimarra (Chimarrita) merengue Blahnik, 1997: 210. Flint & Sikora, 2004: 51 *DR > TL: Dajabon, 1.3 km S Loma de
Cabrera, 400 m
Chimarra (Chimarrita) palaenova† Wichard, 2007: 17 [Fossil in Dominican amber]
Chimarra (Curgia) braconoides (Walker, 1860: 179). Flint & Sikora, 2004: 50 *DR, *H > TL: Described without locality data
Chimarra (Curgia) gilvimacula Flint, 1998: 52. Flint & Sikora, 2004: 50 *DR > TL: La Vega Prov., Río Camú, 19 km NE
Jarabacoa
Chimarra dommeli† Wichard, 1983: 142 [Fossil in Dominican amber]
Chimarra (Otarrha) koki Botosaneanu, 1996: 11. Flint & Sikora, 2004: 51 *DR > TL: Arroyo Manuel Estrella, near main
entrance to Parque Nacional Armando Bermúdez, 1,200 m
Chimarra (Otarrha) redonda Blahnik, 2002: 98. Flint & Sikora, 2004: 51 *DR > TL: La Palma, 12 km E of El Río
Chimarra (Otarrha) spinulifera baoruco Flint & Sikora, 2004: 51 *DR > TL: Barahona Prov., San Rafael, 30 m
Chimarra (Otarrha) spinulifera galalcha Botosaneanu, 1996: 11. Flint & Sikora, 2004: 51 *DR > TL: Arroyo El Dulce,
sección Manabao-Los Dajaos of Jarabacoa
Chimarra (Otarrha) spinulifera spinulifera Flint, 1968: 152. Flint & Sikora, 2004: 51 *H > TL: Roche Croix, Mt. La Hotte,
4000 ft.
Chimarra palaeodominicana† Wichard, 1983: 142 [Fossil in Dominican amber]
Chimarra resinae† Wichard, 1983: 142 [Fossil in Dominican amber]
Chimarra succini† Wichard, 1983: 4 [Fossil in Dominican amber]
Chimarra weitschati† Wichard, 1983: 139 [Fossil in Dominican amber]

Superfamily PSYCHOMYIOIDEA
Family ECNOMIDAE
Austrotinodes Schmid, 1955
Austrotinodes labiatus Flint & Sykora, 2004: 4 *DR > TL: Pedernales prov., Río Mulito, 13 km N Pedernales, 230 m

Family POLYCENTROPODIDAE
Subfamily POLYCENTROPODINAE
Antillopsyche Banks, 1941
Antillopsyche auricula† Wichard, 2007: 27 [Fossil in Dominican amber]
Antillopsyche demma Botosaneanu, 1996: 13. Flint & Sikora, 2004: 52 *DR > TL: Pedernales Prov., Río Mulito, at 2 km from
Mencia

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Antillopsyche digitus† Wichard, 2007: 30 [Fossil in Dominican amber]
Antillopsyche oliveri† Wichard, 1985: 118 [Fossil in Dominican amber]
Cernotina Ross, 1938
Cernotina danieli Flint & Sykora, 2004: 52 *DR > TL: Pedernales Prov., Río Mulito, 13 km N Pedernales, 230 m
Cernotina pulchra† Wichard, 2007: 32 [Fossil in Dominican amber]
=Cernotina sp.†—Poinar & Poinar (1999: 206) [Fossil in Dominican amber]
Polycentropus Curtis, 1835
Polycentropus domingensis Banks, 1941a: 399. Flint & Sikora, 2004: 54 *DR > TL: Loma Rucilla, 5 to 10,000 ft.
Polycentropus jeldesi Flint, 1976: 237. Flint & Sikora, 2004: 54 *DR > TL: Convento, 12 km S of Constanza
Polycentropus marcanoi Flint, 1976: 238. Flint & Sikora, 2004: 54 *DR > TL: Dajabon, Prov., 13 km S of Loma de Cabrera,
400 m
Polycentropus pedernales Flint & Sykora, 2004: 54 *DR > TL: Pedernales Prov., Río Mulito, 13 km N Pedernales, 230 m
Polycentropus vanderpooli Flint, 1976: 237. Flint & Sikora, 2004: 55 *DR > TL: Convento, 12 km S of Constanza

Family XIPHOCENTRONIDAE
Xiphocentron Brauer, 1870
Xiphocentron (Antillotrichia) haitiense (Banks, 1941a: 402). Flint & Sikora, 2004: 55
Xiphocentron sp.—Flint & Sykora (2004: 56) *DR > TL: Monseñor Nouel Prov., near Jima, 670 m
Note: The species Limnephilus toussainti Banks, 1924 was described based on a male holotype labelled as being from Haiti.
This is apparently an error. Flint (1967: 16) reported specimens of this species from northwestern Mexico.

Superfamily HYDROPSYCHOIDEA
Family HYDROPSYCHIDAE
Calosopsyche Ross & Unzicker, 1977
Calosopsyche batesi (Flint, 1962: 25). Ross & Unzicker, 1977: 309; Flint & Sikora, 2004: 18 *H > TL: La Visite and vicinity,
La Selle Range, 5–7000 ft.
Calosopsyche bohio (Botosaneanu, 1991a: 310). Flint & Sikora, 2004: 18 *H > TL: Département de l’Ouest, Ville Bonheur, Le
Saut d’Eau
Calosopsyche carinifera (Flint, 1962: 27). Flint & Sikora, 2004: 18 *DR > TL: foothills Cordillera Central, South of Santiago
Calosopsyche domingensis (Banks, 1941a: 398). Flint & Sikora, 2004: 19 *DR > TL: Constanza to Jarabacoa, 2 to 4,000 ft.
Calosopsyche palaeoelegans† Wichard, 2007: 20 [Fossil in Dominican amber]
Leptonema Guérin-Méneville, 1844
Leptonema sp.†—Wichard (1987: 68) [Fossil in Dominican amber]
Macronema Stephens, 1829
Macronema sp.—Flint & Sykora (2004: 19) *H > TL: Département du Sud, Rivière du Cavaillon, Saut Mathurine
Macronema hispaniola† Wichard, 2007: 22 [Fossil in Dominican amber]
Palaehydropsyche† Wichard, 1986
Palaehydropsyche fossilis† Wichard, 1986: 192 [Fossil in Dominican amber]
Smicridea MacLachlan, 1871
Smicridea (Smicridea) banksi Flint, 1967: 13. Botosaneanu, 1996: 15; Flint & Sikora, 2004: 24 *H, *DR > TL: La Visite &
vic., La Selle Range, 5–7000 ft.
=Smicridea unicolor Banks, 1938: 303; Flint & Sikora, 2004: 24
Smicridea (Smicridea) brunnescens Flint & Sykora, 2004: 22 *DR > TL: La Vega Prov., Río Jimenoa, Jarabacoa
Smicridea (Smicridea) comma Banks, 1924. Botosaneanu, 1991a: 133; Flint & Sikora, 2004: 23
Smicridea (Smicridea) completa Banks, 1941a: 398 *DR > TL: Villa Altagracia, 3 to 4,000 ft.
Smicridea (Smicridea) duarte Flint & Sykora, 2004: 24 *DR > TL: La Vega Prov., Río Baiguate, 1–2 km S Jarabacoa, 520 m
Streptopsyche Ross & Unzicker, 1977
Streptopsyche antilles (Ross & Palmer, 1946: 184). Flint & Sikora, 2004: 20 *DR > TL: City of Santo Domingo
Streptopsyche davisorum Ross & Unzicker, 1977: 308. Flint & Sikora, 2004: 20 *DR > TL: La Estrelleta Prov., 4 km SE Río
Limpio, ca. 760 m
Streptopsyche parander (Botosaneanu, 1996: 19). Flint & Sikora, 2004: 20 *DR > TL: Pedernales Prov., Río Mulito, at 2 km
from Mencia
Streptopsyche praecipua Flint & Sykora, 2004: 22 *H > TL: Département de l’Ouest, Manneville, 60 ft.
Streptopsyche rawlinsi Flint & Sykora, 2004: 20 *H > TL: Département du Sud, S slope Morne Formon, Ville Formon, 1405 m

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 387
Suborder INTEGRIPALPIA
Superfamily GLOSSOSOMATOIDEA
Family GLOSSOSOMATIDAE
Campsiophora Flint, 1964
Campsiophora pedophila Flint, 1964. Botosaneanu, 1991: 116; Flint & Perez-Gelabert, 1999: 36; Flint & Sikora, 2004: 5
Cariboptila Flint, 1964
Cariboptila aurulenta Flint, 1974: 8. Flint & Perez-Gelabert, 1999: 36; Flint & Sikora, 2004: 5 *DR > TL: Convento, 12 km S
of Constanza
Cariboptila caab Botosaneanu, 1996: 8. Flint & Perez-Gelabert, 1999: 36; Flint & Sikora, 2004: 5 *DR > TL: cachones of La
Descubierta
Cariboptila calcigena Flint, 1974: 8. Flint & Perez-Gelabert, 1999: 36; Flint & Sikora, 2004: 6 *DR > TL: La Palma, 12 km E
of El Río
Cariboptila hispaniolica Flint, 1974: 8. Flint & Perez-Gelabert, 1999: 36; Flint & Sikora, 2004: 6 *DR > TL: La Palma, 12 km
E of El Río
Cariboptila mathisi Flint & Sykora, 2004: 6 *DR > TL: Monseñor Nouel prov., 8.7 km W Bonao, 890 m
Cariboptila paradoxa Flint & Sykora, 2004: 7 *DR > TL: Independencia Prov., La Descubierta, 0 m
Cubanoptila Sykora, 1973
Cubanoptila grimaldii† Wichard, 1995: 160 [Fossil in Dominican amber]
Cubanoptila longiscapa† Wichard, 2007: 7 [Fossil in Dominican amber]
Cubanoptila mederi† Wichard, 1989: 93 [Fossil in Dominican amber]
Cubanoptila poinari† Wichard, 1989: 92 [Fossil in Dominican amber]

Superfamily HYDROPTILOIDEA
Family HYDROPTILIDAE
Alisotrichia Flint, 1964
Alisotrichia aglae Botosaneanu, 1991a: 118. Flint & Sikora, 2004: 26 *H > TL: Département de l’Ouest, Ville Bonheur, Le
Saut d’Eau
Alisotrichia aquaecadentis Botosaneanu, 1991a: 116. Flint & Sikora, 2004: 27 *H > TL: Département du Sud: Saut Mathurine
Alisotrichia arcana Botosaneanu, 1991a: 124. Flint & Sikora, 2004: 27 *H > TL: Département du Sud, près de Camp Perrin:
Resurgence du Moreau
Alisotrichia arizela† Wells & Wichard, 1989: 43 [Fossil in Dominican amber]
Alisotrichia bisetosa Flint & Sykora, 2004: 27 *DR > TL: Independencia Prov., Río Guayabal, 4.5 km N Postrer Río, 150 m
Alisotrichia euphrosyne Botosaneanu, 1991a: 118. Flint & Sikora, 2004: 27 *H > TL: Département de l’Ouest, Ville Bonheur,
Le Saut d’Eau
Alisotrichia hirudopsis aitija Botosaneanu, 1995: 22. Flint & Sikora, 2004: 27 *DR > TL: Arroyo Los Guineos, a tributary of
Río Nagua, from southern slopes of Loma Quita Espuela, 930 m
Alisotrichia hispaniolina Botosaneanu, 1991a: 116. Flint & Sikora, 2004: 27 *H > TL: Département de l’Ouest, Rivière Tombe
à Mirebalais
Alisotrichia thalia Botosaneanu, 1991a: 120. Flint & Sikora, 2004: 29 *H > TL: Département de l’Ouest, Ville Bonheur, Le
Saut d’Eau
Alisotrichia ultima Flint & Sykora, 2004: 29 *DR > TL: Azua Prov., Río Las Cuevas, 8 km NE Padre Las Casas, 580 m
Alisotrichia woodruffi Flint & Sykora, 2004: 29 *DR > TL: Monseñor Nouel Prov., 68 km NW Rt. 1 on road to Constanza
Hydroptila Dalman, 1819
Hydroptila ditalea Flint, 1968. Botosaneanu, 1995: 27; Flint & Sikora, 2004: 31
Hydroptila dominicana Botosaneanu, 1995: 27. Flint & Sikora, 2004: 31 *DR > TL: Arroyo Los Dajaos, sección Manabao-Los
Dajaos of Jarabacoa, 1000–1100 m
Hydroptila medinai Flint, 1964. Botosaneanu, 1991a: 130; Flint & Sikora, 2004: 31
Leucotrichia Mosely, 1934
Leucotrichia adela† Wells & Wichard, 1989: 42 [Fossil in Dominican amber]
Leucotrichia gomezi Flint, 1970: 7. Flint & Sikora, 2004: 32 *DR > TL: La Palma, 12 km E of El Río
Leucotrichia tubifex Flint, 1964. Botosaneanu, 1991: 116; Flint & Sikora, 2004: 32
Metrichia Ross, 1938
Metrichia cafetalera Botosaneanu, 1980: 110. Flint & Sikora, 2004: 32

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Metrichia fontismoreaui (Botosaneanu, 1991a: 125). Flint & Sikora, 2004: 32 *H > TL: Département du Sud, près de Camp
Perrin: Resurgence du Moreau
Metrichia kumanskii (Botosaneanu, 1991a: 128). Flint & Sikora, 2004: 33 *H > TL: Département de l’Ouest, Ville Bonheur, Le
Saut d’Eau
Metrichia longispina Flint & Sykora, 2004: 33 *DR > TL: La Vega Prov., Convento, 12 km S Constanza
Metrichia sp.—Flint & Sykora (2004: 33) *H > TL: Département de l’Ouest, Parc National La Visite, Pic La Visite, 2100 m
Metrichia squamigera Flint, 1992. Flint & Sikora, 2004: 33
Neotrichia Morton, 1905
Neotrichia bifurcata Harris in Flint & Sykora, 2004: 34 *DR > TL: Pedernales Prov., Río Mulito, 13 km N Pedernales, 230 m
Neotrichia iridescens Flint, 1964. Botosaneanu, 1991a: 128; Flint & Sikora, 2004: 34
Neotrichia pequenita Botosaneanu, 1977. Botosaneanu, 1991a: 128; Flint & Sikora, 2004: 36
Neotrichia sp.—Botosaneanu (1991a: 130)
Ochrotrichia Mosely, 1934
Ochrotrichia aldama† (Mosely, 1937). Wells & Wichard, 1989: 46 [[Fossil in Dominican amber specimen, presumably an
extant species]
Ochrotrichia aliciae† Wichard, 2000: 242 [Fossil in Dominican amber]
Ochrotrichia baorucoensis Flint & Sykora, 2004: 36 *DR > TL: Barahona Prov., San Rafael, 30 m
Ochrotrichia brodzinskyi† Wells & Wichard, 1989: 45 [Fossil in Dominican amber]
Ochrotrichia cachonera Botosaneanu, 1995: 23. Flint & Sikora, 2004: 36 *DR > TL: Arroyo Los Verros, a tributary of Río
Comatillo, S from Parque Los Haitises, 200 m
Ochrotrichia chaulioda† Wells & Wichard, 1989: 46 [Fossil in Dominican amber]
Ochrotrichia denaia† Wells & Wichard, 1989: 47 [Fossil in Dominican amber]
Ochrotrichia doehleri† Wichard, 1981: 161. Wells & Wichard, 1989: 48 [Fossil in Dominican amber]
Ochrotrichia ingloria Botosaneanu, 1995: 25. Flint & Sikora, 2004: 37 *DR > TL: Arroyo Los Verros, a tributary of Río
Comatillo, S from Parque Los Haitises, 200 m
Ochrotrichia larimar Flint & Sykora, 2004: 38 *DR > TL: Barahona Prov., Larimar mine, nr. Filipinas
Ochrotrichia obovata Flint & Sykora, 2004: 38 *DR > TL: La Vega Prov., 20 km S Constanza
Ochrotrichia seiba Flint & Sykora, 2004: 38 *DR > TL: El Seibo Prov., Pedro Sánchez, small stream
Ochrotrichia serra Botosaneanu, 1991a: 125. Flint & Sikora, 2004: 39 *H > TL: Département de l’Ouest, Ville Bonheur, Le
Saut d’Eau
Ochrotrichia sp. A—Flint & Sykora (2004: 40) *DR > TL: Barahona Prov., Larimar mine, nr. Filipinas
Ochrotrichia sp. B—Flint & Sykora (2004: 40) *DR > TL: Barahona Prov., Larimar mine, nr. Filipinas
Ochrotrichia verda Flint, 1968. Flint & Sikora, 2004: 40
Orthotrichia Eaton, 1873
Orthotrichia aegerfasciella (Chambers, 1873). Botosaneanu, 1991a: 130; Flint & Sikora, 2004: 40
Orthotrichia cristata Morton, 1905. Flint & Sikora, 2004: 40
Oxyethira Eaton, 1873
Oxyethira (Dampfitrichia) cirrifera Flint, 1964. Botosaneanu, 1991a: 130; Flint & Sikora, 2004: 43
Oxyethira (Dampfitrichia) mirebalina Botosaneanu, 1991a: 130. Flint & Sikora, 2004: 41 *H > TL: Département de l’Ouest,
Grande rivière de l’Artibonite à Mirebalais
Oxyethira (Dampfitrichia) simulatrix Flint, 1968. Flint & Sikora, 2004: 44
Oxyethira (Dampfitrichia) tega Flint, 1968. Flint & Sikora, 2004: 45
Oxyethira (Loxotrichia) janella Denning, 1948. Botosaneanu, 1991a: 130; Flint & Sikora, 2004: 43
Oxyethira (Loxotrichia) puertoricensis Flint, 1964. Botosaneanu, 1991a: 130; Flint & Sikora, 2004: 44
Oxyethira (Mesotrichia) albaeaquae Botosaneanu, 1995: 30. Flint & Sikora, 2004: 41 *DR > TL: Arroyo Los Dajaos, sección
Manabao-Los Dajaos of Jarabacoa, 1000–1100 m
Oxyethira (Mesotrichia) geminata Flint & Sykora, 2004: 41 *DR > TL: La Vega Prov., 11.5 km S of Constanza, 1,410 m
Oxyethira (Mesotrichia) ortizorum Botosaneanu, 1995: 29. Flint & Sikora, 2004: 41 *DR > TL: Arroyo El Dulce, sección
Manabao-Los Dajaos of Jarabacoa
Oxyethira (Mesotrichia) scopulina Flint & Sykora, 2004: 43 *DR > TL: Peravia Prov., 3 km SW La Nuez, upper Río Las
Cuevas, 1850 m
Oxyethira (Mesotrichia) sp.—Flint & Sykora (2004: 45) *DR > TL: La Vega Prov., spring brook near Salto Aguas Blancas

Superfamily RHYACOPHILOIDEA

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 389
Family HYDROBIOSIDAE
Atopsyche Banks, 1905
Atopsyche batesi Banks, 1938: 304. Flint & Sikora, 2004: 14 *H > TL: La Visite, La Selle Range
Atopsyche conventica Flint, 1974: 4. Flint & Sikora, 2004: 14 *DR > TL: Convento, 12 km S of Constanza
Atopsyche davisorum Flint, 1974: 4. Flint & Sikora, 2004: 14 *DR > TL: 4 km SE of Río Limpio, 760 m
Atopsyche hinnulus Flint & Sykora, 2004: 14 *DR > TL: Pedernales Prov., Río Mulito, 21 km N Pedernales, 280 m
Atopsyche lilicae Botosaneanu, 1991a: 114. Flint & Sikora, 2004: 15 *H > TL: Départament de l’Ouest, Ville Bonheur (Ville
Saut d’Eau)
Atopsyche orientalis Flint & Sykora, 2004: *DR > TL: El Seibo Prov., 15 km S Miches, ca. 500 m
Atopsyche peravia Flint & Sykora, 2004: 16 *DR > TL: Peravia Prov., 3 km SW La Nuez, upper Río Las Cuevas, 1880 m
Atopsyche sp.—Flint & Sykora (2004: 18) *H, *DR > TL: Département du Sud, Ville Formon, s slope Morne Formon, Massif
de la Hotte, 1405 m
Atopsyche perlucida† Wichard, 2007: 4 [Fossil in Dominican amber]
=Atopsyche sp.†—Wichard (1987: 68); Poinar & Poinar (1999: 206) [Fossil in Dominican amber]
Atopsyche taina Flint, 1974: 4. Flint & Sikora, 2004: 16 *DR > TL: Convento, 12 km S of Constanza
Atopsyche thomasi Flint & Sykora, 2004: 16 *H > TL: Département de l’Oueste, saddle between Fe Noir and Enfer, 1700 m

Infraorder BREVITENTORIA
Superfamily LEPTOCEROIDEA
Family CALAMOCERATIDAE
Phylloicus Müller, 1880
Phylloicus iridescens Banks, 1941a: 397. Flint & Perez-Gelabert, 1999: 36; Flint & Sikora, 2004: 3 *DR > TL: Valle Nuevo to
Constanza, 3 to 7,000 ft.
Phylloicus pulchrus Flint, 1964. Flint & Sikora, 2004: 3
Phylloicus velteni† Wichard, 2007: 35 [Fossil in Dominican amber]

Family LEPTOCERIDAE
Nectopsyche Müller, 1879
Nectopsyche cubana (Banks, 1938). Botosaneanu, 1996: 19; Flint & Sikora, 2004: 45
Oecetis MacLachlan, 1877
Oecetis haitises Flint & Sykora, 2004: 46 *DR > TL: Hato Mayor Prov., Parque Los Haitises, 3 km Cueva de Arena 20 m
Oecetis inconspicua (Walker, 1852). Botosaneanu, 1991a: 134; Flint & Sikora, 2004: 46
Setodes Rambur, 1842
Setodes anomalus Flint & Sykora, 2004: 47 *DR > TL: Peravia Prov., 3 km SW La Nuez, tributary to Río Las Cuevas, 1870 m
Setodes aureoinclusa† Wichard, 1995: 170 [Fossil in Dominican amber]
Setodes resinocapta† Wichard, 1995: 168 [Fossil in Dominican amber]

Family ODONTOCERIDAE
Marilia Müller, 1880
Marilia gracilis Banks, 1938: 297. Flint & Sikora, 2004: 49 *H > TL: La Visite & vic., La Selle Range, 5–7000 ft.
Marilia nigrescens Banks, 1941a: 397. Flint & Sikora, 2004: 49 *DR > TL: Valle Nuevo., southeast of Constanza, 7,000 ft.
Marilia valga Flint & Sykora, 2004: 49 *DR > TL: Peravia Prov., 3 km SW La Nuez, tributary to Río Las Cuevas, 1870 m

Superfamily SERICOMASTOIDEA
Family HELICOPSYCHIDAE
Helicopsyche Siebold, 1856
Helicopsyche electra† Johanson & Wichard, 1996: 199 [Fossil in Dominican amber]
Helicopsyche (Feropsyche) altercoma Botosaneanu & Flint, 1991a: 178. Flint & Sikora, 2004: 8 *DR > TL: Elías Piña Prov., 4
kn SE Río Limpio, 760 m
Helicopsyche (Feropsyche) dominicana Botosaneanu & Flint, 1991b: 200. Flint & Sikora, 2004: 8 *DR > TL: La Vega Prov.,
Convento, 12 km S of Constanza
Helicopsyche hageni Banks, 1938. Botosaneanu, 1996: 21; Flint & Perez-Gelabert, 1999: 37; Flint & Sikora, 2004: 8
Helicopsyche (Feropsyche) haitiensis (Banks, 1938: 296). Botosaneanu & Flint, 1991a: 134; Flint & Sikora, 2004: 9 *H > TL:
La Visite & vicinity, La Selle Range, 5–7000 ft.

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Helicopsyche (Feropsyche) kalaom Botosaneanu, 1996: 22. Flint & Sikora, 2004: 9 *DR > TL: Springbrook with aquatic
vegetation, 150 m from Salto Aguas Blancas
Helicopsyche (Feropsyche) lutea (Hagen, 1861: 271). Flint & Sikora, 2004: 9 *DR > TL: St. Domingo
Helicopsyche (Feropsyche) melanochaeta Flint & Sykora, 2004: 11 *DR > TL: Pedernales Prov., Río Mulito, 21 km N Pedernales, 280 m
Helicopsyche (Feropsyche) nigrisensilla (Botosaneanu & Flint, 1991b: 210). Flint & Sikora, 2004: 10 *DR > TL: La Vega
Prov., 20 km S of Constanza
Helicopsyche (Feropsyche) parahageni Flint & Sykora, 2004: 9 *DR > TL: Barahona Prov., San Rafael, 8.3 km S of Baoruco, 30 m
Helicopsyche (Feropsyche) poliochaeta Flint & Sykora, 2004: 11 *DR > TL: Pedernales Prov., Río Mulito, 21 km N Pedernales, 280 m
Helicopsyche (Feropsyche) septifera Flint & Sykora, 2004: 13 *DR > TL: Pedernales Prov., Río Mulito, 21 km N Pedernales, 280 m
Helicopsyche scaloida† Johanson & Wichard, 1996: 197 [Fossil in Dominican amber]
Helicopsyche voigti† Wichard, 1996: 201 [Fossil in Dominican amber]

Order ZORAPTERA
Family ZOROTYPIDAE
Zorotypus Silvestri, 1913
Zorotypus absonus† Engel, 2008: 131 [Fossil in Dominican amber]
Zorotypus barberi Gurney, 1938. Hubbard, 1990: 49
Zorotypus goeleti† Engel & Grimaldi, 2000: 155 [Fossil in Dominican amber]
Zorotypus mnemosyne† Engel, 2008: 129 [Fossil in Dominican amber]
Zorotypus palaeus† Poinar, 1988: 254 [Fossil in Dominican amber]

LITERATURE REFERENCES

Aalbu, R.L., Andrews, F.G. & Pollock, D.A. (2005) Revision of western hemisphere Dacoderinae with a phylogenetic analysis of world taxa
(Coleoptera: Salpingidae). The Coleopterists’ Society Monographs, 4, 1–40.
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Aalbu, R.L., Triplehorn, C.A., Campbell, J.M., Brown, K.W., Somerby, R.E. & Thomas, D.B. (2002) Family 106. Tenebrionidae Latreille
1802. In: Arnett, R.H. Jr., Thomas, M.C., Skelley, P.E. & Frank, J.H. (Eds.), American Beetles. Vol. II. Polyphaga: Scarabaeoidea
through Curculionoidea. CRC Press LLC, Boca Raton, Florida, pp. 463–509.
https://doi.org/10.1201/9781420041231.ch7
Aaron, E.M.B. (1893) New localities for Papilio homerus. The Canadian Entomologist, 25, 258–259.
Abele, L.G. (1992) A review of the grapsid crab genus Sesarma (Crustacea: Decapoda: Grapsidae) in America, with the description of a new
genus. Smithsonian Contributions to Zoology, 527, 1–60.
https://doi.org/10.5479/si.00810282.527
Abo-Shnaf, R.I.A., Sánchez, L. & De Moraes, G.J. (2016) Plant-inhabiting Gamasina mites (Acari: Mesostigmata) from the Dominican
Republic, with descriptions of four new species of Lasioseius (Blattisociidae) and complementary descriptions of other species.
Systematic & Applied Acarology, 21, 607–646.
https://doi.org/10.11158/saa.21.5.5
Abreu C., D. (1996) "...medita y ora pequeño saltamontes". El Nacional, Santo Domingo, 23 de Abril, 7A.
Abreu I., J.S. (1991) Evaluación de Efectividad R de la Ivermectina (Ivomec) en el Tratamiento de Miasis Cutánea en Bovinos. Veterinary
Medicine Thesis, Universidad Autónoma de Santo Domingo, Santo Domingo, unknown pagination. [unpublished]
Abreu P., J., Arias L., L.L. & Mejía S., F. (1990) Determinación del Momento Óptimo de Aplicación del Nim en el Control del Gusano
Cogollero (Spodoptera frugiperda J. E. Smith) en el Cultivo de Maíz. Agronomic Engineering Thesis, Universidad Central del Este, San
Pedro de Macorís, 111 pp. [unpublished]
Abud R., I.M. (1993) Evaluación del Acaricida en el Uso Agrícola en Animales Domésticos. Veterinary Medicine Thesis, Universidad
Autónoma de Santo Domingo, Santo Domingo, unknown pagination. [unpublished]
Abud, A. & Alvarez, P. (1995a) Situación de la mosca blanca Bemisia tabaci Genn. (Homoptera). Workshop on Survey Techniques for Tomato
Yellow Leaf Curly Virus in Dominican Republic, Santo Domingo, Dominican Republic, 1995, unknown pagination.
Abud, A. & Alvarez, P. (1995b) Medidas legales de control de la mosca blanca en la República Dominicana. Workshop on Survey Techniques
for Tomato Yellow Leaf Curly Virus in Dominican Republic, Santo Domingo, Dominican Republic, unknown pagination.
Abud, A.J. & Marcano, E. de J. (1981) Nueva plaga de los pastos. Naturalista Postal, 30/77, 89. [Antonina graminis]
Abud, A.J. & Marcano, E. de J. (1987) Identificación de una nueva cochinilla en el mango. Naturalista Postal, 11/87, no pagination.
[unpublished]
Abud, A.J. & Marcano, E. de J. (1988a) Un nuevo trípido, Thrips palmi en la República Dominicana. Naturalista Postal, 6/88, no pagination.
[unpublished]
Abud, A.J. & Marcano, E. de J. (1990) Insecto de reciente introducción. Naturalista Postal, 4/90, no pagination. [unpublished]
Abud, A.J. & Marcano, E. de J. (1993) Orthezia praelonga Doug., cochinilla acanalada nueva para el país. Naturalista Postal, 2/93, no
pagination. [unpublished]
Abud, A.J. & Torres, D.D. (1983) Técnicas entomológicas para formar una colección de insectos. Facultad de Ciencias Agronómicas y
Veterinarias, UASD, Santo Domingo, unknown pagination.

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 391
Abud, A.J. (1980) Ophion control biológico de Noropsis. Naturalista Postal, 8/80, no pagination. [unpublished]
Abud, A.J. (1981a) Hormiga cortadora de hojas en Barahona. Naturalista Postal, 11/76, 21. [Trachymyrmex jamaicensis]
Abud, A.J. (1981b) Polilla plaga del maní. Naturalista Postal, 13/76, 23. [Stegasta]
Abud, A.J. (1981c) Salta-hoja plaga recien introducida. Naturalista Postal, 16/76, 26.
Abud, A.J. (1981d) Hiede vivo plaga de leguminosas. Naturalista Postal, 19/76, 29. [Acrosternum]
Abud, A.J. (1981e) Nueva plaga de Gelechiidae en batata. Naturalista Postal, 24/76, 34. [Trichotaphe]
Abud, A.J. (1981f) Coreidae en los cultivos de Cucurbitáceas. Naturalista Postal, 19/77, 78. [Anasa]
Abud, A.J. (1981g) Oruga defoliadora del flamboyán. Naturalista Postal, 24/77, 83. [Lyncestis]
Abud, A.J. (1981h) El áfido amarillo de las orquídeas. Naturalista Postal, 32/77, 91. [Sitobeum luteum]
Abud, A.J. (1981i) Nueva plaga del cultivo de frijol. Naturalista Postal, 36/77, 95. [Picturaphis brasiliensis]
Abud, A.J. (1981j) Afidos en el pino. Naturalista Postal, 2/79, 146. [Cinara carolina]
Abud, A.J. (1981k) Plaga de las cucurbitáceas. Naturalista Postal, 5/79, 149. [Diaphania]
Abud, A.J. (1981l) Nueva plaga de Stegobium paniceum (L.) en almacen. Naturalista Postal, 16/81, no pagination. [unpublished]
Abud, A.J. (1981m) Hallazgo de un nuevo Pyralidae. Naturalista Postal, 26/81, no pagination. [unpublished]
Abud, A.J. (1981n) Plaga en cítricos. Naturalista Postal, 38/81, no pagination. [unpublished]
Abud, A.J. (1982) Plagas de insectos en Elaeis guineensis Jacq. Naturalista Postal, 2/82, no pagination. [unpublished]
Abud, A.J. (1983a) Insectos en Elaeis guineensis Jacq. II. Naturalista Postal, 1/83, no pagination. [unpublished]
Abud, A.J. (1983b) Insectos en Elaeis guineensis Jacq. III. Naturalista Postal, 5/83, no pagination. [unpublished]
Abud, A.J. (1984a) Insectos en Elaeis guineensis Jacq. IV. Naturalista Postal, 1/84, no pagination. [unpublished]
Abud, A.J. (1984b) Picudo negro en leguminosas. Naturalista Postal, 2/84, no pagination. [unpublished]
Abud, A.J. (1984c) Diabrotica undecimpunctata Barker. Naturalista Postal, 6/84, no pagination. [unpublished]
Abud, A.J. (1984d) Plagas en el cultivo de sorgo. Naturalista Postal, 7/84, no pagination. [unpublished]
Abud, A.J. (1985) Araecerus fasciculatus (De Geer) nueva plaga detectada en la República Dominicana. Naturalista Postal, 4/85, no
pagination. [unpublished]
Abud, A.J. (1986) Membracidae plaga de leguminosas. Naturalista Postal, 4/86, no pagination. [unpublished]
Abud, A.J. (1987a) Plaga de importancia en el cultivo del Ramio. Naturalista Postal, 5/87, no pagination. [unpublished]
Abud, A.J. (1987b) Nueva plaga para el girasol. Naturalista Postal, 6/87, no pagination. [unpublished]
Abud, A.J. (1990) Corynothrips stenopterus Williams en República Dominicana. Naturalista Postal, 2/90, no pagination. [unpublished]
Abud, A.J. (1991a) Comején en árboles frutales y madereros. Naturalista Postal, 5/91, no pagination. [unpublished]
Abud, A.J. (1991b) Presence of the avocado lace bug, Pseudacysta perseae (Heidemann) (Hemiptera: Tingidae) in Dominican Republic. In:
Primera Jornada de Protección Vegetal, Universidad Autónoma de Santo Domingo, Santo Domingo, pp. 4. [abstract]
Abud, A.J. (1992a) Afido plaga nueva en el país. 1991. Naturalista Postal, 1/92, no pagination. [unpublished]
Abud, A.J. (1992b) Afido vector de la tristeza de los cítricos en República Dominicana. Naturalista Postal, 1/92, no pagination. [unpublished]
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Perez-Gelabert, D.E. (2006) Mydidae (Diptera) de las Indias Occidentales. Cocuyo, 16, 35–36.
Perez-Gelabert, D.E. (2007) New records of Paramusonia cubensis (Saussure, 1869) (Dictyoptera: Mantodea) and notes on its behavior in the
Dominican Republic. Boletín de la Sociedad Entomológica Aragonesa, 40, 220.
Perez-Gelabert, D.E. (2008) Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography. Zootaxa, 1831 (1), 1–
530.
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Perez-Gelabert, D.E. (2008) Salto!! Para la diversidad biológica. Atajo, 7, 16–17.
Perez-Gelabert, D.E. (2009) Synonymy in Caribbean Tetrigidae (Orthoptera). Proceedings of the Entomological Society of Washington, 111,
900–901.
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Perez-Gelabert, D.E. (2011) Distribución de las luciérnagas (Coleoptera: Lampyridae) en la Hispaniola y su correspondencia con las
paleoislas. Novitates Caribaea, 4, 126–130.
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Perez-Gelabert, D.E. (2011) Diversidad y endemismo de los escarabajos (Insecta: Coleoptera) en la Hispaniola, Antillas Mayores. Novitates
Caribaea, 4, 65–79.
https://doi.org/10.33800/nc.v0i4.154
Perez-Gelabert, D.E. (2011) Primeros registros de acrocéridos (Diptera: Acroceridae) para la fauna actual de República Dominicana.
Novitates Caribaea, 4, 123–125.
https://doi.org/10.33800/nc.v0i4.163
Perez-Gelabert, D.E. (2012) Corrigenda a los escarabajos (Coleoptera) en la lista de artrópodos de la Hispaniola. Novitates Caribaea, 5, 114–
119.
https://doi.org/10.33800/nc.v0i5.141
Perez-Gelabert, D.E. (2012) New records of Dicrepidius ramicornis (Coleoptera: Elateridae: Elaterinae) from Dominican Republic,
Hispaniola. Novitates Caribaea, 5, 110–113.
https://doi.org/10.33800/nc.v0i5.140
Perez-Gelabert, D.E. (2014) Diversidad de saltamontes tetrígidos cladonótinos (Orthoptera: Tetrigidae: Cladonotinae) en las Antillas.
Resúmenes VIII Congreso de la Biodiversidad Caribeña, Santo Domingo, República Dominicana, 29 Enero–1 Febrero 2014, 111.
Perez-Gelabert, D.E. (2014) Inventario de las luciérnagas (Coleoptera: Lampyridae) de la Hispaniola. Resúmenes VIII Congreso de la
Biodiversidad Caribeña, Santo Domingo, República Dominicana, 29 Enero–1 Febrero 2014, 111.
Perez-Gelabert, D.E. (2014) Reporte sobre el conocimiento actual de los insectos en la Hispaniola. Resúmenes VIII Congreso de la
Biodiversidad Caribeña, Santo Domingo, República Dominicana, 29 Enero–1 Febrero 2014, 111–112.
Perez-Gelabert, D.E. (2014) Two new records of katydids (Orthoptera: Tettigoniidae: Conocephalinae) from the Dominican Republic,
Hispaniola. Novitates Caribaea, 7, 37–43.
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Perez-Gelabert, D.E. (2015) Ficha técnica de Espagnola darlingtoni J. A. G. Rehn y J. W. Rehn, 1939. Bioma, La Naturaleza en sus Manos, 3
(29), 59–61.
Perez-Gelabert, D.E. (2015) Entomofauna de la Reserva Científica Ebano Verde, Cordillera Central, República Dominicana. Novitates
Caribaea, 8, 61–81.
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Perez-Gelabert, D.E. (2016) Exploring the taxonomic diversity of the West Indian grasshopper genus Dellia Stål, 1878 (Acrididae:
Copiocerinae). Metaleptea, Special Issue, Abstracts 12 th International Congress of Orthopterology, Ilheus, Bahia, Brazil, 30 October–3
November 2016, 64.
Perez-Gelabert, D.E. (2016) New Hispaniola locality record for the endemic beetle Nicrophorus hispaniola Sikes & Peck, 2000 (Coleoptera:
Silphidae: Nicrophorinae). Novitates Caribaea, 10, 92–95.
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Perez-Gelabert, D.E. (2016) Ampliación de la distribución conocida de Dellia viridissima (Orthoptera: Acrididae) y notas sobre la historia
natural de Dellia en República Dominicana. Boletín de la Sociedad Entomológica Aragonesa, 59, 297–298.
Perez-Gelabert, D.E. (2019) Brachybaenus domingensis sp. nov. of gryllacridids (Orthoptera: Stenopelmatoidea: Gryllacrididae) from

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 Dominican Republic, Hispaniola. Novitates Caribaea, 13, 64–73.
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Perez-Gelabert, D.E. (2019) Discovery of a new fossil species of New World resin assassin bugs (Heteroptera: Reduviidae: Harpactorinae:
Apiomerini: Apicrenus). 8th International Conference on Fossil Insects, Arthropods & Amber, Talks, Abstracts, 62–63.
Perez-Gelabert, D.E. (2019) The invasive armored scale Aulacaspis yasumatsui Takagi (Hemiptera: Coccoidea: Diaspididae) in the
Dominican Republic. Proceedings of the Entomological Society of Washington, 121, 525–528.
https://doi.org/10.4289/0013-8797.121.3.525
Perez-Gelabert, D.E., Abud-Antún, A. & Nolasco, C. (2019) First record of the introduced Asian Plataspid Brachyplatys subaeneus
(Westwood) (Hemiptera: Heteroptera: Plataspidae) in the Dominican Republic. Proceedings of the Entomological Society of Washington,
121, 135–138.
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Perez-Gelabert, D.E. & Alba Suriel, F. (2016) Primer registro de ácaros acuáticos (Arrenurus sp.) como ectoparásitos de odonatos en la
Hispaniola. Novitates Caribaea, 10, 96–99.
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Perez-Gelabert, D.E. & Bastardo, R.H. (2005) Synoditella bisulcata (Kieffer) (Hymenoptera: Scelionidae) parasitizing Orphulella punctata
(DeGeer) (Orthoptera: Acrididae) in the Dominican Republic. Proceedings of the Entomological Society of Washington, 106, 932–933.
Perez-Gelabert, D.E., Bastardo, R.H. & Hierro, B. (2010) A lubber grasshopper Xyleus discoideus rosulentus (Stål, 1878) (Orthoptera:
Romaleidae: Romaleinae) in the Dominican Republic. Novitates Caribaea, 3, 77–80.
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Perez-Gelabert, D.E., Bastardo, R.H. & Medrano, S. (2011) Entomofauna del Parque Nacional Loma Nalga de Maco y alrededores, Prov.
Elías Piña, República Dominicana. Novitates Caribaea, 4, 80–90.
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Perez-Gelabert, D.E., Dominici, G. & Hierro, B. (1995) Jaragua: New genus and two new species of American pyrgomorphids (Orthoptera:
Pyrgomorphidae) from Hispaniola, West Indies. Annals of the Entomological Society of America, 88, 31–38.
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Perez-Gelabert, D.E., Dominici, G., Hierro, B. & Otte, D. (1995) New grasshopper genera and species from the Dominican Republic
(Hispaniola) (Acridoidea: Acrididae). Transactions of the American Entomological Society, 121, 153–171.
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Perez-Gelabert, D.E., Dominici, G., Hierro, B. & Otte, D. (1997) New eumastacid grasshopper taxa (Orthoptera: Eumastacidae: Episactinae)
from the Dominican Republic, including a fossil new genus and species from Dominican amber. Journal of Orthoptera Research, 6,
139–151.
Perez-Gelabert, D.E. & Floriano, C.F.B. (2016) New records of the water strider Platyvelia brachialis (Stål 1860) (Hemiptera: Heteroptera:
Veliidae) from the Dominican Republic and Haiti, with a checklist of the West Indian Veliidae. Proceedings of the Entomological Society
of Washington, 118, 471–476.
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Perez-Gelabert, D.E. & Edgecombe, G.D. (2013) Scutigeromorph centipedes (Chilopoda: Scutigeromorpha) of the Dominican Republic,
Hispaniola. Novitates Caribaea, 6, 36–44.
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Perez-Gelabert, D.E. & Flint, O.S. Jr. (2001) Annotated list of the Neuroptera of Hispaniola, with new faunistic records of some species.
Journal of Neuropterology, 3, 9–23.
Perez-Gelabert, D.E. & Grogan, W.L. Jr. (1999) Forcipomyia (Microhelea) tettigonaris (Diptera: Ceratopogonidae) parasitizing katydids
(Orthoptera: Tettigoniidae) in the Dominican Republic. Entomological News, 110, 311–314.
Perez-Gelabert, D.E., Hierro, B. & Otte, D. (1997) Espagnolopsis and Espagnolina, two new genera of eumastacid grasshoppers (Orthoptera:
Eumastacidae: Episactinae) from Hispaniola. Journal of Orthoptera Research, 6, 153–160.
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Perez-Gelabert, D.E., Hierro, B. & Otte, D. (1998) New genera and species of Greater Antillean grouse-locusts (Orthoptera: Tetrigidae:
Cladonotinae). Journal of Orthoptera Research, 7, 189–204.
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Perez-Gelabert, D.E. & Otte, D. (1999) Dos nuevas especies de saltamontes del género Dellia Stål (Orthoptera: Acrididae) de la República
Dominicana. Novitates Caribaea, 1, 1–13.
Perez-Gelabert, D.E. & Otte, D. (2000) Duartettix montanus, a new genus and species of high mountain grasshopper (Acrididae:
Melanoplinae) from Dominican Republic. Journal of Orthoptera Research, 9, 129–134.
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Perez-Gelabert, D.E. & Otte, D. (2003a) A new genus and species of tetrigid (Orthoptera: Tetrigidae) from Dominican Republic, Hispaniola.
Journal of Orthoptera Research, 12, 111–114.
https://doi.org/10.1665/1082-6467(2003)012[0111:ANGASO]2.0.CO;2
Perez-Gelabert, D.E. & Otte, D. (2003b) Melanoplus sanguinipes (Fabricius) (Orthoptera: Acrididae: Melanoplinae), a pest grasshopper
introduced to Dominican Republic, Hispaniola. Transactions of the American Entomological Society, 129, 497–502.
Perez-Gelabert, D.E. & Otte, D. (2003c) Tergoceracris, a new genus and six new species of montane grasshoppers (Orthoptera: Acrididae:
Ommatolampinae) from Dominican Republic and Puerto Rico. Zootaxa, 155 (1), 1–31.
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Perez-Gelabert, D.E. & Otte, D. (2012) Two new species of Dellia Stål, 1878 grasshoppers (Orthoptera: Acrididae) from Sierra de Bahoruco,
Dominican Republic. Novitates Caribaea, 5, 7–16.
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Perez-Gelabert, D.E. & Rowell, C.H.F. (2006) Further investigations of Hispaniolan eumastacoid grasshoppers (Espagnolinae: Episactidae:
Orthoptera). Journal of Orthoptera Research, 15, 241–249.
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Perez-Gelabert, D.E. & Thompson, F.C. (2006) A new genus and species of Richardiidae (Diptera) from Hispaniola. Zootaxa, 1259, 25–31.
Perez-Gelabert, D.E. & Yong, S. (2017) Erechthis gundlachi Bolívar, 1888 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini) preying
on a mollusk in Dominican Republic. Proceedings of the Entomological Society of Washington, 119, 286–289.
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Podenas, S. & Poinar, G.O. Jr. (2001) New crane flies (Diptera: Tipulidae, Limoniidae) from Dominican and Mexican amber. Proceedings of
the Entomological Society of Washington, 103, 863–878.
Poe, S.L. (1973) Tomato pinworm, Keiferia lycopersicella (Walsingham) (Lepidoptera: Gelechiidae) in Florida. Florida Department of
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Pogue, M.G. (2009) A review of the Tripudia quadrifera (Zeller) (Lepidoptera: Noctuidae) species complex. Proceedings of the
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Bohartillidae & Myrmecolacidae). Mitteilungen aus dem Geologisch-Palaeontologischen Institut der Universitaet Hamburg, 78, 197–
209.
Poinar, G.O. Jr. (1981) Fossil dauer rhabditoid nematodes. Nematologica, 27, 466–467.
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Poinar, G.O. Jr. (1982a) Feature photograph. Male preserved in amber, first record of Stylopoidea (Coleoptera) from New World. Annals of the
Entomological Society of America, 75(6). [unpaginated]
Poinar, G.O. Jr. (1982b) Sealed in amber. Natural History, 91, 26–32.
Poinar, G.O. Jr. (1984a) First fossil record of parasitism by insect parasitic Tylenchida (Allantonematidae: Nematoda). Journal of
Parasitology, 70, 306–308.
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Poinar, G.O. Jr. (1984b) Heidenius dominicus n. sp. (Nematoda: Mermithidae), a fossil parasite from the Dominican Republic. Journal of
Nematology, 16, 371–375.
Poinar, G.O. Jr. (1985) Fossil evidence of insect parasitism by mites. International Journal of Acarology, 11, 37–38.
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Poinar, G.O. Jr. (1986) Fossil evidence of spider parasitism by Ichneumonidae. Journal of Arachnology, 14, 399–400.
Poinar, G.O. Jr. (1988a) Zorotypus palaeus n. sp., a fossil Zoraptera (Insecta) in Dominican amber. Journal of the New York Entomological
Society, 96, 253–259.
Poinar, G.O. Jr. (1988b) The amber ark. Natural History, 97, 42–46.
Poinar, G.O. Jr. (1991a) Praecoris dominicana gen. n., sp. n. (Hemiptera: Reduviidae: Holoptilinae) from Dominican amber, with an
interpretation of past behavior based on functional morphology. Entomologica Scandinavica, 22, 193–199.
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Poinar, G.O. Jr. (1991b) The mycetophagous and entomophagous stages of Iotonchium californicum n. sp. (Iotonchidae: Tylenchida). Revue
de Nematologie, 14, 565–580.
Poinar, G.O. Jr. (1992a) Fossil evidence of resin utilization by insects. Biotropica, 24, 466–468.
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Poinar, G.O. Jr. (1992b) Life in Amber. Stanford University Press, Stanford, 350 pp.
Poinar, G.O. Jr. (1993a) Insects in amber. Annual Review of Entomology, 46, 145–159.
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Poinar, G.O. Jr. (1993b) Still life in amber—Time capsules of DNA from 40,000,000 B. C. The Sciences, 33, 34–38.
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Poinar, G.O. Jr. (1994a) Bees in fossilized resin. Bee World, 75, 71–77.
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Poinar, G.O. Jr. (1994b) The range of life in amber: significance and implications in DNA studies. Experientia, 5, 536–542.
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Poinar, G.O. Jr. (1995a) Discovering the Mysteries of Amber. Geofin, Udine, 67 pp.
Poinar, G.O. Jr. (1995b) First fossil soft ticks, Ornithodorus antiquus n. sp. (Acari: Argasidae) in Dominican amber with evidence of their
mammalian host. Experientia, 51, 384–387.
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Poinar, G.O. Jr. (1995c) Fleas (Insecta: Siphonaptera) in Dominican amber. Medical Science Research, 23, 789.

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Poinar, G.O. Jr. (1996a) A fossil stalk-winged damselfly, Diceratobasis worki spec. nov., from Dominican amber, with possible ovipositional
behavior in tank bromeliads (Zygoptera: Coenagrionidae). Odonatologica, 25, 381–385.
Poinar, G.O. Jr. (1996b) Fossil velvet worms in Baltic and Dominican amber: Onychophoran evolution and biogeography. Science, 273, 1370–
1371.
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Poinar, G.O. Jr. (1998a) Cenozoic fauna and flora in amber. World Congress on Amber Inclusions, Alava, Spain, 20–23 October, 1998,
Abstract p. 55.
Poinar, G.O. Jr. (1998b) Palaeontology of amber. Geology Today, 14, 154–160.
https://doi.org/10.1046/j.1365-2451.1998.014004154.x
Poinar, G.O. Jr. (1999a) A fossil palm bruchid, Caryobruchus dominicanus sp. n. (Pachymerini: Bruchidae) in Dominican amber.
Entomologica Scandinavica, 30, 219–224.
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Poinar, G.O. Jr. (1999b) Paleochordodes protus n. g., n. sp. (Nematomorpha, Chordodidae), parasites of a fossil cockroach, with a critical
examination of other fossil hairworms and helminthes of extant cockroaches (Insecta: Blattaria). Invertebrate Biology, 118, 109–115.
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Poinar, G.O. Jr. (1999c) Paleoeuglossa melissiflora gen. n., sp. n. (Euglossinae: Apidae), fossil orchid bees in Dominican amber. Journal of
the Kansas Entomological Society, 71, 29–34.
Poinar, G.O. Jr. (2000a) Extinction of tropical insect lineages in Dominican amber from Plio-Pleistocene cooling events. Russian
Entomological Journal, 8, 1–4.
Poinar, G.O. Jr. (2000b) Fossil onychophorans from Dominican and Baltic amber: Tertiapatus dominicanus n. g., n. sp. (Tertiapatidae n. fam.)
and Succinipatopsis balticus n. g., n. sp. (Succinipatopsidae n. fam.) with a proposed classification of the subphylum Onychophora.
Invertebrate Biology, 119, 104–109.
https://doi.org/10.1111/j.1744-7410.2000.tb00178.x
Poinar, G.O. Jr. (2001) Lagerstaetten. Dominican amber. In: Briggs, D.E.G. & Crowther, P.R. (Eds.), Paleobiology II. Blackwell Science,
Oxford, pp. 362–364.
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Poinar, G.O. Jr. (2003) Trends in the evolution of insect parasitism by nematodes as inferred from fossil evidence. Journal of Nematology, 35,
129–132.
Poinar, G.O. Jr. (2004) Evidence of parasitism by Strepsiptera in Dominican amber. BioControl, 49, 239–244.
https://doi.org/10.1023/B:BICO.0000025393.15395.b6
Poinar, G.O. Jr. (2005a) Culex malariager, n. sp. (Diptera: Culicidae) from Dominican amber: The first fossil mosquito vector of Plasmodium.
Proceedings of the Entomological Society of Washington, 107, 548–553.
Poinar, G.O. Jr. (2005b) Review of Fossil Spiders in Amber and Copal by Joerg Wunderlich. Journal of Arachnology, 33, 878–879.
https://doi.org/10.1636/0161-8202(2005)033[0878:BR]2.0.CO;2
Poinar, G.O. Jr. (2005c) Triatoma dominicana sp. n. (Hemiptera: Reduviidae: Triatominae), and Tripanosoma antiquus sp. n. (Stercoraria:
Trypanosomatidae), the first fossil evidence of a triatomine-trypanosomatid vector association. Vector Borne and Zoonotic Diseases, 5,
72–81.
https://doi.org/10.1089/vbz.2005.5.72
Poinar, G.O. Jr. (2006) Feroseta priscus (Neuroptera: Mantispidae), a new genus and species of mantidflies in Dominican amber. Proceedings
of the Entomological Society of Washington, 108, 411–417.
Poinar, G.O. Jr. (2009) Dominibrentus leptus, n. gen., n. sp. (Curculionoidea, Brentidae, Cyphagoginae, Dominibrentini, n. tribe), a straight-
snouted weevil in Dominican amber. Historical Biology, 21, 51–55.
https://doi.org/10.1080/08912960903122807
Poinar, G.O. Jr. (2009) Lutzomyia adiketis sp. n. (Diptera: Phlebotomidae), a vector of Paleoleishmania neotropicum sp. n. (Kinetoplastida:
Trypanosomatidae) in Dominican amber. Parasites & Vectors, 1, 1–8.
https://doi.org/10.1186/1756-3305-1-22
Poinar, G.O. Jr. (2009) Meloe dominicanus n. sp. (Coleoptera: Meloidae) phoretic on the bee Proplebeia dominicana (Hymenoptera: Apidae)
in Dominican amber. Proceedings of the Entomological Society of Washington, 111, 145–150.
https://doi.org/10.4289/0013-8797-111.1.145
Poinar, G.O. Jr. (2010) Palaeoecological perspectives in Dominican amber. Annales de la Société Entomologique de France, (N. S.), 46, 23–52.
Poinar, G.O. Jr. (2011) A walking stick, Clonistria dominicana n. sp. (Phasmatodea: Diapheromeridae) in Dominican amber. Historical
Biology, 23, 223–226.
https://doi.org/10.1080/08912963.2010.522405
Poinar, G.O. Jr. (2011) Descriptions of a broad-nosed weevil (Eudiagogini: Curculionidae) and false ladybird beetle (Nilionini: Nilionidae) in
Dominican amber. Historical Biology, 23, 231–235.
https://doi.org/10.1080/08912963.2010.527159
Poinar, G.O. Jr. (2012) Minyscapheus dominicanus n. gen., n. sp. (Hemiptera: Cicadidae), a fossil cicada in Dominican amber. Historical
Biology, 24, 329–333.
https://doi.org/10.1080/08912963.2011.603421
Poinar, G.O. Jr. (2011) New Termitaphididae and Aradidae (Hemiptera) in Mexican and Dominican amber. Palaeodiversity, 4, 51–62.
Poinar, G.O. Jr. (2011) Paleotrichius dominicanus n. gen., n. sp. (Coleoptera: Cetoniidae), a flower beetle in Dominican amber. Historical
Biology, 23, 109–113.
https://doi.org/10.1080/08912963.2010.482417
Poinar, G.O. Jr. (2011) The origin of insect-borne human diseases as revealed in amber. American Entomologist, 57, 170–178.
https://doi.org/10.1093/ae/57.3.170
Poinar, G.O. Jr. (2011) Vetufebrus ovatus n. gen., n. sp. (Haemosporida: Plasmodiidae) vectored by a streblid bat fly (Diptera: Streblidae) in
Dominican amber. Parasites & Vectors, 4 (229), 1–5.

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 https://doi.org/10.1186/1756-3305-4-229
Poinar, G.O. Jr. (2012) A fungal-like organism associated with a wasp (Hymenoptera: Pteromalidae) in Dominican amber. Journal of
Invertebrate Pathology, 110, 132–134.
https://doi.org/10.1016/j.jip.2012.02.013
Poinar, G.O. Jr. (2012) Desiomorphs in amber. American Entomologist, 58, 214–222.
https://doi.org/10.1093/ae/58.4.214
Poinar, G.O. Jr. (2012) Fossil gregarines in Dominican and Burmese amber: examples of accelerated development? Palaeodiversity, 5, 1–6.
Poinar, G.O. Jr. (2012) Paleorhodococcus dominicanus n. gen., n. sp. (Actinobacteria) in a faecal droplet of Triatoma dominicana (Hemiptera:
Reduviidae: Triatominae) in Dominican amber. Historical Biology, 24, 219–221.
https://doi.org/10.1080/08912963.2011.615931
Poinar, G.O. Jr. (2013) A snail-killing fly, Dominimyza tanycaena n. gen. n. sp. (Diptera: Sciomyzidae) in Dominican amber. Historical
Biology, 26, 428–432.
https://doi.org/10.1080/08912963.2013.784905
Poinar, G.O. Jr. (2013) Discoclavata dominicana n. gen., n. sp. (Coleoptera: Bostrichidae) and Lissantauga epicrana n. gen., n. sp.
(Coleoptera: Eucnemidae) in Dominican amber. Historical Biology, 25, 107–113.
https://doi.org/10.1080/08912963.2012.712518
Poinar, G.O. Jr. (2013) Panstrongylus hispaniolae sp. n. (Hemiptera: Reduviidae: Triatominae), a new fossil triatomine in Dominican amber,
with evidence of gut flagellates. Palaeodiversity, 6, 1–8.
Poinar, G.O. Jr. (2013) Stenaspidiotus microptilus n. gen., n. sp. (Coleoptera: Chrysomelidae: Chrysomelinae) in Dominican amber, with
evidence of tachinid (Diptera: Tachinidae) oviposition. Historical Biology, 25, 101–105.
https://doi.org/10.1080/08912963.2012.710443
Poinar, G.O. Jr. (2016) A sphere-forming scarab beetle (Ceratocanthinae: Hybosoridae) in Dominican amber. Historical Biology, 28, 433–437.
https://doi.org/10.1080/08912963.2014.985668
Poinar, G.O. Jr. (2015) A new genus of fleas with associated microorganisms in Dominican amber. Journal of Medical Entomology, 52, 1234–
1240.
https://doi.org/10.1093/jme/tjv134
Poinar, G.O. Jr. (2015) Pulchellaranea pedunculata n. gen. n. sp. (Araneae: Araneidae), a new genus of spiders with a review of araneid
spiders in Cenozoic Dominican amber. Historical Biology, 27, 103–108.
https://doi.org/10.1080/08912963.2013.869799
Poinar, G.O. Jr. (2015) Spirochete-like cells in a Dominican amber Amblyomma tick (Arachnida: Ixodidae). Historical Biology, 27, 565–570.
https://doi.org/10.1080/08912963.2014.897699
Poinar, G.O. Jr. (2016) Orchid pollinaria (Orchidaceae) attached to stingless bees (Hymenoptera: Apidae) in Dominican amber. Neues
Jahrbuch fuer Geologie und Palaeontologie Abhandlungen, 279, 287–293.
https://doi.org/10.1127/njgpa/2016/0556
Poinar, G.O. Jr. (2017) Ancient termite pollinator of milkweed flowers in Dominican amber. The American Entomologist, 63, 52–56.
https://doi.org/10.1093/ae/tmx011
Poinar, G.O. Jr. (2017) Fossilized mammalian erythrocytes associated with a tick reveal ancient piroplasms. Journal of Medical Entomology,
54, 895–900.
https://doi.org/10.1093/jme/tjw247
Poinar, G.O. Jr. (2018) Pygmy mole crickets (Orthoptera: Tridactylidae) in Dominican and Burmese amber. Historical Biology. [published
online]
https://doi.org/10.1080/08912963.2018.1483362
Poinar, G. O., Jr. (2019) Associations between fossil beetles and other organisms. Geosciences, 9, 184. 1–14.
https://doi.org/10.3390/geosciences9040184
Poinar, G.O. Jr. & Ballerio (2017) Remarks on some Ceratocanthinae (Coleoptera: Hybosoridae) in Dominican amber. Zootaxa, 4286, 125–
128.
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Poinar, G.O. Jr., Curcic, B.P.M. & Cokendolpher, J.C. (1998) Arthropod phoresy involving pseudoscorpions in the past and present. Acta
Arachnologica, 47, 79–96.
https://doi.org/10.2476/asjaa.47.79
Poinar, G.O. Jr., Brown, A.E. & Legalov, A.A. (2016) First record of the genus Pseudopilolabus Legalov, 2003 (Coleoptera: Attelabidae) in
Dominican amber. Fossil Record, 19, 11–16.
https://doi.org/10.5194/fr-19-11-2016
Poinar, G.O. Jr. & Brodzinsky, J. (1986) Fossil evidence of nematode (Tylenchida) parasitism in Staphylinidae (Coleoptera). Nematologica,
31, 353–355.
https://doi.org/10.1163/187529285X00427
Poinar, G.O. Jr. & Brown, J.W. (1993) A new fossil tortricid (Lepidoptera, Tortricidae) from Dominican amber. Entomologica Scandinavica,
24, 25–30.
https://doi.org/10.1163/187631293X00028
Poinar, G.O. Jr. & Brown, A. (2012) The first fossil streblid bat fly, Enischnomyia stegosoma n. g., n. sp. (Diptera: Hippoboscoidea:
Streblidae). Systematic Parasitology, 81, 79–86.
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Poinar, G.O. Jr. & Brown, A. (2015) New genera and species of Jumping Ground Bugs (Hemiptera: Schizopteridae) in Dominican and
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Organic Agriculture Movements (IFOAM), São Paulo, Brazil, 1992, pp. 173–180.
Serra, C.A. (1992d) Untersuchungen zum Einsatz von Niemsamenextrakten im Rahmen integrierter Ansätze zur Bekämpfung von
Tomatenschadlingen in der Dominikanishen Republik. Ph. D. Thesis, Universität Gieβen, Gieβen, 189 pp. [unpublished]
Serra, C.A. (1992f) Untersuchungen zur Wirkung von Niem- (Azadirachta indica A. Juss.) Extrakten auf den Blattminierer Keiferia
lycopersicella Wals. (Lepidoptera: Gelechiidae) und wichtigen Gegenspielern in der Dominikanischen Republik. Phytomedizin, 1992,
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Serra, C.A. (1993) Extractos de semillas del árbol Nim (Azadirachta indica A. Juss.) para el manejo integrado de minadores de hoja del
tomate. Proceedings of the Interamerican Society of Tropical Horticulture, 37, 54–61.
Serra, C.A. (1994a) Effects of selective insecticides, synthetics and from the neem tree (Azadirachta indica A. Juss.), on populations
of Bemisia tabaci (Genn.) (Homoptera: Aleyrodidae). In: Technical Center for Agricultural and Rural Co-operation (CTA)
(Eds.), CTA Seminar Proceedings: Integrated Pest Management—New Strategies for the Caribbean Farmer, Santo Domingo,
1994, pp. 185–193.
Serra, C.A. (1994b) Perspectivas y limitantes para el uso de insecticidas de nim (Azadirachta indica A. Juss.) en el cultivo de tomate industrial en
la República Dominicana. In: Brechelt, A. & Hellpap, C. (Eds.), Memorias 1er Congreso Latinoamericano y del Caribe sobre NIM y otros
Insecticidas Vegetales. Santo Domingo República Dominicana, Instituto Politécnico Loyola, San Cristóbal, pp. 227–238.
Serra, C.A. (1994c) Preliminary results of an intended host-free period in different North-western Dominican tomato growing areas on the
population dynamics of whiteflies and the incidence of geminivirus. Second annual review. Silverleaf Whitefly (formerly Sweetpo tato
Whitefly, Strain B) 1994 Supplement to the Five-Year National Research and Action Plan. U. S. Department of Agriculture, Orlando,
Florida, 17 pp. [poster abstract]
Serra, C.A. (1996a) Biología de moscas blancas. In: Hilje, L. (Ed.), Metodologías para el diagnóstico, investigación y manejo de moscas blancas
y geminivirosis. CATIE, Turrialba, pp. 11–21.
Serra, C.A. (1996b) Insecticidas de Nim (Azadirachta indica A. Juss.) para el manejo integrado de plagas del tomate—perspectivas para la
República Dominicana. In: Sabillón, A. & Bustamante, M. (Eds.), Memoria del 1er Taller Latinoamericano Bio-Plaguicidas ¿Mito,
Placebos o una alternativa en la agricultura sostenible? Escuela Agrícola Panamericana El Zamorano, Honduras, pp. 9.
Serra, C.A. (1996c) Muestreo de moscas blancas. In: Hilje, L. (Ed.), Metodologías para el diagnóstico, investigación y manejo de moscas blancas
y geminivirosis. CATIE, Turrialba, pp. 22–29.
Serra, C.A. (1996e) Whitefly problems and research in Latin America. Proceedings of the XX International Congress of Entomology,
Florence, Italy, 1996, 461.
Serra, C.A. (1998a) El control biológico de ‘moscas blancas’ (Homoptera: Aleyrodidae) ¿un componente viable para el manejo integrado? 3ra
Reunión Científica Tecnológica de la SODIAF, Santo Domingo, República Dominicana, 1998, unknown pagination.
Serra, C.A. (1998b) Population Dynamics of Bemisia in the Dominican Republic and Implications for Control. 2nd International Workshop on
Bemisia and Geminiviral Diseases, San Juan, Puerto Rico, 1998, L-27. [abstract]
Serra, C.A. (2000a) El control biológico de ‘Moscas Blancas’ (Homoptera: Aleyrodidae)- un recuento de investigaciones realizadas en la
República Dominicana. Memorias 40ma Reunión de la American Phytopathological Society-Caribbean Division (APS-CD) y 4 ta Reunión de
la SODIAF, Santo Domingo, República Dominicana, unknown pagination.
Serra, C.A. (2000b) Insecticidas botánicos de Nim y microbiológicos para la producción orgánica de tomates–Potencial y limitantes. In: Foro
Regional en Agricultura Orgánica ‘Limitaciones tecnológicas y oportunidades para la Región del Caribe‘. JAD-UNPHU-CEDAF,
Santo Domingo, unknown pagination.
Serra, C.A. (2001) Riesgos ambientales del control biológico clásico y de organismos genéticamente manipulados—una reflexión. 5ta Jornada
Científica, Sociedad Dominicana de Investigadores Agropecuarios y Forestales (SODIAF), Santo Domingo, 33. [poster abstract]
Serra, C.A. (2005) Update on invasive species initiatives in the Dominican Republic. Proceedings of the Caribbean Food Crops Society

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 (CFCS), 41, 110–124.
Serra, C.A. (2006) Manejo Integrado de Plagas de Cultivos: Estado Actual y Perspectivas para la República Dominicana. Proyecto AGORA,
Fundación Kellogg/CEDAF, Santo Domingo, 176 pp.
Serra, C.A. (2007) Distribución e impacto preliminar del ácaro rojo de las palmeras (Raoiella indica Hirst, Prostigmata: Tenuipalpidae) en la
República Dominicana. Memorias 3er Congreso Bianual, SODIAF, Santo Domingo, 2007, 12. [abstract]
Serra, C.A. (2014) Seguimiento de artrópodos exóticos invasores establecidos en la República Dominicana y planteamiento de estrategias para
su manejo. Resúmenes VIII Congreso de la Biodiversidad Caribeña, Santo Domingo, República Dominicana, 2014, 132. [abstract]
Serra, C.A., Cayetano, X., Féliz, A., Ferreira, M., García, S., Godoy, G., Halpay, M., Martínez, R.T., Méndez, R.M., Moya, J. de D., Silverio,
L. & Matos, L. (2011) Impacts of recently emerged invasive exotic species and major threats to the Dominican agriculture. Proceedings
of the Caribbean Food Crops Society, 47, 146–156.
Serra, C.A., Concepción, S. & Román, R. (1994) Resultados preliminares de un intento de período libre de hospederos en diferentes áreas
tomateras de la República Dominicana sobre la dinámica de población de "Moscas blancas" y la incidencia del geminivirus. V Congreso
Internacional de Manejo Integrado de Plagas, San José, Costa Rica, 1994, 209. [abstract]
Serra., C., Concepción, C., Román, R., Núñez, J., Ortiz, M. & Benoit, P. (1995) Effects of a host-free period on whitefly populations and the
spread of TYLCV-DR in northwestern Dominican tomato fields. Workshop on Survey Techniques for Tomato Yellow Leaf Curl Virus,
Santo Domingo, Dominican Republic, 1995, 155 [abstract]
Serra, C.A., Ferreira, M., García, S., Santana, L., Castillo, M., Nolasco, C., Morales, P., Holler, T., Roda, A., Aluja, M. & Sivinski, J. (2011)
Establishment of the West Indian fruit fly (Diptera: Tephritidae) parasitoid Doryctobracon areolatus (Hymenoptera: Braconidae) in the
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Serra, C.A., García, S. & Ferreira, M. (2005) Evaluación de Mangifera indica, Spondias spp. (Anacardiaceae) y Psidium guajava (Myrtaceae),
hospederos de moscas de las frutas, Anastrepha spp. (Diptera: Tephritidae), en cuanto a la presencia de parasitoides en diferentes zonas
de la República Dominicana. 51 Reunión Anual, Sociedad Interamericana para Horticultura Tropical (ISTH), Boca Chica, República,
2005, 155. [abstract]
Serra, C.A., García, S., Arias, J. & Segura, Y. (2006) Plantas hospederas de la mosca asiática del guandul en la República Dominicana.
Memoirs Caribbean Food Crops Society (CFCS), 42, 113–120.
Serra, C.A., García, S., Ferreira, M. & Batista, O. (2005) Comparación de tipos de trampas para el trampeo de moscas de las frutas,
Anastrepha spp. (Diptera: Tephritidae) en frutales en la República Dominicana. 51 Reunión Anual, Sociedad Interamericana para
Horticultura Tropical (ISTH), Boca Chica, República Dominicana, 2005, 156. [abstract]
Serra, C.A., García, S., Ferreira, M.A., Batista, O., Epsky, N.D. & Heath, R.R. (2005) Comparación de atrayentes para el trampeo de moscas
de las frutas, Anastrepha spp. (Diptera: Tephritidae) en frutales en la República Dominicana. Proceedings of the Caribbean Food Crops
Society (CFCS), 41, 524–532.
Serra, C.A., García, S., Segura, Y. & Arias, J. (2003) Dramatic impact of the recently introduced Asian pigeon pea podfly, Melanagromyza obtusa
(Malloch) (Diptera: Agromyzidae), in the Dominican Republic. Proceedings of the Caribbean Food Crops Society (CFCS), 39, 40. [poster abstract]
Serra, C.A., Jorge, P.E., Abud-Antún, A.J., Alvarez, P. & Peguero, B. (2003) Invasive alien species in the Dominican Republic: Their impacts,
and strategies to manage introduced pests. Simposio sobre Especies Invasoras en el Caribe, 39 Reunión Anual 2003, Grenada,
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Serra, C.A., Nuñez, C.A. & García, S. (2004) El control natural y biológico clásico de la cochinilla rosada de los hibiscus (CRH),
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Agropecuarias y Forestales (IDIAF), Santo Domingo, República Dominicana, 2004, 19.
Serra, C.A., Núñez, J.B. & Ramírez, W. (1997) Perspectivas del control biológico de moscas blancas (Homoptera: Aleyrodidae) en cultivos de
tomates y ornamentales. Memorias VI Taller Latinoamericano y del Caribe sobre Moscas Blancas y Geminivirus, Santo Domingo,
República Dominicana, 1997, 20.
Serra, C.A., Nuñez, J.B., Ortíz, M. (1995) Perspectivas de Cyrtopeltis spp. (Hemiptera: Miridae) como depredadores de Bemisia tabaci
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Serra, C.A. & Ogando, F.R. (2015) Improvement of ethological methods for integrated management of fruit flies, Anastrepha spp. (Diptera:
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Serra, C.A., Ortíz, M. & Nuñez, J.B. (1995) Distribución preliminar de 'moscas blancas' (Homoptera: Aleyrodidae) y sus enemigos naturales
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Serra, C.A., Ortíz, M., Nuñez, J B. & Benoit, P.F. (1995) Survey of indigenous natural enemies of whiteflies in the Dominican Republic. 4th
Annual progress review of the 5-year national research and action plan for development of management and control methodology fo r
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Serra, C.A., Ortíz, M., Nuñez, J.B. & Benoit, P.F. (1997) Inventario de parasitoides de Bemisia tabaci (Genn.) en República Dominicana.
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Serra, C.A., Ortíz, M., Nuñez, J.B., Schulz, A. & Benoit, P.F. (1995) Potential of entomophagous fungi to control whiteflies on Dominican
vegetable and ornamental crops. 4th Annual progress review of the 5-year national research and action plan for development of
management and control methodology for silverleaf whitefly, San Antonio, Texas, 19995, 142. [poster abstract]
Serra, C.A., Ortíz, M., Nuñez, J.B., Schulz, A. & Benoit, P.F. (1996a) Natural enemies of whiteflies in the Dominican Republic—their
prospects for biocontrol in tomato and ornamental crops. Proceedings of the XX International Congress of Entomology, Firenze, Italy,
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to native Opuntia floras in Mexico and the United States of America. Diversity and Distributions, 6, 259–269.
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Zippel, K.C., Powell, R. & Smith, D.D. (1996) The distribution of larval Eutrombicula alfreddugesi (Acari: Trombiculidae) infesting Anolis
lizards (Lacertilia: Polychrotidae) from different habitats on Hispaniola. Caribbean Journal of Science, 32, 43–49.
Zompro, O. (2001) A generic revision of the insect order Phasmatodea: The New World genera of the stick insect subfamily Diapheromeridae:
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Zrzavy, J. & Nedved, O. (1999) Evolution of mimicry in the New World Dysdercus (Hemiptera: Pyrrhocoridae). Journal of Evolutionary
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Zuparko, R.L. & Poinar, G.O. Jr. (1997) Aivalykus dominicanus (Hymenoptera: Braconidae), a new species from Dominican amber.
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CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 573
INDEX TO FAMILIES, GENERA AND SPECIES

?begini (Diglyphus), 293 absonus (Parathysanoes), 178 achrysa (Aaroniella), 381

Aaroniella, 381 absonus† (Zorotypus), 391 Achrysocharoides, 292
Abaeis, 338 abstrusum (Cycloptilum), 376 Achryson, 149
abagus (Horismenus), 293 absumens (Metallata), 354 Achyra, 324
Abanaxipha†, 375 abudi (Heteroctenus), 82 acicula (Stenotrupis), 170
abares (Cyrtoxipha), 375 abudi (Monobelus), 270 aciculata (Cressonomyia), 232
Abaristophora, 214 abudi (Odo), 70 aciculaticollis (Rhyssomatus), 176
abatos (Amphiacusta), 373 abudi (Tityus), 82 aciculus (Ataenius), 120
abbotti (Ortholestes), 362 abydata (Macaria), 333 aciculus (Transeius), 53
abbotti (Polyancistrus), 368 Abyssocypris, 26 acidaliata (Macaria), 333
abbreviata† (Holocompsa), 187 Abyssocythere, 27 Acilius, 100
abbreviatus (Diaprepes), 173 Acacesia, 65 acinaces (Zeadolopus), 110
abbreviatus (Eurypanopeus), 44 acaciae (Tetraleurodes), 273 Acinia, 222
abbreviatus (Neoscapteriscus), 376 Acalitus, 48 Acinopterus, 265
abdita† (Proplebeia), 317 Acallepitrix, 156 acis petioni (Strymon), 339
abditus (Prostemmiulus), 93 Acalymma, 159 ackleianus (Pachycheles), 40
abdominale (Allotrichoma), 232 Acamptus, 170 Acladocera, 129
abdominale (Dactylosternum), 109 Acanalonia, 258 Acmaeodera, 125
abdominale (Zeta), 309 ACANALONIIDAE, 258 Acmopolynema, 295
abdominalis (Mezira), 248 Acanthaegilips, 289 acolum (Toxorhynchus), 167
abdominalis (Microcephalothrips), 384 Acanthepeira, 65 Acontia, 349
abdominalis (Pissonotus), 259 Acanthinus, 149 acontiaria (Omyza?), 333
abdominalis (Tramea), 364 Acanthocerus, 253 acontioides (Melipotis), 353
abdominalis abdominalis (Balclutha), Acanthocheila, 242 Acratodes, 335
266 Acanthognathus, 302 Acrepidopterum, 154
abdominalis colona (Eumenes), 309 Acanthonyx, 42 ACRIDIDAE, 366
abdominalis hispaniolae (Zethus), 309 Acanthoscelides, 163 acridophagoides (Blaesoxipha), 219
abdominaria (Thysanopyga), 334 Acanthostichus, 298 Acridurus, 367
abductus† (Electromyrmococcus), 279 Acanthotomicus, 179 acrita (Agalliopsis), 263
abeja (Cladochaeta), 229 acanthura (Cirolana), 31 Acrocercops, 320
abeja (Grallipeza), 221 acanthuri (Anilocra), 31 ACROCERIDAE, 206
abeja (Heterosmaitia), 339 acanthurus (Macrobrachium), 37 Acrocyum, 156
abejanus (Euconnus), 116 Acaricoris, 248 ACROLOPHIDAE, 319
abejas (Hapithus), 369 ACARIDAE, 51 Acrolophus, 319
Abelona, 377 acarophilus† (Dryophthorus), 172 Acropyga, 302
aberrans (Clastopteromyia), 229 Acarus, 51 Acroricnus, 287
aberrans (Eoromus), 90 acastusalis (Bleptina), 356 acroschismata (Curtara), 267
abia (Wyeomyia), 201 aceitillar (Phyllophaga), 121 Acrosticta, 223
abiecta (Scaptosciara), 205 aceitillari (Euconnus), 116 Acrotaenia, 222
Ablabesmyia, 199 Aceratagallia, 263 Actenodes, 124
Ablechroides, 136 acerba (Bradysia), 203 actiniae (Heteromysis), 29
Ablechrus, 136 ACERENTOMIDAE, 96 aculatus (Bromeloecia), 228
ablunaris (Achaea), 351 Aceria, 48 aculeata (Omoteina), 160
abnormalis (Cosmopterix), 322 Acerophagus, 291 aculeata (Pitho), 42
abnormeantennatus (Euconnus), 116 Acerotella, 283 Aculepeira, 65
abnormis (Aleuropleurocelus), 272 acestealis (Apogeshna), 324 aculicaput† (Loxosceles), 57
abnormis (Leptomastidea), 292 acevedoi (Cryptorhopalum), 133 Aculops, 48
abnormis† (Paralicus), 170 Achaea, 351 acuminata (Chalcosicya), 162
abraxasella abraxasella (Ethmia), 321 Achaearanea, 61 acuminata (Kilifia), 276
abrupta (Cladochaeta), 229 Achelous, 43 acuminata (Melanophila), 125
abruptus (Paratachys), 107 achemon (Cosmosoma), 348 acuminata (Solenocera), 35
absconditaria (Drepanodes), 335 Achenomorphus, 113 acuminatus (Euschistus), 255
absconditum (Cycloptilum), 376 acheronta semele (Historis), 342 acuminiforceps (Lachesilla), 381
absitaria (Cidaria), 335 ACHILIDAE, 258 acuminipennis acuminipennis (Cyarda),
absoluta (Tuta), 322 Achromoporus, 87 260

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 574
acuminipennis haitensis (Cyarda), 260 adventitius (Typhloseiopsis), 54 affinis (Tetracha), 101
acupunctatus (Scyphophorus), 177 Aedeomyia, 200 affinis (Xyleborus), 180
acuta (Adelphe), 310 Aedes, 199 affirmata (Triphosa), 335
acutangula (Anasa), 253 Aedmon, 156 afflicta hispaniolana (Manduca), 344
acutangula (Coproica), 228 aegerfasciella (Ochrotrichia), 389 africanus (Hypothenemus), 178
acutangulus (Euarestoides), 223 AEGIDAE, 31 Agalenocosa, 68
acutangulus (Robopus), 131 aegisthus (Mania), 336 Agallia, 263
acuticauta (Agalliopsis), 263 Aegithus, 139 Agalliana, 263
acuticornis (Mithrax), 42 aegypti (Stegomyia), 200 Agalliopsis, 263
Acuticythereis, 27 Aeletes, 109 AGAONIDAE, 289
acutilabella (Drosophila), 229 Aellopos, 344 Agapostemon, 315
acutipenne (Steirastoma), 154 aelops (Beskia), 220 agarithe antillia (Phoebis), 338
acutipennis (Tetracha), 101 aemete (Stygiomysis), 29 agassizi (Xestocephalus), 269
acutispina (Silis), 132 aenea (Apraea), 157 agassizii (Nephropsis), 38
acutissimus (Coccus), 275 aenea (Paridris), 284 agastos (Hapithus), 370
acutissimus (Tylospilus), 255 aenea (Rhinoprosopa), 216 agasusaria (Hyperythra?), 334
acutus (Flatoidinus), 261 Aeneolamia, 262 Agathidium, 110
acutus (Oecleus), 259 aeneomicans (Leucocera), 161 Agathispa, 159
acutus† (Ischnocolinopsis), 56 aenescens (Acylophorus), 117 Agathodes, 324
Acylomus, 138 aeneus (Pectinifer), 231 agavealis (Herpetogramma), 325
Acylophorus, 117 aeneus (Sicoderus), 176 Agelaia, 309
Adaina, 330 aeneus aeneus (Aspathines), 143 AGELENIDAE, 69
adamesi (Eurycotis), 184 aeneus ovatus (Aspathines), 143 ageneius (Hemipteroseius), 52
adansoni (Hasarius), 75 aenigmatica (Sibyllina), 308 Ageniaspis, 291
adaptos (Hapithus), 370 aenobarbus (Strategus), 123 Ageniella, 307
adarca (Polytrichophora), 232 aeola (Jadera), 254 Agenodesmus, 89
addens (Eulepidotis), 352 AEOLOTHRIPIDAE, 383 agens (Polytrichophora), 232
addita (Semiclivina), 107 Aeolus, 128 agerochus (Anthonomus), 169
adecton (Cycloptilum), 376 aequale (Trogoxylon), 134 agilis (Belonuchus), 117
adectos (Hispanogryllus), 372 aequalis (Enochrus), 108 agilis (Cyrtopholis), 56
adela† (Leucotrichia), 388 aequalis (Micrathyria), 364 aglae (Alisotrichia), 388
Adelencyrtus, 291 aequalis (Renodes), 354 aglandis (Ischnoptera), 186
Adelina, 146 aequatus (Lophoturus), 86 Aglaocoris, 248
Adelpha, 341 aequinoctialis (Pheropsophus), 101 Aglyptinus, 110
Adelphe, 310 aequinoctialis (Platynus), 104 Agobardus, 74
adelphe (Teleonemia), 243 aequinoctialis (Scyllarides), 38 agoiadiomensis‡ (Hemicyprideis), 22
adenolepus† (Paraulosomus), 171 aequipars (Nystalea), 347 agona (Parthenope), 43
Adesmus, 154 aequus‡ (Nephrops), 38 agonellus (Platynus), 104
adiketis† (Pintomyia), 195 aereus (Lachnopus), 174 Agonoscelis, 256
adipaloides (Diacme), 325 aerialis (Amblyseius), 53 AGONOXENIDAE, 322
adnexa (Coryphaeschna), 363 Aeroppia, 47 Agonum, 104
adona (Madoniella), 135 AESHNIDAE, 363 AGORISTENIDAE, 77
adonidum (Pseudococcus), 280 aestiva (Rhagovelia), 240 Agoristenus, 77
adorabilis (Anasaitis), 74 aetherea (Hexatoma), 193 agraphodactylus (Leioptilus), 330
adorabilis (Hapithus), 370 aetheria (Amphiacusta), 373 Agraulis, 341
adorea (Myelois), 328 aethiopica (Lachesilla), 381 Agrenocythere, 27
adrastus† (Procoilodes), 119 aethiops (Throscinus), 127 agri† (Trentepohila), 193
adstrictosetus† (Epidapus), 204 Aethosia, 349 Agrilus, 124
adusta (Caenurgia), 352 AETHRIDAE, 40 Agriognatha, 64
adustum (Allotrichoma), 232 aff. cinquaginta (Hexabdella), 48 Agrius, 344
adustum (Pterogramma), 228 aff. fiseri (Proctolaelaps), 52 Agroiconota, 159
adustus† (Eugonus), 165 affinis (Empyreuma), 347 Agromyza, 224
advena (Ahasverus), 138 affinis (Hadropenaeus), 35 AGROMYZIDAE, 2224
advena (Cariblatta), 186 affinis (Neoconocephalus), 368 agrorum (Palpada), 215
advena (Coccotrypes), 179 affinis (Passalus), 118 Agrotis, 357
advena (Hydrocanthus), 99 affinis (Prostemmiulus), 93 aguanueva (Cyrtoxipha), 375

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agujas (Syspira), 70 albipes (Xenasphondylia), 202 Aleurolobus, 272
Aguna, 337 albipunctata (Telmatoscopus), 195 Aleuroplatus, 272
Ahasverus, 138 albiscapus (Horismenus), 293 Aleuropleurocelus, 272
ahenea (Heliozela), 319 albisignis (Mormidea), 257 aleurothrixi (Neopomphale), 293
Ahotta, 77 albistramineus (Camponotus), 299 Aleurothrixus, 272
aicen (Paracanace), 226 albitacta (Gonitis), 351 Aleurotrachelus, 272
ainigma (Calisto), 342 albitegula (Lophocampa), 358 Aleurotuberculatus, 272
aitia (Proaspis), 89 albiterga (Grallipeza), 221 alexae† (Dynamine), 340
aitkeni (Pseudosciara), 205 alboannulatus (Anthonomus), 169 Alexandrina, 369
Aivalykus, 285 albocapitis (Anthonomus), 169 alexbrowni† (Quizqueiplana), 258
Alabagrus, 285 albocincta (Neopamera), 251 ALEYRODIDAE, 272
Alabama, 351 albofasciatus (Microrhagus), 127 aleyrodis (Signiphora), 296
alacris (Thermesia), 354 albogilvus (Pericales), 136 alfreddugesi (Trombicula), 50
aladecoris (Decorosa), 135 albolatus (Anadenobolus), 92 alia (Efferia), 207
Alaetrinus, 145 albomaculata (Chrysso), 61 aliada (Phyllophaga), 121
Alagoasa, 157 albomaculata (Cyphomyia), 206 aliceae (Basileunculus), 214
alametallus† (Dryinus), 311 albomaculata (Paratrikona), 161 aliciae† (Ochrotrichia), 389
Alaptus, 295 albomaculatus (Lachnopus), 174 Alienates, 238
alardus heriul (Astraptes), 337 albomarginata (Paroxacis), 148 alienus (Acanthaegilips), 289
alas (Horismenus), 293 albonotata (Howardina), 199 alifer (Docodesmus), 90
alata‡ (Perissocytheridea), 22 albonotata (Microvelia), 240 Alima, 28
Alathetus, 255 albonotata (Spectralia), 125 Alisalia, 111
Alatuncusia, 324 alboornatus (Lygaeus), 249 Alismobates, 46
Alaus, 127 albopalpis (Erythrolychnia), 130 Alisotrichia, 388
alaya (Hoplisoides), 312 albopicta (Stegomyia), 200 aliter† (Drasyllinus), 72
alayoi (Agapostemon), 315 albopictum (Amblyomma), 52 alius (Heterophotinus), 131
alayoi (Kapogea), 67 albopictus (Zethus), 309 allardi (Antisolabis), 183
alayoi (Tachysphex), 313 albopunctatus (Arvelius), 256 allardi (Oecanthus), 373
alayoni (Centruroides), 81 albosquamosus (Exophthalmus), 173 Allaudomyia, 196
alayoni (Pseneo), 313 albovenosus (Pissonotus), 259 allecta (Calycomyza), 225
alazon (Dysmicoccus), 279 albugo (Phaops), 173 Allecula, 145
alba (Siphonocybe), 91 albula (Spodoptera), 356 alleni† (Antillipeltis), 135
albaeaquae (Oxyethira), 389 albula (Tethina), 226 alleni† (Canthochilum), 121
albicans (Elousa), 352 Albunea, 38 Allocyclosa, 66
albicincta (Aedmon), 156 ALBUNEIDAE, 38 Allodardanus, 39
albicollis (Omophoita), 158 albuquerquensis (Mastigodiaptomus), 16 Allodia, 202
albicornis (Isotes), 159 albus (Aonidomytilus), 278 Allograpta, 216
albicornis (Onthophagus), 120 albutius (Pachyneuron), 296 Allonothrus, 48
albicornis albicornis (Onthophagus), 120 Alcaeorrhynchus, 255 allosetosipennis (Euconnus), 116
albicornis capitatus (Onthophagus), 120 Alcidion, 153 Allosmaitia, 339
albida (Nesara), 358 Alcimobolus, 92 Allostemma, 283
albidula (Agallia), 263 Alcimosphenus, 64 allotestaceus (Euconnus), 116
albidula (Zale), 355 alcinus (Valenzuela), 381 Allothenemus, 178
albidus (Macrocyclops), 16 alcoa (Calliscelio), 283 Allothrips, 385
albidus (Rhopalogaster), 209 alcoa (Phyllophaga), 121 Allotingis, 242
albifasciatus† (Dasymutilla), 306 Alconeura, 269 Allototua, 63
albifemur (Pteroptrix), 290 alcyone (Augochlora), 315 Allotrichoma, 232
albifrons (Palpada), 215 aldama† (Ochrotrichia), 389 Allotropa, 283
albifrons (Symmerista), 347 alejandroi (Phrynus), 54 almandoensis (Euconnus), 116
albigena (Spalangiopelta), 296 Aleochara, 111 Almodes, 335
albigera (Condica), 350 aleptos (Cyrtoxipha), 375 alnus (Horismenus), 293
albimaculata (Curtara), 267 Alethaxius, 162 alope alope (Erinnyis), 345
albimanus (Anopheles), 200 Aleurocanthus, 272 alpestris (Corthylus), 181
albiornatella (Varneria), 330 Aleurodicus, 273 alpha† (Azteca), 298
albipes (Doru), 183 Aleurodothrips, 385 ALPHEIDAE, 36
albipes (Homoeotarsus), 114 Aleuroglandulus, 272 Alpheopsis, 36

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 576
Alpheus, 36 ambigua (Euborellia), 183 Amoea, 358
Alphitobius, 144 ambiguus (Dendrocranulus), 179 amoena (Mesovelia), 239
alta‡ (Parakrithe), 24 ambiguus† (Anochetus), 304 amoenaria astylusaria (Euchlaena), 333
altagracia (Nesocyrtosoma), 147 Amblycerus, 163 amoenus (Pseudothysanoes), 178
altagraciae (Centruroides), 81 Amblydromalus, 53 amomidis (Eriococcus), 279
altamira (Emblyna), 69 Amblygnathus, 102 amone (Platynus), 105
altercoma (Helicopsyche), 390 Amblyomma, 52 Amorbia, 332
alternans (Scolopendra), 85 Amblypsilopus, 212 Amphelictogon, 89
alternans (Selenophorus), 102 Amblyseius, 53 Amphiacusta, 373
alternata (Atheta), 111 Amblystira, 242 Amphibalanus, 16
alternata (Cyrtaphe), 88 Amblytropidia, 367 amphichloe diasia (Hamadryas), 340
alternata (Tinearia), 195 Amboakis, 135 Amphicranus, 181
alternatus (Sicoderus), 176 Ambocythere, 27 Amphidraus, 74
Althos, 253 amboinensis (Lysmata), 36 AMPHIENTOMIDAE, 380
alticolus (Ardistomis), 106 Ambradolon†, 211 amphioceanica† (Aphaenogaster), 303
alticolus (Stenotabanus), 207 ambrosiae (Uroleucon), 274 amphioculus† (Toxorhynchus), 167
altifluminis (Platynus), 105 Ambrosiodmus, 179 AMPHIPSOCIDAE, 380
altila‡ (Paracytheridea), 25 ambrosion (Cycloptilum), 376 amphitrite‡ (Amphibalanus), 16
altimontana (Callopisma), 130 Amegarthria, 328 ampliata (Enocomia), 262
altioculata† (Dipoenata), 61 AMEIRIDAE, 17 Amplicephalus, 266
Altisessor, 139 americana (Callitroga), 217 amplicollis (Colaspis), 162
altithronus (Tityus), 82 americana (Carcinophora), 183 amplioculatus† (Nasutitermes), 189
altivagans (Camponotus), 299 americana (Cotesia), 286 amplioculus† (Dryotribus), 171
altus (Derallus), 108 americana (Dunhevedia), 14 ampliplaga (Hampsonodes), 350
altus† (Heterotmarus), 73 americana (Peckhamia), 76 amplus (Cnesinus), 177
Alujamyia, 223 americana (Periplaneta), 184 ampulla (Cassida), 160
Alumeda†, 245 americana antilleana (Bembix), 312 ampullocera (Leptosciarella), 205
alumnus (Philonthus), 117 americana† (Forcipomyia), 197 Amyelois, 328
alveolatus† (Cephalotes), 301 americanum (Gnathophyllum), 37 Anabasis, 328
ALYDIDAE, 252 americanus (Dineutus), 99 Anacaena, 108
alzadae† (Prorhinopsenius), 118 americanus (Euchirograpsus), 41 Anacaona, 368
alzatei (Rogeria), 304 americanus (Faiditus), 62 anacaona (Atomosia), 208
amarilla (Rachispoda), 228 americanus (Folsomides), 95 anacaona (Notocyphus), 306
amaroides (Metacerylon), 140 americanus (Laccophilus), 100 anacaona (Selenops), 73
amatella (Dioryctria), 329 americanus (Myrmecophilus), 376 Anachrostomus, 249
AMAUROBIIDAE, 70 americanus ambiguus (Anoplius), 307 anacrostichalis† (Stenomicra), 226
Amaurochrous, 257 americanus† (Brachypogon), 196 Anadelosemia, 328
Amauromyza, 225 Americhernes, 80 Anadenobolus, 92
amaurota (Neohadria), 265 Ametallon, 292 Anaea, 340
amazili (Anax), 363 amethystina (Macrohaltica), 158 Anagrus, 295
amazona (Sirthenea), 246 amethystinus amethystinus (Anoplius), Anagyrus, 292
ambarensis† (Microtityus), 82 307 Anaides, 119
ambarensis† (Sceloenopla), 161 amica (Amblystira), 242 anaitisalis (Aluaca), 352
ambarinus† (Paleomastacris), 366 amica (Anaxipha), 375 anale (Cyparium), 116
amberae† (Megaselia), 214 amiculus (Sympherobius), 360 analis (Centris), 317
amberdominica† (Mezira), 248 amigdala‡ (Aurila), 23 analis (Exomalopsis), 317
amberestris† (Chrysobothris), 125 Amitermes, 190 analis (Oxybelus), 312
amberica† (Glyphembia), 234 Amithao, 124 ananassae (Drosophila), 230
ambericum† (Cryptorhopalum), 133 Amitus, 283 ananatis (Anagyrus), 292
ambericus† (Attagenus), 133 Ammalo, 348 Ananca, 148
ambericus† (Culicoides), 196 ammon noeli (Cyclargus), 339 anansi† (Micromalthus), 99
ambericus† (Ornebius), 376 AMMOTHEIDAE, 45 anaphus anausis (Astraptes), 337
ambericus† (Syndesus), 124 Ammotrechella, 83 ANAPIDAE, 63
amberifera† (Periscelis), 226 AMMOTRECHIDAE, 83 Anaplecta, 186
Amberobyrsa, 242 Ammotrechinus, 83 Anartia, 342
Amberoderma†, 133 Amnestus, 254 Anasa, 253

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Anasaitis, 74 angusta (Apogonalia), 264 Anomalon, 287
anasilviae (Rowlandius), 81 angusta (Cheguevaria), 132 anomalus (Setodes), 390
anasilviae (Tityus), 82 angusta (Efferia), 207 Anomis, 351
Anastatus, 294 angusta‡ (Occulocythereis), 27 Anommatelmis, 126
anastera (Eustala), 66 angustata (Mormidea), 257 Anoncia, 322
Anastrepha, 223 angustata (Phaleria), 147 anonella (Cerconota), 321
anastrephae (Utetes), 286 angustatus (Neoheterocerus), 127 anonyma (Monochaetoscinella), 227
Anastrus, 337 angusticercus (Lepidosina), 228 Anopheles, 200
anatifera (Lepas), 15 angusticollis (Ectmesopus), 159 Anopliomorpha, 151
Anaulacomera, 368 angustiformis (Diochus), 117 Anoplischius, 128
Anaulacus, 102 angustipes (Haplocyclodesmus), 91 Anoplius, 307
Anax, 363 angustostriatus† (Cladoctonus), 177 Anoplodactylus, 45
Anaxipha, 375 angustum (Plesiomma), 210 Anoplotermes, 189
anceps (Portunus), 43 angustus (Docodesmus), 90 anoplus (Tenuipalpus), 50
Anchastomorphus, 129 angustus† (Eugonus), 165 Anopsilana, 31
Anchonoderus, 103 angustus† (Veterator), 72 anormalis (Agobardus), 74
Anchonus, 176 Anicetus, 291 anormalis (Microthyris), 326
Ancylostomia, 328 Anicla, 367 anormalis montanus (Agobardus), 74
ancyra (Agalliopsis), 263 Anilocra, 31 ANOSTOSTOMATIDAE, 376
Andaspis, 278 animatus (Tricolus), 181 Anotylus, 113
anderseni† (Miohebrus), 239 animosum (Cycloptilum), 376 ansa (Caribecandona), 20
anderseni‡ (Cushmanidea), 21 ANISEMBIIDAE, 234 anserifera (Lepas), 15
andersoni (Apodrosus), 175 Anisepyris, 309 Antachara, 350
Andrallus, 255 Anisolabis, 183 antaeus antaeus (Cocytius), 344
andreae (Dysdercus), 252 Anisomorpha, 378 Antapistis, 352
andremona (Hypocala), 353 Anisopilothrips, 384 antarcticus (Parribacus), 38
ANDRENIDAE, 314 ANISOPODIDAE, 201 antecognatus† (Episinus), 62
Andrenosoma, 208 Anisostena, 159 antennalis (Scopaeus), 114
andrewi (Strymon), 340 ankeri (Lysmata), 36 antennata (Distigmoptera), 157
androgeus epidaurus (Heraclides), 338 Anlemmus†, 171 antennatum (Platydema), 146
androsensis (Elaphidion), 151 annae (Typhlocypris), 21 antennatus (Dipropus), 128
androw (Phyllophaga), 121 annae† (Okamninus), 134 antennatus (Elateropsis), 156
andujari (Achromoporus), 87 annecta (Thaumasia), 68 Anteon, 310
andujari (Pozonia), 68 annectens (Galendromus), 53 Anteos, 338
Anegcephalesis, 328 annexa (Agrotis), 357 antestiae (Corioxenos), 383
Anelaphus, 151 annonae (Eriophyes), 48 Anthanassa, 342
Anelosimus, 61 annonae (Euxesta), 223 ANTHICIDAE, 149
anembatos (Ectotrypa), 371 annonae (Tenuipalpus), 50 Anthicus, 149
Anetia, 341 annosa† (Lobopoda), 145 Anthocoptes, 48
Aneurus, 247 annulata (Brachymeria), 291 ANTHOCORIDAE, 245
angelia (Electrostymon), 339 annulata (Careospina), 235 ANTHOMYIIDAE, 216
angelica (Boenasa), 349 annulatus (Anicetus), 291 Anthomyza, 225
angelicus (Pseudaphycus), 292 annulicornis (Estola), 154 ANTHOMYZIDAE, 225
angelicus (Sciapus), 212 annulifera (Mouralia), 357 Anthonomus, 169
angelo (Omalodes), 109 annulipes (Amphiacusta), 373 Anthophora, 316
anglicanus (Tutaibo), 64 annulipes (Craspedochaeta), 225 Anthophorula, 317
anguinus (Lispinus), 112 annulipes (Leptostylopsis), 153 anthracinus (Derancistrus), 156
angulata (Poecilosomella), 228 annulipes (Lethocerus), 240 Anthrax, 210
angulatus (Acinopterus), 265 annulipes (Pagurus), 39 anthrenoides (Trogoderma), 133
angulatus (Aeolus), 128 annulus (Eucatops), 111 ANTHRIBIDAE, 164
angulatus (Aglyptinus), 110 ANOBIIDAE, 134 ANTHURIDAE, 31
angulatus (Alpheus), 36 Anoblepsis†, 286 Antianthe, 270
angulatus (Modisimus), 58 Anochetus, 304 Antias, 243
angulatus (Ormiscus), 165 anodontos (Apotomoderes), 173 Antiblemma, 351
angulatus† (Lorelus), 143 anodontotus† (Lemmasomus), 171 antica (Jadera), 254
angulatus‡ (Caudites), 24 anomala (Atheta), 111 antica† (Leptopharsa), 242

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 578
Anticarsia, 351 antilleanus† (Culicoides), 196 antiquum† (Nesocyrtosoma), 147
Anticedusa, 260 Antillena, 56 antiquum† (Pison), 313
anticella (Oncolabis), 329 Antillengis, 139 antiquus† (Ornithodorus), 52
antictypos (Anaxipha), 375 antillensis (Clibanarius), 39 antiquus† (Pseudomyrmex), 306
anticua† (Pheidole), 303 antillensis (Dicranocentrus), 95 antiquus† (Trinemurodes), 97
anticus† (Diaprepes), 173 antillensis (Enoplometopus), 37 Antisolabis, 183
antidupos (Anaxipha), 375 antillensis (Laccobius), 108 Antonina, 279
antigone (Buenoa), 241 antillensis (Metadorylas), 215 anulatus (Oedudes), 155
antigonus (Hemerobius), 360 antillensis (Microphrys), 42 Anurogryllus, 372
antiguensis (Cyptocephala), 256 antillensis (Phthitia), 228 anurum (Belostoma), 240
antiguensis (Neostasina), 73 antillensis (Synalpheus), 36 Anyphaena, 70
antiguus† (Cenocephalus), 182 antilleorum (Anthicus), 149 ANYPHAENIDAE, 70
antiguus† (Phloeotribus), 177 antilleorum (Lithurgus), 316 Anyphaenoides, 71
Antillacris, 366 antilles (Streptopsyche), 387 Anypsipyla, 328
antillalis (Herpetogramma), 325 Antillesia, 17 Aonidiella, 276
antillana (Chrysopa), 359 antilleus (Hemipteroseius), 52 aonidium (Chrysomphalus), 277
antillana (Dorymyrmex), 298 antillia (Diaphania), 325 Aonidomytilus, 278
antillana (Dysderina), 59 antilliana (Typhlocypris), 21 Apachekolos, 209
antillana (Eperigone), 64 antilliana† (Alumeda), 245 Apallates, 226
antillana (Hentzia), 76 Antilliclodes, 374 Apalonychus, 119
antillana (Leptogenys), 305 antillicus (Hyporhagus), 143 Apanteles, 286
antillana (Lyroda), 312 antillicus (Pityophthorus), 181 apantelivorus (Horismenus), 293
antillana† (Oxyidris), 304 antillicus (Sphenoceros), 181 Apate, 134
antillanum (Theridion), 63 antillicus† (Cenocephalus), 182 Apenes, 103
antillanus (Lyssomanes), 76 antilliensis (Anastrepha), 223 aper (Dipropus), 128
antillanus (Myopsocus), 382 Antillipeltis, 135 aperta (Caribovia), 264
antillanus (Stenocaecilius), 381 Antillisa, 346 aperta (Iscadia), 358
antillarum (Anopliomorpha), 151 Antilliscaris, 107 Aphaenogaster, 303
antillarum (Boreofairchildia), 194 Antillitermes, 189 Aphaereta, 285
antillarum (Callimantis), 190 Antillocladius, 198 APHALARIDAE, 271
antillarum (Chaetodactylus), 51 Antillocoris, 250 Aphanisticus, 124
antillarum (Drepanaporus), 307 Antillognatha, 64 aphanton (Cycloptilum), 376
antillarum (Epierus), 110 Antilloides, 57 Apharosoma†, 171
antillarum (Eulimnadia), 14 Antillolpium, 79 Apharus, 115
antillarum (Onthophagus), 120 Antillonirvana, 267 aphaura (Anaxipha), 375
antillarum (Panchlora), 186 Antillopsyche, 387 apheles‡ (Cytheromorpha), 25
antillarum (Paratyndaris), 126 Antillosciocoris, 257 APHELINIDAE, 289
antillarum (Parvitermes), 189 Antillosorius, 112 Aphelinus, 289
antillarum (Tetyra), 258 Antillostenochrus, 81 aphelofacies† (Pityophthorus), 181
antillarum (Tytthonyx), 132 Antillotettix, 365 Aphelopus, 310
antillarum† (Mirhipipteryx), 365 Antillotrecha, 83 aphenges (Hapithus), 370
Antillattus, 74 antillummagnorum (Culex), 200 APHIDIDAE, 273
Antillea, 342 antillus (Blissus), 248 Aphis, 273
antillea† (Atlantitermes), 189 Antillusa, 111 Aphrastomyia, 202
antillea† (Mycetobia), 201 antimimon (Cycloptilum), 376 Aphrissa, 338
antillea‡ (Bairdia), 19 antiphates (Ocyptamus), 216 Aphthona, 157
antillea‡ (Loxoconcha), 25 antiphonos (Hapithus), 371 Aphylla, 363
antillea‡ (Paranesidia), 19 antiqua antiqua (Pyrrhocalles), 336 Aphytis, 289
antillea‡ (Quadracythere), 26 antiqua† (Bradysia), 203 apian (Huaca), 169
antillea‡ (Xestoleberis), 28 antiqua† (Dominiperla), 378 apiata (Cyrtoxipha), 375
antilleana (Moodna), 329 antiqua† (Problepharoneura), 222 apicale sulphurea (Myzine), 308
antilleana† (Dasyhelea), 197 antiqua† (Pterosclerogibba), 311 apicalis (Acanthoscelides), 163
antilleana† (Forcipomyia), 197 antiquarius† (Microcorthylus), 181 apicalis (Acrosticta), 223
antilleana† (Stilobezzia), 197 antiquarius† (Pityophthorus), 181 apicalis (Agrotis), 357
antilleanus (Dryinus), 311 antiquua† (Coniopteryx), 360 apicalis (Centris), 316
antilleanus (Nesepyris), 310 antiquum† (Cryptocheiridium), 79 apicalis (Coproporus), 118

579 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

apicatus (Brachyvatus), 100 ARANEIDAE, 65 ARGASIDAE, 51
apicecoloratus (Ataenius), 120 araneiformis (Neoclytus), 150 argentata (Argiope), 66
apicicornis (Clerada), 250 araneiformis araneiformis (Lagocheirus), argentata (Cicindela), 101
Apicrenus†, 246 153 argentata (Porphyrochroa), 212
apiculata (Ghilianella), 246 Araneometa, 66 argentatus (Anthonomus), 169
APIDAE, 316 Araneus, 66 argentatus (Apodrosus), 175
Apiomerus, 246 arapha (Chalcis), 291 argentatus (Leptostylopsis), 153
APIONIDAE, 167 Araptus, 180 argentea (Adelphe), 310
Apionion, 167 Aratus, 41 argenteosetosa (Silis), 132
Apis, 316 arawakan (Elaver), 71 argentifolii (Bemisia), 272
apithanon (Ectotrypa), 372 Arawakis†, 135 argentina (Fundella), 329
Apleurotropis, 292 arboreum (Planctolpium), 79 argentiventris (Cephalotes), 301
Aplopododesmus, 88 arboreus (Microcerotermes), 190 argentum† (Tainopompilus), 308
Apocephalus, 214 arcana (Alisotrichia), 388 ARGIDAE, 282
Apodrosus, 175 arcana† (Archicratyna), 203 argillacea (Alabama), 351
Apogeshna, 324 arcanum (Trichapion), 167 argillaceus (Acanthoscelides), 163
Apogonalia, 264 arcas (Calisto), 342 Argilloecia, 26
apolithomenus† (Neoxantholinus), 117 arcasalis (Acrolophus), 319 Argiope, 66
Apollophanes, 73 Archaeoellipes, 365 Argon, 336
apoplanos (Amphiacusta), 373 Archaeopolietes†, 218 Argopistes, 157
apopnus† (Tetrapus), 289 Archaeoprepona, 341 Argoravinia, 219
Aporinellus, 307 Archaeopristocera†, 309 argus (Panulirus), 38
Apotoforma, 332 archaia† (Milichiella), 227 argutus (Lygaeus), 249
Apotomoderes, 173 archboldi (Typopsilopa), 234 argyra (Agriognatha), 64
Apoweckelia, 29 Archeatelura†, 97 argyra (Leucauge), 65
appendiculata (Corethrella), 199 archebates (Calisto), 342 Argyrodes, 61
appendiculata (Turpilia), 368 archeodominica (Copidosoma), 291 Argyrogramma, 357
appendiculatus (Enicospilus), 288 Archicratyna†, 203 argyrosoma† (Eulechriops), 170
appendigaster (Evania), 288 Archimestra, 340 argyrostilba (Glenognatha), 65
applanatus (Antillatus), 74 Archiokrisna, 266 Argyrotaenia, 332
approxima (Phyllophaga), 121 ARCHIPSOCIDAE, 381 Arhopalus, 149
Apraea, 157 Archipsocus, 381 Ariadna, 59
apricaria (Chaetocnema), 157 Archisepsis, 224 Ariamnes, 61
Aprostocetus, 294 Archytas, 220 ariasae (Colapteroblatta), 185
Apsectus, 134 arcifer (Toxomerus), 216 aridula (Balclutha), 266
apseudes (Anaxipha), 375 arctella (Anegcephalesis), 328 aridus (Carpelimus), 113
Apsilophrys, 291 Arctodiaptomus, 16 aridus (Scolytodes), 179
apsilutea (Craspedochaeta), 225 Arctosa, 68 Arilophia, 333
aptera (Chalcosicya), 162 arcus (Osbornellus), 267 arimao (Schizogenius), 107
Apterostigma, 300 Ardeutica, 331 arioles (Priocnemis), 307
Aptilotella, 228 Ardistomis, 106 arispilos (Hoplisoides), 312
Aptunga, 328 arena (Anaxipha), 375 ARISTEIDAE, 35
aquaecadentis (Alisotrichia), 388 Arenaeus, 43 aristodemus aristodemus (Heraclides),
Aquarius, 239 arenalis (Glympis), 352 338
aquatile (Pterogramma), 228 arenaria (Halocoryza), 106 aristor (Heraclides), 338
arabesca (Neoscona), 67 arenata (Verrucosa), 68 Aristotelia, 321
arabidis (Asterolecanium), 276 arenosa (Parabezzia), 196 arizela† (Alisotrichia), 388
Arachnocoris, 245 arenosa (Phurys), 352 Armases, 41
Arachnolithulus†, 57 areolata (Entylia), 271 armasi (Arucillus), 77
aracynthia (Myscelia), 340 areolatus (Doryctobracon), 286 armasi (Ataeniopsis), 120
ARADIDAE, 247 areolatus (Paramacrobiotus), 14 armasi (Caribeacarus), 52
Aradophagus, 283 Areva, 349 armata (Archisepsis), 224
Araecerus, 164 arganoi (Mexicolaophonte), 17 armata (Ceratophysella), 94
araguaiana (Sthenaridea), 244 Arganotus, 78 armatipennis (Cercyon), 109
Aralichus, 51 argante rorata (Phoebis), 338 armatipes (Dieuches), 252
Araneagryllus†, 374 Argas, 51 armatus (Alpheus), 36

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 580
armatus (Euscelus), 166 asperatus (Caribbomerus), 150 atlanticus (Chramesus), 177
armatus (Jollas), 76 aspericornis (Mesocyclops), 16 atlanticus (Phloetribus), 177
armatus (Odontobracon), 286 Asphondylia, 202 atlanticus (Scolytodes), 179
armatus (Petrolisthes), 40 Aspidiella, 277 atlanticus (Theoborus), 180
armatus (Tytthonyx), 133 Aspidiotus, 277 atlantidis (Solenocera), 35
armatus (Zanysson), 312 aspidistrae (Pinnaspis), 278 Atlantitermes, 189
armigera (Acanthocheila), 242 Aspidoglossa, 106 atlas atlas (Polyancistrus), 368
armila (Calliscelio), 283 Aspiduchus, 185 atlas pilanus (Polyancistrus), 368
armillata (Microsepsis), 224 Aspisoma, 129 atomaria (Phanerota), 112
armipes (Cophes), 171 assimile (Sceliphron), 314 Atomosia, 208
arnaudi (Leschenaultia), 220 assimilis assimilis (Gryllus), 372 Atomothenemus, 178
arndti (Geogryllus), 372 assis (Apleurotropis), 292 atomus (Hypothenemus), 178
arndti (Leptoteleia), 284 assoi (Diastolinus), 145 Atoniomyia, 208
Arocera, 256 assoi (Hypselodesmus), 88 Atopocixius, 261
aroidephagus (Crenidorsum), 273 Astata, 312 ATOPOMELIDAE, 51
Arphthicarus, 47 Asteia, 225 Atopopsyllus†, 382
arragensis (Rotundabaloghia), 54 ASTEIIDAE, 225 Atopsyche, 390
Arrenurus, 49 Astenus, 114 atra (Athyroglossa), 232
arsinotus† (Geratozygops), 170 Asteriza, 159 atra† (Brachymetroides), 239
Artema, 57 Asterocampa, 340 Atractocerus, 135
Artemia, 14 asterolecanii (Coccophagus), 290 Atractomerus, 169
ARTEMIIDAE, 14 ASTEROLECANIIDAE, 276 atramentarius (Lachnopus), 174
Arthmius, 115 Asterolecanium, 276 atrata (Campsomeris), 308
Arthromastix, 324 asthenicus (Horistonotus), 128 atratus (Culex), 200
Arthrostictus, 102 Astraptes, 337 atratus (Exitianus), 266
Arthrovertex, 48 asuai (Hologaeus), 104 atratus (Monodesmus), 155
arthuri† (Myopsocus), 382 asymmetrica† (Gnophomyia), 192 atratus (Zophobas), 145
articulatus (Geron), 211 asymmetrica‡ (Cardobairdia), 19 atrechoensis (Achromoporus), 87
articulatus (Selenaspidus), 278 ata (Halysidota), 348 atriantenus† (Toxorhynchus), 167
Artipus, 174 atabey (Sipho), 369 atriartus† (Toxorhynchus), 167
artus (Apodrosus), 175 Ataeniopsis, 120 Atrichobrunettia, 195
Arucillus, 77 Ataenius, 120 atrichomatus (Phormictopus), 56
Arvelius, 256 atalanta rubria (Vanessa), 342 Atrichopogon, 197
asander haitensis (Aguna), 337 Atalopedes, 336 atricolor (Letis), 353
Asca, 52 Atanygnathus, 117 atricornis (Pluto), 313
Ascalapha, 351 atavus† (Lispinomimus), 112 atrifacies (Soberocephala), 226
ASCALAPHIDAE, 358 Atax, 49 atrimana (Palpada), 215
Ascia, 339 Ataxia, 155 atriscuta (Nostima), 234
ASCIDAE, 52 ATEMNIDAE, 80 atriventris armstrongi (Thyreodon), 288
Asciodes, 324 ater (Altisessor), 139 atrocrocea (Lychnacris), 130
ascita (Allograpta), 216 ater (Dermestes), 133 atrohumeralis atrohumeralis (Pamphan-
asclepia (Micromyrmex), 172 ater (Hydrophilus), 108 tus), 249
Ascorhynchus, 45 ater (Styraxodesmus), 90 atrohumeralis dominicanus (Pamphan-
ascutum† (Veterator), 72 aterrimus (Pogonomyrmex), 303 tus), 249
ASELLIDAE, 32 athanes (Anaxipha), 375 atromaculatus (Caligus), 18
Asellus, 32 Atherigona, 218 Atropacarus, 47
asellus (Oniscus), 33 Atheta, 111 atropicta (Ozophora), 251
ashei† (Caulophilus), 170 Atholus, 109 atropos† (Geratozygops), 170
ashmeadi (Neorileya), 295 Athyroglossa, 232 atrosus (Ommatius), 209
ashmeadi (Pseudoxenos), 383 Atimiola, 154 atrovittatus (Styloleptus), 153
ASILIDAE, 207 Atissa, 233 atrox (Zilus), 141
asini (Haematopinus), 379 atlanta (Artema), 57 Attagenus, 133
Asiomorpha, 89 ATLANTASELLIDAE, 32 attala (Erythemis), 363
Aspathines, 143 Atlantasellus, 32 Attalomimus, 136
asper (Paectes), 355 Atlantea, 342 ATTELABIDAE, 166
asperata (Mecaphesa), 74 atlantica (Plectana), 66 Attelabus, 166

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attenboroughi† (Electrotettix), 365 auspicatus (Pityophthorus), 181 Bahadzia, 29
attenuata (Elachiptera), 227 australasiae (Periplaneta), 184 bahamaensis (Guttipsilopa), 231
attenuatella (Cosmopterix), 322 australe (Theridion), 63 bahamensis (Culex), 200
Atteva, 322 Australephestiodes, 328 bahamensis (Lygaeus), 249
Attheyella, 17 australica (Melittobia), 294 bahamensis (Paraglenanthe), 232
attritus (Ochthebius), 110 australis (Acrolophus), 319 bahamensis (Petrochirus), 39
Atya, 36 australis (Haldorus), 266 bahamica (Exomalopsis), 317
ATYIDAE, 36 australis (Hydrometra), 239 bahamicus (Decadiomus), 141
atymnusalis (Bleptina), 356 australis (Tauriphila), 364 bahoruco (Agobardus), 74
audanti (Oecobius), 60 Australolimnophila, 193 bahoruco (Calisto), 342
audanti (Philonthus), 117 Austrophthiracarus, 47 bahoruco (Charinus), 54
audanti (Phyllophaga), 121 Austrotinodes, 386 bahoruco (Tityus), 82
audanti (Stilomedon), 115 Autoplusia, 357 Bahorucocricus, 92
audanti (Xantholinus), 118 avernus (Pseudospirobolellus), 93 bahorucoensis (Beatadesmus), 88
auge (Cosmosoma), 348 averrhoae (Phytoseius), 53 bahorucoensis (Lychnacris), 130
Augesmus, 89 avidus (Ctenus), 69 Bahorucotettix, 365
Augochlora, 315 avitinquilinus† (Termitaradus), 275 Bairdia, 19
Augocoris, 258 avitus† (Hypothenemus), 178 bairdii (Cryptosoma), 40
augustei (Camponotus), 299 avitus† (Pseudomyrmex), 306 bairdii‡ (Cryptosoma), 40
Aulacaspis, 278 avium (Strebla), 217 BAIRDIIDAE, 19
Aulacigaster, 225 avrilensis (Phalloniscus), 33 Bairdoppilata, 19
AULACIGASTRIDAE, 225 avus† (Electropteron), 129 bakeri (Anthocoptes), 48
Aulacorthum, 274 ayiti (Erechthis), 368 bakeri (Oxysarcodexia), 219
aulicus (Lachnopus), 174 ayitiensis (Nemocephalus), 168 bakeri (Tenuipalpus), 50
aulicus (Oncopeltus), 250 ayti (Cerceris), 313 Bakeriella, 309
Auplopus, 307 azari† (Tityus), 82 BALANIDAE, 15
aurantianum (Gymnandrosoma), 331 Azeta, 351 Balanus, 15
aurantii (Aonidiella), 276 azia (Ministrymon), 339 Balclutha, 266
aurantii (Toxoptera), 274 Azilia, 64 Balega, 115
aurantius (Paraphidippus), 76 Azochis, 324 Baliomydas, 211
aurata (Lampetis), 125 Azteca, 298 balli (Agalliopsis), 263
aurata domingoensis (Psiloptera), 125 aztecus (Farfantepenaeus), 35 balli (Dictyla), 242
aurea (Argoravinia), 219 azua (Calisto), 342 balma (Hypogrammodes), 353
aureoinclusa† (Setodes), 390 azuae (Hoplisoides), 312 baloghi (Arthrovertex), 48
aureolus (Euscelus), 166 azuensis (Diastolinus), 144 balteata (Macrodiplax), 363
aureomicans (Psiloptera), 125 azuensis (Erythrolychnia), 130 balteatus (Cryptocephalus), 161
aureomixtus (Hormotrophus), 173 azurea (Silis), 132 balteatus (Leptoglossus), 253
aureopilosus (Trichoprionus), 156 azureus† (Tyrtaeus), 146 bambusae (Bambusaspis), 276
aureus (Melanopleurus), 249 Azygopleon, 32 bambusae (Leptodictya), 242
auricoma (Elaphidion), 151 babyrussus† (Eocader), 242 Bambusaspis, 276
auricomus (Cephalotes), 301 bacca (Elytrogona), 160 Banasa, 256
auricosta (Epiphragma), 193 bacharenge (Oosternum), 109 Bandara, 265
auricula† (Antillopsyche), 387 bachata (Drepanaporus), 307 banesensis‡ (Loxoconcha), 25
auricula‡ (Aurila), 23 bachata (Modisimus), 58 bani (Centruroides), 81
auricularia (Caribecandona), 20 bacoa (Mimapsilopa), 232 bani (Metepeira), 67
aurifera (Lampetis), 125 Baconia, 109 bani (Selenops), 73
auriferalis (Tortyra), 322 Bactra, 331 Baniana, 351
Aurila, 23 bacula (Mesoplophora), 47 baniensis (Ceratocanthus), 119
auronotatus (Actenodes), 125 Badister, 104 baniensis (Tytthonyx), 133
auropunctata (Wasmannia), 301 baehri† (Riegerochterus), 241 banii (Euconnus), 116
aurora (Hemeroplanis), 353 Baeodasymyia, 196 Banisia, 323
aurora† (Epiphragma), 193 Baeotettix†, 365 banksi (Anasaitis), 74
auroseta (Ozophora), 251 BAETIDAE, 235 banksi (Eutetranychus), 50
aurulenta (Cariboptila), 388 Baetis, 235 banksi (Smicridea), 387
aurulentus (Toxomerus), 216 Baeus, 283 banksi (Thymoites), 63
aurulentus aurulentus (Epitragus), 144 Bagisara, 350 banksi (Ululodes), 358

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 582
banksiana (Geranomyia), 194 basifulvus (Leptostylopsis), 153 bella (Frontinella), 64
baorucae (Kiskeya), 158 basilaris (Madoniella), 135 bella (Gymnoclasiopa), 231
baorucensis (Canthonella), 121 basilense (Nesocyrtosoma), 147 bella (Utetheisa), 347
baorucensis (Platynus), 105 basilensis (Peripatus), 13 bellator (Anasa), 253
baorucensis (Poecilantrops), 228 Basileunculus, 214 bellator (Miogryllus), 372
baorucensis (Psorolyma), 140 basilica (Polymorphomyia), 222 bellatrix (Anacaona), 368
baoruco (Cyphomyia), 206 basillaris (Thermonectus), 100 bellimanus (Amblypsilopus), 212
baoruco (Jurinella), 220 basiplaga (Oraesia), 354 bellingeri (Indotritia), 47
baoruco (Micropholcus), 57 basitestaceus (Tylocerus), 132 bellipulex‡ (Costa), 27
baoruco (Neostasina), 73 Basslerites, 27 Bellitudo, 272
baoruco (Phyllophaga), 121 bastardoae (Achromoporus), 87 bellorum (Brasiella), 101
baoruco (Tilloclytus), 150 bastardoae (Malacomorpha), 378 bellulus (Actenodes), 125
baorucoensis (Lophocampa), 348 bastardoae (Robopus), 131 bellulus (Misumenops), 74
baorucoensis (Microdiphot), 131 bastardoae (Scapsipedus), 372 bellulus (Tityus), 82
baorucoensis (Notaepytus), 139 bastardoanus (Ommatius), 209 bellus (Auplopus), 307
baorucoensis (Ochrotrichia), 389 bastardoi (Robopus), 131 bellus (Conoderus), 128
baracoa loma (Polites), 336 bastardoi (Scapsipedus), 372 bellus (Hapithus), 370
barahona (Microtityus), 82 batangensis (Anoplodactylus), 45 bellus (Lachnopus), 174
barahona (Stenotabanus), 207 batatae (Euscepes), 171 bellus (Osbornellus), 267
barahona (Yoyuteris), 374 batatas (Spartocera), 254 bellus (Scolytodes), 179
barahonae (Heteronebo), 83 batesi (Atopsyche), 390 Belonuchus, 117
barahonaensis (Tytthonyx), 133 batesi (Calisto), 342 Belopherus, 168
barahonense (Metachroma), 162 batesi (Calosopsyche), 387 Belorhynchus, 168
barahonensis (Dominicanocoris), 319 batesi (Monobia), 308 Belostoma, 240
barahonensis (Eurycotis), 184 batesi (Ochlodes), 336 BELOSTOMATIDAE, 240
barahonensis (Leptolycus), 129 batesi (Shannonomyia), 193 belusalis (Botys), 325
barahonensis (Synalpheus), 36 batesi (Stenotabanus), 207 Belvosia, 220
barba (Polytrichophora), 232 Bathydema, 250 Bema, 328
barbarae (Macrobiotus), 13 Batina, 351 Bembidion, 107
barbata (Homola), 41 Battus, 338 Bembix, 312
barbata (Pseudolycoriella), 205 bauri (Odontomachus), 305 Bemisia, 272
barbatus (Caribothyrus), 54 bautistai (Sicoderus), 176 beneficus (Ufens), 297
barbatus (Chalepides), 123 bayaguana (Achromoporus), 87 benneti (Phytoseius), 53
barberi (Limnichoderus), 126 bayahibe (Dellia), 366 bennetti (Deuterixys), 286
barberi (Pamphantus), 249 bayahibe (Tainonia), 58 bennetti (Horismenus), 293
barberi (Puto), 280 bazochii gundlachianus (Strymon), 339 bennetti† (Caulophilus), 170
barberi (Zorotypus), 391 Bdella, 48 bensoni‡ (Cyprideis), 21
barbosai (Culicoides), 196 BDELLIDAE, 48 Bephratelloides, 294
barbuti (Antillisa), 346 beali† (Amberoderma), 133 bequaerti (Leptoconops), 197
Baris, 169 Beameromyia, 209 bequaerti (Incisitermes), 188
baros† (Pseudomyrmex), 306 Beatadesmus, 88 berac (Modisimus), 58
barrera (Curtara), 267 beattyi (Ataenius), 120 berenice (Erythrodiplax), 363
Barroeuplectoides, 115 beattyi (Carpelimus), 113 bergrothi (Rheumatobates), 239
Barronopsis, 69 beattyi (Xantholinus), 118 berlandi (Cyclosa), 66
barrosa (Phyllophaga), 121 beauvoisi (Chelonarium), 127 berlesei (Encarsia), 290
barrowsi (Barronopsis), 69 beauvoisii (Discoderus), 102 bermudensis (Mothocya), 31
bartletti (Enochrus), 108 bechlyi† (Carventius), 248 bermudezi (Antilliclodes), 374
Barychara, 111 beckae (Polypedilum), 198 bermudezi (Neostasina), 73
BARYCHELIDAE, 56 beckeri (Araptus), 180 bermudezi crebripustulosa‡ (Trachyleb-
Baryconus, 283 beckeri† (Stichotrema), 383 eris?), 27
Barylaus, 105 Beckeriella, 232 bermudezi‡ (Munseyella), 26
Baryscapus, 294 Beckerina, 214 bernsteinae† (Megaselia), 214
basalis (Morellia), 218 beckii (Lepidosaphes), 278 Berosus, 108
basalis (Systena), 159 beebei (Teleophrys), 42 Bertamyia, 215
basalis (Xyonysius), 250 Belaphopsocus, 380 bertrandae (Caribarctia), 347
Basalys, 288 BELIDAE, 168 BERYTIDAE, 248

583 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Beskia, 220 Bidessonotus, 100 bipunctata (Opharus), 349
bestiola (Nesocyrtosoma), 147 bidigita (Agalliopsis), 263 bipunctatum (Simulium), 201
BETHYLIDAE, 309 Bidoma, 77 bipunctatus (Nephotettix), 267
beugelorum† (Nilakantha), 76 bifasciata (Caribovia), 264 bipunctatus (Xestocephalus), 269
beynai† (Centruroides), 81 bifasciata (Comperiella), 291 bipunctella (Graminella), 266
beynai† (Selenops), 73 bifasciatus (Laccophilus), 101 bipustulata (Paraclivina), 107
Bezzia, 197 biferana (Hemerophila), 322 biramosus (Platynus), 105
BIANTIDAE, 77 bifibulus (Euschistus), 256 biramosus transcibao (Colpodes), 105
Biaporus, 87 bifida (Edessa), 255 Birandra, 155
biarmatus (Antillosorius), 112 bifidalis (Achyra), 324 birgei (Arrenurus), 49
bibadia (Nylanderia), 300 bifidus (Elcaribe), 212 biscaynensis (Culex), 200
BIBIONIDAE, 202 biformis (Furcaspis), 277 biscutatum (Homoschema), 158
Biblis, 340 bifoveatum (Monarthrum), 181 biseriatum (Menopon), 379
bibullus† (Piesocorynus), 164 bifoveatus (Conoderus), 128 biseriatus (Dicranocentrus), 95
Bicalcasura†, 172 bifrons (Bifronsina), 228 bisetosa (Alisotrichia), 388
bicarinatum (Tetramorium), 304 bifrons (Spelobia), 229 bisetosa (Eustala), 66
bicavatus (Dissomphalus), 310 Bifronsina, 228 bisignata (Argyrotaenia), 332
bicelliforma‡ (Perissocytheridea), 22 bifurca (Allocyclosa), 66 bispatha (Coproica), 228
biceps (Neoathyreus), 119 bifurcata (Cyclosa), 66 bispinatus (Xyleborus), 180
bichensis‡ (Quadracythere), 26 bifurcata (Neotrichia), 389 bispinosus (Dysmicoccus), 279
Bicilia, 324 bifurcatus‡ (Psygmophthalmus), 44 bispinus† (Paleophthorus), 182
bicincta (Prosapia), 262 bigibber† (Meioneta), 64 bistictus (Pureleon), 361
bicinctum (Trogoderma), 133 bigibbosa (Chrysometa), 65 bistriatus (Cafius), 117
bickleyi (Proctolaelaps), 52 bignoniae (Horistonotus), 128 bistriga (Bolina), 353
biclavis (Howardia), 278 biguttatum (Callidium), 150 bisulcata (Synoditella), 284
bicolor (Alagoasa), 157 biguttatus (Monobelus), 270 Bitoma, 143
bicolor (Bryoporus), 118 biimpresa (Hypogena), 145 bituberculatum (Hovorodon), 155
bicolor (Drapetes), 129 biimpresus (Pycnomerus), 143 bituberculatus (Charaphloeus), 138
bicolor (Hypselodesmus), 88 bijuduvali‡ (Cytheropteron), 22 bituberculatus (Epialtus), 42
bicolor (Mesocheira), 317 bilbisaria (Caberodes), 334 bituberculatus† (Mimetus?), 60
bicolor (Microchrysa), 206 bilimeki (Pheidole), 303 Bituryx, 348
bicolor (Monomorium), 303 bilineata (Tapinopa), 64 biunguiculatus (Brachycarpus), 37
bicolor (Neostasina), 73 bilineata (Zygothrica), 231 biustus (Styloleptus), 153
bicolor (Rileya), 295 bilineatus (Metacynortoides), 78 bivittata (Melipotis), 353
bicolor (Schizophragma), 295 bilobata (Neopamera), 251 bivittata (Soberocephala), 226
bicolor (Stirellus), 266 bilobus (Nettarhinus), 176 bivittatum (Acalymma), 159
bicolor (Symploce), 186 bilyi (Neotrachys), 124 bivittatus (Glyphonyx), 129
bicolor (Trachelas), 72 bilyi (Polycesta), 126 bivittatus (Menemerus), 76
bicolor (Trichrous), 152 bimaculata (Aleochara), 111 BLABERIDAE, 185
bicolor (Trichrous), 152 bimaculata (Oedancala), 250 Blaberus, 185
bicolor† (Corthylites), 182 bimaculatus (Caccodes), 132 blackwelderi (Anthicus), 149
bicolorata† (Proanaxipha), 373 bimaculatus (Tetranychus), 50 Blaesoxipha, 219
biconicus† (Cenocephalus), 182 bimarginatus (Osbornellus), 267 blaini (Aellopos), 344
biconigerus† (Chrosiothes), 61 bimini (Cyptocephala), 256 blakeae (Asteriza), 160
bicornata (Orekthophora), 270 binotata (Tramea), 364 blancoi (Curtara), 268
bicornis (Phrygionis), 334 binotatus (Colymbetes), 100 blanda (Meta), 66
bicornis (Pseudolycoriella), 205 binotatus (Platyscytus), 244 blandita (Blaesoxipha), 219
bicornutus (Microphrys), 42 binotatus (Rhantus), 100 blandus (Nops), 59
bicuspidatus (Haplopus), 377 binotatus (Suphisellus), 100 Blapstinus, 144
Bicyrtes, 312 Biocrypta, 114 Blaste, 382
bidegradata (Hemipenthes), 210 biorbis (Fecelia), 256 Blastopsocus, 382
bidens (Adelina), 146 biovata (Soberocephala), 226 Blatta, 184
bidens (Elaphoidella), 17 bipartita (Erythrolychnia), 130 Blattella, 186
bidens (Toxonotus), 165 bipartites (Sabethes), 201 BLATTIDAE, 184
bidens bidens (Gonodonta), 352 bipenicilla (Cryptaspasma), 331 BLATTISOCIDAE, 52
bidentatus (Plectromerus), 150 bipunctalis (Herpetogramma), 325 blechi (Liriomyza), 225

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 584
Bledius, 113 Bothriomyrmex, 298 brandaoi† (Micropygomyia), 194
Bleptina, 356 botosaneanui (Anommatelmis), 126 brandeni (Haliplus), 99
BLISSIDAE, 248 botosaneanui (Tubomixdea), 49 brandti† (Amberobyrsa), 242
Blissus, 248 botydata (Ptychamalia), 335 branhami (Robopus), 131
Bloodiella, 297 bouceki† (Gummilumpus), 297 branneri (Callianassa), 38
bloosi† (Cephalotes), 301 bourkei (Rekoa), 339 Brasa, 264
bocacanadensis (Selenops), 73 bourreriae (Dendrosinus), 177 Brasiella, 101
Bocchoropsis, 324 bousfieldi (Synalpheus), 36 brasiliensis (Atractocerus), 135
Bocchus, 311 bouvieri (Dasymutilla), 306 brasiliensis (Cerataphis), 275
BOCHICIDAE, 79 Bovicola, 379 brasiliensis (Hecamede), 232
Boenasa, 349 bovis (Bovicola), 379 brasiliensis (Microparsus), 274
Boettcheria, 219 bowditchi (Yingaresca), 159 brasiliensis (Picturaphis), 274
Bogidiella, 29 bowmani (Renocila), 32 brasiliensis (Plesiochrysa), 359
BOGIDIELLIDAE, 29 bowreyi (Pseudischnaspis), 278 brasiliensis (Prospaltella), 290
bogotensis (Anisepyris), 309 bozzolai (Endroedianibe), 123 brasiliensis (Protalebrella), 268
Bohartilla, 383 brachialis (Platyvelia), 240 brassicae (Brevicoryne), 274
BOHARTILLIDAE, 383 Brachinus, 101 brassicae (Liriomyza), 225
bohio (Calosopsyche), 387 Brachybaenus, 377 BRATRACHEDRIDAE, 322
bohio† (Anoplotermes), 189 Brachycamacus†, 175 brazilorum† (Pintomyia), 195
boisduvallii (Diaspis), 278 Brachycarpus, 37 bredini (Allodardanus), 39
Bolacothrips, 384 Brachycentrotus, 270 bredini (Neogonodactylus), 28
Boletina, 202 Brachychara, 206 brenthus (Anthonomus), 169
Bolivaridia, 96 Brachycorynus, 165 BRENTIDAE, 167
Bolostromus, 56 Brachycythere, 27 Brentus, 167
Bolothrips, 385 Brachydeutera, 233 Brephidium, 339
BOMBYLIIDAE, 210 Brachyglutina, 115 Brepholoxa, 257
BOMOLOCHIDAE, 18 Brachymeria, 291 breva (Neokrisna), 267
Bonagota, 332 Brachymesia, 363 brevicauda (Radaweckelia), 30
Bonchis, 327 Brachymetroides†, 239 breviceps (Cymodema), 249
bondi (Centrolimnesia), 49 Brachymyrmex, 300 brevicercus (Apocephalus), 214
bondi (Eucyclops), 16 Brachyodina, 206 brevicollis (Ataenius), 120
bondi (Exosphaeroma), 33 Brachyomus, 173 brevicollis (Dicoelotrachelus), 159
bondi (Geogryllus), 372 Brachyplatys, 257 brevicornis (Glyptoscelis), 162
bondi (Spelaeala), 369 Brachypogon, 196 Brevicoryne, 274
bonettii (Heteroctenus), 82 Brachypremna, 194 brevicostata (Spinilimosina), 229
bonfils (Phyllophaga), 121 Brachypsectra, 127 brevida (Cubrasa), 264
Bonfilsus, 157 BRACHYPSECTRIDAE, 127 brevidactylus (Pagurus), 39
boninensis (Pseudococcus), 280 brachypterus (Haplopus), 377 brevidentatus† (Anochetus), 304
boninsis (Dysmicoccus), 279 brachypygaia‡ (Quadracythere), 26 breviembolus† (Nephila), 67
boops (Habroscelimorpha), 101 brachyrhinus† (Semnorhynchus), 172 breviforceps (Gonatopus), 311
Booralana, 31 Brachystomella, 94 brevinervis (Henicospilus), 288
BOPYRIDAE, 32 BRACHYSTOMELLIDAE, 94 Brevipalpus, 50
Bopyrione, 3s brachystyla (Coproica), 228 brevipalpus† (Episinus), 62
bordoni (Laccomimus), 100 Brachyufens, 297 brevipennis (Antiblemma), 351
Boreofairchildia, 194 brachyurum (Diaphanosoma), 14 brevipes (Dysmicoccus), 279
Borinquena, 235 Brachyvatus, 100 brevipes† (Pholcophora), 58
borinqueni (Culicoides), 196 Bracon, 285 breviptera (Espagnolopsis), 366
borinquinensis (Epinysson), 312 BRACONIDAE, 285 brevis (Cryptotermes), 188
bornoi bornoi (Pseudochrysops), 339 braconoides (Chimarra), 386 brevis (Ethirothrips), 385
Borophaga, 214 Bradleya, 26 breviscapus† (Stenotrupis), 170
bosci (Caenochrysis), 310 bradleyi (Munidopsis), 39 Brevisensoria, 248
boscii (Nesogenes), 258 bradycellinus (Tachys), 107 brevisensoriata† (Piezobarra), 165
Bosquetina, 27 Bradycellus, 102 brevisetosus (Cladoctonus), 177
BOSTRICHIDAE, 134 bradyi‡ (Hemicytherura), 23 brevispineus† (Cephalotes), 301
BOTHRIDERIDAE, 139 Bradysia, 203 brevistigmus (Conobregma), 287
Bothriocera, 258 Bradysiopsis, 204 brevistipitis† (Tetragoneura), 203

585 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

brevistriatum (Metachroma), 162 brunneus (Habronattus), 75 BYTHOCYTHERIDAE, 19
brevitarsus (Agobardus), 74 brunneus (Lyctus), 134 caab (Cariboptila), 388
brevitubus (Osbornellus), 267 brunneus (Megischus), 282 Cabares, 337
brevitubus (Trachythrips), 385 brunneus (Tytthonyx), 133 caborojensis (Erythrolychnia), 130
brevivena torrida (Dicranomyia), 193 brunnifrons† (Glabellula), 211 caborojo (Pelonium), 136
briarea briarea (Anetia), 341 brunnipennis (Turbopsebius), 206 caborojo (Zeadolopus), 110
brighti† (Electroborus), 182 bryantae (Agalenocosa), 68 caborojoanus (Euconnus), 116
broccai (Corythalia), 75 bryantae (Araneus), 66 caborojoensis (Ischnoptera), 186
broccus (Planophileurus), 124 Bryocamptus, 17 caborojoi (Euconnus), 116
brochymenae (Trissolcus), 285 Bryocyclops, 16 cabralense (Hapithus), 370
brodzinskii† (Brachypremna), 194 Bryodelphax, 13 cabrerensis (Haplaxius), 259
brodzinskyi† (Alienates), 238 Bryoporus, 118 cacata (Ephyrodes), 352
brodzinskyi† (Cessator), 129 Bubaces, 251 Caccodes, 132
brodzinskyi† (Culicoides), 196 bucarensis† (Kallyntrosternidius), 153 Caccoleptus, 133
brodzinskyi† (Dacne), 139 buccata (Pseudocycnoides), 18 cachonera (Ochrotrichia), 389
brodzinskyi† (Docodesmus), 90 Bucculatrix, 323 cachote (Achromoporus), 87
brodzinskyi† (Ochrotrichia), 389 bucidae (Colomerus), 48 cachote (Neostasina), 73
brodzinskyi† (Periscelis), 226 bucki (Soberocephala), 226 cachote (Ochyrocera), 57
brodzinskyi† (Stenotabanus), 207 buenae (Gonaives), 159 cachotensis (Truncattus), 76
brodzinskyorum (Canthochilum), 121 Buenoa, 241 cacique† (Anoplotermes), 189
brodzinskyorum† (Calisiopsis), 247 bulbipenis (Soberocephala), 226 Cacocharis, 331
brodzynskyi† (Caenocholax), 383 bulbus† (Velocitermes), 189 Cacodemonius, 80
bromeliae (Diaspis), 278 Bulia, 351 cacti (Chilocorus), 140
bromeliae (Drosophila), 230 bulla (Elytrogona), 160 Cactoblastis, 328
bromelicola (Corythalia), 75 bulla† (Anyphaenoides), 71 cactorum (Cactoblastis), 328
bromelicolus (Nesocyrtosoma), 147 bullalis (Madoniella), 135 cacumena (Ozophora), 251
Bromeloecia, 228 bullata (Carposina), 319 cadabra (Carylla), 369
bromia (Stenogryllus), 371 bullata (Efferia), 207 cadenati (Paguristes), 39
bromiliae (Melanaspis), 277 bullatum (Monarthrum), 181 cadma torrebia (Lucinia), 340
Brontaea, 218 bullatus (Dilophops), 89 caeciliana† (Borinquena), 235
Brontes, 137 BUPRESTIDAE, 124 CAECILIUSIDAE, 381
brontes haitiensis (Manduca), 344 Buprestis, 125 Caecophloeus, 146
broodbakkeri (Cyathura), 31 Bura, 141 CAECULIDAE, 48
browneanus (Bidessonotus), 100 Burca, 337 Caeculus, 48
browni† (Anthonomus), 169 burdettellus (Paedus), 329 caecus† (Ilemomyrmex), 304
browni† (Oeclidius), 261 burguersi (Uca), 42 caelata‡ (Cytherella), 21
bruchelus† (Cyptoxenus), 164 burmeisteri (Ozophora), 251 caelatipennis (Copelatus), 100
Bruchus, 163 Burtinus, 253 caelatus (Orthoporus), 93
bruesi (Ageniella), 307 burungae (Neohydatothrips), 384 caelatus‡ (Archaeopilumnus), 44
bruesi (Liobracon), 286 busccalis (Diaphania), 325 caementarium (Sceliphron), 314
bruesi (Microdon), 215 buscki (Antillosorius), 112 CAENIDAE, 235
brumpti (Hippelates), 227 buscki (Epiphragma), 193 Caenis, 235
bruneri (Frankliniella), 384 buscki (Homoschema), 158 Caenocholax, 383
brunettii (Stenotabanus), 207 buscki (Marietta), 289 Caenochrysis, 310
brunnea (Alisalia), 111 buscki (Pyragropsis), 183 Caenodelphax, 259
brunnea (Anasaitis), 74 buscki (Toxomerus), 216 caenozoica† (Mesembrinella), 219
brunnea (Flatoides), 261 busckiellum (Lasioglossum), 315 Caenurgia, 352
brunnea (Periplaneta), 184 bushi (Rhinton), 336 caerula (Nylanderia), 300
brunnea (Prytanes), 251 busu (Aculepeira), 65 caerulea (Aptilotella), 228
brunnea (Pseudothamiaraea), 112 BUTHIDAE, 81 caerulea (Scutellista), 296
brunnea (Rogeria), 304 buxi (Pinnaspis), 278 caeruleus (Entypus), 307
brunnescens (Smicridea), 387 Byrsochernes, 80 caeruleus (Sepedomerus), 224
brunnescens brunnescens (Chaetoc- Bythinogaster, 115 caerulum (Eumecosoma), 209
nema), 157 Bythoceratina, 19 cafetalera (Metrichia), 389
brunneum (Smodicum), 152 Bythocrotus, 75 Cafius, 117
brunneus (Brachinus), 101 Bythocypris, 19 cahobensis‡ (Perissocytheridea), 22

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 586
caicus (Phrixus), 345 Callida, 103 Camptopteroides, 295
cajanus (Pediobius), 293 Callidium, 150 camptus† (Caulophilus), 170
cajennense (Amblyomma), 52 calliergon calliergon (Dicranomyia), 193 Campylocythere, 27
cala† (Dipoenata), 61 calliergon polygrapha (Dicranomyia), Campylothorax, 95
CALAMOCERATIDAE, 390 193 Canacea, 226
Calappa, 40 Callimantis, 190 CANACIDAE, 226
Calappella, 40 Callinectes, 43 canaliculatus (Attelabus), 166
CALAPPIDAE, 40 calliomenae (Callionima), 345 canalinasus† (Apodrosus), 175
calathinus (Platynus), 105 Callionima, 345 canalis (Alatuncusia), 324
calcar† (Modisimus), 58 calliope (Cicadetta), 263 canariensis (Pseudolynchia), 217
calcarata (Grammonota), 64 Calliphora, 216 canavalia (Empoasca), 269
calcaroides† (Modisimus), 58 CALLIPHORIDAE, 216 Cancer, 40
calceatus (Artipus), 174 Callipogon, 155 cancer (Deinocerites), 201
calceolaris (Plusia), 357 Calliscelio, 283 cancerides (Phormictopus), 56
calcigena (Cariboptila), 388 callisto (Euxestonotus), 283 CANCRIDAE, 40
Calcinus, 39 Callopisma, 130 cancriformis (Gasteracantha), 66
calcitrans (Stomoxys), 218 Callopistria, 350 Cancrincola, 17
calcitrapa‡ (Paracytheridea), 25 callosipennis (Euscelus), 166 canda (Empoasca), 269
caldarium (Tetramorium), 304 Callosobruchus, 163 candacalis (Diaphantania), 325
caldwelli (Balclutha), 266 callosus (Canthon), 121 candida (Lymire), 348
caldwelli (Nilaparvata), 259 Calocosmus, 154 Candona, 20
Calepitrimerus, 48 Calolampra, 185 CANDONIDAE, 20
Cales, 289 Calosoma, 101 Candonopsis, 20
Caliadurgus, 307 Calosopsyche, 387 canescens (Lachnopus), 174
calidata (Alydda?), 333 Calosota, 294 caniceps (Ilythea), 234
Calidota, 348 Calotelea, 283 caninus caninus (Dermestes), 133
calidus (Rhantus), 100 Calpodes, 336 Cannaphila, 363
californica (Mecaphesa), 74 calura (Melanaspis), 277 canoa (Phyllophaga), 121
californicus (Anagyrus), 292 calva (Aedmon), 156 canofusa (Spragueia), 350
californicus (Brevipalpus), 50 calva (Micrutalis), 271 CANTHARIDAE, 132
californicus (Cyclodinus), 149 calvata† (Dohrniphora), 213 Canthidium, 120
californicus (Dynaspidiotus), 277 calverti (Hydrellia), 234 CANTHOCAMPTIDAE, 17
californicus (Xyonysius), 250 calverti (Tramea), 364 Canthochilum, 121
californicus manni (Stenomorphus), 103 calverti (Zeros), 234 Canthon, 121
CALIGIDAE, 18 calvovis (Cladochaeta), 229 Canthonella, 121
caligineus (Cyrtocapsus), 243 Calycomyza, 225 cantor (Pteronemobius), 373
caliginosa (Ozophora), 252 calypso (Paululusus), 175 caonabo (Selenops), 73
caliginosus (Heraeus), 251 Calyptolana, 31 caonao (Zanysson), 312
caliginosus (Leptostylopsis), 153 Cambalomma, 93 caparti (Chirnyssoides), 51
caligonoides (Valenzuela), 381 CAMBARIDAE, 38 capitata (Ceratitis), 222
caligonus (Valenzuela), 381 cambridgei (Antillatus), 74 capitatus (Onthophagus), 120
Caligus, 18 cameroni (Ablechrus), 136 capitella (Stegasta), 321
Calinda, 272 cameroni (Cryptocephalus), 161 CAPONIIDAE, 59
Caliothrips, 384 cameroni (Orus), 114 caprae (Bovicola), 379
caliptera caliptera (Erioptera), 192 cameroni (Spalangia), 296 capreolus (Ischnura), 362
CALISCELIDAE, 258 cameroni (Stamnoderus), 114 Capricephala, 247
Calisiopsis, 247 Camillina, 72 capsiformis (Nabis), 245
Calisto, 342 campalis (Botys), 325 capsum (Hydrochasma), 231
callani (Probethylus), 311 Camponotus, 299 capucinus (Dryocoetoides), 180
Callianassa, 38 campostegus† (Semnorhynchus), 172 carabeus† (Platycorypha), 272
CALLIANASSIDAE, 38 Campsiophora, 388 CARABIDAE, 101
Calliasmata, 36 Campsomeris, 308 Carabodes, 46
Callicentrus, 270 Camptocercus, 14 CARABODIDAE, 46
callicerus (Callicentrus), 270 Camptodes, 137 Carabunia, 291
Callichirus, 38 Camptoprosopella, 224 caradrinalis (Bleptina), 356
Calliclytus, 152 Camptoptera, 295 caraiba (Aphylla), 363

587 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

caraiba (Hydrometra), 239 caribeanus (Lyctus), 134 Carposina, 319
caraibea (Euborellia), 183 Caribecandona, 20 CARPOSINIDAE, 319
caraibianus (Treseolobus?), 90 caribensis (Diplosmittia), 198 carsusi (Cautethia), 346
caraibica (Cynortoides), 78 caribensis (Sclerocoelus), 229 cartaba (Phyllophaga), 121
caraibus (Hydrovatus), 100 caribeus (Dactylokepon), 32 carvalhoi† (Reuteroscopus), 245
carbonarius (Dendrocranulus), 179 caribiana (Scaptosciara), 205 Carventius, 248
Carcelia, 220 caribicus (Byrsochernes), 80 caryae (Halysidota), 349
Carcha, 327 caribicus (Euconnus), 116 caryae (Perigonia), 345
carcinophila (Drosophila), 230 caribicus† (Cephalotes), 301 Caryedon, 163
carcinophilus (Culex), 200 Caribitermes, 189 Carylla, 369
Carcinophora, 183 Caribitroides, 30 Caryobruchus, 163
CARCINOPHORIDAE, 182 Cariblatta, 186 caryochron (Pteronemobius), 373
Carcinops, 109 Caribo, 256 casabito (Neowadotes), 69
carcinus (Macrobrachium), 37 Caribochernes, 80 casabito (Rowlandius), 81
carcinus (Neoblattella), 187 cariboea (Odopoea), 263 Casandria, 352
Cardiastethus, 245 Caribojosia, 346 casca† (Gnamptogenys), 299
cardinalis (Caribarctia), 347 Caribophyllum, 368 casca† (Sialis), 358
cardinalis (Rodolia), 140 Cariboptila, 388 casis (Upogebia), 38
cardini (Drosophila), 230 Caribothyrus, 54 Cassida, 160
cardini (Metaleurodicus), 273 Caribovia, 264 cassida‡ (Bairdia), 19
Cardiocondyla, 302 caribus (Stethorus), 141 cassidoides (Diolcus), 258
Cardisoma, 40 caricae (Davara), 328 cassius theonus (Leptotes), 339
cardisomae (Antillesia), 17 carinata (Bitoma), 143 castaneum (Tribolium), 145
Cardobairdia, 19 carinata (Corythaica), 242 castaneus (Callicentrus), 270
cardui (Vanessa), 342 carinata (Entylia), 270 castaneus (Neotermes), 188
Carectocultus, 327 carinata (Macroteleia), 284 castaria (Semaeopus), 335
Careospina, 235 carinator (Ataenius), 120 castelloides (Heteroonops), 59
carib (Paratachys), 107 carinatus (Dynameniscus), 33 Castianeira, 71
carib† (Anoplotermes), 189 carinatus (Podiscodesmus), 88 CASTNIIDAE, 331
cariba (Pseudosciara), 205 carinatus (Sierratettix), 365 Catabena, 350
caribaeus (Rhombognathus), 49 carinifera (Calosopsyche), 387 catacrotos (Hapithus), 371
Caribanosis, 144 cariniger (Neodiachipteryx), 102 catacustes (Miogryllus), 372
Caribarctia, 347 cariniscutum (Conotrachelus), 176 cataulacus (Arphthicarus), 47
caribbaea (Loxosceles), 57 Carinisphindus, 136 catenalis (Rhodaria), 325
caribbaeus† (Dolichoderus), 298 cariniventris (Eurygerris), 239 Catenes, 251
caribbea (Nymphocixia), 259 Carinozetes, 48 Catharthosilvanus, 138
caribbeana (Allaudomyia), 196 Carios, 51 Catharthus, 138
caribbeana (Parabezzia), 196 Caristanius, 328 catherinae (Encarsia), 290
caribbeana (Pseudomorpha), 108 caritatis† (Technomyrmex), 299 Cativella, 27
caribbeana‡ (Typhlocypris), 21 caritatis† (Temnothorax), 302 Catogenus, 138
caribbeanae (Mononychellus), 50 carloorum (Curtara), 268 Catolethrus, 170
caribbeanensis (Pseudothysanoes), 178 carmelita (Pachybrachis), 162 Catonia, 258
caribbeanus (Bledius), 113 carmodyae (Culex), 200 Catorhintha, 254
caribbeanus (Symphylus), 258 carnegie (Phyllophaga), 121 caudata (Paracerceis), 33
caribbee (Ozophora), 252 CARNIDAE, 226 caudatus (Faiditus), 62
Caribbella, 26 carnifex (Helvibotys), 325 Caudellia, 328
caribbeus (Manduca), 344 carnifex (Pyrausta), 326 Caudites, 24
caribbeus (Mithrax), 42 caroli (Bryocyclops), 16 Caularis, 350
caribbeus (Pilumnus), 44 caroli (Cyclosa), 66 Caulophilus, 170
caribbiana† (Lygistorrhina), 202 caroli (Ozophora), 252 Caupolicana, 314
Caribbomerus, 150 carolina (Cinara), 273 cauta (Bradysia), 203
caribea (Drosophila), 230 carpa (Rugosana), 268 cautella (Ephestia), 329
caribea (Formicilabia), 183 Carpelimus, 113 Cautethia, 346
caribea (Glenanthe), 232 Carpilius, 44 cautis (Antillosorius), 112
caribea (Stenacris), 367 carpophagus (Coccotrypes), 179 cavagnaroi (Echiniscus), 13
Caribeacarus, 52 Carpophilus, 137 Cavariella, 274

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 588
cavatica (Tugana), 70 Ceraticelus, 64 ceuthonymus† (Ceraphron), 297
cavera (Curtara), 268 Ceratina, 318 cf. africanus (Synalpheus), 36
cavernicola (Strandesia), 21 Ceratitis, 222 cf. barbara (Thalassozetes), 48
cavifrons (Cryptotermes), 188 Ceratobaeus, 283 cf. inexpectatus (Alismobates), 46
cavina (Madoniella), 135 CERATOCANTHIDAE, 119 cf. jamaicensis (Tropocyclops), 17
cavipennis (Premnobius), 179 Ceratocanthus, 119 cf. jimhensoni (Fimbricyclops), 16
cayennensis hispaniolae (Micropygo- Ceratocapsidea, 244 cf. kingsleyii (Candonopsis), 20
myia), 194 Ceratochrysis, 310 cf. mangrovi (Carinozetes), 48
cayman (Catogenus), 138 Ceratoclasis, 324 cf. platensis (Hyalella), 30
caymanensis (Colliuris), 104 Ceratoconcha, 16 chacei (Pylocheles), 40
cazabitae (Euconnus), 116 Ceratodacus, 222 Chaetanaphothrips, 384
cazabitanus (Microscydmus), 116 Ceratogramma, 297 Chaetocalirrhoe, 220
Cazierius, 83 Ceratoneura, 294 Chaetoclusia, 225
CECIDOMYIIDAE, 202 Ceratophysella, 94 Chaetocnema, 157
cecivora (Tropicharis), 293 Ceratopogon, 196 CHAETODACTYLIDAE, 51
Cedusa, 260 CERATOPOGONIDAE, 196 Chaetodactylus, 51
ceibaensis (Microcyclops), 17 Ceratothoa, 31 chaetodactylus (Metaniphargus), 30
celadonum† (Nesocyrtosoma), 147 ceratus (Bledius), 113 chaetodontis (Anilocra), 31
celatum (Cycloptilum), 376 cerbatana (Hapithus), 370 Chaetophloeus, 178
celatus (Dicranocentrus), 95 cerberus (Ligyra), 210 Chaetosciara, 204
celeno (Macrothemis), 363 Cerceris, 313 Chaetosiphon, 274
celer (Mecaphesa), 74 Cerconota, 321 CHALCIDIDAE, 290
celerans (Hapithus), 370 CERCOPIDAE, 262 chalciope (Nyridela), 348
celeris (Rhysida), 84 Cercyon, 109 chalcipes undata (Gymnetis), 124
celida (Nesiostrymon), 339 cerea (Cyclocephala), 123 Chalcis, 291
celida baorucoensis (Nesiostrymon), 339 cerealella (Sitotroga), 321 Chalcodermus, 177
Celiptera, 352 cereella (Nola), 357 chalcodermus (Stator), 164
Cenocephalus, 182 ceresalis (Diaphantania), 325 Chalcolepidius, 127
Cenophorus, 144 Ceresium, 150 Chalcophora, 125
Centralaphthona, 157 Ceriodaphnia, 15 Chalcosicya, 162
centralis (Altisessor), 139 Ceriomydas, 211 Chalepides, 123
centralis (Euchroa), 106 Cernotina, 387 Chalepus, 160
centratus (Agapostemon), 315 CEROCOCCIDAE, 275 chalicophora (Eugnosta), 332
Centris, 316 Cerococcus, 275 chalybaeus (Selenophorus), 102
Centrocorisa, 241 ceroplastae (Coccidoctonus), 292 chalybea (Cyphomyia), 206
Centrolimnesia, 49 ceroplastae (Coccophagus), 290 chalybea (Discocerina), 231
Centruroides, 81 Ceroplastes, 275 chalybea (Lonchaea), 222
Cephalalges, 175 Ceropsilopa, 232 chalybeum (Andrenosoma), 208
cephalica (Frankliniella), 384 Ceropsylla, 272 chamberlini (Lygromma), 72
cephalonica (Corcyra), 328 Cerotainia, 208 championana (Hydrometra), 239
Cephalonomia, 309 Cerotoma, 159 championi (Sicoderus), 176
Cephaloplopus, 377 ceroton (Cophella), 374 chapini (Thinocharis), 115
Cephalops, 214 cerritus (Yoyuteris), 374 Characoma, 357
Cephalosphaera, 214 cerverai (Eremoleon), 361 Charaphloeus, 138
Cephalotes, 301 cerverinus (Eremoleon), 361 Chararica, 328
cephalotes (Attalomimus), 136 cervicalis (Amegarthria), 328 Charidotella, 160
cephalotes (Bythocrotus), 75 cerviculina (Madoniella), 136 chariedris (Apotomoderes), 173
Ceracis, 142 cervina (Flatoides), 261 Chariesterus, 253
Ceraeochrysa, 359 cervinalis (Murgisca), 327 charinae (Urgleptes), 154
CERAMBYCIDAE, 149 cervinistrigalis (Hypochalcia), 329 CHARINIDAE, 54
ceramica (Argyrotaenia), 332 cervinus (Naupactus), 175 Charinus, 54
Ceranisus, 292 Cerylcautomus, 139 charitonia churchi (Heliconius), 341
Ceraphron, 297 CERYLONIDAE, 139 charitopis (Anaxipha), 375
CERAPHRONIDAE, 297 Cesonia, 72 charlesi (Auplopus), 307
cerasis (Pereirapis), 315 cesonia cynops (Zerene), 338 charynae (Nesiosphaerion), 151
Cerataphis, 275 Cessator, 129 chasei (Cryptotermes), 188

589 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

chaudhrii (Lasioseius), 52 Chlorotettix, 265 CHTHONIIDAE, 79
chaulioda† (Ochrotrichia), 389 chloroticus (Cryptocephalus), 161 chthonus (Rugilus), 114
Cheguevaria, 132 Chobata, 349 chydaeus (Horismenus), 293
CHEIRACANTHIIDAE, 70 chocoensis (Achromoporus), 87 CHYDORIDAE, 14
Cheiracanthium, 70 Choeras, 286 Chydorus, 14
CHEIRIDIIDAE, 79 Chondrocera, 253 Chymomyza, 229
chejela (Leucania), 355 Chondromorpha, 89 Ciba, 69
CHELIFERIDAE, 80 Chondrotropis, 88 cibaensis (Aeolus), 128
CHELODESMIDAE, 87 Chonocephalus, 214 ciboney (Canthochilum), 121
CHELONARIIDAE, 127 choprae (Synsynella), 32 CICADELLIDAE, 263
Chelonarium, 127 Choragus, 164 Cicadettana, 263
Chelonus, 285 Choranthus, 336 CICADIDAE, 262
Chelymorpha, 160 CHOREUTIDAE, 322 Cicadulina, 266
Chemmisomma, 71 Chramesus, 177 cicatricosus (Hyporhagus), 143
Chernes, 80 Chremon, 371 cicero (Cazierius), 83
CHERNETIDAE, 80 christophe (Platynus), 105 ciceroana (Dellia), 366
Chersodromia, 212 christophei (Camponotus), 299 Cicindela, 101
chevrolati (Calocosmus), 154 christophei (Leucochrysa), 359 cienaga (Metazygia), 67
chevrolati (Colaspis), 162 christophei (Pintomyia), 195 cienaga (Modisimus), 58
chevrolati (Cryptocephalus), 161 christophei (Strymon), 340 cienaga (Tainonia), 58
chevrolati (Thonalmus), 129 christyi (Astraptes), 337 cienagaensis (Lychnacris), 130
chiapensis (Amblyseius), 53 Chroantha, 256 ciferrii (Rhyscotoides), 33
chiaroscuro (Capricephala), 247 chromis (Anilocra), 31 ciferrii (Temnothorax), 302
chichimecum (Chlorion), 314 Chrosiothes, 61 ciguaya (Atomosia), 208
chierchiae (Siriella), 29 Chrysanthrax, 210 CIIDAE, 142
Chilaphrodesmus, 89 chrysaoros (Calisto), 342 ciliata (Oxyurella), 15
childi (Pallenopsis), 45 chrysaoros chrysaoros (Calisto), 342 ciliata (Pseudosquilla), 28
Chilicola, 314 chrysaoros galii (Calisto), 342 ciliatus (Anaulacus), 102
Chilocorus, 140 chryselectrus (Jobertus), 244 ciliocera (Bradysia), 203
Chimarra, 386 Chrysendeton, 323 Cillaeus, 137
Chinaria, 263 CHRYSIDIDAE, 310 Cimex, 245
Chinavia, 256 Chrysis, 310 CIMICIDAE, 245
chinensis (Callosobruchus), 163 Chrysobothris, 125 cimitarra (Syspira), 70
Chioides, 337 Chrysocharis, 293 Cinara, 273
Chiomara, 337 CHRYSOMELIDAE, 156 cincta (Ceraeochrysa), 359
Chionaspis, 278 Chrysometa, 65 cinctellus (Hormathus), 152
Chirnyssoides, 51 chrysomphali (Aphytis), 289 cinctimanus (Mithrax), 42
chirographus (Lachnopus), 174 Chrysomphalus, 277 cinctipes (Halysidota), 348
chiron chiron (Marpesia), 340 Chrysomya, 217 cinctipes (Metachroma), 162
chiron necchus (Xylophanes), 346 Chrysonotomyia, 293 cinctiventris (Astenus), 114
CHIRONOMIDAE, 197 Chrysopa, 359 cinctiventris (Euplectrus), 294
Chironomus, 197 Chrysoperla, 359 cinctus (Alagoasa), 157
Chirothrips, 384 CHRYSOPIDAE, 358 cinctus (Anadenobolus), 92
Chirotica, 287 Chrysopilus, 206 cinerascens (Exophthalmus), 173
Chlaenius, 102 Chrysopodes, 359 cinerea (Milichiella), 227
Chlamisus, 161 Chrysops, 206 cinerea (Nauphoeta), 185
chlorescens (Leptostylopsis), 153 chrysopus (Exophthalmus), 173 cinerea (Parlatoria), 277
Chlorida, 153 chrysopyga (Tachytes), 313 cinerea (Protheca), 134
Chlorogonalia, 265 chrysorrhaeella (Cydosia), 349 cinerea (Stenopoda), 247
chlorophanus (Lachnopus), 174 chrysosuccinea† (Forcipomyia), 197 cinereinota (Geranomyia), 194
chlorophyla (Edessa), 255 Chrysotus, 212 cinereus (Anelaphus), 151
CHLOROPIDAE, 226 Chrysso, 61 cinereus (Conocephalus), 368
Chloroprocta, 217 chrysura (Xanthaciura), 223 cinereus (Neohybos), 213
chlorosticta (Chrysobothris), 125 chrysura (Yingaresca), 159 cineticon (Cycloptilum), 376
Chlorostrymon, 339 CHTHAMALIDAE, 16 Cinetorhynchus, 44
Chlorotabanus, 207 Chthamalus, 16 cingulata (Phyllovates), 190

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 590
cingulatus (Agrius), 344 clarki (Sambomorpha), 124 clypeatus (Coenobita), 39
cingulifer (Oncopeltus), 250 clarki (Xylophanes), 346 clypeatus† (Genus indet.), 62
cinnabarinus (Tetranychus), 50 clarkii (Procambarus), 38 Clystea, 347
cinnamomeus (Ommatius), 209 clarus (Prostemmiulus), 93 Clystopsenella, 311
cinnamomeus (Orthomegas), 155 Clasiopella, 231 Cnemonyx, 177
cioides (Neomida), 146 Clastoptera, 262 Cnesinus, 177
cionus† (Atopopsyllus), 382 CLASTOPTERIDAE, 262 c-nigrum (Dictyla), 242
circiimperfecta (Chararica), 328 Clastopteromyia, 229 Cnodalon, 147
circuita (Condica), 350 claudia claudia (Euptoieta), 341 coarctata (Asiomorpha), 89
Circulifer, 267 claudialis (Chrysendeton), 323 coarctatus (Diastolinus), 144
circulus (Halticoptera), 296 CLAUSIIDAE, 18 Cobanus, 75
circumcinctus (Chlaenius), 102 clava (Efferia), 207 Cobubatha, 349
circumdata (Siphonella), 227 Clavaspis, 277 COCCIDAE, 275
circumductus† (Palaeolathys?), 70 clavata (Dioprosopa), 216 coccidarum (Diadiplosis), 202
circumscriptus (Thermonectus), 100 clavata (Heza), 246 coccidivorus (Anagyrus), 292
circumvagans (Phoetalia), 185 clavata (Rhysotritia), 47 coccidivorus (Eupelmus), 294
circumveha (Zygothrica), 231 claveri (Ceraeochrysa), 359 coccinea (Florinda), 64
Cirolana, 31 clavicornis (Aegithus), 139 COCCINELLIDAE, 140
CIROLANIDAE, 31 clavicornis (Nausibius), 138 Coccobius, 290
cirrifera (Oxyethira), 389 clavicornis (Phanocerus), 126 Coccoelasma, 89
cirripediformis (Ceroplastes), 275 clavigera† (Ephuta), 306 coccolobae (Euscelus), 166
Cirrospilus, 294 Clavilispinus, 112 coccolobae (Melanaspis), 277
Cis, 142 clavipes (Aeollopos), 345 Coccophagoides, 290
ciseruditus (Araptus), 180 clavipes (Apachekolos), 209 Coccophagus, 290
Cissites, 148 clavipes (Nephila), 67 Coccotrypes, 179
Cistogaster, 220 clavipes (Prostemmiulus), 93 Coccus, 275
Citharacanthus, 56 claviscapus occultus (Camponotus), 299 cochinchinae seminulum (Aphanisticus),
citrella (Phyllocnistis), 320 clebschi (Doryctopambolus), 285 124
citri (Dialeurodes), 273 clenchi (Calisto), 342 Cochliomyia, 217
citri (Diaphorina), 271 clenchi (Tabanus), 207 cochrusalis (Marasmia), 326
citri (Panonychus), 50 Cleogonus, 176 Cochylis, 332
citri (Planococcus), 280 cleophile (Danaus), 341 cockerelli (Lopholeucaspis), 279
citri (Unaspis), 278 Clerada, 250 cockerelli (Nilakantha), 76
citricidus (Toxoptera), 274 CLERIDAE, 135 cockerellii (Lecaniobius), 294
citricola (Aphis), 274 clerodendri (Colpoptera), 261 cococolus (Aleuroplatus), 272
citricola (Cyrtophora), 66 Cletocamptus, 17 cocois (Aleurodicus), 273
citrifolii (Singhiella), 273 Clibanarius, 39 Cocytius, 344
citrina (Encarsia), 290 Cligenes, 250 coecilialis (Botys), 325
citrina (Tholymis), 364 Climaciella, 360 coecum (Enallagma), 362
citrus (Horismenus), 293 Clinidium, 108 coelebs (Allosmaitia), 339
civile (Enallagma), 362 Cliniodes, 324 Coelioxys, 316
CIXIIDAE, 258 clisaulis† (Pseudomoides), 171 Coelophora, 140
CLADIOPSOCIDAE, 380 clitellaria (Agalliopsis), 264 Coelosia, 203
Cladiopsocus, 380 Clivina, 106 Coelostathma, 332
Cladochaeta, 229 Cloeodes, 235 COENAGRIONIDAE, 362
Cladoctonus, 177 Clonistria, 377 Coenipeta, 352
Claduncaria, 332 Cloridopsis, 28 Coenobita, 39
clancularia (Jabulania), 371 clorinde (Anteos), 338 COENOBITIDAE, 39
clandestina (Walkerana), 369 Closterocerus, 293 Coenonica, 111
clandestinum (Cycloptilum), 376 CLUBIONIDAE, 71 Coenosopsites, 216
clara (Anisomorpha), 378 cluentius (Neococytius), 344 coenosus (Hister), 109
clara (Bradysia), 204 CLUSIIDAE, 225 coeruleovittata (Chlorogonalia), 265
clara (Wendilgarda), 63 Clusiodes, 225 coffeacola (Ormenina), 261
clarescens (Leucania), 355 clydoniata (Calisto), 342 coffeae (Geococcus), 279
claripennis (Tabanus), 207 clypealis (Selenophorus), 102 coffeae (Saissetia), 276
clarki (Erythrolychnia), 130 clypeata† (Dipoenata), 61 coffeella (Perileucoptera), 323

591 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

cognata (Atheta), 111 communis (Camptodes), 137 concolor (Achromoporus), 87
cognata (Corimelaena), 255 communis (Frontinella), 64 concolor (Anax), 363
cognatus (Prostemmiulus), 93 communis (Gomphochernes), 80 concolor (Glomeridesmus), 86
cognita (Graminella), 266 communis (Myriophora), 214 concolor (Heraeus), 251
cohiba (Tethina), 226 communis (Tominotus), 255 concolor (Leptoglossus), 253
cokendolpheri (Kimula), 77 comosa (Aedmon), 156 Condica, 350
Colapteroblatta, 185 comosus (Steneotarsonemus), 50 conducta (Anomis), 351
Colaspis, 162 compacta (Pachygrontha), 250 Condylostylus, 212
Colaspoides, 162 compacta† (Scelianoma), 174 condylum (Pelonium), 136
colei (Ascorhynchus), 45 compactus (Xylosandrus), 180 condylus‡ (Bairdia), 19
Coleomegilla, 140 comperei (Aonidiella), 276 conficita (Thermesia), 354
Coleosoma, 61 Comperiella, 291 configurata (Agallia), 263
colini (Renocila), 32 complanatus (Hemiphileurus), 123 confiniaria (Tephrina), 334
Collaria, 244 complanatus (Hister), 109 confinis (Atholus), 109
collaris (Atopoxicius), 261 complectalis (Hypenula), 356 confirmans (Bulia), 351
collaris (Chrysopodes), 359 complectus (Oliarus), 259 conformis (Agalliopsis), 264
collaris (Drepanaporus), 307 completa (Smicridea), 387 confragosa‡ (Radimella), 24
collaris (Neosmerinthothrips), 385 complicata (Nepheloleuca), 333 confraternus (Hadrosomus), 249
collaris (Ochrimnus), 249 Composia, 348 confusa (Calisto), 342
collaris (Rhagovelia), 240 compositoides (Melanagromyza), 224 confusa (Hoplistocelis), 245
collaris (Tropisternus), 109 compressa (Aphthona), 157 confusa (Penestregania), 268
COLLETIDAE, 314 compressipenne (Ceresium), 150 confusa debarriera (Calisto), 343
collinsae (Parvitermes), 189 compressipenne (Elaphidion), 151 confusella (Ethmia), 321
collinum (Monarthrum), 181 compressus (Parachernes), 80 confusum (Elaphidion), 151
Colliuris, 104 compsa (Metepeira), 67 confusum (Tribolium), 145
Collodes, 42 Compsocryptus, 287 confusus (Dactylopius), 276
Collomena, 355 Compsodecta, 75 confusus (Leptoglossus), 253
Collops, 136 Compsodes, 187 confusus (Lophoscutus), 247
Colobura, 341 compta (Parathiodina), 76 confusus sequestus (Pityophthorus), 181
colombo (Corinna), 71 compta (Psilopa), 233 congeae (Amblydromalus), 53
Colomerus, 48 comptalis (Scopula), 357 congemmalis (Syllectra), 355
colona (Cedusa), 260 Compterosmittia, 198 congestocostata‡ (Coquimba), 24
colona (Lobopoda), 145 comptus (Isotomus), 150 congruata (Geometra), 334
colonus (Unerus), 266 Comstockiella, 278 congruus (Pityophthorus), 181
colorata† (Palaeometopia), 137 comtea (Rhysotritia), 47 Conicera, 213
coloratus (Achromoporus), 87 Conalia, 142 conicocystata‡ (Ceratoconcha), 16
coloratus neiba (Achromoporus), 87 Concana, 352 CONIOPTERYGIDAE, 359
coloritubus (Osbornellus), 267 concavus eseptatus‡ (Balanus), 15 Coniopteryx, 360
colossea (Colossendeis), 45 concentricus (Peridinetus), 170 conisquamis† (Anochetus), 304
COLOSSENDEIDAE, 45 concepcioni (Pseudopachybrachius), 251 conjuncta (Galerucella), 159
Colossendeis, 45 CONCHASPIDIDAE, 276 conjuncta (Iomoides), 90
Colpoptera, 261 Conchaspis, 276 conjunctus (Urgleptes), 154
columbi (Hypothenemus), 178 Conchylodes, 324 connectans (Telenomus), 284
columbina (Eunomia), 347 conciliatus (Epidapus), 204 connectens (Hyperaspis), 140
columbusi (Metophthalmus), 142 concinna (Megachile), 316 connexalis (Botys), 326
columella (Strymon), 340 concinna (Stobaera), 260 connexionis (Xanthaciura), 223
Colydium, 143 concinnata (Acidalia), 335 connexum (Stilomedon), 115
Comanopa, 268 concinnulalis (Hypena), 357 connexus (Tinus), 68
comatus† (Toxonotus), 165 concinnulus (Monanus), 138 Conobregma, 286
combinataria (Hyperetes), 333 concinnus (Acanthinus), 149 Conocephalus, 368
comma (Disonycha), 157 concinnus (Gnatholeptus), 180 Conoderus, 128
comma (Smicridea), 387 concinnus (Oecobius), 60 conopa (Oeclidius), 261
commelinae (Prodenia), 356 concinnus (Spirobolellus), 91 Conophthorus, 180
commixtus (Tabanus), 207 concisa (Condica), 350 CONOPIDAE, 221
commune (Crenidorsum), 273 concisa (Epidromia), 352 Conostigmus, 297
communicans (Hamataliwa), 68 concludens (Chobata), 349 Conotelus, 137

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 592
Conotrachelus, 176 Copelatus, 100 cornutus (Metadorylas), 215
consentaneus (Lachnopus), 174 coperniciae (Mycococcus), 276 cornutus (Rhipidandrus), 144
consequens (Poaphila), 354 Copestylum, 215 cornutus† (Episinus), 62
conservatus† (Homoconnus), 116 Cophella, 374 cornutus† (Proneoproxius), 248
consimilis (Hemiphileurus), 123 Cophes, 171 coronai (Corythalia), 75
consimilis (Hydrometra), 239 Copicerus, 259 coronatus (Macrobiotus), 13
consimilis (Telephanus), 137 Copidosoma, 291 correctus (Carpelimus), 113
consoleae (Mnemosyne), 259 Coproica, 228 corrugata (Aedmon), 156
conspersa (Chrysometa), 65 Coptarthria, 328 Corthylites†, 182
conspersum (Elaphidion), 151 Coptodera, 103 Corthylus, 181
conspersus (Homocloeus), 165 Coptotermes, 188 corticarium (Strongylaspis), 155
conspicua† (Chrysso?), 61 coquilletti (Ceropsilopa), 232 Corticatus, 75
constans (Canthonella), 121 coquilletti (Scaptomyza), 231 Corticeus, 146
constans (Caribovia), 264 Coquimba, 24 corticina (Phalaenomorpha), 261
constanza (Geranomyia), 194 CORALLANIDAE, 31 cortico (Nemobiopsis), 373
constanzae (Aedmon), 156 corallinus (Carpilius), 44 coruscus† (Pseudomyrmex), 306
constanzae (Chalcosicya), 162 corallinus (Trigoniulus), 93 CORYDIIDAE, 187
constanzae (Heterophotinus), 131 corayi† (Anochetus), 304 CORYLOPHIDAE, 141
constanzae (Paululusus), 175 corbis (Clinidium), 108 Corynoptera, 204
constanzae (Syphraea), 158 Corcyra, 328 Corynothrips, 384
constanzae (Yingaresca), 159 cordatus (Ucides), 42 Coryphaeschna, 363
constanzanus (Euconnus), 116 cordiae (Conchaspis), 276 coryphe (Mithrax), 42
constellata (Joguina), 359 cordiae (Omegalebra), 268 Coryssocnemis, 57
Constrictotermes, 189 cordifera (Limnesia), 49 Corythaica, 242
consuelo (Microtityus), 82 cordiformis (Agobardus), 74 Corythalia, 75
consueta (Melanagromyza), 224 cordobensis (Lispinus), 112 Corythucha, 242
consutus (Cubobolus?), 93 Cordyla, 203 COSMETIDAE, 77
contacta (Poaphila), 354 COREIDAE, 253 cosmiata (Semiothisa), 334
Contarinia, 202 Corethrella, 199 cosmioides (Gonitis), 351
contectum (Cycloptilum), 376 CORETHRELLIDAE, 199 Cosmocomoidea, 295
contenta (Phurys), 358 coriaceipennis (Tytthonyx), 133 Cosmopolites, 177
conterminus cressoni (Episyron), 307 coriaceus (Palaminus), 114 COSMOPTERIGIDAE, 322
conterminus posterus (Episyron), 308 coriaceus (Tylocerus), 132 Cosmopterix, 322
conthorhinus† (Neoulosomus), 171 coriaria (Dalotia), 111 Cosmosoma, 348
continua (Seirocastnia), 350 Corimelaena, 255 COSSIDAE, 330
continuata (Cyrtaphe), 88 CORIMELAENIDAE, 255 Cossonus, 170
contorhinus† (Semnorhynchus), 172 Corinna, 71 Costa, 27
contorta (Melipotis), 353 CORINNIDAE, 71 costalis (Nasutitermes), 189
contortus (Calocosmus), 154 Corinthiscus, 136 costalis (Ponometia), 349
contrarium† (Theridion), 63 CORIOXENIDAE, 383 costaricana (Paramycodrosophila), 230
contricticeps (Platynus), 105 Corioxenos, 383 costaricensis (Orasema), 292
Conura, 291 CORIXIDAE, 241 costata (Diaphania), 325
conventica (Atopsyche), 390 Corizus, 254 costata (Hypasclera), 148
convergens (Hippodamia), 140 Cormocephalus, 84 costata‡ (Scylla), 38
convexa (Chalcosicya), 162 cornelli† (Heterothops), 117 costatus (Exophthalmus), 173
convexifrons (Anthonomus), 169 cornica (Priocnemis), 307 costatus (Zophobas), 145
convexinotatus (Heterotermes), 188 corniceps (Procryptotermes), 188 costicollis (Palaeopus), 171
Convexocoleus, 160 corniger (Culex), 200 costipenne (Elaphidion), 151
convexoculus† (Toxorhynchus), 167 corniger (Nasutitermes), 189 costipennis (Chlorion), 314
convexus (Apallates), 226 cornuta (Ceriodaphnia), 15 costipennis (Diastolinus), 144
convexus (Epinannolene), 93 cornuta (Chrysometa), 65 costura (Phyllophaga), 121
convexus† (Toxorhynchus), 167 cornuta (Curtara), 268 Cotesia, 286
cooki (Uranotaenia), 201 cornuta (Dohrniphora), 213 Cothonaspis, 289
Copaeodes, 336 cornuta (Edessa), 255 cottalis (Zebronia), 327
copal† (Empicoris), 246 cornuta (Orekthophora), 270 coursetiae (Tanaostigma), 297
copalis† (Palaeolathys), 70 Cornutidion†, 61 coxalis† (Pensacolatus), 76

593 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Crabro, 312 cressoni (Metadorylas), 215 Cryptobables, 328
CRABRONIDAE, 312 cressoni (Mimapsilopa), 233 Cryptocarenus, 178
crabroniformis (Cryptocephalus), 161 Cressonomyia, 232 Cryptocephalus, 161
craccivora (Aphis), 273 cretella (Tinea), 320 Cryptocheiridium, 79
craesus (Ectemnius), 312 Creugas, 72 Cryptoflata, 260
craigi† (Syringogaster), 222 crewi (Metazygia), 67 Cryptolaemus, 141
CRAMBIDAE, 323 crewi (Oxyopes), 68 Cryptolestes, 138
crambidoides (Diatraea), 323 crewii (Singa), 67 crypton (Cycloptilum), 376
crameri (Erinnyis), 345 cribraria (Chelymorpha), 160 CRYPTOPHAGIDAE, 138
cranekeyensis‡ (Hemicytherura), 23 cribrarius (Arenaeus), 43 Cryptophagus, 138
Crangoweckelia, 29 cribrarius (Euscelus), 166 CRYPTOPIDAE, 84
craniifer (Blaberus), 185 cribratum (Plectrocerum), 152 Cryptops, 84
Craspedisia, 61 cribricollis (Ips), 179 cryptops (Listronotus), 172
Craspedochaeta, 225 Cricotopus, 198 Cryptopygus, 95
crassa (Dunhevedia), 15 criniticornis (Pagurus), 39 Cryptorhopalum, 133
crassa (Tugana), 70 crinitus (Anthicus), 149 Cryptorhynchus, 171
crassartus (Haplocyclodesmus), 91 crinitus crinitus (Polistes), 309 Cryptosoma, 40
crassicaudus (Witica), 68 crisia (Pyrgus), 337 Cryptotermes, 188
crassicera (Bradysia), 204 crispula (Aedmon), 156 Cryptothelea, 320
crassicornis (Ectmesopus), 159 crispulum (Anelaphus), 151 cryptus (Pseudococcus), 280
crassicornis (Umbonia), 270 cristata (Homalopoda), 291 CTENIDAE, 69
crassifemoralis† (Modisimus), 58 cristata (Ochrotrichia), 389 Ctenocolum, 163
crassimanus (Tityus), 82 cristata (Symploce), 186 Ctenolepisma, 97
crassipatellaris† (Faiditus), 62 cristatus (Dryocoetoides), 180 Ctenophilus, 86
crassipes (Lepiselaga), 207 cristatus (Leptostylopsis), 153 Ctenucha, 347
Crassomicrodus, 285 cristatus† (Pseudoarmadillo), 33 Ctenuchidia, 348
crassus (Apocephalus), 214 cristinae (Ferrisia), 279 Ctenus, 69
crassus (Bahorucocricus), 92 cristobalensis (Erythrolychnia), 130 ctilos (Cyrtoxipha), 375
crassus (Euschistus), 256 croceipes (Thinodromus), 113 cuadro (Modisimus), 58
crassus (Jollas), 76 Crocidomera, 328 Cubachelifer, 80
Craterodesmus, 87 Crocidosema, 331 cubaensis (Coccophagus), 290
Cratyna, 204 croizati (Compterosmittia), 198 cubaensis (Pseudomyrmex), 306
Creagrophorus, 111 Cronius, 43 cubana (Augochlora), 315
credibilis (Thermesia), 352 Cropia, 350 cubana (Ceraeochrysa), 359
credula (Composia), 348 Crossocerus, 312 cubana (Climaciella), 361
credula† (Cedusa), 260 crotonis (Parlatoria), 277 cubana (Exoprosopa), 210
cregyos (Anaxipha), 375 Crovettia, 311 cubana (Heteropsylla), 271
Crematogaster, 302 crucians (Anopheles), 200 cubana (Leucocera), 161
Cremnops, 285 crucifer (Acanthocerus), 253 cubana (Loxosceles), 57
crenator (Euschistus), 256 cruciferum (Amblyomma), 52 cubana (Microvelia), 240
crenatus (Metaniphargus), 30 cruciger (Acanthocerus), 253 cubana (Nectopsyche), 390
crenatus (Spasalus), 118 crudiae (Hypothenemus), 178 cubana (Neoconis), 360
crenatus (Trox), 119 crudus (Haplaxius), 259 cubana (Newportia), 85
crenellatus (Penteporus), 90 cruenta (Acmaeodera), 125 cubana (Nusalala), 360
Crenidorsum, 273 cruentata (Goniopsis), 41 cubana (Onychotillus), 136
Crenulaspidiotus, 277 crumena‡ (Triebelina), 19 cubana (Prosierola), 309
crenulatum (Macrobrachium), 37 cruraluma† (Anthonomus), 169 cubana‡ (Costa), 27
crenulatum (Nesocyrtosoma), 147 cruzi (Domingoecius), 51 Cubanella, 258
crenulatus (Plectromerus), 150 crybelum (Macrobrachium), 37 Cubanochirus, 51
creolus (Temnothorax), 302 crypta (Calisto), 343 Cubanops, 59
creonalis (Botys), 326 cryptadia (Atlantea), 342 Cubanoptila, 388
Creophilus, 117 cryptadia (Cyrtoxipha), 375 cubanus (Campylothorax), 95
cresbios (Hapithus), 370 cryptambra† (Mycetobia), 201 cubanus (Enicocephalus), 238
cresera‡ (Cytherura), 23 Cryptaspasma, 331 cubanus (Enosmaeus), 150
cressoni (Enicospilus), 287 Crypticerya, 279 cubanus (Loxandrus), 106
cressoni (Liobracon), 286 crypticus (Bythocrotus), 75 cubanus (Mamurius), 253

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 594
cubanus (Oncerotrachelus), 247 curta (Gypona), 268 Cyclopoliarus, 258
cubanus (Pseudaptinus), 102 curta (Megachile), 316 Cyclopterum, 260
cubanus ubancus (Selenophorus), 102 curta‡ (Cyprideis), 21 Cycloptilum, 376
Cubaris, 32 curtalis (Asopia), 325 Cyclosa, 66
cubensis (Bephratelloides), 294 Curtara, 267 cyclosignatus (Notaepytus), 139
cubensis (Chaetophloeus), 178 curtellus (Crambus), 323 Cydia, 331
cubensis (Cladoctonus), 177 curtior (Anadenobolus?), 92 CYDNIDAE, 254
cubensis (Clibanarius), 39 curtipennis (Urgleptes), 154 Cydosia, 349
cubensis (Coleomegilla), 140 Curtomerus, 151 Cylas, 168
cubensis (Encarsia), 290 CURTONOTIDAE, 229 Cylindera, 101
cubensis (Frankliniella), 384 Curtonotum, 229 cylindricornis (Raphirhynchus), 168
cubensis (Gonatopus), 311 curtulus (Xylosandrus), 180 cylindricus (Ocyptamus), 216
cubensis (Haliplus), 99 curtus (Ceracis), 142 Cylindromyrmex, 298
cubensis (Hololepta), 109 curva‡ (Phlyctocythere), 25 Cylindroxystus, 114
cubensis (Idiataphe), 363 curvicauda (Anastrepha), 223 cyllarus (Psocus), 382
cubensis (Lymaenon), 295 curvicauda (Paralabella), 183 Cymaenes, 336
cubensis (Micracis), 178 curvicauda (Sterrha), 336 cymatilis (Madoniella), 136
cubensis (Paramusonia), 190 curvicaudatus (Eudorylas), 215 Cymatothes, 144
cubensis (Pylocheles), 40 curvifrons (Conoderus), 128 CYMIDAE, 249
cubensis (Scolytodes), 179 curvimedia (Pseudolycoriella), 205 Cymindis, 103
cubensis (Teredon), 282 curvipes (Lachnopus), 174 Cymodema, 249
cubensis cubensis‡ (Hemicyprideis), 22 curvipubens (Rogeria), 304 Cymoninus, 249
cubensis‡ (Trachyleberidea), 27 curvirostris (Anthonomus), 169 Cymophyes, 250
Cubobolus, 93 curvispinosus† (Chrosiothes), 61 Cymothoa, 31
Cubrasa, 264 Cushmanidea, 21 CYMOTHOIDAE, 31
cubrosa (Mormidea), 257 CUSHMANIDEIDAE, 21 cymotoma (Cacocharis), 331
cucullata (Fulgoraecia), 330 cyanea (Holocompsa), 187 Cymotricha, 321
cucumeris (Epitrix), 157 cyanea (Leucocera), 161 cynocephala (Placopsidella), 232
cucurucho (Hapithus), 370 cyanea (Scutellista), 296 Cynorta, 78
Culex, 200 cyanella (Cydosia), 349 Cynortoides, 78
CULICIDAE, 199 cyanella (Psorolyma), 140 Cyparium, 116
culicinus (Modisimus), 58 cyaneolimbatus (Semiardistomis), 107 cyperi (Coccotrypes), 179
Culicoides, 196 cyaneopacus (Discoderus), 102 cypha (Empoasca), 269
Culiseta, 200 cyaneus (Pyropus), 172 Cyphoceratops, 262
culmenicola (Aptunga), 328 cyanipennis (Omophoita), 158 Cyphoderus, 95
cultellator (Remosa), 262 cyanipes (Polycesta), 126 Cyphomyia, 206
cultellatum (Neoerythromma), 362 cyanipes (Rhammatocerus), 367 Cyphomyrmex, 301
cuneatus (Monobeloides), 270 cyanoptera (Anisostena), 159 Cyphotylus, 89
cuniculus (Aphodius), 120 cyanura (Xenochalepus), 161 Cypretta, 20
cuniculus (Tomarus), 123 Cyarda, 260 Cypria, 20
cuphos (Hapithus), 370 cyathicollis (Corythaica), 242 Cypriacis, 125
Cupiennius, 69 Cyathura, 31 Cyprideis, 21
cuprascens (Colpodes), 105 CYBOCEPHALIDAE, 137 CYPRIDIDAE, 20
cuprea (Bura), 141 Cybocephalus, 137 Cypridopsis, 20
cupreorutilans† (Tyrtaeus), 146 cychrinus (Platynus), 105 CYPSELOSOMATIDAE, 221
cuprescens (Elaphria), 355 Cyclargus, 339 Cyptocephala, 256
cuprina (Lucilia), 217 cyclata (Pachydia), 334 Cyptoxenus, 164
cupripennis (Dyschromus), 106 Cyclocena, 324 Cyrsylus, 157
cupripennis (Euchroa), 106 Cyclocephala, 123 Cyrtaphe, 88
curacaoensis (Neogonodactylus), 28 Cyclocypris, 20 CYRTAUCHENIIDAE, 56
CURCULIONIDAE, 169 Cyclodinus, 149 Cyrtinus, 154
Curius, 150 Cyclograpsus, 41 Cyrtocapsus, 243
currani (Heriades), 316 Cyclomia, 333 Cyrtognatha, 65
currani (Liohippelates), 227 Cycloneda, 140 Cyrtopholis, 56
cursarius (Electropterum), 129 Cyclophora, 335 Cyrtophora, 66
cursor (Cyrtopholis), 56 CYCLOPIDAE, 16 Cyrtoxipha, 375

595 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Cytherella, 21 darlingtoni (Anthicus), 149 Dasyllis, 208
CYTHERELLIDAE, 21 darlingtoni (Antillatus), 74 Dasymutilla, 306
Cytherelloidea, 21 darlingtoni (Antillipeltis), 135 Dasyodontus, 89
Cytheretta, 22 darlingtoni (Antilliscaris), 107 Dasypyga, 328
CYTHERETTIDAE, 22 darlingtoni (Arachnocoris), 245 dasypyga (Coptarthria), 328
cythereus (Haplopus), 377 darlingtoni (Asteriza), 160 dativaria (Lobocleta), 335
CYTHERIDAE, 21 darlingtoni (Ataenius), 120 datona (Metepeira), 67
CYTHERIDEIDAE, 21 darlingtoni (Caecophloeus), 146 Davara, 328
Cytheridella, 24 darlingtoni (Canthochilum), 121 davidi (Leberis), 15
cytheridellaformis‡ (Perissocytheridea), darlingtoni (Ceriomydas), 211 davidsoni (Agalliopsis), 264
22 darlingtoni (Chalcosicya), 162 davidsoni (Apenes), 103
CYTHEROMATIDAE, 22 darlingtoni (Chilaphrodesmus), 89 davidsoni (Brasiella), 101
Cytheromorpha, 25 darlingtoni (Colapteroblatta), 185 davidsoni (Cryptorhopalum), 133
Cytheropteron, 22 darlingtoni (Colaspis), 162 davidsoni (Neodiachipteryx), 102
Cytherura, 23 darlingtoni (Ctenus), 69 davidsoni (Phyllophaga), 121
CYTHERURIDAE, 22 darlingtoni (Cubanops), 59 davidsoni (Trogoderma), 133
czernyi (Heteromeringia), 226 darlingtoni (Cylindromyrmex), 298 davisi† (Didymia), 282
Dacne, 139 darlingtoni (Decarloa), 240 davisorum (Atopsyche), 390
dactyliperda (Coccotrypes), 179 darlingtoni (Desmopachria), 100 davisorum (Streptosyche), 387
Dactylokepon, 32 darlingtoni (Dicoelotrachelus), 159 Dawnarioides, 260
dactylopii (Leptomastix), 292 darlingtoni (Ectmesopus), 159 debarriera (Calisto), 343
DACTYLOPIIDAE, 276 darlingtoni (Espagnola), 366 debililimbata (Silis), 132
Dactylopius, 276 darlingtoni (Eugamandus), 153 debilis (Brontaea), 218
Dactylosternum, 109 darlingtoni (Faiditus), 62 debilis (Enochrus), 108
Dadaya, 14 darlingtoni (Heikertingerella), 157 debilis (Holisus), 117
daemon (Antillosorius), 112 darlingtoni (Hemerobius), 360 debilis† (Micromalthus), 99
Dagbertus, 244 darlingtoni (Hoplicnema), 141 debordae (Crenidorsum), 358
Dagus, 233 darlingtoni (Isotes), 159 Decadiomus, 141
Dahlimosina, 228 darlingtoni (Leiopleura), 124 Decarloa, 240
daidaleos (Heuretes), 330 darlingtoni (Lobopoda), 145 decemvittata (Yingaresca), 159
Daimon, 270 darlingtoni (Lomaita), 64 deceptor (Megacerus), 163
daira palmira (Eurema), 338 darlingtoni (Malleoceps), 115 deceptrix (Megachile), 316
dajaboni (Enicospilus), 287 darlingtoni (Metachroma), 162 decernens (Agrotis), 357
Dalbulus, 267 darlingtoni (Neoxantholinus), 117 decipiens (Paralimna), 233
Dallasiellus, 254 darlingtoni (Nesocyrtosoma), 147 decipiens (Peridinetus), 170
Dalotia, 111 darlingtoni (Novantinoe), 153 decipiens (Thermocyclops), 17
dama (Spragueia), 350 darlingtoni (Ocrepeira), 67 decisa (Orphulella), 367
damiensis (Chrysopa), 359 darlingtoni (Oomorphus), 163 decliva (Neokrisna), 267
damiensis (Nusalala), 360 darlingtoni (Ozophora), 252 declivis (Telephanus), 137
danae (Callinectes), 43 darlingtoni (Parawixia), 66 declivis‡ (Callinectes), 43
Danaus, 341 darlingtoni (Peripatus), 13 decolor (Ectomyelois), 329
danforthi (Pinophilus), 114 darlingtoni (Pheidole), 303 decolorata (Centris), 316
danieli (Cernotina), 387 darlingtoni (Polyancistrus), 368 decoloratus (Cephalotes), 301
danieli (Triepeolus), 317 darlingtoni (Proctacanthus), 208 decora (Corythalia), 75
danielperezi (Coelioxys), 316 darlingtoni (Sunius), 115 decora (Hypercompe), 347
danum (Enyo), 345 darlingtoni (Syphraea), 158 decoralis (Dichogama), 324
dapanera (Amphiacusta), 373 darlingtoni (Tabanus), 207 decorata (Notiphila), 234
daphne (Orsa), 78 darlingtoni (Thinodromus), 113 decoratum (Microcentrum), 368
Daphnia, 15 darlingtoni (Xantholinus), 118 decoratus (Caribbomerus), 150
DAPHNIIDAE, 15 darlingtoni (Zatania), 300 decoratus (Phrynus), 54
dardana (Garcialdretia), 381 darlingtoniana (Brasiella), 101 decoregens (Notaepytus), 139
Dardanus, 39 darlingtoniana (Tipula), 194 Decorosa, 135
darlingtoni (Aedmon), 156 Darwinula, 23 decorum (Agonum), 104
darlingtoni (Alethaxius), 162 DARWINULIDAE, 23 decorus (Gampsocoris), 248
darlingtoni (Anchonoderus), 103 Dasiops, 222 decrepitaria decrepitaria (Erastria), 333
darlingtoni (Anisepyris), 309 Dasyhelea, 197 defecta† (Loxosceles), 57

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 596
defectus† (Oxyopes), 68 denticulata (Xantho), 44 destructus† (Bolostromus), 56
defensata (Pterocypha), 335 denticulatum† (Proceratium), 305 detersa (Neoblattella), 187
deficiens (Ninyas), 250 dentifer (Chydorus), 14 detexta (Epimecis), 333
deformans (Synsynella), 32 dentifer (Perileptus), 107 detonnancouri (Sicoderus), 176
deformis† (Loxosceles), 57 dentifera (Orchestina), 59 detrahens (Ophisma), 354
deformis‡ (Procythereis?), 27 dentifrons (Epaphrodita), 190 deturbata (Pterocypha), 335
degener (Melanolestes), 246 dentiger (Mermessus), 64 Deuterixys, 286
degiacomii (Odopoea), 263 dentilis (Typhlodromips), 53 Deuterolita, 327
Deinocerites, 201 dentipes (Chrysobothris), 125 devexulus (Ambrosiodmus), 180
DEINOPIDAE, 60 dentipes (Euscelus), 166 devia (Eustala), 66
Deinopis, 60 dentipes (Lachnopus), 174 Dexosarcophaga, 219
deitersi (Cletocamptus), 17 dentipes (Megachile), 316 diabolica (Coproica), 228
dejecta (Cladochaeta), 229 dentipes (Philothermus), 140 Diachus, 162
deklei (Cerococcus), 275 Deopalpus, 220 Diacme, 325
delacruzi (Cryptorhopalum), 133 depilatus (Dicoelotrachelus), 159 Diadegma, 288
delamarei (Troglopedetes), 96 deplanata (Cerotothoa), 31 diadelos (Stenogryllus), 371
delasmata (Eustala), 66 depressa (Plagusia), 41 diadema (Melanagromyza), 224
delclosi† (Tetrapus), 289 depressa‡ (Kangarina), 23 Diadiplosis, 202
delecta (Neohadria), 265 depressicollis (Exophthalmus), 173 diadromos (Nemobiopsis), 373
delicatula (Cariblatta), 186 depressifrons (Portunus), 43 Dialectica, 320
delicatulus (Anthrax), 210 Depressonotum†, 229 Dialeurodes, 273
delicatus (Yelicones), 287 depressus (Aeolus), 128 dianae (Metacyclops), 17
delimitalis (Ceratoclasis), 324 depressus (Garyops), 79 diaperinus (Alphitobius), 144
delineata (Pterocypha), 335 depressus (Metopaulias), 41 Diaperis, 146
deliriosa (Elaphria), 355 Derallus, 108 diaphana (Euthlastoblatta), 187
Dellia, 366 Derancistrachroma, 156 diaphana calimete (Greta), 341
Delocrania, 160 Derancistrus, 156 diaphana charadra (Greta), 341
DELPHACIDAE, 259 DERBIDAE, 260 diaphana galii (Greta), 341
Delphacodes, 259 Derelomus, 172 Diaphania, 325
delphacoides (Lomagenes), 261 Dermacentor, 52 Diaphanosoma, 14
Delphastus, 141 Dermestes, 133 Diaphantania, 325
delphinicauda (Phyllophaga), 121 DERMESTIDAE, 133 diaphanum (Crenidorsum), 273
deltus (Hemiptiloceroides), 329 dermoidea† (Statira), 143 diaphanus (Hyalopsallus), 320
demissus (Hapithus), 370 derogata (Anomis), 351 diaphanus quisqueya (Greta), 341
demma (Antillopsyche), 387 Derophthalma, 244 Diaphorina, 271
demoleus demoleus (Papilio), 338 Derovatellus, 100 Diaprepes, 173
demophoon insulicola (Archaeoprepona), Descangeles†, 75 DIAPRIIDAE, 288
341 Descanso, 75 DIAPTOMIDAE, 16
Demotispa, 160 descriptalis (Ommatospila), 326 Diaptomus, 16
denaia† (Ochrotrichia), 389 deserticola (Opopaea), 59 diaspidicola (Encarsia), 290
Dendrobiella, 134 desertorum (Xestocephalus), 269 DIASPIDIDAE, 276
Dendrocerus, 298 DESIDAE, 69 Diaspidiotus, 277
Dendrocranulus, 179 designalis (Agathodes), 324 diaspidis (Aphytis), 289
Dendrodesmus, 88 desmata (Polytrichophora), 232 Diaspis, 278
Dendrosinus, 177 Desmia, 324 Diastolinus, 144
Dendrothereua, 84 Desmometopa, 227 Diatraea, 323
Dendrothripoides, 384 Desmopachria, 100 diatraeae (Apanteles), 286
dengizicus (Microcyclops), 17 desotoi (Perigonia), 345 diatraeae (Lixophaga), 220
denia (Selenops), 73 desotoi (Schizuroides), 346 dibolius† (Dolichoderus), 298
denotatum (Pelonium), 136 despaignei (Ananca), 148 Dibothrocricus, 92
densa (Biocrypta), 114 despiciens (Hydroma), 49 dicellus (Osbornellus), 267
densicollis (Thinocharis), 115 dessalinesi (Selenophorus), 102 Diceratobasis, 362
dentata† (Platythyrea), 305 destituta (Scaptosciara), 205 Dicerca, 125
dentatum (Metachroma), 162 destructor (Aspidiotus), 277 Dichogama, 324
dentatus (Ablechrus), 136 destructor (Trichomyrmex), 302 Dichomeris, 321
dentatus (Telephanus), 137 destructor (Varroa), 54 Dichrogaster, 287

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dichroma (Agalliopsis), 264 dimidiatus (Ocyptamus), 216 discolor (Lobrathium), 114
dichroma (Elateropsis), 156 diminutalis (Sufetula), 327 discopunctatus (Selenophorus), 102
Diclidophlebia, 271 dimorpha‡ (Paranesidia), 19 discordis (Hypothenemus), 178
Dicoelotrachelus, 159 dina mayobanex (Pyrisitia), 338 Discothyrea, 305
dicrana (Beameromyia), 209 Dinattus, 75 discoviridis (Tylocerus), 132
Dicranocentrus, 95 Dineutus, 99 disjectus (Stenogryllus), 371
Dicranomyia, 193 Dinoderus, 134 disjuncta† (Valeseguya), 196
Dicrepidius, 128 Dinurothrips, 384 dislocatus (Pseudothysanoes), 178
Dicrocheles, 52 Diochus, 117 Dismorphia, 339
Dicromantispa, 361 diogenes (Petrochirus), 39 Disonycha, 157
Dicrotendipes, 197 DIOGENIDAE, 39 Disorygma, 289
Dicrotingis, 242 Diolcogaster, 286 dispansalis (Pyralis), 327
Dictyla, 242 Diolcus, 258 dispar (Dohrniphora), 213
DICTYNIDAE, 69 Diomus, 141 dispar (Hemiphileurus), 123
dictyospermi (Chrysomphalus), 277 Dioprosopa, 216 dispar (Toxomerus), 216
dictys (Tangella), 262 Dioryctria, 329 disparodigitus (Synalpheus), 36
dicyrta (Empoasca), 269 Diostracus, 212 dispersus (Aleurodicus), 273
DICYRTOMIDAE, 96 diotimealis (Scopula), 324 dispertitum (Linepithema), 298
didactylus (Neoscapteriscus), 376 Dioxyna, 222 dispilus (Enicospilus), 287
didicostae† (Antilloides), 57 Diphleps, 244 Displotera, 142
didyma (Eburia), 151 diphteralis (Conchylodes), 324 disputabilis (Apanteles), 286
didyma (Micrathyria), 364 Diphthera, 352 dissectus (Cryptocephalus), 161
Didymia, 282 Diphuia, 232 disseverans (Mocis), 354
Diestota, 111 dipilus (Osbornellus), 267 dissimile (Amblyomma), 52
dieteri† (Cephalotes), 301 diplacanthum (Macrocoeloma), 42 dissimilis (Amphiacusta), 373
dieteri† (Paraponera), 304 Diplochaetus, 107 dissimilis (Anastrepha), 223
Dieuches, 252 Diplonevra, 213 dissimilis (Prytanes), 251
differalis (Pyralis), 324 Diplosmittia, 198 dissimilis (Robopus), 131
difficilis (Placosternus), 150 Diplozona, 243 dissimilis† (Pintomyia), 195
diffissa (Geometra), 334 DIPLURIDAE, 56 dissita (Megistops), 158
diffluens (Mocis), 354 Dipoena, 61 dissocians (Micrathyria), 364
difformis (Cyarda), 260 Dipoenata, 61 Dissodactylus, 43
diffundens (Toxonprucha), 355 Dipogon, 307 Dissomphalus, 310
dificilis (Valenzuela), 381 Dipropus, 128 distans (Dicranomyia), 193
digestalis (Nazuda), 351 dirce wolcotti (Colobura), 341 Distigmoptera, 157
digitalis (Dicrotingis), 242 dircealis (Asopia), 327 distincta (Bdella), 48
Digitonthophagus, 120 directa (Kakopoda), 352 distincta (Paraleucolepidopa), 39
digitus† (Antillopsyche), 387 Dirhinus, 291 distincta (Parvomygale), 56
Diglyphus, 293 disatra (Nylanderia), 300 distinctus (Cleogonus), 176
dignus (Pseudapanteles), 286 discalis (Chobata), 349 distinctus (Cligenes), 250
Digrammia, 333 discalis (Toonglasa), 248 distinctus (Coccotrypes), 179
DILARIDAE, 360 discifer (Kuschelaxius), 176 distinctus (Plectromerus), 150
dilatata (Odopoea), 263 Disclisioprocta, 335 distinctus (Tachytes), 313
dilatiforceps (Lachesilla), 381 discludellus (Microcrambus), 323 distinctus (Valtissius), 251
dilatus (Trachelas), 72 Discocerina, 231 distinctus distinctus (Merocoris), 253
dilitara (Empoasca), 269 Discoclavata†, 134 distracta (Eutelia), 355
Dilophops, 89 Discocoris, 241 distributa (Lacinipolia), 355
Dilophus, 202 Discoderus, 102 ditalea (Hydroptila), 388
dilucidum (Theridion), 63 discoidalis (Acontia), 350 diva (Cophella), 374
dimalea (Amphiacusta), 373 discoidalis (Blaberus), 185 divaricata (Dicerca), 125
dimidiata (Agathispa), 159 discoideus (Philonthus), 117 divaricata (Dohrniphora), 213
dimidiata (Ponometia), 350 discoideus rosulentus (Xyleus), 368 divaricata (Ozophora), 252
dimidiata† (Bradysia), 204 DISCOLOMATIDAE, 141 divaricatus (Holobomolochus), 18
dimidiatipennis (Trichrous), 152 Discolomopsis†, 141 divarichela† (Promecops), 173
dimidiatus (Alagoasa), 157 discolor (Archisepsis), 224 divergens (Mordella), 142
dimidiatus (Carpophilus), 137 discolor (Caribitermes), 189 divergens (Paragonatas), 251

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 598
divergentis (Neibamastax), 366 domingensis (Amphiacusta), 373 dominicana (Curiosa), 150
diversa (Camptoprosopella), 224 domingensis (Brachybaenus), 377 dominicana (Derophthalma), 244
diversa (Neohadria), 265 domingensis (Calosopsyche), 387 dominicana (Dipoena), 61
diversipes (Leiocricus), 92 domingensis (Centris), 317 dominicana (Encarsia), 290
diversus (Microcyclops), 17 domingensis (Chrysopa), 359 dominicana (Euphasiopteryx), 220
dives (Eucosmophora), 320 domingensis (Cyrtaphe), 88 dominicana (Eusarsiella), 19
divina (Omphale), 293 domingensis (Dolichopeza), 194 dominicana (Falagria), 111
divinatoria (Liposcelis), 380 domingensis (Eucypris), 20 dominicana (Glyphembia), 234
divisa (Caribovia), 264 domingensis (Geranomyia), 194 dominicana (Hadria), 265
divisa (Dicranomyia), 193 domingensis (Hapithus), 370 dominicana (Helicopsyche), 390
divisalis (Desmia), 325 domingensis (Hemerobius), 360 dominicana (Hydroptila), 388
divisata (Coremia), 333 domingensis (Hexatoma), 193 dominicana (Metepeira), 67
divisata (Xanthorhoe), 333 domingensis (Odopoea), 263 dominicana (Mimapsilopa), 233
divisus (Heuretes), 330 domingensis (Parancistrocerus), 309 dominicana (Parmenonta), 154
divisus (Macrosteles), 267 domingensis (Polycentropus), 387 dominicana (Peckia), 219
divisus (Photinus), 131 domingensis (Rhantus), 100 dominicana (Polycesta), 126
divisus (Styloleptus), 154 domingensis (Stagmomantis), 190 dominicana (Proctacanthus), 208
Dixella, 201 domingensis (Tenuipalpus), 50 dominicana (Protoparce), 362
DIXIDAE, 201 domingensis (Toxorhina), 193 dominicana (Ptenothrix), 96
Docodesmus, 89 domingensis‡ (Euphylax), 43 dominicana (Rotundabaloghia), 54
doehleri† (Ochrotrichia), 389 Domingoecius, 51 dominicana (Silis), 132
doesburgi (Passalus), 118 domingoensis (Blapstinus), 144 dominicana (Telebasis), 362
dohmi‡ (Costa), 27 domingoensis (Plinthocoelium), 150 dominicana (Trigonuropoda), 54
dohrni (Holopyga), 310 domingoensis (Xixuthrus), 155 dominicana (Trogoderma), 134
dohrnii (Paromius), 251 domingonis (Dupo), 346 dominicana (Xyalaspis), 289
Dohrniphora, 213 domingonis (Erinnyis), 345 dominicana† (Alumeda), 245
DOLABELLOPSOCIDAE, 380 domingonis (Hemiceras), 346 dominicana† (Anticedusa), 260
dolabrutum (Hydrochasma), 231 domingonis (Lauron), 348 dominicana† (Baeodasymyia), 196
Dolerocypris, 20 domingonis (Torycus), 349 dominicana† (Boreofairchildia), 194
Dolicheremaeus, 48 domingonus (Nettarhinus), 176 dominicana† (Ceratochrysis), 310
dolichodeira‡ (Krithe), 24 dominguensis (Dioryctria), 329 dominicana† (Chersodromia), 212
Dolichoderus, 298 Dominibrentus†, 168 dominicana† (Chrysonotomyia), 293
Dolichopeza, 194 dominica (Anartia), 341 dominicana† (Clonistria), 377
DOLICHOPODIDAE, 212 dominica (Astata), 312 dominicana† (Dasyhelea), 197
dolichops (Pediobius), 293 dominica (Cistogaster), 220 dominicana† (Didymia), 282
dolichoptera (Methia), 152 dominica (Rhizoperta), 134 dominicana† (Discoclavata), 134
Dolichorhinotermes, 188 dominica† (Anaxipha), 375 dominicana† (Discolomopsis), 141
dolichos (Spodoptera), 355 dominica† (Dasymutilla), 306 dominicana† (Eickwortapis), 315
dolichosoma† (Odontamera), 171 dominicae basilensis (Peripatus), 13 dominicana† (Ellipes), 364
dolichosurstyla† (Milichiella), 227 dominicae darlingtoni (Peripatus), 13 dominicana† (Hemipenthes), 210
Dolichotetranychus, 50 dominicae haitiensis (Peripatus), 13 dominicana† (Heteromyia), 197
Dolichowithius, 80 dominicae lachauxensis (Peripatus), 13 dominicana† (Hypselosoma), 239
Dolichoxenus†, 113 dominicana (Acanthoscelides), 163 dominicana† (Leptosalda), 238
dolionatis (Myriophora), 214 dominicana (Adelina), 146 dominicana† (Lindneromyia), 215
doloma (Caribovia), 264 dominicana (Adelphe), 310 dominicana† (Megaselia), 214
dolosum (Nesocyrtosoma), 147 dominicana (Antias), 243 dominicana† (Mesovelia), 239
domballah (Neolimonia), 194 dominicana (Antillusa), 111 dominicana† (Milichiella), 227
domestica (Musca), 218 dominicana (Anyphaena), 70 dominicana† (Mixorthezia), 280
domestica domestica (Rhipidia), 194 dominicana (Atrichobrunettia), 195 dominicana† (Mythicomyia), 211
domibicolor† (Forcipomyia), 197 dominicana (Biocrypta), 114 dominicana† (Neotanypeza), 222
domicambarae† (Abaristophora), 214 dominicana (Bonagota), 332 dominicana† (Nilio), 143
dominataria (Anaitis), 335 dominicana (Brasiella), 101 dominicana† (Orchestina), 59
domingalis (Phaedropsis), 326 dominicana (Callopisma), 130 dominicana† (Palaeometopia), 137
domingellus (Crambus), 323 dominicana (Carcinops), 109 dominicana† (Pieza), 211
domingensis (Ageniella), 307 dominicana (Centralaphthona), 157 dominicana† (Pintomyia), 195
domingensis (Alcimobolus), 92 dominicana (Cladochaeta), 229 dominicana† (Praecoris), 246

599 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

dominicana† (Prisciba), 262 dominicanus (Melanopleuroides), 249 dominicensis (Phytoseius), 53
dominicana† (Proplebeia), 317 dominicanus (Messatoporus), 287 dominicensis (Ramosella), 270
dominicana† (Scaptomyza), 231 dominicanus (Metacrangonyx), 30 dominicensis (Solenoptera), 156
dominicana† (Sceloenopla), 161 dominicanus (Microscydmus), 116 dominicensis (Synalpheus), 36
dominicana† (Schizoptera), 239 dominicanus (Nemomydas), 211 dominicensis (Thonalmus), 129
dominicana† (Scutigerella), 94 dominicanus (Ninyas), 250 dominicensis† (Caenocholax), 383
dominicana† (Semaeomyia), 288 dominicanus (Opius), 286 dominicensis† (Procaeculus), 48
dominicana† (Stilobezzia), 196 dominicanus (Opsablechrus), 136 Dominicicada†, 263
dominicana† (Styringomyia), 193 dominicanus (Parachelifer), 80 Dominicophus, 374
dominicana† (Triatoma), 247 dominicanus (Parachernes), 80 dominicus (Blapstinus), 144
dominicana† (Velatis), 176 dominicanus (Passalus), 118 dominicus (Episinus), 62
dominicana‡ (Cytherella), 21 dominicanus (Plectromerus), 150 dominicus (Eurygerris), 239
dominicana‡ (Cytheretta), 22 dominicanus (Scopaeus), 114 dominicus (Halictophagus), 383
dominicanae (Parvitermes), 189 dominicanus (Scydmaenus), 117 dominicus (Heteronebo), 83
dominicanense (Ametallon), 383 dominicanus (Stratus), 116 dominicus (Notapictinus), 248
dominicanensis (Ceratocapsidea), 244 dominicanus (Syneches), 213 dominicus (Pocadius), 137
dominicanensis (Charidotella), 160 dominicanus (Tachysphex), 313 dominicus (Poecilantrops), 228
dominicanensis (Eudorylas), 215 dominicanus (Trichobius), 217 dominicus† (Belaphopsocus), 380
dominicanensis (Ignelater), 128 dominicanus (Truncattus), 76 dominicus† (Cymatothes), 144
dominicanensis (Microtityus), 82 dominicanus† (Aivalykus), 285 dominicus† (Dolichorhinotermes), 188
dominicanensis (Oedichirus), 114 dominicanus† (Anopheles), 200 dominicus† (Neoliodes), 47
dominicanensis (Paracarniella), 243 dominicanus† (Atrichopogon), 197 dominicus† (Tenuicornus), 296
dominicanensis (Porphyrochroa), 213 dominicanus† (Brachypogon), 196 Dominimyza, 224
dominicanensis† (Nacaeus), 112 dominicanus† (Caryobruchus), 163 Dominiperla†, 378
Dominicanocoris, 243 dominicanus† (Discocoris), 241 dominitica (Cladochaeta), 229
Dominicanopsis†, 64 dominicanus† (Echmepteryx), 379 Dominocephala†, 296
dominicanum (Anteon), 310 dominicanus† (Femurraptor), 62 Dominopteron, 129
dominicanum (Lathrobium), 114 dominicanus† (Meloe), 148 domitia (Perithemis), 364
dominicanum (Medon), 114 dominicanus† (Miagrammopes), 60 dommeli† (Chimarra), 386
dominicanum (Mexobisium), 79 dominicanus† (Mindazerius), 275 dommeli† (Nephila), 67
dominicanum† (Cryptorhopalum), 133 dominicanus† (Minisporus), 255 dorafeliciangeliae† (Micropygomyia),
dominicanum† (Proceratium), 305 dominicanus† (Minyscapheus), 263 194
dominicanum† (Stichotrema), 383 dominicanus† (Neomachilellus), 97 dorantes cramptoni (Urbanus), 337
dominicanum† (Trypoxylon), 313 dominicanus† (Paleohemilophus), 155 Dorcasta, 154
dominicanum‡ (Cytheropteron), 22 dominicanus† (Paleotrichius), 124 dorcyi (Rutela), 122
dominicanus (Acanthoscelides), 164 dominicanus† (Porrerus), 361 doriai (Zimiris), 72
dominicanus (Agrilus), 124 dominicanus† (Pseudarachnocephalus), dorsalis (Arctodiaptomus), 16
dominicanus (Anisepyris), 309 376 dorsalis (Hedriodiscus), 206
dominicanus (Atlantasellus), 32 dominicanus† (Scopoides), 72 dorsalis (Leptotrachelus), 102
dominicanus (Bryodelphax), 13 dominicanus† (Selenops), 73 dorsalis (Leucania), 355
dominicanus (Caccodes), 132 dominicanus† (Temnothorax), 302 dorsalis (Lithocharis), 114
dominicanus (Campylothorax), 95 dominicanus† (Termitaradus), 275 dorsalis (Neolema), 161
dominicanus (Charinus), 54 dominicanus† (Tertiapatus), 13 dorsalis (Odontota), 160
dominicanus (Cletocamptus), 17 dominicanus† (Toxorhynchus), 167 dorsalis (Paralabella), 183
dominicanus (Dagus), 233 Dominicaphis†, 275 dorsalis (Pseudaptinus), 102
dominicanus (Doryctopambolus), 285 dominicata (Semiothisa), 333 dorsalis (Scirtothrips), 384
dominicanus (Euconnus), 116 dominicense (Omophron), 106 dorsalis (Sphex), 314
dominicanus (Eupsenius), 115 dominicensis (Cryptoflata), 260 dorsata (Campsomeris), 308
dominicanus (Euscelus), 166 dominicensis (Dellia), 366 dorsolinea (Caribovia), 264
dominicanus (Gastrothrips), 385 dominicensis (Episcepsis), 347 dorsotuberculata‡ (Loxoconcha), 25
dominicanus (Heraeus), 251 dominicensis (Hemipeplus), 148 Doru, 183
dominicanus (Hoplisoides), 312 dominicensis (Lasioseius), 52 Doryctobracon, 286
dominicanus (Lagynochthonius), 79 dominicensis (Metadorylas), 215 Doryctopambolus, 285
dominicanus (Lapethinus), 140 dominicensis (Mnemosyne), 259 Dorymyrmex, 298
dominicanus (Lophoscutus), 247 dominicensis (Myrmecoderus), 149 Doryphoribius, 13
dominicanus (Mediocampus), 256 dominicensis (Nelasa), 141 dosenos (Geogryllus), 372

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 600
doublierii (Diaprepes), 173 dubiosa (Ehagua), 265 Echidnophaga, 382
douglundbergi† (Thionia), 261 dubitabilis (Microcyclops), 17 echinatus (Apocephalus), 214
dowii (Ageniella), 307 dubitata (Acropyga), 302 Echinisca, 15
dowlingi (Seira), 96 dubius (Amaurochrous), 257 Echinischus, 13
Doxocopa, 340 dubius (Minilentus), 13 ECHINISCIDAE, 13
doyeni (Psorolyma), 140 ducassei (Paracentrobia), 297 echinocacti (Diaspis), 278
dozieri (Amblystira), 242 ducens (Feltia), 357 Echinocythereis, 27
dozieri (Brasa), 264 ducoudrayi (Rowlandius), 81 Echmepteryx, 379
dozieri (Ceroplastes), 275 dufai (Metacerylon), 140 eclectica (Syspira), 70
dozieri (Chrysoperla), 359 dulus (Argas), 51 eclipta (Trupanea), 223
dozieri (Marietta), 289 Dunhevedia, 14 ECNOMIDAE, 386
dozieri (Signiphora), 296 dunklei (Protoneura), 362 economoi (Strumigenys), 302
dozieri (Stizocera), 152 duo‡ (Xestoleberis), 28 ecplecticos (Hapithus), 371
dozieri (Styloleptus), 153 duodecimstriata (Stenocrepis), 104 Ectaetia, 195
dozieri (Trichophoroides), 152 duorarum notialis (Farfantepenaeus), 35 Ectemnius, 312
drakei (Aglaocoris), 325 Duplaspidiotus, 277 Ectmesopus, 159
dramba† (Voltinia), 340 duplicator (Culex), 200 ECTOBIIDAE, 186
Drapetes, 129 duponchel (Cocytius), 344 Ectocyclops, 16
Drapetis, 212 Duta, 283 Ectomyelois, 329
Drassyllinus†, 72 dyari (Mansonia), 200 ectopus† (Neivamyrmex), 299
drasticor (Cophella), 374 dylani† (Araneagryllus), 374 Ectotrypa, 371
Drepanaporus, 307 Dynamenella, 33 ectrocta (Aethosia), 349
drepanon (Pheidole), 303 Dynameniscus, 33 Ecyrus, 155
DREPANOSIPHIDAE, 275 Dynamine, 340 edentata‡ (Cyprideis), 21
DROMIIDAE, 40 Dynaspidiotus, 277 Edessa, 255
droogeri‡ (Pseudoceratina), 20 Dypterygia, 350 editha (Ctenucha), 347
Drosophila, 229 dyscheres (Geogryllus), 372 editha (Phoebis), 338
drosophilae (Spalangia), 296 Dyschirius, 106 edithae (Paracerceis), 33
DROSOPHILIDAE, 229 Dyscinetus, 123 editrix (Anomis), 351
Druciatus, 228 Dysdercus, 252 edmistoni (Hydrochasma), 231
drusilla boydi (Glutophrissa), 339 Dyseuaresta, 223 edwardsi (Ataenius), 120
dryalis (Psara), 326 Dysimia, 260 edwardsi (Lymire), 348
Dryas, 341 Dysmicoccus, 279 Efferia, 207
dryas (Bacteria), 377 Dythemis, 363 effossus† (Pachychernes), 80
Drycothaea, 154 DYTISCIDAE, 100 effrenata (Ravinia), 219
DRYINIDAE, 310 Eantis, 337 egena (Autoplusia), 357
Dryinus, 311 earinusparsus (Apodrosus), 175 eggersi (Osoriellus), 113
Drymusa, 57 eatoni (Hagenulus), 235 eggersi (Strumigenys), 302
DRYMUSIDAE, 57 ebalea (Nystalea), 347 eggersianus (Pityophthorus), 181
Dryocoetoides, 180 ebano (Thaloe), 71 eggerti (Vella), 361
Dryomites†, 182 Ebanodesmus, 88 Ehagua, 265
Dryophthorus, 172 ebanoverde (Achromoporus), 87 Eickwortapis†, 315
DRYOPIDAE, 126 ebanoverde (Metabus), 65 eickworti† (Oligochlora), 315
Dryotribus, 171 ebanoverde (Tergoceracris), 367 eickworti† (Protandrena), 315
duaguttata (Nostima), 234 ebanoverde (Tityus), 82 Eidmannella, 61
dualis (Prairiana), 268 ebena (Madoniella), 136 eidmanni (Sciocharis), 114
duan (Selenops), 73 ebena† (Didymia), 282 Einfeldia, 198
duarte (Smicridea), 387 ebenina (Conalia), 142 Eiphosoma, 288
duarteiensis (Lophocampa), 348 ebeninum (Monomorium), 303 ejuncidum (Anomalum), 287
Duartettix, 367 ebeninus (Chalcodermus), 177 Elabra, 268
dubia (Athyroglossa), 232 ebuleata (Allyda?), 333 elachia (Aedmon), 156
dubia (Ceriodaphnia), 15 Eburia, 151 elachia (Aphthona), 157
dubia (Cloridopsis), 28 Eburiola, 151 Elachiptera, 227
dubia (Metazygia), 67 ecalcarata (Archisepsis), 224 eladio (Phyllophaga), 121
dubia† (Chemmisomma), 71 eccritus (Osbornellus), 267 Elaphidion, 151
dubia† (Chrysso?), 61 Echiaster, 114 Elaphoidella, 17

601 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Elaphria, 355 elegans (Diaphania), 324 elusivus (Ommatius), 209
Elaphropus, 107 elegans (Glutoxys), 217 elutella (Ephestia), 329
Elasmopalpus, 329 elegans (Hylocorus), 178 Elytrogona, 160
Elasmus, 294 elegans (Leucophenga), 231 emarginata (Coproica), 228
ELATERIDAE, 127 elegans (Mixochlorus), 125 emarginatus† (Germarostes), 119
Elateropsis, 156 elegans (Oxycopis), 148 Embidobia, 283
elathea elathea (Eurema), 338 elegans (Rhagovelia), 240 Embioctonus, 283
Elaver, 71 elegans (Valenzuela), 381 Emblyna, 69
Elcaribe, 211 elegantissima (Anasaitis), 74 Embola, 322
electa (Siloca), 75 elegantissima‡ (Paranesidia), 19 EMBOLEMIDAE, 311
electella (Homeosoma), 329 elegantula (Leptopharsa), 242 Embolemus, 311
electra† (Agelaia), 309 elegantula (Nostima), 334 embrikstrandi (Actenodes), 125
electra† (Helicopsyche), 390 eleleus (Calisto), 343 emelyanovi (Bothriocera), 258
electra† (Microvelia), 240 ELENCHIDAE, 383 Emerita, 39
electra† (Neocorynura), 315 Elephantomyia, 193 Emersonella, 293
electra† (Zatania), 300 eleuchea dospassosi (Marpesia), 340 emersoni (Morlockia), 15
electrae (Megamelus), 259 elevans (Colpoptera), 261 emeryi (Cardiocondyla), 302
electrans† (Ctenolepisma), 97 elevata (Polygrammodes), 326 Emesa, 245
electrica† (Phaonia), 218 elevatus (Argyrodes), 61 Emesopsis, 246
electricus† (Amnestus), 254 elevatus (Euwallacea), 180 eminens (Anaxipha), 375
electrina† (Cyrtoxipha), 375 elii (Tityus), 82 emmabaconae (Wattius), 145
electrina† (Oligocixia), 259 elimatus (Trissolcus), 285 emmae (Strumigenys), 302
electrina† (Plectoptera), 187 ELIPSOCIDAE, 381 Empicoris, 246
electrina† (Strumigenys), 302 Elipsocus, 381 EMPIDIDAE, 212
Electrobates, 239 elisae (Hapithus), 370 Empiploiariola†, 246
Electroborus†, 182 elisae (Pseudococcus), 280 Empoasca, 269
electroconstrictus† (Constrictotermes), elisaensis (Ataenius), 120 empoascae (Anagrus), 295
189 elizabethae (Araneus), 66 empsychos (Hapithus), 370
electrodominica† (Chilicola), 314 ellacombei (Calliomma), 345 Empyreuma, 347
electrodominica† (Halovelia), 240 Ellipes, 365 emulgatus† (Chrosiothes), 61
electrodominicana† (Amoea), 358 Ellipodesmus, 88 Enacrosoma, 66
electrodominicanus† (Quedius), 118 ellipticus (Mesocyclops), 16 Enallagma, 362
electrodominicum† (Curtonotum), 229 ello ello (Erinnyis), 345 Encarsia, 290
electrodominicus† (Alienates), 238 ellops (Anurogryllus), 372 encollos (Walkerana), 369
electrodominicus† (Mastotermes), 188 ELMIDAE, 126 ENCYRTIDAE, 291
Electrognostus†, 134 elongata (Megachile), 316 Endeitoma, 143
electrohispaniolana† (Ablabesmyia), 199 elongata (Plesiochrysa), 359 Enderleinella, 381
Electromyrmococcus†, 279 elongata (Pseudosymploce), 187 endius (Spalangia), 296
electron† (Attagenus), 133 elongata (Pulvinaria), 276 ENDOMYCHIDAE, 141
electron† (Caccoleptus), 133 elongata (Tetragnatha), 65 Endroedianibe, 123
electron† (Coproporus), 118 elongata (Xiphocaris), 37 energos (Hapithus), 370
electron† (Limiattagenus), 133 elongata‡ (Acuticythereis), 27 engombe (Callopisma), 130
electronasutus† (Nasutitermes), 189 elongatula (Asca), 52 engombe (Rowlandius), 81
electropilosum† (Apterostigma), 301 elongatum (Belostoma), 240 engonatus (Beatadesmus), 88
Electropteron†, 129 elongatum (Strongylium), 147 Engytatus, 243
electrosimus† (Scolytodes), 179 elongatum† (Cornutidion), 61 ENICOCEPHALIDAE, 238
Electrosminthuridia†, 96 elongatus (Ataenius), 120 Enicocephalus, 238
Electrostrymon, 339 elongatus (Caribbomerus), 150 Enicospilus, 287
Electrotettix†, 365 elongatus (Carios), 51 enigmaticum (Peckidium), 288
electrum† (Antillotettix), 365 elongatus (Elcaribe), 212 enigmaticum (Trichapion), 167
electrum† (Trypoxylon), 313 elongatus (Notaepytus), 139 enigmaticus (Bothriomyrmex), 298
electrus† (Planinasus), 226 elongatus (Paratemnoides), 80 Enischnomyia, 217
electus (Antillatus), 75 elongatus† (Caulophilus), 170 ennery (Thaloe), 71
elegans (Achromoporus), 87 elongatus† (Tyrtaeus), 146 enneryensis (Achromoporus), 87
elegans (Augochlora), 315 Elousa, 352 enneryensis (Haplocyclodesmus), 91
elegans (Derancistrus), 156 Elpidium, 24 Enochrus, 108

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 602
Enocomia, 262 Epilobocera, 44 erigoniformis (Steatoda), 62
ENOPLOMETOPIDAE, 37 epimeces (Stenogryllus), 371 Erinnyis, 345
Enoplometopus, 37 Epimecis, 333 ERIOCOCCIDAE, 279
Enosmaeus, 150 epimeles (Hapithus), 371 Eriococcus, 279
enriquillo (Modisimus), 58 epimonon (Cycloptilum), 376 Eriophora, 66
enriquillo (Selenops), 73 Epinannolene, 93 eriophorus (Ceratopogon), 196
ensifer duvali (Hydrophilus), 108 Epinysson, 312 Eriophyes, 48
enstates (Walkerana), 369 epionata (Patalene), 334 ERIOPHYIDAE, 48
Entedononecremnus, 293 Epiphragma, 193 Erioptera, 192
ENTHOCYTHERIDAE, 23 Epiplatea, 222 Eriosoma, 275
Entocythere, 23 Epipompilus, 307 Eriphia, 44
ENTOMOBRYIDAE, 95 EPIPSOCIDAE, 380 Eris, 75
Entylia, 270 Epipsocus, 380 erithacus (Euxestus), 140
Entypus, 307 EPIPYROPIDAE, 330 eritheles (Amphiacusta), 373
Enyo, 345 EPISACTIDAE, 366 ernestoi (Nops), 59
Eobates, 239 Episcepsis, 347 ernsti ernsti (Newportia), 85
Eocader, 242 Episcirrus†, 171 erosa (Anomis), 351
Eohomopterus, 106 Episimus, 331 erosa (Epiplatea), 222
Eophrixus, 32 Episinus, 62 erosus (Glyptoxanthus), 44
Eoromus, 90 Episyron, 307 EROTYLIDAE, 139
EOSENTOMIDAE, 96 Epitausa, 352 errabundus (Hapithus), 370
Eosentomon, 96 Epitragus, 144 errans (Cyrtoxipha), 375
Eospilopsyllus, 382 Epitrix, 157 erraticus (Culex), 200
eowilsoni† (Apterostigma), 301 epochthidius (Antillengis), 139 Erucoblatta†, 187
epagogos (Hapithus), 371 Eprhopalotus, 293 eruditus (Hypothenemus), 178
Epaphrodita, 190 Epyris, 310 eruptor (Metaphycus), 292
EPAPHRODITIDAE, 190 equi (Gasterophilus), 219 Erynnis, 337
Epargyreus, 337 equi (Werneckiella), 379 erysimi (Lipaphis), 274
epaxia (Margarettia), 369 equinoxialis (Drosophila), 230 Erythemis, 363
epaxia (Velapia), 373 Erastria, 333 Erythmelus, 295
epepye (Modisimus), 58 eratos (Nemobiopsis), 373 ERYTHRAEIDAE, 49
eperatos (Anaxipha), 375 eratus (Osbornellus), 267 erythrocephala (Deuterixys), 286
ephelipticus (Hepatus), 40 Ercta, 325 erythrocephala (Ornithoctona), 217
ephemeraeformis (Thyridopteryx), 320 EREBIDAE, 347 erythrocera (Leptocera), 228
Ephestia, 329 erebus (Corythalia), 75 erythrocera (Notiphila), 234
ephippium (Bromeloecia), 228 Erechthis, 367 Erythrodiplax, 363
ephippium ephippium (Myzine), 308 erecta (Anthophila), 351 Erythrolychnia, 130
epholcos (Hapithus), 370 erectoides† (Theridion), 63 erythrolytris (Robopus), 131
Ephuta, 306 erectum† (Theridion), 63 erythropus (Piestus), 115
Ephydra, 233 erectus (Hypothenemus), 178 erythroscelis (Loxostethus), 146
EPHYDRIDAE, 231 erectus (Trachelas), 72 erythrotermus (Spirobolellus), 91
Ephyriades, 337 Erelieva, 329 Espagnola, 366
Ephyrodes, 352 Eremaeozetes, 46 Espagnoleta, 366
epia (Hygronemobius), 373 EREMAEOZETIDAE, 46 Espagnolina, 366
Epialtus, 42 Eremochrysa, 359 Espagnolopsis, 366
Epiblema, 331 Eremoleon, 361 espanola (Cyrtognatha), 65
Epicorsia, 325 Eremonidia, 346 esperanza (Nylanderia), 300
epicrana† (Lissantauga), 127 eresimus tethys (Danaus), 341 Espeson, 112
Epidapus, 204 Eretmocerus, 290 espina (Phyllophaga), 121
epidendrana (Mordellistena), 142 Eretmocoris, 248 espinii (Psallodia), 263
epidendri (Asterolecanium), 276 ergaticos (Miogryllus), 372 espoloni (Diastolinus), 144
epidendrios (Hapithus), 370 Eriastichus, 294 esquinada (Phyllophaga), 122
Epidromia, 352 eribombos (Ectotrypa), 372 estebani (Diastolinus), 144
Epierus, 110 eribombos (Hapithus), 370 estebani (Taeniogonalos), 282
Epilampra, 185 ericata (Syllectra), 355 Estenorhinus, 168
epilepsis (Petrusa), 261 eridania (Spodoptera), 356 Estola, 154

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estradai (Chrysopa), 359 Euides, 259 Eurytoma, 295
estuariensis (Alpheus), 36 Eukoenenia, 78 EURYTOMIDAE, 294
esulcatus (Spirobolellus), 91 EUKOENENIIDAE, 78 Eusarsiella, 19
ethelae (Phylolestes), 362 Eulechriops, 170 Euscelus, 166
Ethirothrips, 385 Eulepidotis, 352 Euscepes, 171
ethlius (Calpodes), 336 Eulepte, 325 Euschistus, 255
Ethmia, 321 Eulimnadia, 14 Euseius, 53
Etiella, 329 Eulimnichus, 126 Eusicyonia, 35
etiennei (Ceratogramma), 297 EULOPHIDAE, 292 Eustala, 66
Euacidalia, 335 Eumacrodes, 335 eustrophoides (Amblycerus), 163
Eualebra, 268 eumeces† (Azteca), 298 Eutelia, 355
Euaresta, 223 Eumecosoma, 209 euterpiformis (Pyrisitia), 338
Euarestoides, 223 Eumecosomyia, 223 Eutetranychus, 50
Eublemma, 349 Eumenes, 308 Euthlastoblatta, 187
eubolialis (Glympis), 352 eumenes (Antillacris), 366 EUTYPHLOCIIDAE, 235
Euborellia, 183 Eumorpha, 346 Euwallacea, 180
Eubule, 254 Eunica, 340 Euxesta, 223
eubule (Conura), 291 Eunomia, 347 Euxestonotus, 283
Eucaerus, 103 Euodynerus, 308 Euxestus, 140
Eucalymnatus, 276 eupatorivora (Calycomyza), 225 evadne (Parhaplopus), 377
Eucatops, 111 EUPELMIDAE, 294 evai‡ (Proteoconcha?), 27
Eucereon, 347 Eupelmus, 294 evanescens† (Stizocera), 152
Euceroplatus, 203 Euphasiopteryx, 220 Evania, 288
Euceropsylla, 271 euphorbiae (Macrosiphum), 274 evanidus (Hapithus), 371
euchara† (Geranomyia), 194 Euphorticus, 103 EVANIIDAE, 288
eucharis (Phrynus), 54 euphrosyne (Alisotrichia), 388 evansi (Orasema), 292
eucharis† (Trypoxylon), 313 Euphyes, 336 evansi (Tetranychus), 50
EUCHARITIDAE, 292 Euphyia, 335 evarete (Junonia), 342
Euchirograpsus, 41 Euphylax, 43 evectus (Typhlodromus), 53
Euchlaena, 333 eupithecioides (Renodes), 354 evelinae (Doryphoribius), 13
euchlorus (Cephaloplopus), 377 Euplatypus, 182 everiata (Macaria), 333
Euchroa, 105 Euplectrus, 294 evides (Cophella), 374
euclastos (Anaxipha), 375 Eupogonius, 154 evoplon (Cophella), 374
EUCNEMIDAE, 127 Eupsenius, 115 evsyunini† (Leptopharsa), 242
Euconnus, 116 Euptoieta, 341 ex gr. asperrima‡ (Henryhowella), 27
Eucosmophora, 320 Eupyrrhoglossum, 345 ex gr. complexa‡ (Eucytherura), 23
eucrines (Cyrtoxipha), 375 Eurema, 338 ex gr. dictyon‡ (Bradleya), 26
Eucyclops, 16 Eurhabdus, 209 ex gr. dreikanter‡ (Pontocypris), 26
Eucypris, 20 Eurhinus, 169 ex gr. johnsoni‡ (Cytherura), 23
Eucytherura, 23 Eurhopalothrix, 301 ex. gr. angulata‡ (Candona), 20
eudami (Horismenus), 293 Eurocalia, 260 exacerbans (Amphiacusta), 373
Euderus, 293 europaeum (Pseudectroma), 292 exacta (Anomis), 351
Eudocima, 352 Europs, 137 exacta (Efferia), 208
Eudorylas, 214 euryaspus† (Semnorhynchus), 172 exallos (Stenogryllus), 371
Eueides, 341 Euryconus, 271 exaltata (Espagnolopsis), 366
Eugamandus, 153 Eurycotis, 184 exarata (Bitoma), 143
eugenii (Anthonomus), 169 Eurycyde, 45 exaratus (Cryptocephalus), 161
Eugeonemus, 174 Eurydice, 31 exasperatus (Callinectes), 43
Euglossa, 317 Eurygerris, 239 exasperatus (Thinobius), 113
Eugnosta, 332 Eurypanopeus, 44 excavans (Bolina), 353
Eugonops, 165 Eurypedus, 160 excavata (Manidia), 333
Eugonus, 165 euryrhina† (Apharosoma), 171 excavata (Toxonprucha), 355
eugraphiata (Fidonia), 333 Euryscelis, 150 excavator (Geogryllus), 372
eugraphis (Prionapteryx), 323 eurystegus† (Semnorhynchus), 172 excavatum (Platydema), 146
Euhybus, 213 eurysternus (Haematopinus), 379 excavatum (Trypoxylon), 313
Euhyloptera, 260 Eurytium, 44 excavatus (Hortoonops), 59

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 604
excavatus (Neolargulus), 252 extenta (Amphiacusta), 373 fascifrons (Macrosteles), 267
excavatus group (Tanytarsus), 198 exterior (Chrysopa), 359 fascinans (Amphiacusta), 373
excellens (Phoebolampta), 368 externa (Chrysoperla), 359 fasciolaris (Melipotis), 353
excelsa† (Araneometa), 66 externata (Euacidalia), 335 fastidiosum (Bembidion), 107
exceptus (Hypothenemus), 178 extersignata (Lema), 161 fastigata† (Meioneta), 64
excisus (Anisepyris), 309 extincta† (Achaearanea), 61 fauroti (Micropholcus), 57
excisus (Aspidiotus), 277 extrema (Agalliopsis), 264 fausta (Tettigella), 264
excisus (Teloplatypus), 182 fabae (Aphis), 274 faustinum (Macrobrachium), 37
excitans (Oraesia), 354 fabalis (Empoasca), 269 fauveli (Cyrtinus), 154
Excorallana, 31 fabricii (Pachycoris), 258 fauveli (Osorius), 112
excoriata (Edessa), 255 fabricii (Selenophorus), 102 favorabilis (Pityophthorus), 181
Exechia, 203 faciale (Hydrochasma), 231 fax (Signiphora), 296
Exechiopsis, 203 factaria (Geometra), 334 Fecelia, 256
Exelastis, 330 Fageicera, 57 Felicola, 379
exigua (Ponometia), 349 Faiditus, 62 felisana (Argyrotaenia), 332
exigua (Spodoptera), 356 Falagria, 111 Feltia, 357
exiguus (Clavilispinus), 112 falcaorum† (Pintomyia), 195 Feltiella, 202
exiguus (Halicyclops), 16 falcaria (Gnamptogenys), 299 feminea† (Grossoxipha), 373
exiguus (Zeadolopus), 110 falcarius (Haplocyclodesmus), 91 femoralis (Fannia), 218
exilipes† (Acamptus), 170 falcata (Fulakora), 298 femoralis (Menudo), 256
exilis (Smicrips), 137 Falconia, 244 femoralis (Oncopeltus), 250
exilis (Stereodermus), 168 falini† (Caulophilus), 170 femorata (Conura), 291
exilis isophthalma (Brephidium), 339 falli (Oxycopis), 148 femoratus (Elateropsis), 156
exilis‡ (Ambocythere), 27 Fallia, 141 femoratus (Modisimus), 58
eximia (Disonycha), 157 famelica (Masebia), 352 Femurraptor, 62
eximia (Lucilia), 217 famelica (Melipotis), 353 fenestra (Stenotabanus), 207
eximius (Apodrosus), 175 familiaris (Tinea), 320 fenestralis (Zeros), 234
eximius (Phytocoris), 244 Fannia, 218 fennahi (Baryscapus), 294
exis (Plectromerus), 150 FANNIIDAE, 218 Fennahiella, 243
Exitianus, 266 fantastica (Trogoderma), 134 Fennellia, 90
exitiosus (Exitianus), 266 Farfantepenaeus, 35 Fennerosquilla, 28
exoleta† (Bradysia), 204 farri (Hispaniolara), 126 fera† (Dicranomyia), 193
Exoma, 260 fascialata (Mangora), 67 ferdinandi (Leptoteleia), 284
Exomalopsis, 317 fascianota† (Periscelis), 226 fernaldi (Enicospilus), 287
Exophthalmus, 173 fasciapenna (Plecia), 202 fernandezi (Amblyseius), 53
exophthalmus (Enicocephalus), 238 fasciapennis (Aleurodothrips), 385 Feroseta†, 361
Exoprosopa, 210 fasciata (Apenes), 103 ferox (Psorophora), 201
Exosphaeroma, 33 fasciata (Paraclivina), 107 ferreri (Lasioglossum), 315
exotera (Chrysoperla), 359 fasciatai (Pseudoxenos), 383 ferreus (Phrygionis), 334
exotica (Ligia), 32 fasciatum (Sceliphron), 314 ferrigeralis (Salia), 357
exotica† (Ogcodes), 206 fasciatus (Alagoasa), 157 ferrilinealis (Megatomis), 356
expansa (Amphiacusta), 373 fasciatus (Caribo), 256 Ferrisia, 279
expansa (Asteia), 225 fasciatus (Dagbertus), 244 ferrocana (Casandria), 352
explicatrix (Antillacris), 366 fasciatus (Monobelus), 270 ferruginea (Bothriocera), 258
exprimens (Ephyrodes), 352 fasciatus (Ocyptamus), 216 ferruginea (Neomida), 146
exquisitus (Allothenemus), 178 fasciatus (Oedionychus), 159 ferruginea (Nyctiplantus), 159
exquisitus† (Cenocephalus), 182 fasciatus (Oncopeltus), 250 ferruginea (Platydema), 146
exscindens (Mocis), 354 fasciatus (Osbornellus), 267 ferruginea (Procula), 140
exsculpta (Ochthera), 232 fasciatus (Polypsocus), 381 ferruginea (Sethenira), 254
exsertus (Stenodontes), 155 fasciatus (Sanctanus), 266 ferruginea (Strabala), 158
exsertus† (Choragus), 164 fasciatus (Sargus), 206 ferruginea group (Orthemis), 364
exstincta† (Hemitrinemura), 97 fasciatus (Tylozygus), 265 ferrugineus (Callicentrus), 270
exstinctus† (Anochetus), 399 fasciatus fasciatus (Eumorpha), 346 ferrugineus (Cryptolestes), 138
exstriatum (Phaenonotum), 109 fascicularis (Pherhimius), 127 ferrugineus (Xyleborus), 180
extenda (Agalliopsis), 264 fasciculatus (Araecerus), 164 ferrugineus bicolor (Tachypompilus), 308
extensus (Eucyclops), 16 fasciculus (Wulfila), 71 ferrugineus ferrugineus (Scolopocryp-

605 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

 tops), 85 flagellata (Symploce), 186 flavoantennatus† (Tyrtaeus), 146
fervens (Dysdercus), 252 flagellifera† (Selenyphantes), 64 flavoapicalis (Leptolycus), 129
fervida (Erythrodiplax), 363 flammea (Calappa), 40 flavofemorata (Silis), 132
fervida (Myophora), 220 flammea‡ (Calappa), 40 flavolobata (Silis), 132
fervidalis (Parachmidia), 327 flammicincta (Anthophila), 349 flavomaculata (Acrosoma), 67
fessonia lapitha (Adelpha), 341 flammipennis (Pompilus), 307 flavopicta (Spilochalcis), 291
festina (Anaxipha), 375 FLATIDAE, 260 flavosa (Banasa), 256
festina (Lycosa), 68 Flatoides, 261 flavoscutellatus (Enicospilus), 287
festina (Tetragnatha), 65 Flatoidinus, 261 flavostigma (Enicospilus), 287
festinus (Spissistilus), 271 Flatormenis, 261 flavothoracicus (Epidapus), 204
festiva (Chlorida), 153 flava (Anomis), 351 flavus (Curtomerus), 151
festiva (Coptodera), 103 flava (Eris), 75 flavus (Doryphoribius), 13
festiva (Diphthera), 351 flava (Signiphora), 296 flavus (Enicospilus), 376
festiva (Hyperaspis), 140 flava (Sipha), 273 flavus (Truncattus), 76
festivus (Eurhinus), 169 flavedana (Platynota), 332 flegia (Palpita), 326
ficorum (Gynaikothrips), 385 flavens (Pheidole), 303 flesalis (Hypenula), 356
ficticia (Amphiacusta), 374 flaventis (Apachekolos), 209 fletcheri (Chonocephalus), 214
fictilis (Zale), 355 flaveola (Mansonia), 200 Fletcherimyia, 219
fictilium (Rhompaea), 62 flaveolus (Anagrus), 295 flexa (Drosophila), 230
Fictotama, 60 flaveolus (Parvitermes), 189 flexa (Perasia), 354
ficus (Pachylia), 345 flavescens (Acanthoscelides), 163 flexilobata (Silis), 132
ficus (Planococcus), 280 flavescens (Balclutha), 266 flinti (Diphuia), 232
Fidiobia, 283 flavescens (Linepithema), 298 flinti (Stenognatha), 348
FIGITIDAE, 289 flavescens (Ommatius), 209 floccosus (Aleurothrixus), 272
figularis (Conoderus), 128 flavescens (Pantala), 364 floccosus (Geopemphigus), 275
filamentarius (Tytthonyx), 133 flavibrunnea (Tripudia), 355 flocculus (Tabanus), 207
filamentosus (Nipaecoccus), 279 flaviceps (Pseudomethoca), 306 flora (Pyrellia), 218
filicornis (Gondwanocrypticus), 146 flaviceps (Sphaerocera), 229 floralis (Omonadus), 149
filidigitus (Synalpheus), 36 flavicollis (Coproporus), 118 floralis (Toxomerus), 216
filifera (Collomena), 355 flavicoma (Aedmon), 156 florella (Syngamia), 327
filiformis (Leptocorisa), 253 flavicorne (Dactylosternum), 109 florentissima (Pseudolycoriella), 205
filipalpis† (Pintomyia), 195 flavicornis (Asteriza), 160 floricola (Monomorium), 303
FILISTATIDAE, 57 flavicosta (Planicephalus), 266 Floricomus, 64
filum (Scopaeus), 114 flavicrus (Spalangia), 296 floridae puertoricensis (Delphacodes),
filum brevialatum (Spalacopsis), 154 flavida (Drapetis), 212 259
filum filum (Spalacopsis), 154 flavida (Nostima), 234 floridana (Bakeriella), 309
fimbriata (Titanogrypa), 219 flavida (Pseudophorellia), 223 floridana (Balclutha), 266
fimbriatus chiltoni (Paracyclops), 17 flavidulus (Pseudomyrmex), 306 floridana (Eurhopalothrix), 301
fimbriatus† (Ommatius), 209 flavidulus (Temnothorax), 302 floridana (Leucochrysa), 359
Fimbricyclops, 16 flavidus (Monobelus), 270 floridana (Lindingaspis), 277
fimbrillus† (Ommatius), 209 flavidus (Ophion), 288 floridana‡ (Puriana), 24
finduilasi (Anisepyris), 309 flavigaster (Cephalotes), 301 floridanum (Coleosoma), 61
finneganae (Persephona), 41 flavilabris ubancus (Selenophorus), 103 floridanus (Dolichotetranychus), 50
Fiorinia, 278 flavipennis (Barychara), 111 floridanus (Geocoris), 249
fioriniae (Fiorinia), 278 flavipes (Brachymeria), 291 floridanus (Pilumnus), 44
fischeri‡ (Loxoconcha), 25 flavipes (Carpelimus), 113 floridanus (Tenebroides), 135
fisheri (Eburia), 151 flavipes (Discocerina), 231 floridanus (Tyrannochelifer), 80
fisheri (Polycesta), 126 flavipes (Euplectrus), 294 floridata (Nepheloleuca), 333
fisheri (Trichrous), 152 flavipes (Karnyothrips), 385 floridata (Pterocypha), 335
Fissicrambus, 323 flavipes (Liohippelates), 227 floridensis (Callopistria), 350
fissiradiellus (Fissicrambus), 323 flavipes (Pinophilus), 114 floridensis (Ceroplastes), 275
fissuralis (Crocidomera), 328 flavipes (Zeros), 234 floridensis (Melanagromyza), 224
fistulosum† (Plaumannion), 284 flavipes flavipes (Pentagonica), 104 floridensis (Trialeurodes), 273
fithianae‡ (Eucytherura), 23 flavipes picipes (Pentagonica), 104 Florinda, 64
flabellatus (Ablechroides), 136 flavitarsis (Notaepytus), 139 florissantensis† (Metachroma), 162
flageliformis† (Corinna), 71 flaviventris (Lymaenon), 295 florum (Thrips), 385

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 606
fluviatile (Diaphanosoma), 14 fractilateralis (Platynus), 105 fuesslini (Theridion), 63
foliacea (Antianthe), 270 fractilinea (Platynus), 105 fugitans (Antilliclodes), 374
Folsomides, 95 fractus (Achromoporus), 87 FUHRMANNODESMIDAE, 89
fontana (Lemalelmis), 126 fragaefolii (Chaeosiphon), 274 Fulakora, 298
fontanai (Cochliomyia), 217 fragilella (Lepyrotica), 320 fulgens (Apraea), 157
fontismoreaui (Metrichia), 389 fragilis (Jocara), 327 fulgens (Entedononecremnus), 293
foraminosus† (Lorelus), 143 fragilis (Wulfila), 71 fulgida (Erythrolychnia), 130
forbesi (Aphis), 274 fragistyla (Pseudosciara), 205 fulgidus (Anoplius), 307
forceps (Lupella), 43 francescellus (Microcrambus), 323 Fulgoraecia, 330
forceps (Mithrax), 42 francisca (Apate), 134 fuliginosa (Cassida), 160
forcipata (Micrathena), 67 franciscana (Artemia), 14 fuliginosa (Zale), 355
forcipitigerus (Brentus), 168 franciscana (Nostima), 234 fuliginosus (Cryptocephalus), 161
Forcipomyia, 197 franciscanus (Limnogonus), 239 fullonium (Elaphidion), 151
forda‡ (Loxoconcha), 25 franciscanus (Oliarus), 259 fulva (Leptocera), 228
forficifera (Pallenopsis), 45 franciscoi (Calisto), 343 fulva (Nylanderia), 300
forficula (Lestes), 362 Frankliniella, 384 fulva (Phaleria), 147
FORFICULIDAE, 183 Franklinothrips, 384 fulvibarbis (Efferia), 208
formicaria (Licracantha), 153 fratellus (Antillosorius), 112 fulviguttata (Atteva), 323
formicarius (Cylas), 168 fratellus (Cleogonus), 176 fulvipes (Carpelimus), 113
formicarius (Micromyrmex), 172 frater (Ataenius), 120 fulvipes (Piestus), 115
FORMICIDAE, 298 frater† (Leptopharsa), 242 fulvipes (Podium), 313
Formicilabia, 183 fraterna (Antillotrecha), 83 fulvitarsis (Notaepytus), 139
formoculus† (Apionion), 167 fraterna (Aphthona), 157 fulviventris (Centris), 317
formosa (Encarsia), 290 fraterna (Chalcosicya), 162 fulvus (Ogulnius), 63
formosa (Prytanes), 251 fraternalis (Renia), 334 fulvus (Psalistops), 56
formosoides (Anachrostomus), 249 fraternus (Anelosimus), 61 fumata (Coniopteryx), 360
formosus (Anachrostomus), 249 fraternus (Megadytes), 100 fumator (Mimogonus), 112
formosus (Descanso), 75 fratrellus (Titanophilus), 85 fumidus fraterculus (Camponotus), 299
formula (Robinsonia), 349 fraudans (Hapithus), 370 fumidus illitus (Camponotus), 299
formularis (Lesmone), 353 frazari (Chrysops), 206 fumidus imbecillus (Camponotus), 299
fornicatus (Euscelus), 166 freemani (Chrysopa), 359 fumidus soulouquei (Camponotus), 299
fortipes (Sarcophaga), 219 fremens (Stenogryllus), 371 fumipennis (Heteromeringia), 226
fortis (Efferia), 208 frenatus (Exophthalmus), 173 fumoalata ( Cratyna), 204
fortunei (Adesmus), 154 frequens (Trichobius), 217 Fundella, 329
FORTUYNIIDAE, 46 Frevillea, 41 funebris (Anthrax), 210
fossatus (Gymnochthebius), 110 freygessneri (Paratettix), 365 funebris† (Stomatomiris), 244
Fossilaraneus†, 66 freytagi (Antillonirvana), 267 funeralis (Desmia), 325
fossilis† (Apicrenus), 246 friabilis‡ (Limnocythere), 25 funeratus† (Holisus), 117
fossilis† (Floricomus), 64 frisia (Anthanasa), 342 funesta (Thermesia), 354
fossilis† (Homalometa), 65 fritzmuelleri (Synalpheus), 36 furca† (Nephila), 67
fossilis† (Neuraphes), 117 frivolus (Aeolus), 128 Furcaspis, 277
fossilis† (Palaehydropsyche), 387 froeschneri (Myodocha), 251 furcata (Brachymesia), 363
fossilis† (Pycnosinga), 68 Froeschneria, 251 furcata (Caribovia), 264
Fossilopaea, 59 frontalis (Cyclocephala), 123 furcata (Eutelia), 355
fossor† (Gnathotrichus), 182 frontalis (Euscepes), 171 furcata (Heterobelba), 47
fossoria (Phyllophaga), 122 frontalis (Leucophenga), 231 furcata (Lepyronura), 262
fossula (Scarabaeus), 123 frontalis (Pagiocerus), 177 furcata (Microsepsis), 224
fossulatella (Oryctometopia), 329 frontalis (Scapanea), 363 furcata (Popcornella), 76
fossulatus† (Paleosinus), 182 frontalis (Tangyria), 262 furcata (Stenocionops), 42
fotuto (Margarettia), 369 fronterizus (Truncotettix), 365 furcatus (Argyrodes), 61
Fountainea, 340 Frontinella, 64 furcatus (Cyphoceratus), 262
foveolatus (Helodytes), 175 frosti (Spintharus), 62 furcatus (Neospintharus), 62
foxi (Asca), 52 frugiperda (Spodoptera), 356 furcatus (Orthobelus), 270
foxi (Culicoides), 196 frustrana (Rhyacionia), 332 furcatus (Styraxodesmus), 90
foxi (Glenognatha), 65 frutetosuccinea† (Forcipomyia), 197 furcianus (Anadenobolus), 92
fractifera (Gonitis), 351 fuentesi (Trichogramma), 297 furcianus (Spirobolellus), 91

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furcifera (Stenocoris), 253 Ganaspidium, 289 Geogryllus, 372
furcipes (Achromoporus), 87 Gangamocytheridea, 21 geometricus (Latrodectus), 62
furcula (Pentatax), 49 Ganyra, 339 GEOMETRIDAE, 333
furens (Culicoides), 196 garcia (Ischnothele), 56 Geopemphigus, 275
furfurosa (Solenoptera), 156 garciai (Culex), 200 GEOPHILIDAE, 85
furthi (Homoschema), 158 Garcialdretia, 381 geophilus (Dominicodesmus), 90
furtiva (Anaxipha), 375 garciamarquezi† (Archiokrisna), 266 georgei† (Spalangiopelta), 296
furva (Chaetoclusia), 225 Gargaphia, 242 georgiae (Morion), 104
furva illicta (Banisia), 323 garnoti (Perasia), 354 Geotrupes, 119
fusca (Heteropsylla), 271 garrota (Phyllophaga), 122 GEOTRUPIDAE, 119
fusca (Scytodes), 57 Garyops, 79 Geranomyia, 193
fusca (Spartocera), 254 Gasteracantha, 66 Geratozygops†, 170
fuscaspecula (Nylanderia), 300 Gasterophilus, 219 geratus† (Tityus), 82
fuscifascia (Bendis), 353 GASTERUPTIIDAE, 288 germanica (Blattella), 186
fuscinervis (Rachispoda), 229 Gasteruption, 288 Germarostes, 119
fuscipennis (Amitus), 283 gastralis (Eulepte), 325 Geron, 211
fuscipennis (Aphytis), 289 gatunensis‡ (Puriana), 24 GERRIDAE, 239
fuscipennis (Lymeon), 287 gavisa (Hapithus), 371 gerstaeckeri (Ghilianella), 246
fuscipennis (Ozophora), 252 gaweini (Tethysbaena), 29 gerstaeckeri (Trachypus), 313
fuscitarsis (Amblystira), 242 gazella (Digitonthophagus), 120 Gerstaeckeria, 171
fuscivenosa (Pseudolycoriella), 205 Gea, 66 gertschi (Meriosfera), 77
fusciventris (Ornithoctona), 217 Geaya, 78 gertschi (Nops), 59
fuscocorpus† (Toxorhynchus), 167 GECARCINIDAE, 40 gerulus (Polyancistrus), 368
fuscosignum (Peridinetus), 170 Gecarcinus, 41 Gesta, 337
fuscovittata (Eustala), 66 geiseri† (Tytthonyx), 133 gesta (Gesta), 337
fusculus (Glyphonyx), 129 geisteri‡ (Caribbella), 26 Ghilianella, 246
fuscum (Psomizopelma), 294 gelabertae (Rhagoletotrypeta), 223 ghiliani (Spongovostox), 183
fuscus (Araptus), 180 gelania gelania (Adelpha), 341 Ghinallelia, 246
fuscus (Chrysanthrax), 210 GELECHIIDAE, 321 gibba (Lebia), 103
fuscus (Modisimus), 58 gelidalis (Pyralis), 324 gibba (Lepidosina), 228
fusilinea (Thermesia), 351 gelinasus (Scalaventer), 175 gibba (Octhispa), 160
fusilineata (Pyrinia), 334 gemina (Chalcosicya), 162 gibberosa (Zatania), 300
fusiparamera† (Forcipomyia), 197 geminata (Eburiola), 151 gibberum† (Proceratium), 305
Fustiger, 115 geminata (Oxyethira), 389 gibbesi (Albunea), 39
gabbi‡ (Portunus), 43 geminata (Solenopsis), 304 gibbesi (Percnon), 41
Gabronthus, 117 geminus (Ommatius), 210 gibbicollis (Thionia), 261
gagates (Belonuchus), 117 gemmata (Elytrogona), 160 gibbipennis (Leiopleura), 124
gagates (Myrmelachista), 300 gemmatalis (Anticarsia), 351 gibbosa (Neralsia), 289
gaitoi (Hemerobius), 360 gemmatus (Lithophorus), 139 gibbosus (Mucrotettix), 365
gala (Aprostocetus), 294 genalis (Porphyrochroa), 213 giesberti (Hormathus), 152
GALATHEIDAE, 39 genaroi (Blapstinus), 144 giffardi (Dirhinus), 291
Galendromus, 53 genaroi (Caribitroides), 30 giganteus (Blaberus), 185
Galerita, 102 genialis (Metazygia), 67 giganteus (Ceroplastes), 275
galerita‡ (Aurila), 23 geniculata (Paroxacis), 148 giganteus (Megadytes), 100
Galethalea, 347 geniculata (Zosis), 60 gigas† (Discothyrea), 305
Galgupha, 255 geniculatus (Engytatus), 243 gilippus cleothera (Danaus), 341
galianoae† (Lyssomanes), 76 geniculatus (Lutrochus), 127 gilva (Chaetosciara), 204
galii choneupsilon (Calisto), 342 genoveva (Junonia), 342 gilvimacula (Chimarra), 386
galii galii (Calisto), 342 gentilis (Ophiomyia), 225 gilvipes (Nostima), 234
galipote (Masteria), 56 gentilis (Stenus), 118 gilvum (Sympetrum), 364
gallegosi (Enicospilus), 288 gentilis suavis (Laccophilus), 101 gimmeli (Cercyon), 109
Galleria, 328 Genus near Fallopia, 49 giovannolii (Nostima), 234
gallinacea (Echidnophaga), 382 Geococcus, 279 girschneri (Psilopa), 233
gallinae (Menopon), 379 GEOCORIDAE, 249 glabalus (Lipurometriocnemus), 198
gallus (Calappa), 40 Geocoris, 249 Glabellula, 211
Gampsocoris, 248 Geograpsus, 41 glaber (Calliscelio), 283

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glaber (Miraces), 159 Glyptapanteles, 286 grabhami (Lysmata), 36
glaber (Scolytodes), 179 Glyptobairdia, 19 graciella (Cydosia), 349
glabra (Caribovia), 264 Glyptoscelis, 162 gracilenta (Aedmon), 156
glabra (Chalcosicya), 162 Glyptotermes, 188 gracilicornis (Chariesterus), 253
glabra (Iomoides), 90 Glyptoxanthus, 44 Gracilidris, 298
glabra‡ (Cardobairdia), 19 Gnamptogenys, 299 graciliforceps (Calinda), 272
glabratus (Enicospilus), 288 Gnaphosa, 72 gracilior (Siphonophora), 91
glabricristata (Nephrotoma), 194 GNAPHOSIDAE, 72 gracilipalpis tenuis (Atax), 49
glabriuscula (Lysiosquilla), 28 Gnatholeptus, 180 gracilipes (Podochela), 42
glabroguttata (Aedmon), 156 GNATHOPHYLLIDAE, 37 gracilipes (Prostemmiulus), 93
glabrus (Elcaribe), 212 Gnathophyllum, 37 gracilipes (Wulfila), 71
gladiator (Diastolinus), 144 Gnathopleura, 285 gracilis (Antillatus), 75
gladioli (Taeniothrips), 385 Gnathotrichus, 182 gracilis (Ataenius), 120
glaesaria† (Acropyga), 302 Gnathotrupes, 181 gracilis (Baliomydas), 211
glaesaria† (Chrysopa?), 359 gnomalis (Botys), 326 gracilis (Dorcasta), 154
glaesaria† (Leucospis), 295 gnomus (Nanolycus), 129 gracilis (Ephydra), 233
glaesaria† (Spiloconis), 360 Gnophomyia, 192 gracilis (Holepyris), 310
glaesaria† (Thaumatosoma), 316 godartiana godartiana (Aphrissa), 338 gracilis (Leucopodella), 216
glaesarius† (Glyptotermes), 188 godmani (Pantomorus), 175 gracilis (Marilia), 390
glaesi† (Myrmecholax), 383 godmani (Thiotricha), 321 gracilis (Oxidus), 89
glaesum (Ignelater), 128 Goeldichironomus, 198 gracilis (Pachygrapsus), 41
glaphyropus (Athyroglossa), 232 goeleti† (Zorotypus), 391 gracilis (Pheneps), 127
glareosus (Anotylus), 113 Goetheana, 293 gracilis (Tricolus), 181
glauca glauca (Oreodera), 154 gomezi (Leucospis), 295 gracilis (Xyleborinus), 180
glaucescens (Flatormenis), 261 gomezi (Leucotrichia), 389 gracilis† (Chilicola), 314
glaucescens (Perigonia), 345 gomezi (Pseudoapseudomorpha), 29 gracilis† (Pholcophora), 58
glaucipennis (Bolina), 353 gomezmenori (Orthobelus), 270 gracilis† (Troctopsocoides), 380
glaucocephala (Wyeomyia), 201 GOMPHIDAE, 363 gracilispina (Silis), 132
glauculalis (Diaphania), 325 Gomphochernes, 80 graciliventris (Horismenus), 293
glaucus (Heterophotinus), 131 gonagra (Eriphia), 44 GRACILLARIDAE, 320
glaucus (Modisimus), 58 gonagra (Leptoglossus), 253 gradata (Homoptera), 355
gleditsiae (Caryobruchus), 163 gonagrus (Melalgus), 134 gradatus† (Anisepyris), 309
Glenanthe, 232 Gonaives, 159 Grallipeza, 221
Glenognatha, 65 Gonatista, 190 Graminella, 266
Glipa, 142 Gonatopus, 311 gramineus (Agobardus), 74
globosum (Sphaerognathium), 146 Gondwanocrypticus, 146 graminis (Antonina), 279
globosus (Physocyclus), 58 GONEPLACIDAE, 41 graminum (Schizaphis), 274
globosus† (Neoxenus), 164 gongyla (Empoasca), 269 Grammonota, 64
globularia (Solenopsis), 304 Gonionota, 321 granarium (Trogoderma), 134
globularia borinquensis (Solenopsis), 304 Goniopsis, 41 granarius (Sitophilus), 177
globularia steinheili (Solenopsis), 304 GONODACTYLIDAE, 28 grandella (Hypsipyla), 329
globulus (Haplocyclodesmus), 91 Gonodonta, 352 grandicollis (Ips), 179
globulus† (Dipoenata), 61 Gonomyia, 192 grandiflavus (Apocephalus), 214
glochium (Hydrochasma), 231 Gonoporomiris, 244 grandis (Ablechrus), 136
GLOMERIDESMIDAE, 86 gonygaster (Theridula), 63 grandis (Aceratagallia), 263
Glomeridesmus, 86 gonzalezi (Calisto), 343 grandis (Aglyptinus), 110
gloriosa (Zicca), 254 gordialis (Asciodes), 324 grandis (Anthonomus), 169
GLOSSOSOMATIDAE, 388 gorgon (Chrysanthrax), 210 grandis (Chaetocalirrhoe), 220
gloveri (Tetranychus), 50 gossypii (Acalitus), 48 grandis (Licodia), 376
gloverii (Lepidosaphes), 278 gossypii (Aphis), 274 grandis (Pintaweckelia), 30
Glutophrissa, 339 gossypii (Corythucha), 242 grandis (Pseudacteon), 214
Glutoxys, 217 gossypii (Empoasca), 269 grandis† (Megalostrata), 72
Glympis, 352 gossypii (Phenacoccus), 280 grandjeani (Mochloribatula), 46
Glyphembia, 234 gowdeyi (Haplothrips), 385 granicollis (Lachnopus), 174
Glyphodes, 325 gowdeyi (Loxostethus), 146 grannus amazona (Calisto), 343
Glyphonyx, 129 grabhami (Anopheles), 200 grannus dilemma (Calisto), 343

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grannus dystacta (Calisto), 343 GRYLLIDAE, 369 guttiger (Galgupha), 255
grannus grannus (Calisto), 343 gryllipes (Ammotrechinus), 83 Guttipsilopa, 231
grannus micheneri (Calisto), 343 Gryllodes, 372 guttula (Catorhintha), 254
grannus micrommata (Calisto), 343 GRYLLOTALPIDAE, 376 guttularis (Erinnyis), 345
grannus phoinix (Calisto), 343 Gryllus, 372 gwenae (Ommatius), 210
grannus sommeri (Calisto), 343 Gryon, 284 Gymnandrosoma, 331
granosus (Lobodesmus), 90 guaba (Yabisi), 60 Gymnetis, 124
granulata (Endeitoma), 143 guadalupensis (Thoracophorus), 113 Gymnochthebius, 110
granulata (Mnemosyne), 259 guadeloupensis (Cnesinus), 177 Gymnoclasiopa, 231
granulata (Parthenope), 43 guajanae (Balclutha), 266 Gymnometopa, 199
granulofrons (Poratia), 90 guajey (Margarettia), 369 gymnopta (Hapithus), 371
granulosa (Jeekelia), 90 guanajayensis‡ (Messinella), 22 gymnorhynchus (Tabanus), 207
graphica (Hemilactica), 158 guanhumi (Cardisoma), 40 Gynacantha, 363
grapholithoides (Tripudia), 355 guanica (Phantyna), 69 Gynaikothrips, 385
Graphomya, 218 guanicae (Thymoites), 63 Gypona, 268
GRAPSIDAE, 41 guanicanus (Cremnops), 285 Gyranusoidea, 292
Grapsus, 41 guanita (Anthonomus), 169 Gyretes, 99
grapsus (Grapsus), 41 guanyangi (Sicoderus), 176 GYRINIDAE, 99
grassii (Dysmicoccus), 279 guarnacciana (Ceratina), 318 Gyrinus, 99
grata† (Elephantomyia), 193 guatemalensis (Enicospilus), 288 gyrommatus† (Brachycamacus), 175
gratalis (Petrophila), 323 guatemalensis (Nylanderia), 300 Gyrophaena, 111
gratiosus (Neopisinus), 62 guatemalensis (Paraputo), 280 Gyrostigma, 219
gratus (Pityophthorus), 181 guatemalensis itinerans (Nylanderia), habanensis (Yabisi), 60
gratus (Retiro), 70 300 habilitator (Culex), 200
gravidus (Haliplus), 99 gudmannella (Tildenia), 322 Habrobracon, 285
grayi (Paguristes), 39 guerini (Gerris), 239 Habronattus, 75
Grazia, 256 guerrero (Selenops), 73 habros (Hapithus), 370
greeni (Anagyrus), 292 guerreroi (Nesocyrtosoma), 147 Habroscelimorpha, 101
gregalis (Metazygia), 67 guerreronis (Aceria), 48 Habroxenus, 164
Greta, 341 guiana (Episimus), 331 hades† (Philonthus), 117
Grimalantha, 216 guianae (Neodryocoetes), 180 hadratum† (Nesocyrtosoma), 147
grimaldii (Aulacigaster), 225 guildingi (Cormocephalus), 84 Hadria, 265
grimaldii (Dagus), 233 guildingii (Mecistocephalus), 85 Hadropenaeus, 35
grimaldii† (Ambradolon), 211 guildingii (Sphendononema), 84 hadroprion (Apotomoderes), 174
grimaldii† (Clinidium), 108 guildinguii (Pselliodes), 84 Hadrosomus, 249
grimaldii† (Cubanoptila), 388 guildinii (Piezodorus), 257 HADZIIDAE, 29
grimaldii† (Dominicanopsis), 64 gulliver (Aeletes), 109 Haematobia, 218
grimaldii† (Dryinus), 311 gulosus (Creugas), 72 haematobiae (Spalangia), 296
grimaldii† (Forcipomyia), 197 Gummilumpus†, 297 Haematopinus, 379
grimaldii† (Glabellula), 211 gundlachi (Acmaeodera), 125 haematoxyli (Tanaostigmodes), 297
grimaldii† (Glyptotermes), 188 gundlachi (Erechthis), 368 haemorrhoidalis (Cryptocerus), 301
grimaldii† (Heterotrioza), 272 gundlachi (Strumigenys), 302 haemorrhoidalis (Heliothrips), 384
grimaldii† (Microvelia), 240 gundlachiana (Neolema), 161 haemorrhoidalis (Salbia), 327
grimaldii† (Oligochlora), 315 gundlachii (Lasioglossum), 315 haemorrhoidalis haemorrhoidalis
grimaldii† (Plectromerus), 150 gundlachii (Palinurellus), 38 (Myzine), 308
grimaldii† (Protortalotrypeta), 222 guraboense‡ (Cytheropteron), 22 haemuli (Anilocra), 31
grisea (Neohadria), 265 guraboensis‡ (Hepatus), 40 haetianum (Strongylium), 147
griseus (Docodesmus), 90 guttata (Aedmon), 157 haetianus (Tityus), 82
grisselli (Anisepyris), 309 guttata (Allototua), 63 hagedorni (Ambrosiodmus), 180
grollei† (Palaeopatasson), 295 guttata (Cassida), 160 hageni (Helicopsyche), 391
grossa (Steatoda), 62 guttata (Flatoides), 261 hagenii (Micrathyria), 364
grossacavis† (Baris), 169 guttata (Hispanognatha), 65 Hagenulites†, 235
Grossoxipha†, 373 guttatus (Heraeus), 251 Hagenulus, 235
Grotiusomyia, 294 guttatus (Marshallius), 176 HAHNIIDAE, 69
grouveillei (Xanthocomus), 138 guttatus (Panulirus), 38 haina (Ctenus), 69
GRYLLACRIDIDAE, 377 gutticollis (Rhantus), 100 haita (Clonistria), 377

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haitensis (Agoristenus), 77 haitianus (Mesethodesmus), 91 haitiensis (Nesomylacris), 186
haitensis (Ariamnes), 61 haitianus (Metaniphargus), 30 haitiensis (Occemyia), 221
haitensis (Blapstinus), 144 haitianus (Omalodes), 109 haitiensis (Ommatius), 210
haitensis (Chaerocampa), 346 haitianus (Paracarnus), 244 haitiensis (Orthoporus), 93
haitensis (Choranthus), 336 haitianus (Psephenops), 127 haitiensis (Parabezzia), 196
haitensis (Cyarda), 260 haitianus (Selenophorus), 103 haitiensis (Pellobunus), 78
haitensis (Efferia), 208 haitianus (Stenolemus), 246 haitiensis (Peripatus), 13
haitensis (Epilampra), 185 haitianus (Telephanus), 137 haitiensis (Phyllophaga), 122
haitensis (Epinannolene), 93 haitiaria (Sphacelodes), 334 haitiensis (Polycricus), 85
haitensis (Euscelus), 166 haitica (Strongylopsalis), 183 haitiensis (Pselliophora), 84
haitensis (Gryllacris), 377 haitica (Trogolaphysa), 96 haitiensis (Quadrastichus), 294
haitensis (Haitobolus), 92 haitiellus (Hymenorus), 145 haitiensis (Stenus), 118
haitensis (Hydraena), 110 haitiellus (Osoriellus), 113 haitiensis (Tabanus), 207
haitensis (Lobopoda), 145 haitiense (Clinidium), 108 haitiensis (Taphrocerus), 124
haitensis (Notaepytus), 139 haitiense (Diadegma), 377 haitiensis (Tethysbaena), 29
haitensis (Shannonomyia), 193 haitiense (Eiphosoma), 288 haitiensis (Tetragnatha), 65
haitensis (Statira), 143 haitiense (Rhopalosoma), 308 haitiensis (Tytthonyx), 132
haitensis (Trichopoda), 220 haitiense (Xiphocentron), 387 haitiensis (Urgleptes), 154
haitensis (Vestitecola), 77 haitiensis (Acaricoris), 248 haitiensis (Vella), 361
haitensis (Xeromelecta), 317 haitiensis (Acrocyum), 156 Haitimera, 77
haitensis haitensis (Sphingonotus), 367 haitiensis (Alathetus), 255 haitises (Antilloides), 57
haitensis‡ (Perissocytheridea?), 22 haitiensis (Amphiacusta), 374 haitises (Oecetis), 390
haitensis‡ (Portunus), 43 haitiensis (Aneurus), 247 haitius (Hymenorus), 145
haiti (Cyclosa), 66 haitiensis (Ataxia), 155 haitius (Xantholinus), 118
haiti (Tiphia), 308 haitiensis (Augochlora), 315 Haitobolus, 92
haitia (Biocrypta), 114 haitiensis (Catonia), 258 HALACARIDAE, 49
Haitiana, 258 haitiensis (Centruroides), 81 Haldorus, 266
haitiana (Amphiacusta), 374 haitiensis (Chlamisus), 161 halia cleobaea (Lycorea), 341
haitiana (Bothriocera), 258 haitiensis (Chrysobothris), 125 halia halia (Lycorea), 341
haitiana (Euhyloptera), 260 haitiensis (Chrysopa), 359 HALICTIDAE, 315
haitiana (Ghinallelia), 246 haitiensis (Coniopteryx), 360 HALICTOPHAGIDAE, 383
haitiana (Glyphembia), 234 haitiensis (Cryptocephalus), 161 Halictophagus, 383
haitiana (Heza), 246 haitiensis (Ctenus), 69 halictophagus (Nesosphecodes), 315
haitiana (Hispanisella), 115 haitiensis (Cubanella), 258 Halictus, 315
haitiana (Koenikea), 49 haitiensis (Culicoides), 196 Halicyclops, 16
haitiana (Lebia), 103 haitiensis (Cynortoides), 78 HALIPLIDAE, 99
haitiana (Nanolachesilla), 381 haitiensis (Dictyla), 242 Haliplus, 99
haitiana (Neomamersa), 49 haitiensis (Docodesmus), 90 halli (Planococcus), 280
haitiana (Oligota), 112 haitiensis (Ecpantheria), 347 Halocoryza, 106
haitiana (Pelitropis), 262 haitiensis (Empyreuma), 347 Halovelia, 240
haitiana (Schizopera), 18 haitiensis (Encarsia), 290 halticon (Cycloptilum), 376
haitiana (Silis), 132 haitiensis (Euodynerus), 308 Halticoptera, 296
haitianella (Amphiacusta), 374 haitiensis (Eupogonius), 154 Halysidota, 348
Haitianotettix, 365 haitiensis (Exophthalmus), 173 Hamadryas, 340
haitianum (Polynema), 295 haitiensis (Geaya), 78 hamata (Acacesia), 65
haitianum (Rhypasma), 143 haitiensis (Hadria), 265 hamata (Caribovia), 264
haitianum (Zuphium), 106 haitiensis (Helicopsyche), 391 Hamataliwa, 68
haitianus (Ablechroides), 136 haitiensis (Leucochrysa), 359 hamatula (Bogidiella), 29
haitianus (Arvelius), 256 haitiensis (Litoprosopus), 353 hamatus (Atropacarus), 47
haitianus (Ataenius), 120 haitiensis (Lophopterygella), 382 hamatus (Rasahus), 323
haitianus (Cyrtocapsus), 243 haitiensis (Lophoscutus), 247 hamatus (Reuteroscopus), 245
haitianus (Eucaerus), 103 haitiensis (Lumacra), 320 Hambletonia, 292
haitianus (Gonatopus), 311 haitiensis (Melissodes), 317 hamifera (Pardosa), 68
haitianus (Haplocyclodesmus), 91 haitiensis (Micromus), 360 Hamiltoniauletes, 166
haitianus (Hexacylloepus), 126 haitiensis (Mordella), 142 hampei (Hypothenemus), 178
haitianus (Hottettix), 365 haitiensis (Nelasa), 141 Hampsonodes, 350

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hamulus (Cephalotes), 301 hecalialis (Phaedropsis), 326 hemipterus (Carpophilus), 137
hamus† (Piesocorynus), 164 Hecamede, 232 hemipterus (Cimex), 245
hancocki (Calyptolana), 31 hecate (Ircila), 331 hemipterus sericeus (Metamasius), 177
hanno ceraunus (Hemiargus), 339 hedonialis (Conchylodes), 324 Hemiptiloceroides, 329
hanno watsoni (Hemiargus), 339 Hedriodiscus, 206 HEMISCORPIIDAE, 83
hansoni (Eprhopalotus), 293 heeri (Brachymyrmex), 300 hemisphaerica (Saissetia), 276
Hapithus, 369 hegesia hegesia (Euptoieta), 341 Hemisphaerota, 160
Haplaxius, 259 heidemanni (Brepholoxa), 257 hemisurus (Aedes), 199
Haplocyclodesmus, 91 heidti† (Strotarchus), 70 Hemitrinemura, 97
Haplocytheridea, 22 heiei† (Quisqueyaaphis), 273 hemphilli (Synalpheus), 36
HAPLODESMIDAE, 89 Heikertingerella, 157 hendersoni (Calisto), 343
Haplopus, 377 heinekeni (Ataenius), 120 hendersoni (Venezillo), 33
Haplothrips, 385 Heinia, 25 Henicomus, 90
Happlodontus†, 83 helbi (Solenoptera), 156 hennigi† (Milichiella), 227
harlequina (Pheidole), 303 helcitalis (Botys), 326 hennigi† (Proceroplatus), 202
Harmostes, 254 helenae (Ozophora), 252 hennigianus† (Boreofairchildia), 194
harmsworthi coronatus (Macrobiotus), 13 helibionta† (Electrosminthuridia), 96 Henryhowella, 27
Haroldiellus, 119 Heliconius, 341 Hentzia, 75
Harpactosphecion, 311 Helicopsyche, 390 hepaticus (Philonthus), 117
harpyia (Ligyra), 210 HELICOPSYCHIDAE, 390 Hepatus, 40
harrisii (Chrysoperla), 359 Helicoverpa, 356 Hephebocerus, 168
hartii (Aspidiella), 277 HELIODINIDAE, 322 Hepomadus, 35
hartkorni† (Tityus), 82 heliothidata (Digrammia), 333 heptacantha (Fennerosquilla), 28
Hasarius, 75 Heliothis, 356 heptagon (Gea), 66
hasei (Carios), 51 Heliothrips, 384 Heraclides, 338
hassleri (Cynorta), 78 Heliozela, 319 Heraeus, 251
hassleri (Theridion), 63 HELIOZELIDAE, 319 herbacea (Banasa), 256
hastata (Ischnura), 362 Helius, 194 herbaria herbaria (Synchlora), 334
hastosa (Empoasca), 269 Hellula, 324 herbicolus (Amblyseius), 53
hatomayor (Zeadolopus), 110 helmoides (Phanocerus), 126 herbida (Brachymesia), 363
hatuey (Hypolestes), 362 Helodytes, 175 herbsti (Panopeus), 44
hatuey (Litoprosopus), 353 Helops, 144 herculeana (Clavaspis), 277
hatueyi (Unicorniella), 368 helops (Ammalo), 348 Heriades, 316
havanense (Lasioglossum), 315 Helvibotys, 325 hermani (Dagus), 233
havaniensis (Haliplus), 99 HEMEROBIIDAE, 360 Hermanites, 24
havaniensis (Ignelater), 128 Hemerobius, 360 HERMANNIELLIDAE, 46
havaniensis (Philonthus), 117 Hemeroblemma, 353 Hermetia, 206
hawaiiensis (Parhyale), 30 Hemerophila, 322 heros (Corythalia), 75
haxairei (Westindia), 349 Hemeroplanis, 353 Herpetocypris, 20
haytensis (Epilobocera), 44 Hemiargus, 339 Herpetogramma, 325
haytiana (Fustiger), 115 Hemiarthrus, 32 herrlingi† (Araneometa), 66
haytiana (Gnamptogenys), 299 Hemiberlesia, 277 hersilialis (Carcha), 327
haytiana (Hamataliwa), 68 Hemiblabera, 185 HERSILIIDAE, 60
haytianus (Anochetus), 304 Hemiceras, 346 hespera (Amphiacusta), 374
haytianus (Mycetomoellerius), 301 Hemicyprideis, 22 Hesperia, 336
haytianus (Pseudomyrmex), 306 HEMICYTHERIDAE, 23 hesperidum (Coccus), 275
haytianus (Trachymyrmex), 394 Hemicytherura, 23 HESPERIIDAE, 336
haytiellus (Fissicrambus), 323 Hemigymnaspis, 277 Hesperochernes, 80
haytiensis (Compsodecta), 75 Hemilactica, 157 Hesperophasma, 377
hazelae‡ (Agrenocythere), 27 Hemipenthes, 210 Heterachthes, 152
hazelae‡ (Bradleya), 26 Hemipeplus, 148 Heterobelba, 47
hebardi (Eurycotis), 184 Hemipepsis, 307 HETEROCERIDAE, 127
hebe (Balclutha), 266 Hemiphileurus, 123 Heterocerus, 127
hebetata (Anasaitis), 74 HEMIPSOCIDAE, 381 heterochirus (Macrobrachium), 37
hebraealis (Conchylodes), 324 Hemipsocus, 381 Heteroctenus, 82
HEBRIDAE, 239 Hemipteroseius, 52 Heterocypris, 20

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Heterogramma, 356 hirsuta (Hirtasphaera), 158 hispaniolae (Calocosmus), 154
Heteromeringia, 225 hirsuta (Hispanisella), 115 hispaniolae (Chrysobothris), 125
heteromus (Achromoporus), 87 hirsuta (Lemelba), 115 hispaniolae (Coccophagus), 290
Heteromyia, 197 hirsuta† (Hispaniolyna), 69 hispaniolae (Copestylum), 215
Heteromysis, 29 hirsutus (Cis), 142 hispaniolae (Cyclopoliarus), 259
Heteronebo, 83 hirsutus (Hypothenemus), 178 hispaniolae (Cyrsylus), 157
HETERONEMIIDAE, 377 hirsutus (Maconellicoccus), 279 hispaniolae (Epyris), 310
Heteroonops, 59 hirsutus (Stratus), 116 hispaniolae (Gonoporomis), 244
Heterophotinus, 131 hirsutus (Teminius), 70 hispaniolae (Heterops), 152
Heteropoda, 73 hirsutus† (Coptotermes), 188 hispaniolae (Holepyris), 310
Heterops, 152 Hirtasphaera, 158 hispaniolae (Hyporhagus), 143
heterops (Prostemmiulus), 94 hirtellus (Lasiochilus), 245 hispaniolae (Irenangelus), 306
Heteropsylla, 271 hirtifemora (Conura), 291 hispaniolae (Lelegeis), 146
Heterosarus, 314 hirtilabris (Antillosorius), 112 hispaniolae (Leptophysa), 158
Heterostylum, 211 hirtipes (Ecyrus), 155 hispaniolae (Leptostylopsis), 153
Heterotermes, 188 hirtus (Lachnopus), 174 hispaniolae (Methia), 152
Heterothops, 117 hirtus (Opius), 286 hispaniolae (Micrasta), 126
Heterotmarus, 73 hirudopsis aitija (Alisotrichia), 388 hispaniolae (Mimetus), 60
heterotricha (Zygoribatula), 47 Hispanacris, 367 hispaniolae (Myochrous), 163
Heterotrioza, 272 hispanicus (Aphytis), 289 hispaniolae (Nesocyrtosoma), 147
Heuretes, 330 hispanicus (Gnatholeptus), 180 hispaniolae (Nyctiplantus), 159
heveae (Cryptocarenus), 178 Hispaniodirphia, 344 hispaniolae (Ommatius), 210
Hexabdella, 48 hispaniola (Araneus), 66 hispaniolae (Paranapiacaba), 159
Hexacylloepus, 126 hispaniola (Bathydema), 250 hispaniolae (Paucivena), 320
Hexadesmus, 89 hispaniola (Biocrypta), 114 hispaniolae (Paululusus), 175
Hexatoma, 193 hispaniola (Hadria), 265 hispaniolae (Pheidole), 303
hexosetosus (Campylothorax), 95 hispaniola (Hoplocheiloma), 221 hispaniolae (Phrynus), 54
heydoni (Ozophora), 252 hispaniola (Kleidocerys), 249 hispaniolae (Polemius), 132
Heza, 246 hispaniola (Lophocampa), 348 hispaniolae (Pseudodisonycha), 158
Hibana, 71 hispaniola (Malacomorpha), 378 hispaniolae (Psorthaspis), 308
hibernalis (Kukulcania), 57 hispaniola (Micropholcus), 57 hispaniolae (Sibinia), 172
hibisci (Aceria), 48 hispaniola (Nicrophorus), 111 hispaniolae (Squamanotoxus), 149
hiemalis (Ctenus), 69 hispaniola (Nops), 59 hispaniolae (Stenotabanus), 207
hieroglyphica (Diphthera), 352 hispaniola (Ozophora), 252 hispaniolae (Tachyta), 107
hieroglyphicus (Exophthalmus), 173 hispaniola (Petrobia), 50 hispaniolae (Temnothorax), 302
hierroi (Anurogryllus), 372 hispaniola (Sirex), 282 hispaniolae (Termes), 190
hierroi (Eurycotis), 184 hispaniola (Spurostigma), 380 hispaniolae (Tiphia), 308
hierroi (Lychnacris), 130 hispaniola (Yoshimotoana), 296 hispaniolae (Uhleroides), 263
hierroi (Ommatius), 210 hispaniola† (Caribitermes), 189 hispaniolae (Voldroguella), 49
hilaris (Anthophora), 316 hispaniola† (Macronema), 387 hispaniolae† (Dasyhelea), 197
hilaris (Cryptocephalus), 161 hispaniola† (Rhopalosoma), 308 hispaniolae† (Linepithema), 298
hilaris (Glipa), 142 hispaniola† (Subulitermes), 189 hispaniolae† (Panstrongylus), 247
hilaris (Proarna), 263 hispaniolae (Acladocera), 129 hispaniolae† (Schizoptera), 239
Hilarotes, 125 hispaniolae (Agrilus), 124 hispaniolae† (Technomyrmex), 299
Hileithia, 325 hispaniolae (Alethaxius), 162 hispaniolaensis (Leptomantispa), 361
Himatium, 170 hispaniolae (Amphiacusta), 374 hispaniolaensis (Ptinus), 135
hinei (Microvelia), 240 hispaniolae (Anelaphus), 151 hispaniolana (Amblytropidia), 367
hinnulus (Atopsyche), 390 hispaniolae (Anoplius), 307 hispaniolana (Ammotrechella), 83
hinojosai† (Cossonus), 170 hispaniolae (Anthicus), 149 Hispaniolara, 126
Hippelates, 227 hispaniolae (Anthophora), 316 hispaniolensis (Aglyptinus), 110
Hippia, 346 hispaniolae (Arhopalus), 149 hispaniolensis (Aneurus), 247
HIPPIDAE, 39 hispaniolae (Ataenius), 120 hispaniolensis (Ardistomis), 106
HIPPOBOSCIDAE, 217 hispaniolae (Bellitudo), 272 hispaniolensis (Blapstinus), 144
Hippodamia, 140 hispaniolae (Bothriocera), 258 hispaniolensis (Corethrella), 199
HIPPOLYTIDAE, 36 hispaniolae (Buprestis), 125 hispaniolensis (Creagrophorus), 111
hirsuta (Frevillea), 41 hispaniolae (Burca), 337 hispaniolensis (Hylaeus), 314

613 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

hispaniolensis (Hymenorus), 145 Hister, 109 hookeri (Dinurothrips), 384
hispaniolensis (Isoplastus), 110 HISTERIDAE, 109 Hoplandria, 111
hispaniolensis (Lobopoda), 145 Historis, 342 Hoplicnema, 141
hispaniolensis (Lophoscutus), 247 histrio (Apogonalia), 264 hoplicnema (Sepsis), 224
hispaniolensis (Parvocyrtusa), 110 histrio (Eurycotis), 184 Hoplisoides, 312
hispaniolensis (Petrolisthes), 40 histrio (Lachnopus), 174 Hoplistocelis, 245
hispaniolensis (Proptomaphaginus), 111 hitchingsi† (Hagenulites), 235 Hoplocheiloma, 221
hispaniolensis (Salmoneus), 36 hiwanneensis‡ (Argilloecia), 26 Hoplophthiracarus, 47
hispaniolensis (Tytthonyx), 132 hiwanneensis‡ (Krithe), 24 Horama, 347
hispaniolensis (Zeadolopus), 110 hoffmani (Aedmon), 157 horatis† (Polyvena), 331
hispaniolensis† (Azilia), 64 hoffmani (Homoschema), 158 Horismenus, 293
hispaniolensis† (Homopterus), 106 hoffmani (Leptopharsa), 242 Horistonotus, 128
hispaniolensis† (Jassosqualus), 267 hoffmani (Leucocera), 161 HORMAPHIDIDAE, 275
hispaniolensis† (Oviedinus), 135 hoffmani (Limatus), 201 Hormathus, 152
hispaniolensis† (Philacarus), 78 hoffmani (Longitarsus), 158 Hormotrophus, 173
hispaniolensis† (Protosinus), 182 hoffmani (Lupraea), 158 hornibrooki‡ (Hermanites), 24
hispaniolensis† (Psalistops), 56 hoffmani† (Siphonophora), 91 hortaria (Epimecis), 333
hispaniolica (Ahotta), 77 hoffmanni (Ecyrus), 155 Hortensia, 265
hispaniolica (Cariboptila), 388 hogardi (Phyllophaga), 122 Hortoonops, 59
hispaniolicus (Agapostemon), 315 hogardii (Sphecius), 312 hortorum (Palpada), 215
hispaniolicus (Arucillus), 77 Hogna, 68 hosios (Cyrtoxipha), 375
hispaniolicus (Chernes), 80 holacanthi (Anilocra), 31 hottae (Pycnomerus), 143
hispaniolicus† (Acanthostichus), 298 Holcobracon, 285 hotteiensis (Araneus), 66
hispaniolicus† (Cephalotes), 301 Holcocephala, 210 hottense (Metachroma), 162
hispaniolicus† (Culicoides), 196 Holepyris, 310 hottensis (Altisessor), 139
hispaniolicus† (Toxorhynchus), 167 Holisus, 117 hottensis (Leptophysa), 158
hispaniolina (Alisotrichia), 388 hollyae (Heraeus), 251 Hottettix, 365
hispaniolum (Cyclopterum), 260 Holobomolochus, 18 hovorei (Ataxia), 155
hispaniolum (Dominopteron), 129 Holocompsa, 187 hovorei (Derancistrus), 156
hispaniolum (Liparthrum), 178 Hologaeus, 104 Hovorodon, 155
hispaniolus (Amphicranus), 181 Hololepta, 109 howardevansi (Cerceris), 313
hispaniolus (Belonuchus), 117 Holoparamecus, 142 howardi (Pteroptrix), 290
hispaniolus (Dasyodontus), 89 Holopothrips, 385 Howardia, 278
hispaniolus (Dawnarioides), 260 Holopyga, 310 Howardina, 199
hispaniolus (Dendrocranulus), 179 holosericeus (Calocosmus), 154 howdeni (Canthonella), 121
hispaniolus (Gnathotrichus), 181 Holostilpna, 164 howdeni† (Anaides), 119
hispaniolus (Neopamphantus), 249 Holoteleia, 284 howdenorum (Enicospilus), 288
hispaniolus (Pinophilus), 114 holoxanthus (Aphytis), 289 howei‡ (Cushmanidea), 21
hispaniolus (Pityophthorus), 181 Homaledra, 322 howei‡ (Microcythere), 25
hispaniolus (Quisquellacricus), 92 Homalometa, 65 howei‡ (Pellucistoma?), 22
hispaniolus (Stenus), 118 Homalopoda, 291 howei‡ (Procythereis?), 27
hispaniolus (Thinodromus), 113 Homalotylus, 292 howei‡ (Quadracythere), 26
hispaniolus† (Trypanaeus), 110 Homeosoma, 329 Huaca, 169
Hispaniolyna†, 69 hominivorax (Cochliomyia), 217 huachuca (Limnellia), 233
Hispanioryctes, 123 homochronon† (Apionion), 167 huasache (Heteropsylla), 271
Hispanisella, 115 Homocloeus, 165 hubbardi (Nasutitermes), 189
Hispanognatha, 65 Homoconnus, 116 hubbardi (Phanocerus), 126
Hispanogryllus, 372 Homoeotarsus, 114 HUBBARDIIDAE, 81
Hispanonesticus†, 61 Homola, 41 hufelandi group (Macrobiotus), 13
Hispanotettix, 367 HOMOLIDAE, 41 huggerti (Omphale), 293
hispanus (Osbornellus), 267 Homophileurus, 124 hugonis (Aedmon), 157
hispida (Encarsia), 290 Homoptera, 358 huidobrensis (Liriomyza), 225
hispidulus† (Toxorhynchus), 167 Homopterus, 106 humanus (Pediculus), 379
hispidum (Pycnarthrum), 179 Homoschema, 158 humanus capitis (Pediculus), 379
hispidus (Iomoides), 90 hondovallensis (Robopus), 131 humanus corporis (Pediculus), 379
hispidus (Stenopus), 35 honesta (Oraesia), 354 humeralis (Chalcosicya), 162

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 614
humeralis (Neohypnus), 117 HYDROPSYCHIDAE, 387 icarus (Mesepipsocus), 380
humeralis (Omiodes), 326 Hydroptila, 388 Icerya, 279
humeralis (Omoteina), 160 HYDROPTILIDAE, 388 ichneumoneus (Sphex), 314
humeralis (Selenisa), 358 Hydrovatus, 100 ICHNEUMONIDAE, 287
humeralis (Sicoderus), 176 Hygronemobius, 373 Icius, 76
humerofulvus (Leptostylopsis), 153 Hylaeus, 314 Idarnes, 289
humidicola (Leucania), 355 Hylastes, 177 Ideobisium, 79
humilis (Delphacodes), 259 Hylephila, 336 Idiataphe, 363
humilis (Neobisnius), 117 Hyles, 346 Idiocerus, 268
humilis (Parydra), 233 hylobates (Yoyuteris), 374 Idioderma, 271
humilis (Plusia), 357 Hylobius, 176 Idiogaryops, 79
humiloides† (Gracilidris), 298 Hylocurus, 178 idioidea (Chloroprocta), 217
hummelincki (Antillolpium), 79 hylonomos (Yoyuteris), 374 Idolia, 110
hummelincki (Candonopsis), 20 hymenaea (Pantala), 364 Idona, 269
hummelincki (Parabetaeus), 36 hymenaea† (Pachygaster), 206 idonearia idonearia (Iridopsis), 333
Hummelinckiolus, 77 hymenaeae (Araptus), 180 Idris, 284
hursti (Trujillina), 69 hymenaeaphilus† (Dryinus), 311 idyja idyja (Asterocampa), 340
huyenae (Melathra), 174 Hymenia, 325 ieva† (Gnophomyia), 192
hyacinthina (Leptogaster), 209 Hymenorus, 145 Ignelater, 128
Hyalella, 30 Hypanartia, 342 ignipennis (Larrada), 312
HYALELLIDAE, 30 Hypargyria, 329 ignitum (Aspisoma), 129
hyalina (Alima), 28 Hypasclera, 148 ignobilis (Fundella), 329
hyalina (Bandara), 265 Hypena, 357 ignotensis (Notaepytus), 139
hyalina (Soberocephala), 226 Hypenula, 356 Ilemomyrmex†, 304
hyalina acuta (Parahydriena), 262 Hyperaspis, 140 Ileopeltus, 266
hyalinata (Diaphania), 325 Hypercompe, 347 ileota (Gypona), 268
hyalinipennis (Oxycarenus), 250 hyperia (Biblis), 340 Iliacantha, 41
hyalinus (Corizus), 254 Hyperlasion, 204 illectans (Hapithus), 370
hyalinus† (Paralienates), 238 Hypocala, 353 illegibilis† (Cyrtoxipha), 375
Hyalistata, 231 Hypochalcia, 329 illex (Hapithus), 370
Hyalochloria, 244 hypoclopon (Cycloptilum), 376 illigeri (Hybosorus), 119
Hyalopsallus, 244 Hypoconcha, 40 illinoisensis (Aphis), 274
Hyalurga, 348 Hypocryphalus, 178 illinoisensis (Hoplophthiracarus), 47
Hybla, 269 Hypoderma, 219 illucens (Hermetia), 206
Hyblaea, 323 HYPOGASTRURIDAE, 94 illudens (Vatiga), 243
HYBLAEIDAE, 323 Hypogena, 145 illuminus (Pityophthorus), 181
HYBOSORIDAE, 119 Hypogeococcus, 279 illustrata (Notioplusia), 358
Hybosorus, 119 Hypogrammodes, 353 illustris (Augocoris), 258
HYBOTIDAE, 213 Hypolestes, 362 illustris (Neoninus), 249
Hydaticus, 100 Hypolimnas, 341 ilosvayi‡ (Cytheridella), 24
Hydraena, 110 Hypomicrogaster, 286 Ilythea, 234
HYDRAENIDAE, 110 Hypoponera, 304 imbella (Perigea), 350
Hydrellia, 234 Hyporhagus, 143 imbellis (Purenleon), 361
hydrillalis (Bleptina), 356 Hypotelus, 115 imbricatus (Ataenius), 120
Hydrobiomorpha, 108 Hypothenemus, 178 imita (Gypona), 268
HYDROBIOSIDAE, 390 Hypselodesmus, 88 imitalis (Margaronia), 325
Hydrocanthus, 99 Hypselosoma, 238 imitata (Crocidomera), 328
Hydrochasma, 231 Hypselosomops, 239 imitator (Rheumatobates), 239
HYDROCHIDAE, 108 HYPSIBIIDAE, 13 imitator (Spalangia), 296
Hydrochus, 108 hypsipetes (Anaxipha), 375 imitatrix (Pteroptrix), 290
Hydroma, 49 Hypsipyla, 329 imitatus (Tyrannochthonius), 79
Hydrometra, 239 Hyptia, 288 immaculata (Conura), 291
HYDROMETRIDAE, 239 hysius (Calisto), 343 immaculata (Microsema), 333
HYDROMIDAE, 49 hysius aleucosticha (Calisto), 343 immaculata immaculata (Erioptera), 192
HYDROPHILIDAE, 108 Hysteroneura, 274 immaculatus (Alpheus), 36
Hydrophilus, 108 iarchasalis (Bicilia), 324 immanis (Pararcte), 354

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immunis (Ptichodis), 354 incongrua† (Abanaxipha), 375 inflatum (Plesiomma), 210
impar (Bendis), 353 inconspicua (Arctosa), 68 inflatus (Melanopleurus), 249
impar (Nacaeus), 113 inconspicua (Leucania), 355 inflatus (Ommatius), 210
imparalis (Lygropia), 326 inconspicua (Oecetis), 390 inflatus (Suphisus), 100
impasta (Anomis), 351 inconstans (Panula), 354 infula (Madoniella), 136
impensum† (Nesocyrtosoma), 147 increta (Semiothisa), 334 infumatus (Alauximus), 70
imperialis (Neoleucinodes), 326 incrustata† (Brevisensoria), 248 infuscata domingensis (Melormenis), 261
imperialis (Peckia), 219 incurva (Gonodonta), 352 infuscatus (Styloleptus), 153
implens (Ephyrodes), 352 incurvus (Pycnoderes), 243 infusella (Salebria), 330
implicata (Elaver), 71 indeclararia (Iodis), 334 ingens (Cydia), 331
impressa (Chrysobothris), 125 indecora (Acrolophus), 319 ingloria (Ochrotrichia), 389
impressicolle (Metaphrenon), 152 indefensa (Dicranomyia), 193 inhibitator (Culex), 200
impressicollis (Nacaeus), 112 indentatus (Rhytidoporus), 254 innocuous (Atya), 36
impressus (Cormocephalus), 84 independencia (Dasypyga), 328 innoxius (Dendrothripoides), 384
imprima (Phyllophaga), 122 independencia (Euchroa), 106 innupta† (Janzenella), 285
imprudens† (Dysimia), 260 independentia (Orthocomotis), 332 Inodesmus, 90
impuralis (Proleucinodes), 326 Indepsylla†, 272 Inopeplus, 149
inaccessibile (Mesoplocia), 235 indiana (Nystalea), 347 inopinatus† (Cylindromyrmex), 298
inaequalis (Coelophora), 140 indica (Diaphania), 325 inopinatus† (Pristapenesia), 311
inaequalis (Exophthalmus), 173 indica (Gyranusoidea), 292 inops (Cycloptilum), 376
inaequalis‡ (Petrochirus), 39 indica (Notonecta), 241 inoptata (Macaria), 333
inagua (Pelonium), 136 indica (Pheidole), 303 inornata (Aonidiella), 276
inanis (Chlorotabanus), 207 indica (Radionaspis), 278 inornatalis (Phlyctaenodes), 324
inarmatus (Microcyclops), 17 indica (Raoiella), 50 Inostemma, 283
incanus (Anthonomus), 169 indicata (Omiodes), 326 inquinatus (Leioptilus), 330
incerta (Brachymeria), 291 indigena (Holcocephala), 210 insecta (Xanthaciura), 223
incertissimus (Pseudothysanoes), 178 indiges (Peruveleon), 361 insertus (Myrmeleon), 361
incertus (Amithao), 124 indigna (Cropia), 350 insidiosus (Orius), 245
incertus (Polistes), 309 indignaria indignaria (Semaeopus), 335 insignata (Lema), 161
incertus† (Fossilaraneus), 66 indigoferestes (Acanthoscelides), 163 insignis (Beatadesmus), 88
incertus† (Stemmops), 62 Indiopsocus, 382 insignis (Culicoides), 196
incertus† (Stenoonops), 60 indistincta (Drycothaea), 154 insignis (Insignorthezia), 280
incipiens (Letis), 355 indivisa (Bidoma), 77 insignitus (Anotylus), 113
incisa (Balclutha), 266 indocera (Pseudolycoriella), 205 Insignorthezia, 280
Incisitermes, 188 indocilia† (Limbopsylla), 272 insolita (Cautethia), 346
incisithorax (Megacerus), 163 indocilis (Nemobiopsis), 373 insolita (Mecampsis), 382
incisum (Hydrochasma), 231 indomita (Melipotis), 353 insolitus (Copelatus), 100
incisura (Coccoelasma), 89 Indotritia, 47 instabilis (Sennius), 164
incisura (Efferia), 208 ineditus (Araptus), 180 instratus (Spirobolellus), 92
incisuralis (Beameromyia), 209 inerma (Metepeira), 67 instructalis (Vurna), 327
incisus (Haplocyclodesmus), 91 inermaria (Lobocleta), 335 insulae (Anastrepha), 223
incisus (Oxytelus), 113 inermis (Strategus), 123 insulae (Pseudochiridium), 80
incisus† (Nasutitermes), 189 inermis† (Empiploiariola), 246 insulalis (Petrophila), 323
incivis (Anicla), 357 infausta (Kakopoda), 353 insulana (Gerstaeckeria), 171
inclinata (Celaena), 350 infausta (Neoblattella), 187 insulana (Leucocera), 161
includens (Pseudoplusia), 357 infecta (Anicla), 357 insulana (Paratheuma), 69
inclusum (Cheiracanthium), 70 infecta (Ptichodis), 354 insulani (Chinavia), 256
inclusum† (Elaphidion), 151 inferior (Cloeodes), 235 insulanus (Anthrax), 210
inclusus† (Electromyrmococcus), 279 infernalis† (Heterothops), 117 insulanus (Ctenus), 69
incognita (Leucania), 355 inficita (Perasia), 354 insulanus (Haplocyclodesmus), 91
incognitus† (Dryomites), 182 infidelis (Ambrosiodmus), 180 insulanus (Hypotelus), 115
incompleta (Tomoplagia), 222 infimus (Loxandrus), 106 insulanus (Mirda), 78
incompletum† (Theridiosoma), 63 infinis (Psorophora), 201 insulanus (Ozophora), 252
incompletus† (Veterator), 72 inflata (Cedusa), 260 insulanus† (Microcerotermes), 190
incomptus (Tricolus), 182 inflata (Pseudomopsis), 172 insulare (Diadegma), 288
inconditus (Lachnopus), 174 inflaticercus (Antillacris), 366 insularia (Psorophora), 201

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 616
insularis (Aeroppia), 47 integer (Progomphus), 363 Ircila, 331
insularis (Agapostemon), 315 integer (Selenophorus), 103 Irenangelus, 306
insularis (Caliotrhrips), 384 integerrimus† (Cephalotes), 301 iridescens (Neotrichia), 389
insularis (Cannaphila), 363 intera (Agalliopsis), 264 iridescens (Phylloicus), 390
insularis (Centris), 317 interdicta (Quatiella), 195 Iridopagurus, 39
insularis (Cephalosphaera), 214 interfracta (Cosmopterix), 322 Iridopsis, 333
insularis (Cercyon), 109 interjecta (Davara), 328 irregularis (Nacaeus), 112
insularis (Chelonus), 285 interjecta (Phymata), 247 irresoluta (Metria), 354
insularis (Chlamisus), 161 intermedia (Neopamera), 251 irritans (Haematobia), 218
insularis (Chrysis), 310 intermedia (Squilla), 28 irritans (Pulex), 382
insularis (Cillaeus), 137 intermedia† (Palaeodyctina), 69 irroratum irroratum (Elaphidion), 151
insularis (Colpoptera), 261 intermedialis (Pseudasopia), 330 irroratus (Cancer), 40
insularis (Conocephalus), 368 intermedius (Brachycentrotus), 270 irroratus (Diolcus), 258
insularis (Eurytoma), 295 intermedius (Haplopus), 377 irroratus (Trichrous), 152
insularis (Feltiella), 202 intermedius (Minibiotus), 13 irroratus irroratus (Copicerus), 259
insularis (Frankliniella), 384 intermedius (Nausibius), 138 isabel (Rowlandius), 81
insularis (Gnamptogenys), 299 intermedius (Trichobius), 217 isabelensis (Achromoporus), 87
insularis (Hemipeplus), 148 intermedius† (Anochetus), 304 isabelicae (Paridris), 284
insularis (Hydrophilus), 108 intermedius† (Dolichoderus), 298 isabella melphis (Eueides), 341
insularis (Leucochrysa), 359 intermedius† (Electrognostus), 134 isabellae (Canthonella), 121
insularis (Limnichoderus), 126 intermittens (Apamea), 350 isabeltorres (Eurycotis), 184
insularis (Lispinus), 112 internitalis (Sathria), 327 isaea (Diglyphus), 293
insularis (Macroperipatus), 13 interposita† (Cratyna), 204 ISAEIDAE, 30
insularis insularis (Macroperipatus), 13 interpunctella (Plodia), 330 Isaloides, 73
insularis (Mimosestes), 163 interrupta (Apogonalia), 264 Iscadia, 358
insularis (Mioptachys), 107 interrupta (Callopisma), 131 Ischionoplus, 174
insularis (Odontomachus), 305 interruptus (Lachnopus), 174 Ischnaspis, 278
insularis (Oebalus), 257 interruptus (Prostemmiulus), 93 Ischnocolinopsis, 56
insularis (Pelonomus), 126 interruptus (Sepedophilus), 118 Ischnodemus, 248
insularis (Planinasus), 226 interruptus (Truncotettix), 365 Ischnoptera, 186
insularis (Prosierola), 309 intersetosus (Xyleborinus), 180 Ischnothele, 56
insularis (Pseudolionothus), 110 interstitialis (Acanthia), 238 Ischnothyreus, 59
insularis (Ptecticus), 206 interstitialis (Berosus), 108 Ischnura, 362
insularis (Pterogyne), 320 interstitialis (Hypothenemus), 178 islacabritosensis (Perdita), 314
insularis (Sarcophaga), 219 interstitialis (Ips), 179 islacolonis (Cariblatta), 186
insularis (Selenops), 73 intestinalis (Gasterophilus), 219 Isodontia, 314
insularis (Sepsis), 224 Inti, 293 Isognathus, 345
insularis (Signiphora), 296 intracta (Letis), 353 Isogona, 353
insularis (Suphisellus), 100 intricata (Catonia), 258 isolata (Alujamyia), 223
insularis (Teminius), 70 intrusus (Xyleborus), 180 isolepus† (Episcirrus), 171
insularis (Tramea), 364 invasus (Apachekolos), 209 Isometrus, 82
insularis (Trombicula), 51 inversum† (Theridion), 63 Isoplastus, 110
insularis (Trypanidius), 154 inversus (Notaepytus), 139 Isotes, 159
insularis (Virbia), 347 inversus (Syneches), 213 Isotoma, 95
insularis insularis (Macroperipatus), 13 io† (Miomyia), 230 ISOTOMIDAE, 95
insularis† (Subulitermes), 189 iocosa† (Corynoptera), 204 Isotomiella, 95
insularum (Leptocera), 228 iodaria (Cyclomia), 333 Isotomurus, 95
insularum (Stenocaecilius), 381 iole (Nathalis), 338 Isotomus, 150
insularum (Stilomedon), 115 Iomoides, 90 israeli (Notiobiella), 360
insulata (Pareuchaetes), 349 Iomus, 90 ISSIDAE, 261
Insulatitan, 212 Iphiseiodes, 53 isthmicus (Lissorhoptrus), 175
insulatus (Conophthorus), 180 ipomaea (Calycomyza), 225 Isthmopsocus, 380
insulicola (Allotingis), 242 ipomoeae (Adaina), 330 itame (Cyrtoxipha), 375
insulicola (Peronaemis), 125 ipomoeae (Frankliniella), 384 Itaquascon, 13
integer (Alethaxius), 162 Ips, 179 Ittys, 297
integer (Cyclograpsus), 41 ipsilon (Agrotis), 357 iulia fucatus (Dryas), 341

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iuloides (Prostemmiulus), 93 jamaicensis jindrai (Dysdercus), 252 Joruma, 269
iuvenis† (Palaeoepeirotypus), 63 jamey (Quisquellacricus), 92 josephi (Leptoteleia), 284
iuvenoides† (Palaeoepeirotypus), 63 janella (Oxyethira), 389 josephi (Plectromerus), 150
ivei (Microtityus), 82 janestanleyae (Brachyodina), 206 josephina josephina (Ganyra), 339
ivei (Zeadolopus), 110 JANIRIDAE, 32 Jubus, 115
iviei (Brasiella), 101 janitor (Culex), 200 jucunda (Melipotis), 353
iviei (Decorosa), 135 jansei† (Cephalotes), 301 judicatalis (Megatomis), 357
iviei (Elaphidion), 151 jantaricum† (Cryptorhopalum), 133 juglandis (Tethysbaena), 29
iviei (Fallia), 141 januaris (Melipotis), 353 Jugosocythereis, 24
iviei (Heteroonops), 59 janus (Calocosmus), 154 juliani (Cryptotermes), 188
iviei (Kormilevida), 247 Janzenella, 285 jungi‡ (Munseyella), 26
iviei (Leiopleura), 124 Japanagromyza, 224 junius (Anax), 363
iviei (Normaltica), 158 jara (Stenoonops), 60 Junonia, 342
iviei (Peplomicrus), 113 jarabacoa (Zeadolopus), 110 juriaani (Tethysbaena), 29
iviei (Stylogaster), 221 Jaragua, 366 jurinei (Zethus), 309
iviei (Tytthonyx), 132 jaragua (Achromoporus), 87 Jurinella, 220
iviei† (Antillipeltis), 135 jaragua (Centruroides), 81 justiniana (Erythrodiplax), 363
ivonneae (Beatadesmus), 88 jaragua (Ctenus), 69 juvenis (Nesidiphilus), 85
ixion (Chioides), 337 jaragua (Eurycotis), 184 kahawaluokalani (Tinocallis), 275
IXODIDAE, 52 jaragua (Nops), 59 Kakopoda, 353
jabase (Nemobiopsis), 373 jaragua (Phyllophaga), 122 kalaom (Helicopsyche), 391
Jabulania, 371 jaraguensis (Bahadzia), 29 kaliki† (Cryptorhopalum), 133
jackbeardsleyi (Pseudococcus), 280 jarbusalis (Synclera), 327 kallimosum† (Apionion), 167
jacksoni (Rhyscotus), 33 jarmilae (Canthonella), 121 Kallyntrosternidius†, 153
jacobsoni (Prosopodesmus), 89 jarmilae (Rowlandius), 81 KALOTERMITIDAE, 188
jaculifera (Feltia), 357 Jassosqualus, 267 kamali (Anagyrus), 292
Jadera, 254 jatrophae saturata (Anartia), 342 Kangarina, 23
jaegeri (Anetia), 341 javanus (Hypothenemus), 178 Kapala, 292
jaegeri (Platynus), 105 javanus (Plaesius), 109 Kapogea, 67
jaegeri (Trichrous), 152 Jeekelia, 90 karlana‡ (Cytheretta), 22
jagua (Atomosia), 208 Jehaia, 32 karlana‡ (Haplocytheridea), 22
jaiba (Gonatista), 190 jeiti (Cerceris), 313 Karnyothrips, 385
jaiba (Margarettia), 369 jeititia (Euodynerus), 308 kasikus (Robopus), 131
jaiqui (Ectotrypa), 372 jeldesi (Polycentropus), 387 kathirithambyi (Stichotrema), 383
jalappae (Scolytogenes), 179 jelineki (Ataenius), 120 kathleena (Cyclargus), 339
Jalysus, 248 jelskii (Pheidole), 303 Keiferia, 321
jamaicae (Haplaxius), 259 jenkinsi (Glomeridesmus), 86 Keilbachia, 205
jamaicana (Symploce), 186 jeremie (Ischnothele), 56 keiteli (Linepithema), 298
jamaicense (Allotrichoma), 232 jima (Modisimus), 58 keiteli flavescens (Linepithema), 298
jamaicense (Sceliphron), 314 jimagua (Atomosia), 208 keiteli subfasciatus (Linepithema), 298
jamaicensis (Bezzia), 197 jimenezi (Phyllophaga), 122 kelliae (Frankliniella), 384
jamaicensis (Cancrincola), 17 jimmyi (Syspira), 70 kennidae (Phrynus), 54
jamaicensis (Epitragus), 144 joachimscheveni†(Bohartilla), 383 kenscoffi (Phyllophaga), 122
jamaicensis (Knemidokoptes), 51 Jobertus, 244 kenscoffi (Tiphia), 308
jamaicensis (Macrohaltica), 158 Jobocricus, 92 kensleyi (Synalpheus), 36
jamaicensis (Metepeira), 67 Jocara, 327 KEROPLATIDAE, 202
jamaicensis (Mezira), 248 jocundus (Anisepyris), 309 Keroplatus, 202
jamaicensis (Mycetomoellerius), 301 Joguina, 359 kheeli (Embolemus), 311
jamaicensis (Neortholomus), 250 johanni (Manduca), 344 kheelorum† (Cryptocephalus), 161
jamaicensis (Pheidole), 303 johnique (Acanthoscelides), 163 Kilifia, 276
jamaicensis (Psorophora), 201 Johnrichardia, 222 killickorum† (Pintomyia), 195
jamaicensis (Sphex), 314 johnsoni (Fountainea), 340 kimballi (Palpita), 326
jamaicensis (Stenotabanus), 207 johnstonii (Psorophora), 201 Kimula, 77
jamaicensis (Trachymyrmex), 394 jole (Tyrannocoris), 255 KIMULIDAE, 77
jamaicensis frontalis (Trachymyrmex), Jollas, 76 kindli (Tityus), 82
301 Jonga, 37 kingmai‡ (Saida?), 26

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 618
KINNARIDAE, 261 kuhniella (Ephestia), 329 laevicula‡ (Aurila), 23
kinzelbachi† (Bohartilla), 383 Kukulcania, 57 laevigata (Cerceris), 313
kirbyi (Ethmia), 321 kulickae† (Oliarus), 259 laevigata (Nyctibora), 186
kirbyi (Oiketicus), 320 kumanskii (Metrichia), 389 laevigatus (Omalodes), 109
Kiskeya, 158 Kunzeana, 269 laevigatus (Prionus), 155
kiskeya (Modisimus), 58 kunzei (Holoparamecus), 142 laevilimbata (Chymomyza), 229
kiskeya† (Micropholcus), 57 Kurzia, 15 laevis (Birandra), 155
Kiskeyana, 247 Kuschelaxius, 176 laevis (Cyclocephala), 123
kiskeyensis (Scolytocis), 142 kymbalion (Ottenwalderia), 30 laevis (Dallasiellus), 254
kittella (Recurvaria), 321 kyphos (Protophthiracarus), 47 laevis (Meloe), 148
Klapperiches, 46 Labarrus, 119 laevis (Parandra), 155
klapperichi (Ablechrus), 136 labeo (Stamnoderus), 114 laevis (Pityophthorus), 181
klapperichi (Adelina), 146 labiatus (Austrotinodes), 386 laevis (Porcellio), 33
klapperichi (Agrilus), 124 labiatus (Liris), 313 laevis (Trientoma), 144
klapperichi (Ambrosiodmus), 180 Labidura, 183 laevissima (Idolia), 110
klapperichi (Ataenius), 120 LABIDURIDAE, 183 laevius (Metachroma), 162
klapperichi (Blapstinus), 144 LABIIDAE, 183 laevus (Ochrimnus), 249
klapperichi (Parazaona), 80 labis (Alpheopsis), 36 Lagocheirus, 153
klapperichi (Pygmephorus), 49 laborium (Omalium), 112 Lagynochthonius, 79
klapperichi (Tylocerus), 132 Labrundinia, 199 lahottensis (Cylloepus), 126
klapperichorum (Catogenus), 138 labruscae labruscae (Eumorpha), 346 laidion (Zamagiria), 330
klapperichorum (Xenorhipis), 125 labyrinthica (Mulsantina), 140 laluzbrillante (Nesolinoceras), 287
Kleidocerys, 249 Laccobius, 108 lambda (Graminella), 266
knabi (Atractomerus), 169 Laccomimus, 100 lambens (Sarcodexia), 219
knabi (Scolytogenes), 179 Laccophilus, 100 lambi‡ (Krithe), 24
Knemidokoptes, 51 lacerum (Omicron), 308 lamellicornis (Callopisma), 130
KNEMIDOKOPTIDAE, 51 lachauxensis (Peripatus), 13 lamentator (Culex), 200
Knyella, 369 lachauxi (Apenes), 103 lamia (Deinopis), 60
kobberti† (Eospilopsyllus), 382 Lachesilla, 381 lamina (Tripudia), 355
Koenikea, 49 LACHESILLIDAE, 381 lamisma (Leucania), 355
kofleri (Amphonyx), 344 Lachnophorus, 103 Lampetis, 125
kohliellus (Agapostemon), 315 Lachnopus, 174 Lamponius, 377
koki (Chimarra), 386 Lacinipolia, 355 Lamproclasiopa, 231
kolophon (Sogatella), 260 laconia (Mecampsis), 382 Lamprosema, 325
Kolourione, 32 lacrymosus (Toxomerus), 216 LAMPYRIDAE, 129
Komisumena, 73 lacteipennis (Milichiella), 227 lanata (Megachile), 316
konokeraia† (Anoblepsis), 286 Laelius, 310 lanceolata (Bituryx), 348
konstantinovi (Lychnacris), 130 LAEMOPHLOEIDAE, 138 lanceolata (Encarsia), 290
Kormilevida, 247 Laemophloeus, 138 lanceolatus (Euconnus), 116
kovariki (Omalodes), 109 laeta (Oxacis), 148 lancifer (Pseudogaurax), 227
kovariki (Ravilops), 83 laetabilis (Epitausa), 352 languidalis (Tachasara), 320
kozari† (Mixorthezia), 280 laetificus (Platynus), 105 lanigerum (Eriosoma), 275
kraemergabi† (Dohrniphora), 213 Laetodon, 215 lanipes (Atya), 36
kraemeri (Empoasca), 269 laetus (Cephaloplopus), 377 lanipes (Centris), 317
krakatauensis (Indotritia), 47 laetus (Exophthalmus), 173 lanius burmeisteri (Gymnetis), 124
Kramerolidia, 265 laetus (Laetodon), 215 Lantanacoccus, 279
Kricogonia, 338 laeve (Cambalomma), 93 lantanae (Oedionychis), 157
Krithe, 24 laevicauda (Opistacanthus), 83 lantiguai (Microtityus), 82
KRITHIDAE, 24 laevicauda (Panulirus), 38 lantiguai (Rowlandius), 81
Krocarites, 268 Laevicephalus, 266 lanuginosa (Strumigenys), 302
kroyeri (Xiphopeneus), 35 laeviceps (Osorius), 113 lanuginosum (Tetramorium), 304
krugi (Hapithus), 371 laeviceps (Platynus), 105 laodamia (Nahublattella), 187
krugii (Nomada), 318 laevicincta (Apenes), 103 LAOMEDIIDAE, 38
krugii (Sericoceros), 282 laevicolle (Phaenonotum), 109 LAOPHONTIDAE, 17
krutaki‡ (Puriana), 24 laevicollis (Dyscinetus), 123 Lapethinus, 139
kuehnei† (Glabellula), 211 laevicollis (Mesonoterus), 99 Lapethus, 140

619 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

lapidaria (Mulona), 349 Lathropus, 138 Lechytia, 79
lapidiscola‡ (Loxoconcha), 25 laticaudis (Neoulosomus), 171 lecontei (Ambrosiodmus), 180
larae (Pyrisitia), 338 laticephala (Ozophora), 252 lecontei (Laemophloeus), 138
LARGIDAE, 252 laticollis (Atanygnathus), 117 lecontei (Neomida), 146
largoensis (Amblyseius), 53 laticollis (Styloleptus), 153 lecontei tenebricosa (Galerita), 102
Largus, 252 laticornis (Elcaribe), 212 lectularius (Cimex), 245
largus (Hypomicrogaster), 375 laticrus (Ommatius), 210 leeuweni (Chaetanaphothrips), 384
larimar (Bahorucotettix), 365 latifascia (Spodoptera), 356 lefebraei (Perigonia), 345
larimar (Ochrotrichia), 389 latifrons (Microcephalops), 215 legorskyi† (Mezira), 248
larimar (Phyllophaga), 122 latifrons (Porphyrochroa), 213 Leia, 203
larimarena (Callopisma), 130 latigidita‡ (Callianassa), 38 Leiocricus, 92
larimarensis (Tytthonyx), 133 latinasus (Caulophilus), 170 LEIODIDAE, 110
larimerius† (Pulex), 382 latior (Selenophorus), 103 Leiopleura, 124
Larinia, 67 latipalpus (Bahadzia), 29 Leioptilus, 330
larochellei (Dyschiriodes), 106 latipennis (Alaus), 127 Lelegeis, 146
larochellei (Dyschirius), 106 latipes (Ascorhynchus), 45 lelex (Cyclocena), 324
larseni (Nesocyrtosoma), 147 latipes (Mocis), 354 Lema, 161
larvatus (Callinectes), 43 latissima (Kurzia), 15 lema† (Dicranomyia), 193
larvigerus maculifrons (Camponotus), latisternus (Acylomus), 138 lemae (Emersonella), 293
300 latiungula (Phyllophaga), 122 lemaireiana (Hispanodirphia), 344
lasabejanus (Euconnus), 116 latiuscula (Leucania), 356 Lemalelmis, 126
lasabejas (Euconnus), 116 latiusculus† (Pseudarachnocephalus), Lembodes, 171
Lasaeola, 62 376 Lemelba, 115
Lascelina, 329 lativentris (Nesothrips), 385 Lemmasomus†, 171
lasciva (Taeniaptera), 221 latoca† (Proanaxipha), 373 Lemnaphila, 234
lasellensis (Tytthonyx), 133 latocanus† (Neoproxius), 248 Lemphus, 136
LASIOCAMPIDAE, 358 latopalpus† (Hispanonesticus), 61 lengi (Heterophotinus), 131
Lasiochilus, 245 latreillei (Birandra), 155 lenticularis (Banasa), 256
Lasioderma, 135 LATRIDIIDAE, 142 lentus (Derovatellus), 100
Lasioglossum, 315 latrodecti (Baeus), 283 leo leo (Polygonus), 337
Lasiomazus, 87 Latrodectus, 62 Leocomia, 262
Lasiomerus, 245 lattini (Acaricoris), 248 Leogorrus, 247
Lasioseius, 52 lattini† (Paleodoris), 241 leonardi (Metachroma), 162
lassauxii (Erinnyis), 345 latus (Corticatus), 75 leonardodicaprioi (Spintharus), 62
lataniae (Aspidiotus), 277 latus (Ogulnius), 63 leonardorum (Ectmesopus), 159
lataniae (Cerataphis), 275 latus (Platycrepidius), 128 LEPADIDAE, 15
lataniae (Hemiberlesia), 277 latus (Polyphagotarsonemus), 50 Lepas, 15
latens (Bothriocera), 258 latus (Spelaeorhynchus), 54 LEPIDOCYRTIDAE, 95
latens (Miagrammopes), 60 lauralis (Arthromastix), 324 Lepidocyrtus, 95
latepunctata (Lophocampa), 348 laurelae (Cyrtoxipha), 375 Lepidopa, 39
laterale (Phaenonotum), 109 Lausus, 71 LEPIDOPSOCIDAE, 379
lateralis (Apotomoderes), 174 lauta (Anyphaena), 70 Lepidosaphes, 278
lateralis (Copidita), 148 lauta (Drosophila), 230 Lepidosina, 228
lateralis (Gecarcinus), 41 lauta (Ochthera), 232 lepidosoma† (Neosibinia), 172
lateralis (Tropisternus), 109 lautus (Microborus), 179 lepidosuccinea† (Forcipomyia), 197
lateralis (Villa), 211 LAUXANIIDAE, 224 Lepiselaga, 207
lateridens (Hexadesmus), 89 lavagaensis (Silidiscodon), 132 Lepisma, 97
laterispinus (Strategus), 123 lavega (Ponsoonops), 59 LEPISMATIDAE, 97
lateroareatus (Opius), 286 lavega (Zeadolopus), 110 leprete (Modisimus), 58
latespargor (Rhinocricus), 92 lavegaensis (Notaepytus), 139 Leptanilloides, 300
latevittata (Charidotella), 160 laxa (Anasaitis), 74 Leptobasis, 362
Latheticomyia, 221 laxatus (Lophocyclus), 91 Leptocera, 228
Latheticus, 145 Leberis, 15 LEPTOCERIDAE, 390
lathrios (Anaxipha), 375 Lebia, 103 Leptoconops, 197
lathripa (Agalliopsis), 264 leboulet (Thaloe), 71 Leptocorisa, 253
Lathrobium, 114 Lecaniobius, 294 leptodactyla (Uca), 42

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 620
Leptodictya, 242 leucographus (Exophthalmus), 173 limatus (Holcobracon), 285
Leptofoenus, 296 leucographus (Lophoscutus), 247 limbata (Lithocharis), 114
Leptogaster, 209 Leucophenga, 231 limbativentris (Tibraca), 257
Leptogenys, 305 Leucopodella, 216 limbatus (Neobisnius), 117
Leptoglossus, 253 leucoproctum (Hydrochasma), 231 limbipennis (Clivina), 106
LEPTOHYPHIDAE, 235 leucops (Cariblatta), 186 Limbopsylla, 272
Leptolycus, 129 leucopterus (Lachnophorus), 103 limenia (Strymon), 340
Leptomantispa, 361 LEUCOSIIDAE, 41 limitatum (Rhopalopterum), 227
Leptomastidea, 292 LEUCOSPIDAE, 295 LIMNADIIDAE, 14
Leptomastix, 292 leucospila (Antiblemma), 351 Limnellia, 233
Leptomyrmex, 298 leucospila (Antipistis), 352 Limnesia, 49
leptonana (Rogeria), 304 Leucospis, 295 LIMNESIIDAE, 49
Leptonema, 387 leucostomus (Euplectrus), 294 Limniattagenus, 133
Leptonesiotes, 159 Leucothoe, 30 LIMNICHIDAE, 126
Leptopharsa, 242 LEUCOTHOIDAE, 30 Limnichites, 126
LEPTOPHLEBIIDAE, 235 leucothorax (Gymnandrosoma), 331 Limnichoderus, 126
Leptophysa, 158 Leucothrips, 384 Limnocythere, 25
LEPTOPODIDAE, 238 Leucotrichia, 388 LIMNOCYTHERIDAE, 24
Leptopsilopa, 232 leucurum (Homoschema), 158 Limnogonus, 239
leptorhinus† (Anlemmus), 171 leucurum (Metachroma), 163 Limonia, 194
leptorhinus† (Stenommatus), 172 Leurolophus, 271 limonus (Aphytis), 289
leptorhinus† (Toxorhynchus), 167 levigatum† (Stenapion), 167 limosum (Eurytium), 44
Leptosalda, 238 levii (Lupettiana), 71 limpioensis (Heterophotinus), 131
Leptosciarella, 205 levimanus? (Pilumnus), 44 linceci (Dendrothereua), 84
leptosomus† (Neoulosomus), 171 levina (Celiptera), 352 linda (Zaraza), 29
leptosphinctos (Carinisphindus), 136 levinates† (Gnamptogenys), 299 lindae† (Pseudochiridium), 80
leptospica (Phyllophaga), 122 levis (Inti), 293 lindborgi (Selenops), 73
Leptostales, 335 levis‡ (Loxoconcha), 25 Lindingaspis, 277
leptostegus† (Semnorhynchus), 172 lewisi (Brevipalpus), 50 Lindneromyia, 215
Leptostylopsis, 153 LIBELLULIDAE, 363 Lindocerus, 312
Leptoteleia, 284 libera (Neokrisna), 267 linearis (Brentus), 168
Leptotes, 339 liberatus (Glyptotermes), 188 linearis (Sitophilus), 177
Leptotrachelus, 102 Liberiana, 112 lineata (Lophocampa), 348
Leptoypha, 242 liberum (Cycloptilum), 376 lineata (Meriosfera), 77
lepturoides (Neopoeciloderma), 152 liboraria (Epione), 334 lineata (Oxycopis), 148
lepturus (Opistacanthus), 83 Libytheana, 341 lineata (Plesiomma), 210
Leptus, 49 lichas (Empyreuma), 347 lineata lineata (Hyles), 346
leptus† (Dominibrentus), 168 lichenosus (Flatoides), 261 lineatum (Hypoderma), 219
Lepyronura, 262 licinus (Alcimosphenus), 64 lineatus (Elateropsis), 156
Lepyrotica, 320 Licodia, 376 lineatus (Leptogaster), 209
lerouxii (Omoteina), 160 Licracantha, 153 lineatus (Proerginus), 78
Leschenaultia, 220 lienaris (Epidromia), 352 lineatus (Spirobolellus), 92
lesieuri (Lophocampa), 348 Ligia, 32 lineola (Pygiopachymerus), 164
Lesmone, 353 LIGIIDAE, 32 lineolata (Gnamptogenys), 299
Lestes, 362 ligius (Haplopus), 377 Linepithema, 298
LESTIDAE, 362 lignigera (Laphygma), 350 lingafelteri (Plectromerus), 150
lestrelleta (Ischnoptera), 186 lignosellus (Elasmopalpus), 329 linguilabris (Chydorus), 14
lethifer (Haitobolus), 92 ligula (Ischnoptera), 186 lingulatum (Homoschema), 158
Lethocerus, 240 ligula† (Lupettiana), 71 linitis (Amboakis), 135
Letis, 353 Ligyra, 210 linki (Oebalus), 257
Leucania, 355 Ligyrocoris, 251 linnei (Philampelus), 346
leucaniella (Crambus), 323 lilacinus (Planococcus), 280 Lintneria, 344
Leucauge, 65 lilicae (Atopsyche), 390 LINYPHIIDAE, 64
leuce memulus (Pyrisitia), 338 LIMACODIDAE, 330 Liobracon, 286
Leucocera, 161 limatula (Decorosa), 135 LIOCRANIDAE, 71
Leucochrysa, 359 Limatus, 201 Liodema, 146

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liogaster (Ataenius), 120 Loberus, 139 longipes (Zeadolopus), 110
Liohippelates, 227 lobiochaeta (Ptilomyia), 233 longipes (Zelus), 246
lioni (Chrysopa), 359 lobipes (Cyrtaphe), 88 longipes longipes (Rhysida), 84
lioni (Megalomus), 360 lobivertex (Epaphrodita), 190 longipes† (Arachnolithulus), 57
Lipaphis, 274 Lobocleta, 335 longipes† (Nanoctenus), 69
Liparthrum, 178 Lobodesmus, 90 longipes† (Veterator), 72
Lipochaeta, 232 Lobonotus, 44 longipunctatum (Metachroma), 163
Lipographis, 329 Lobopoda, 145 longirostris (Ischnaspis), 278
LIPOSCELIDIDAE, 380 lobosternum (Pterogramma), 228 longirostrum† (Proeces), 170
Liposcelis, 380 Lobrathium, 114 longiscapa† (Cubanoptila), 388
Lipurometriocnemus, 198 lobrega (Leucania), 356 longiscapus (Proceratium), 305
liquidambarus (Pityophthorus), 181 locuples (Anasaitis), 74 longiseta (Cancrincola), 17
Lirceus, 32 loecki (Anagyrus), 292 longiseta (Paralimna), 233
Liriomyza, 225 Loewimyia, 225 longisetosa‡ (Bairdia), 19
Liris, 313 Loganius, 177 longisetosus (Meristacarus), 47
lisa euterpe (Pyrisitia), 338 lohsei (Melanophthalma), 142 longisetus (Mesocyclops), 16
Lispe, 218 lomae (Rhopalum), 312 longisetus† (Epidapus), 204
Lispinomimus, 112 Lomagenes, 261 longisoma† (Spintharus), 62
Lispinus, 112 Lomaita, 64 longispina (Metrichia), 389
Lissantauga†, 127 lomata (Saldula), 238 longispina (Morganella), 277
Lissorhoptrus, 175 lomatus (Philonthus), 117 longispina† (Abanaxipha), 375
lissycoleyae† (Mysmenopsis), 64 Lonchaea, 222 longispina† (Palaeodyctina), 69
Listronotus, 172 LONCHAEIDAE, 222 longispinosus (Neoseiulus), 53
Litargus, 143 longembolus† (Nephila), 67 longispinosus† (Chrosiothes), 61
litchivora (Crocidosema), 331 longembolus† (Succinya), 70 longispinus (Anochetus), 304
Lithocharis, 114 longicarpus (Synalpheus), 36 longispinus (Pseudococcus), 280
Lithocharodes, 117 longicaudatus (Triops), 14 longistyla (Agalliopsis), 264
lithochroma (Pammeces), 322 longiceps (Balega), 115 longitarsis (Newportia), 85
Lithophorus, 139 longicera (Pseudosciara), 205 longitarsis stechowi (Newportia), 85
Lithoseopsis, 380 longicollis (Cryptotermes), 188 Longitarsus, 158
Lithurgus, 316 longicorna (Togotolania), 271 longituba (Corethrella), 199
litigiosus (Ligyrocoris), 251 longicornis (Ectmesopus), 159 longivalvata (Protopulvinaria), 276
Litoblatta, 186 longicornis (Eretmocoris), 248 longula (Cubrasa), 264
Litopenaeus, 35 longicornis (Neohybos), 213 longula (Mecidea), 257
Litoprosopus, 353 longicornis (Paratrechina), 300 longula (Neokrisna), 267
litorale (Tapinoma), 298 longicornis (Philonthus), 117 longula (Strandesia), 21
litorea (Glenanthe), 232 longicornis (Tetrapriocera), 134 longulus (Holepyris), 310
litura (Leogorrus), 247 longicornis hagemanni (Paratrechina), longulus (Paromius), 251
liturata (Megistops), 158 300 loomisi (Nesobolus), 92
lituratus (Polycyrtus), 287 longicornis† (Pholcophora), 58 lopesi (Anapunaphyto), 220
lituratus (Tropicus), 127 longilobus (Tytthonyx), 133 Lophoblatta, 187
litus (Pachnaeus), 175 longimanus (Pagurus), 39 Lophocampa, 348
liviae (Macrobiotus), 13 longimanus longimanus (Dineutus), 99 Lophocyclus, 91
lividalis (Hypena), 357 longinasus† (Parvitermes), 189 Lophodesmus, 90
lividipennis (Geocoris), 249 longioculosa (Bradysia), 204 Lopholeucaspis, 279
lividus (Aphodius), 119 longipalpa (Supella), 187 Lophophora, 356
lividus (Closterocerus), 383 longipalpana (Crocidosema), 331 LOPHOPROCTIDAE, 86
lividus (Conoderus), 128 longipalpus (Aglyptinus), 111 Lophopterygella, 382
lividus (Geograpsus), 41 longipalpus (Metaniphargus), 30 Lophoscutus, 247
lividus (Labarrus), 119 longipalpus (Rowlandius), 81 Lophoturus, 86
lividus (Nasutitermes), 189 longipennis (Megistocera), 194 Lorelus, 143
LIVIIDAE, 271 longipes (Beatadesmus), 88 loreta (Ochthera), 232
Lixophaga, 220 longipes (Brachydeutera), 233 loricatus† (Veterator), 72
llinasi‡ (Pterygocythereis), 27 longipes (Microvelia), 240 loripes (Polyancistrus), 368
lobatus (Proctolaelaps), 52 longipes (Phrynus), 54 lottiae† (Baeotettix), 365
lobatus† (Nops), 59 longipes (Scytodes), 57 louisianae (Strumigenys), 302

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 622
louisianae (Trichocorixa), 241 luridula (Anomis), 351 Lyssomanes, 76
louisianica (Niesthrea), 254 luridus (Dicyphus), 243 Lytogaster, 234
lounsburyi (Encarsia), 290 lusaltans (Drosophila), 230 lytrea (Anartia), 342
lourencoi (Microtityus), 82 lusca (Perigonia), 345 mabelae (Ptilomyia), 233
lowii (Uranotaenia), 201 lustrabilis (Teleonemia), 243 maboya† (Anoplotermes), 189
Loxa, 256 lustralis (Margarodes), 325 Mabra, 326
loxa (Apogonalia), 264 lutea (Helicopsyche), 391 mabuya (Hapithus), 371
Loxandrus, 106 lutea (Nostima), 234 Macalla, 327
loxias (Calisto), 343 luteipes (Atissa), 233 Macaria, 333
Loxoconcha, 25 luteola (Hyperaspis), 140 macateei (Canacea), 226
LOXOCONCHIDAE, 25 luteomarginatus (Burtinus), 253 macaya (Malacomorpha), 378
Loxosceles, 57 luteomargo (Ataenius), 120 macaya (Modisimus), 58
Loxostethus, 146 lutescens (Calliomma), 345 macellaria (Cochliomyia), 217
lua (Bothriocera), 258 luteum (Macrosiphum), 274 machaonides (Heraclides), 338
luacesi (Pseudopothea), 245 Lutosa, 376 MACROTHRICIDAE, 15
lualis (Diaphania), 325 LUTROCHIDAE, 127 macieji† (Cryptorhopalum), 133
lubomiri (Leptoteleia), 284 Lutrochus, 127 macleayanus (Ululodes), 358
LUCANIDAE, 124 lutzi (Apallates), 227 macleayanus venezolensis (Ululodes),
lucas (Panoquina), 336 lutzi (Parancistrocerus), 309 358
lucassi (Trigonodes), 355 Lutzomiops, 201 macleayi (Moinodaphnia), 15
lucayanum (Tetramorium), 304 luxurians (Lachnopus), 174 Maconellicoccus, 279
lucayanus (Camponotus), 300 luzziae† (Quasipirhidius), 143 macoris (Calliclytus), 152
lucibilis (Soberocephala), 226 luzzii† (Agapostemon), 315 MACROBIOTIDAE, 13
lucida (Antillognatha), 64 LYCAENIDAE, 339 Macrobiotus, 13
lucida (Balclutha), 266 lyceius (Calisto), 343 Macrobrachium, 37
lucidalis (Ozamia), 329 Lychnacris, 130 Macrocentrus, 286
lucidatus (Ommatius), 210 lychniferus (Pyrophorus), 128 Macrocera, 203
lucidiferata (Ephyra), 334 lychnus (Pyrophorus), 128 macroceria (Rhopobota), 331
lucidula dulculia (Apenes), 103 lycialis (Botys), 325 Macrocoeloma, 42
lucidum (Elaphidion), 151 LYCIDAE, 129 macroconi† (Sulciana), 271
lucidus (Centruroides), 81 lycopersicella (Keiferia), 321 macrocopa (Moina), 15
lucidus† (Amitermes), 190 lycopersici (Aculops), 48 Macrocyclops, 16
lucidus† (Anochetus), 304 Lycorea, 341 MACROCYPRIDIDAE, 25
lucifer (Poecilanthrax), 211 LYCOSIDAE, 68 Macrocyprina, 19
lucifugum (Macrobrachium), 37 Lyctus, 134 Macrocypris, 25
lucigera (Melipotis), 353 LYGAEIDAE, 249 MACRODINYCHIDAE, 51
Lucilia, 217 Lygaeus, 249 Macrodinychus, 51
lucilinea (Heliothis), 356 Lygistorrhina, 202 Macrodiplax, 363
Lucinia, 340 LYGISTORRHINIDAE, 202 Macrohaltica, 158
lucreciae (Hapithus), 370 Lygromma, 72 Macrolophus, 243
luctifera (Megachile), 316 Lygropia, 325 macromalus (Anthonomus), 169
luctuosus (Liris), 313 Lymaenon, 295 Macronema, 387
ludeni (Tetranychus), 50 Lymeon, 287 MACRONYSSIDAE, 52
ludicrus (Collops), 136 LYMEXYLIDAE, 135 macropalpus (Tyrannochelifer), 80
lugubris (Cryptaspasma), 331 Lymire, 348 Macroperipatus, 13
lugubris lugubris (Enyo), 345 lynx (Nephaspis), 141 macrophthalma (Buenoa), 241
Lumacra, 320 LYONETIIDAE, 323 macropilis (Phytoseiulus), 53
luminosus (Ignelater), 128 Lyroda, 312 macrops (Dadaya), 14
lunatus (Messatoporus), 287 Lysathia, 158 macrops† (Pseudomyrmex), 306
lunatus (Pelopaeus), 314 LYSIANASSIDAE, 30 macropunctata‡ (Aurila), 24
lunigerella (Stylopalpia), 330 lyside (Kricogonia), 338 Macrosiagon, 142
Lupella, 43 lysilomae (Merobruchus), 163 Macrosiphoniella, 274
Lupettiana, 71 Lysiosquilla, 28 Macrosiphum, 274
Lupraea, 158 LYSIOSQUILLIDAE, 28 macrospinatus† (Epidapus), 204
lurida (Madoniella), 136 Lysiphlebus, 285 Macrosteles, 267
lurida (Tafalisca), 372 Lysmata, 36 Macroteleia, 284

623 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Macrothemis, 363 magna (Daphnia), 15 Manduca, 344
Macrothrix, 15 magna† (Hispaniolyna), 69 mangiferae (Coccophagus), 290
mactans (Latrodectus), 62 magniclavus (Signiphora), 296 mangiferae (Hypocryphalus), 178
maculata (Antillipeltis), 135 magnifica (Augochlora), 315 mangiferae (Milviscutulus), 276
maculata (Brachychara), 206 magnificus (Calocosmus), 154 mango (Modisimus), 58
maculata (Chrysometa), 65 magnisetae (Crenidorsum), 273 mangoldi (Cryptotermes), 188
maculata (Cissites), 148 magnistylus (Euides), 259 Mangora, 67
maculata (Diaperis), 146 magniventra‡ (Pellucistoma), 22 Mania, 336
maculata (Dysimia), 260 magnocellus† (Nasutitermes), 189 Manidia, 332
maculata (Europs), 137 magnoculus† (Atlantitermes), 189 manifesta† (Leptosciarella), 205
maculata (Holopyga), 310 magnum (Canthochilum), 121 manihotalis (Pyralis), 330
maculata (Hybla), 269 magnus (Achromoporus), 87 mannerheimi (Colpodes), 105
maculata (Neohadria), 265 magnus (Apachekolos), 209 mannerheimi (Exophthalmus), 173
maculata (Nostima), 234 magnus (Descanso), 75 mannerheimi (Hilarotes), 125
maculata (Paralimna), 233 magnus (Oebalus), 257 mannerheimi (Lema), 161
maculata (Signiphora), 296 magnus (Sergiolus), 72 manni (Amphelictogon?), 89
maculata (Trypanalebra), 269 magophila (Cosmopterix), 322 manni (Amphidraus), 74
maculata† (Bicalcasura), 172 magua (Charinus), 54 manni (Cryptops), 84
maculata† (Borinquena), 235 magualis (Hileithia), 325 manni (Ctenus), 69
maculatipennis (Aphytis), 379 Magusa, 358 manni (Entypus), 307
maculativentris (Buprestis), 125 maidis (Dalbulus), 267 manni (Epyris), 310
maculatus (Aeolus), 128 maidis (Peregrinus), 259 manni (Hyptia), 288
maculatus (Belopherus), 168 maidis (Rhopalosiphum), 274 manni (Liobracon), 285
maculatus (Dermestes), 133 maior† (Prototama), 60 manni (Metazygia), 67
maculatus (Isometrus), 82 maissadensis‡ (Cyprideis), 21 manni (Perigonia), 345
maculatus (Nesticus), 61 MAJIDAE, 42 manni (Peripatus), 13
maculatus (Physcius), 149 major (Atopocixius), 261 manni (Siphonophora), 91
maculellus (Amplicephalus), 266 major (Gonatista), 190 Manoa, 198
maculicauda† (Metaleia), 203 major (Pseudaulacaspis), 278 Mansonia, 200
maculicorium (Melanopleurus), 249 major (Rutherfordia), 278 MANTIDAE, 190
maculicornis (Aphytis), 289 major major (Polistes), 309 MANTISPIDAE, 360
maculicornis (Oligota), 112 majus (Placonotus), 138 maoensis‡ (Nephrops), 38
maculicornis (Pentagonica), 104 majusculus (Monobelus), 270 maquayensis‡ (Costa), 27
maculipennis (Contarinia), 202 makandal (Modisimus), 58 maraicola (Styloleptus), 154
maculipennis (Euhyloptera), 260 Malacomorpha, 378 Marasmia, 326
maculipennis (Falconia), 244 Malacopterus, 152 Marava, 183
maculipennis (Morellia), 218 Malacopus, 246 marcaeusalis (Ommatospila), 326
maculiventris (Podisus), 255 malangae (Aleuroglandulus), 272 marcano (Phyllophaga), 122
maculosa (Amauromyza), 225 malariager† (Culex), 200 marcanoi (Centruroides), 81
maculosa (Leucophenga), 231 malcusalis (Petrophila), 323 marcanoi (Erythrolychnia), 130
maculosa (Nemorimomyza), 224 maldonadoi (Antias), 243 marcanoi (Neoscona), 67
maculosa† (Fictotama), 60 maldonadoi (Hadria), 265 marcanoi (Orthobelus), 270
maculosum (Hoplocheiloma), 221 maldonadoi (Prepops), 244 marcanoi (Paranapiacaba), 159
maculosus (Ommatius), 210 mali (Monarthrum), 181 marcanoi (Polycentropus), 387
maculosus (Psalistops), 56 mali (Phyllophaga), 122 marcanoi (Selenops), 73
maddeni (Trichopelma), 57 malitiosa adoceta (Synapte), 336 marcanoi (Stenotabanus), 207
madeirensis (Phenacoccus), 280 Malleoceps, 115 marcella (Miathyria), 364
maderae (Displotera), 142 malpighiae (Anthonomus), 169 marcus (Platynus), 105
maderae (Rhyparobia), 185 malthiniformis (Caccodes), 132 margana (Spragueia), 350
madgesuttonae† (Proanaxipha), 373 maltzani (Jobocricus), 92 margaretae† (Milichiella), 227
Madoniella, 135 malus (Horismenus), 293 Margarettia, 369
madremaestrae‡ (Echinocythereis), 27 mammalicolus† (Culicoides), 196 margaritatus (Centruroides), 81
maerula (Anteos), 338 mammidentata‡ (Trachyleberidea), 27 margaritensis (Iridopagurus), 39
maesites (Chlorostrymon), 339 Mamurius, 253 MARGARODIDAE, 280
maestrica (Calliphora), 217 mandibularis (Hentzia), 76 marginale (Crenidorsum), 273
maestris (Caliadurgus), 307 mandibularis (Sphex), 314 marginalis (Apenes), 103

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 624
marginalis (Batina), 351 martyni† (Gnophomyia), 192 medialuna (Syspira), 70
marginalis (Liriomyza), 225 Maruca, 326 medianus (Aporinellus), 307
marginalis (Rhinotermes), 188 mascula (Gasteracantha), 66 mediator (Anadenobolus?), 92
marginalis (Semidalis), 360 masneri (Adelphe), 310 medinai (Hydroptila), 388
marginalis† (Hypogena), 145 masneri (Anastatus), 294 Mediocampus, 256
marginata (Chinavia), 256 masneri (Cladochaeta), 229 mediocris (Antillosorius), 112
marginata (Cicindela), 101 masneri (Conobregma), 287 medioculatus† (Nasutitermes), 189
marginata (Cynortoides), 78 masneri (Enicospilus), 288 mediodiffusa (Drosophila), 230
marginata (Cyphomyia), 206 masneri (Eriastichus), 294 mediovittata (Gymnometopa), 199
marginata (Pepsis), 307 masneri (Gasteruption), 288 meditabunda (Edessa), 255
marginata (Phymata), 247 masneri (Leptolycus), 129 medium (Pachyolpium), 79
marginata (Pseudozonitis), 148 masneri (Pityophthorus), 181 Medon, 114
marginata (Stagmonantis), 190 masneri (Pseudothysanoes), 178 medranae (Sicoderus), 176
marginata (Tettigonia), 265 masneri (Spalangia), 296 medranoae (Erythrolychnia), 130
marginatus (Carios), 51 masneri (Szelenyia), 297 medranoi (Erythrolychnia), 130
marginatus (Ochrimnus), 250 masneri (Trichomyia), 195 megacephala (Chrysobothris), 125
marginatus (Oxybelus), 312 masneri (Yoshimotoana), 296 megacephala (Chrysomya), 217
marginatus (Paracoccus), 280 Masona, 286 megacephala (Pheidole), 303
marginatus (Pseudothysanoes), 178 Massala, 353 megaceros (Orthobelus), 270
marginatus fabricii (Hyporhagus), 143 Masteria, 56 Megacerus, 163
margineguttatus (Thermonectus), 100 Mastigodiaptomus, 16 Megachile, 316
marginella (Pimpla), 287 Mastigoproctus, 83 MEGACHILIDAE, 316
marginella (Thaumasia), 68 Mastigus, 116 megacornis (Acanthoscelides), 163
marginemaculatus (Phrynus), 54 Mastotermes, 188 Megadytes, 100
marginicollis (Thalpius), 106 MASTOTERMITIDAE, 188 megaholcus† (Neoulosomus), 171
marginicollis fumipes (Pseudaptinus), matasana (Medon), 114 megalognatha (Bohartilla), 383
102 materiarius (Gnathotrichus), 182 megalognatha† (Bohartilla), 383
marginipennis (Paraclivina), 107 mathisi (Cariboptila), 388 Megalomus, 360
maria (Choranthus), 336 mathisi (Dagus), 233 megalopia (Alagoasa), 157
mariana (Leucauge), 65 matilei† (Dilophus), 202 Megalostrata, 72
Marietta, 289 matronaria (Epimecis), 333 megalurus (Arrenurus), 49
Marilia, 390 matsoni‡ (Perissocytheridea), 22 Megamelus, 259
Marimatha, 349 Matsucoccus, 280 Megaphragma, 297
marimutti (Enicocephalus), 238 matthewsi (Canthochilum), 121 MEGAPODAGRIONIDAE, 362
marionae (Urgleptes), 154 mattijoae† (Phatnoma), 242 megapunctatus (Gnathotrupes), 181
mariposa (Calisto), 343 maura (Buprestis), 125 MEGARIDIDAE, 255
mariposa (Lychnacris), 130 mauryae‡ (Mesorhoea), 43 Megaris, 255
mariposas (Modisimus), 58 maxillosus (Antillatus), 75 Megaselia, 214
maritima (Anisolabis), 183 maxillosus (Neoconocephalus), 368 megasomus† (Semnorhynchus), 172
marlatti (Aleurolobus), 272 maximus (Titanophilus), 85 MEGASPILIDAE, 297
marlowei (Dipogon), 307 maya (Trogolaphysa), 96 megastigma (Rileya), 295
marmorata (Emesa), 246 mayaguana (Drosophila), 230 Megischus, 282
marmoratus (Archytas), 220 mayri (Anochetus), 304 Megistocera, 194
Marpesia, 340 Mazax, 72 Megistops, 158
marquettorum† (Oligochlora), 315 mcelfreshi (Phymacista), 242 Megophthalmidia, 203
Marshallella, 270 Mecampsis, 382 megus† (Neoulosomus), 171
Marshallius, 176 Mecaphesa, 73 MEINERTELLIDAE, 96
martiana (Mecynogea), 67 Mecidea, 257 meinerti (Rheumatobates), 239
martinella (Metapenaeopsis), 35 MECISTOCEPHALIDAE, 85 Meioneta, 64
martingarcia (Achromoporus), 87 Mecistocephalus, 85 meiosa (Agalliopsis), 264
martini (Copelatus), 100 Mecynogea, 67 melaleucus (Karnyothrips), 385
martini† (Schildia), 209 medealis (Samea), 325 Melalgus, 134
martiniensis (Euhybus), 213 mederi† (Cubanoptila), 388 melana (Beameromyia), 209
martorelli (Amblycerus), 163 media (Brasa), 264 Melanaethus, 254
martorelli (Macrosiphum), 274 media† (Prototama), 60 Melanagromyza, 224
martorelli (Sitobium), 274 medialis‡ (Caudites), 24 Melanaphis, 274

625 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Melanaspis, 277 menkei† (Pison), 313 Metachroma, 162
melancholica (Dasymutilla), 306 menocrater (Apotomoderes), 174 metacista (Nylanderia), 300
melanocephala (Cyclocephala), 123 MENOPODIDAE, 379 METACRANGONYCTIDAE, 30
melanocephalum (Tapinoma), 299 Menopon, 379 Metacrangonyx, 30
melanochaeta (Helicopsyche), 391 mensuralis (Remigia), 354 Metacyclops, 17
melanochrotum (Metachroma), 163 Menudo, 256 Metacynortoides, 78
melanogaster (Diceratobasis), 362 Meoneura, 226 Metacyrba, 76
melanogaster (Drosophila), 230 mera (Kramerolidia), 265 Metadelphax, 259
Melanolestes, 246 mera (Orasiopa), 231 Metadorylas, 215
melanoleuca (Derancistrachroma), 156 mercator (Lachnopus), 174 Metaleia, 203
Melanophila, 125 mercedes (Eurycotis), 184 Metaleurodicus, 273
Melanophthalma, 142 mercurius sanchesi (Proteides), 337 Metallata, 354
Melanopleuroides, 249 merengue (Chimarra), 386 metallica (Apraea), 157
Melanopleurus, 249 merengue (Pseudomethoca), 306 metallica (Oxycopis), 148
Melanoplus, 367 meridianus (Anoplotermes), 189 metallicus (Acylophorus), 117
melanostictum (Theridion), 63 meridianus (Heterocerus), 127 metallicus (Pandeleteius), 175
melanostoma (Scolopocryptops), 85 merielae (Heterophotinus), 131 metallosomus (Platynus), 105
melanotela (Embola), 322 Meriosfera, 77 Metamasius, 177
melanurus (Calocosmus), 154 Meristacarus, 47 metanasticos (Nemobiopsis), 373
Melathra, 174 meritoria (Encarsia), 290 Metaniphargus, 30
Meleoma, 359 merkli (Nesocyrtosoma), 147 Metapenaeopsis, 35
Melete, 339 Mermessus, 64 Metaphrenon, 152
MELICHARIDAE, 52 Merobruchus, 163 Metaphycus, 292
melidora (Ardeutica), 331 Merocoris, 253 METAPSEUDIDAE, 29
Melipotis, 353 Meromacrus, 215 Metasesarma, 41
melissa (Choranthus), 336 Meropliosepsis, 224 Metasinella, 95
melissia (Cymotricha), 321 merriami (Pasiphaea), 37 metaspilaris (Cobubatha), 349
melissiflora† (Paleoeuglossa), 317 mervini (Attheyella), 17 Metatrichia, 211
Melissodes, 317 Mesembrinella, 219 Metazygia, 67
Melissoptila, 317 MESEMBRINELLIDAE, 219 Metepeira, 67
Melittobia, 294 Mesepipsocoides, 380 Methia, 152
mella (Phyllophaga), 122 Mesepipsocus, 380 meticulosalis (Terastia), 327
mellea (Bradysia), 204 Mesethodesmus, 90 Metopaulias, 41
mellifera (Apis), 316 Mesocheira, 317 Metophthalmus, 142
mellifluus (Pyrophorus), 128 Mesochria, 201 Metopina, 214
mellitae (Dissodactylus), 43 Mesocyclops, 16 Metria, 354
mellitor (Bracon), 285 Mesodixa, 201 Metrichia, 389
mellonella (Galleria), 328 mesogramma apa (Atalopedes), 336 Metriocnemus, 198
melodermus (Phormictopus), 56 Mesolia, 323 Metrobates, 240
Meloe, 148 mesonotalis (Acropyga), 302 mexicana (Diplozona), 243
MELOIDAE, 148 Mesonoterus, 99 mexicana (Dyseuaresta), 223
Melormenis, 261 Mesopenaeus, 35 mexicana (Peosina), 353
melvini (Ollanta), 263 Mesoplia, 317 mexicana (Potimirin), 37
MELYRIDAE, 136 Mesoplocia, 235 mexicana‡ (Cyprideis), 21
MEMBRACIDAE, 269 Mesoplophora, 47 mexicanensis (Tomosvaryella), 215
membranifera (Strumigenys), 302 Mesorhoea, 43 mexicanum (Trypoxylon), 313
membrosa (Cydia), 331 mesosignaria (Argyrotaenia), 332 mexicanus (Chirothrips), 384
Memphis, 340 mesosphaeri (Macrosiphum), 274 mexicanus (Hephebocerus), 168
Menacanthus, 379 Mesovelia, 239 Mexicolaophonte, 17
menalcasalis (Bleptina), 356 MESOVELIIDAE, 239 Mexobisium, 79
mendicus (Amphidraus), 74 Messatoporus, 287 Mezira, 248
mendocinus (Metacyclops), 17 Messinella, 22 mgogoricus (Gabronthus), 117
mendocinus (Microcyclops), 17 mestrei (Lasioglossum), 315 Miagrammopes, 60
Menemerus, 76 Metabus, 65 Miathyria, 363
menes (Ceranisus), 292 Metacanthus, 248 micans (Lobopoda), 145
Menippe, 44 Metacerylon, 140 micans (Oxydrepanus), 107

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 626
micans (Platydema), 146 Microsema, 333 minima (Typhlocybella), 269
micculalis (Heterogramma), 356 Microsepsis, 224 minimus (Chlorotettix), 265
michaeli (Euconnus), 116 microsomus† (Toxorhynchus), 167 minimus (Microtityus), 82
micheneri (Melissoptila), 317 microspinus† (Epidapus), 204 minisculum† (Nesocyrtosoma), 147
micheneri† (Oligochlora), 315 MICROSTIGMATIDAE, 56 minisculus (Zeadolopus), 110
miches (Scaphioides), 60 Microthyris, 326 minisignaria (Argyrotaenia), 332
Micracis, 178 Microtityus, 82 Ministrymon, 339
Micracites†, 182 microtremus† (Dryophthorus), 172 minor (Belostoma), 240
Micrasta, 125 Microvelia, 240 minor (Corythalia), 75
micrata (Scopula), 335 Microxestoleberis, 28 minor (Isotomiella), 95
Micrathena, 67 micrura (Moina), 15 minor (Larinia), 67
Micrathetis, 350 Micrutalis, 271 minor (Megalomus), 360
Micrathyria, 364 miersii (Armases), 41 minor (Planococcus), 280
Micratya, 37 miki† (Archaeopristocera), 309 minor (Polistes), 309
micrepelis (Apocephalus), 214 mikii (Atoniomyia), 208 minor (Purenleon), 361
Microborus, 179 Milichiella, 227 minor† (Prototama), 60
Microcentrum, 368 MILICHIIDAE, 227 minoris† (Dohrniphora), 213
Microcephalops, 215 militaris (Belopherus), 168 Minthea, 134
Microcephalothrips, 384 militaris (Micrathena), 67 minuenda (Idona), 269
microceria (Rhopobota), 331 militaris nigriceps (Dasymutilla), 306 minuscula (Homoptera), 355
Microcerotermes, 190 milleri (Incisitermes), 188 minuta (Adelphe), 310
Microchironomus, 198 milleri (Leptostylopsis), 153 minuta (Aedmon), 157
Microchrysa, 206 millsi (Empoasca), 269 minuta (Antillipeltis), 135
microcirculus (Pseudococcus), 280 millsi (Metasinella), 95 minuta (Corimelaena), 255
Microcorthylus, 181 millsi millsi (Dicranocentrus), 95 minuta (Glenognatha), 65
Microcrambus, 323 MILNESIIDAE, 14 minuta (Minutaleyrodes), 273
Microcyclops, 17 Milnesium, 14 minuta (Nehalennia), 362
Microcythere, 25 Milviscutulus, 276 minuta (Zeugomantispa), 361
MICROCYTHERIDAE, 25 Mimapsilopa, 232 minuta histrionica (Ellipes), 364
microdenticulus (Creagrophorus), 111 mimasalis (Acrolophus), 319 minuta‡ (Coquimba), 24
Microdiphot, 131 Mimeoma, 123 Minutaleyrodes, 273
Microdipoena, 64 mimeticus (Dryotribus), 171 minuticola† (Dasyhelea), 197
Microdon, 215 MIMETIDAE, 60 minutior (Cardiocondyla), 302
microholcus† (Neoulosomus), 171 Mimetus, 60 minutissima (Camptoptera), 295
Microissus, 261 mimica (Melissodes), 317 minutissima (Orasema), 292
MICROMALTHIDAE, 99 mimicum (Canthochilum), 121 minutissima (Phyllophaga), 122
Micromalthus, 99 mimicus (Acanthotomicus), 179 minutissimus (Microcorthylus), 181
micromettleri (Drosophila), 230 Mimogonus, 112 minutissimus (Telenomus), 284
Micromimus, 170 mimosae (Mimosestes), 164 minutulis† (Lorelus), 143
Micromus, 360 Mimoschinia, 324 minutum (Agathidium), 110
Micromyrmex, 172 Mimosestes, 163 minutum (Trichogramma), 297
Micronotus, 365 mimulus mimulus (Dysdercus), 252 minutus (Alaptus), 295
Micropanope, 44 Mindazerius†, 275 minutus (Aleurotuberculatus), 272
Microparsus, 274 mingodo (Amphiacusta), 373 minutus (Ceracis), 142
MICROPEZIDAE, 221 Minibiotus, 13 minutus (Cyphomyrmex), 301
Micropholcus, 57 minikyanos (Electrostrymon), 339 minutus (Dinoderus), 134
Microphrys, 42 Minilentus, 13 minutus (Laemophloeus), 138
microplus (Rhipicephalus), 52 Minilimosina, 228 minutus (Myopsocus), 382
microps (Echiaster), 114 minima (Amphiacusta), 374 minutus (Perileptus), 107
Micropsephodes, 141 minima (Cochliomyia), 217 minutus (Planes), 41
microptera (Espagnoleta), 366 minima (Curtara), 268 minutus‡ (Basslerites), 27
micropterus (Haplopus), 377 minima (Heikertingerella), 157 minyops (Lemalelmis), 126
microptilus† (Stenaspidiotus), 161 minima (Malacomorpha), 378 Minyscapheus†, 263
Micropygomyia, 194 minima (Melanagromyza), 224 Minysporus†, 255
Microrhagus, 127 minima (Neoelmis), 126 miocena† (Pintomyia), 195
Microscydmus, 116 minima (Prytanes), 251 miocena† (Protochymomyza), 230

627 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

miocenecus† (Syringogaster), 221 MOCHLOZETIDAE, 46 Monomacra, 158
miocenica‡ (Callianassa), 38 Mocis, 354 Monomorium, 303
miocenica‡ (Pterygocythereis), 27 moderatus (Adelencyrtus), 291 Mononychellus, 50
miocenicus† (Thaumatodryinus), 311 moderatus (Nemocephalus), 168 Monopelopia, 199
miocenicus† (Trinemurodes), 97 modesta (Anyphaena), 70 monopeteinus (Strymon), 340
Miogryllus, 372 modesta (Baeodasymyia), 196 MONOPHLEBIDAE, 279
Miohebrus, 239 modesta (Teutona), 62 monops (Tabanus), 207
Miomyia, 230 modestior (Anadenobolus?), 92 MONOTOMIDAE, 137
Mionochroma, 150 modestula (Eulepidotis), 352 monotropidia (Dictyla), 242
Miophlugis†, 368 modestus (Agobardus), 74 monstratura (Thermesia), 351
Mioptachys, 107 modestus (Elipsocus), 381 monstruosus (Haitianotettix), 365
mira (Glenognatha), 65 modestus (Engytatus), 243 montana (Azilia), 64
mira (Walkerana), 369 modestus (Notaepytus), 139 montana (Calisto), 343
mira† (Rhipidia), 194 Modisimus, 58 montana (Empoasca), 269
mirabilis (Chondrocera), 253 moerens (Leucauge), 65 montana (Galerita), 102
mirabilis (Hemisphaerodella), 243 moerens (Pheidole), 303 montana (Phloeoxena), 104
mirabilis (Presbyolampis), 132 moerens (Ranzovius), 244 montana (Psychoda), 195
mirabilis† (Clystopsenella), 311 moestus (Coccus), 275 montanus (Achromoporus), 87
Miraces, 159 MOGOPLISTIDAE, 376 montanus (Antillatus), 75
MIRACIIDAE, 18 Moina, 15 montanus (Chaetophloeus), 178
miranda (Nymphodromia), 187 MOINIDAE, 15 montanus (Descanso), 75
Mirda, 78 Moinodaphnia, 15 montanus (Duartettix), 367
mirebalina (Oxyethira), 389 molina (Sogatella), 260 montanus (Haplocyclodesmus), 91
mirella (Urodus), 321 Molinaranea, 67 montensis (Lychnacris), 130
Mirhipipteryx, 365 mollis (Coenipeta), 352 monticellus (Corthylus), 181
miri (Modisimus), 58 mollissima atlantica (Pallenopsis), 45 monticola (Ctenus), 69
MIRIDAE, 243 Molophilus, 192 monticola (Dellia), 366
mirifica (Eremonidia), 346 Molosoma, 112 monticola (Enicospilus), 288
mirificus (Gonoporomiris), 244 mona (Neotermes), 188 monticola (Heteronebo), 83
mirzah (Azeta), 351 monachus (Apate), 134 monticola (Heterophotinus), 131
Mischocyttarus, 309 monachus (Trox), 119 monticola (Iridopsis), 333
misella (Tetracha), 101 monae (Olpiolum), 79 monticola (Micrasta), 126
miserum (Ceresium), 150 monae (Typhlatya), 37 monticola (Tainophasma), 378
misippus (Hypolimnas), 342 Monanus, 138 monticola (Teminius), 70
Misumenops, 74 Monarthrum, 181 montischristi (Exelastis), 330
mitchellii (Wyeomyia), 201 Monelata, 288 montivaga (Chinavia), 256
Mithrax, 42 monicae (Leptoteleia), 284 montrouzieri (Cryptolaemus), 141
mithrax (Chiomara), 337 monilicornis (Anadenobolus), 92 montserratensis (Icerya), 279
mithridates sagra (Achlyodes), 337 monilicornis (Morion), 104 monuste eubotea (Ascia), 339
mitis (Microsepsis), 224 monilicornis (Telenomus), 284 Moodna, 329
mitnicki† (Termitaradus), 275 monilis (Brentus), 168 Moodnopsis, 329
mitnickii† (Molinaranea), 67 monima (Eunica), 340 mopsaria mopsaria (Cyclomia), 333
MITURGIDAE, 70 Monobeloides, 270 morator (Ataenius), 120
Mixochlorus, 125 Monobelus, 270 moratus (Espeson), 112
Mixorthezia, 280 Monobia, 308 mordax (Xylocopa), 318
mixta (Crangoweckelia), 29 Monoblemma, 59 Mordella, 142
mixtalis (Audia), 333 monoceros† (Chrosiothes), 61 MORDELLIDAE, 142
mixtus (Poecilopompilus), 308 monoceros‡ (Bythoceratina), 20 Mordellistena, 142
Mnemosyne, 259 monocerus (Scarabaeus), 123 moreli (Neoscona), 67
mnemosyne† (Zorotypus), 391 Monochaetoscinella, 227 Morellia, 218
mnestor (Brachymeria), 291 monochroma (Apotoforma), 332 Morganella, 277
m-nigrum (Rachispoda), 229 Monocloeus, 165 mori (Brevipalpus), 50
mobilis (Condica), 350 Monodella, 29 morio (Apenes), 103
mocaensis (Cephalosphaera), 214 MONODELLIDAE, 29 morio (Cymindis), 103
Mochloribatula, 46 Monodesmus, 155 morio (Paederus), 114
Mochlozetes, 47 monogrammata (Lobocleta), 335 morio (Zophobas), 145

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 628
Morion, 104 munitalis (Bonchis), 327 Myriophora, 214
morionella (Fletcherimyia), 219 munitus (Lophocyclus), 91 myripristis (Anilocra), 31
morkhoveni‡ (Krithe), 24 munroei (Nefundella), 329 Myrmarachne, 76
Morlockia, 15 Munseyella, 26 Myrmecholax, 383
MORLOCKIIDAE, 15 muralis (Cliniodes), 324 Myrmecoderus, 149
Mormidea, 257 murcialis (Ebulea), 324 MYRMECOLACIDAE, 383
moronei† (Brachypsectra), 127 Murgantia, 257 myrmecophaga (Nemorhinus), 168
moronei† (Dicromantispa), 360 Murgisca, 327 MYRMECOPHILIDAE, 376
moronei† (Euglossa), 317 muricatus (Pseudothysanoes), 178 Myrmecophilus, 376
moronei† (Nesagapostemon), 316 murina (Cubaris), 32 Myrmelachista, 300
morosa (Dipoenata), 62 murinum (Pelonium), 136 Myrmeleon, 361
morrisoni (Amblystira), 242 murinus (Cephalalges), 175 MYRMELEONTIDAE, 361
morrisoni (Eobates), 239 muriscis (Ectomyelois), 329 myrmex (Velapia), 373
morrisoni (Leptoypha), 242 Mursa, 356 Myropsis, 41
morro (Selenops), 73 musae (Amblyseius), 53 myrtacei (Aleurothrixus), 272
morsei (Symploce), 186 musaeperda (Frankliniella), 384 myrto (Eudamus), 337
morsitans (Scolopendra), 85 musarum (Epaphrodita), 190 Myscelia, 340
mortua† (Limbopsylla), 272 Musca, 218 MYSIDAE, 29
mosieri (Metamasius), 177 MUSCIDAE, 218 Mysidium, 29
Mosillus, 232 muta† (Euryconus), 272 MYSMENIDAE, 63
mosquera‡ (Loxoconcha), 25 mutabile (Nesocyrtosoma), 147 Mysmenopsis, 64
motasi (Cyathura), 31 mutabilis (Buenoa), 241 mythica (Amphiacusta), 374
Mothocya, 31 mutabilis (Tytthonyx), 133 Mythicomyia, 211
Mothonica, 322 mutans (Loxandrus), 106 MYTHICOMYIIDAE, 211
Mouralia, 357 mutatum (Anelaphus), 151 mythicos (Hapithus), 370
mucronatus (Podisus), 255 mutica (Pelia), 42 Mythimna, 356
Mucrotettix, 365 muticus caraibeus (Anurogryllus), 372 Myzine, 308
muellenmeisteri† (Azilia), 64 mutilata (Macrosiagon), 142 Myzus, 274
muesebecki (Calepitrimerus), 48 mutilatus (Carpophilus), 137 n. sp. (Ceratina), 318
muesebecki (Laelius), 310 MUTILIIDAE, 306 n. sp. (Hylaeus), 314
mulleri (Drosophila), 230 mycerina (Letis), 353 n. sp. (Parvitermes), 189
Mulona, 349 Mycetaspis, 277 n. sp. 1 (Brachycorynus), 165
mulsanti (Mesovelia), 239 Mycetobia, 201 n. sp. 1 (Eugonops), 165
Mulsantina, 140 MYCETOPHAGIDAE, 143 n. sp. 1 (Monocloeus), 165
multicincta (Glaucopis), 348 Mycetophila, 203 n. sp. 1 (New genus A, nr. Cyptoxenus),
multidentatus (Callicentrus), 270 MYCETOPHILIDAE, 202 164
multifila (Diadiplosis), 202 Mycocepurus, 301 n. sp. 1 (Piesocorynus), 164
multiguttata (Oxycopis), 148 Mycococcus, 276 n. sp. 1 (Ptychoderes), 164
multiguttatus (Cryptocephalus), 161 Mycodrosophila, 230 n. sp. 1 (Trypoxylon), 313
multiguttula (Hexatoma), 193 Mycomya, 203 n. sp. 10 (Homocloeus), 165
multilineatum (Zagrammosoma), 294 MYCTERIDAE, 148 n. sp. 10 (Ormiscus), 165
multipes (Telocricus), 85 MYDIDAE, 211 n. sp. 11 (Homocloeus), 165
multiplicalis (Samea), 327 myersi (Carabunia), 291 n. sp. 11 (Ormiscus), 165
multiporosa (Mochloribatula), 46 Myllaena, 111 n. sp. 12 (Homocloeus), 165
multipunctata (Malacomorpha), 378 Mymar, 295 n. sp. 12 (Ormiscus), 165
multipunctata (Paralimna), 233 MYMARIDAE, 295 n. sp. 13 (Homocloeus), 165
multiseta (Robustagramma), 229 mymaripenne (Megaphragma), 297 n. sp. 13 (Ormiscus), 165
multisetus (Hyperlasion), 205 Myochrous, 163 n. sp. 14 (Homocloeus), 165
multispinosa (Ariadna), 59 Myodocha, 251 n. sp. 14 (Ormiscus), 165
multispinus (Froeschneria), 251 myopa (Pholeomyia), 227 n. sp. 15 (Homocloeus), 165
munda (Napata), 348 Myophora, 220 n. sp. 15 (Ormiscus), 166
mundissima (Concana), 352 MYOPSOCIDAE, 382 n. sp. 16 (Homocloeus), 165
mundulus (Cyclodinus), 149 Myopsocus, 382 n. sp. 16 (Ormiscus), 166
mundus (Lachnopus), 174 Myospila, 218 n. sp. 17 (Homocloeus), 165
mundus (Selenophorus), 103 myrae (Hydrometra), 239 n. sp. 17 (Ormiscus), 166
Munidopsis, 39 myricae (Parabemisia), 273 n. sp. 18 (Homocloeus), 165

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n. sp. 18 (Ormiscus), 166 nana (Lamproclasiopa), 231 Nefundella, 329
n. sp. 19 (Ormiscus), 166 nana (Paectes), 355 negans (Apotoforma), 332
n. sp. 2 (Brachycorynus), 165 nana (Psyllobora), 140 neglecta (Balclutha), 266
n. sp. 2 (Holostilna), 164 nana† (Palaeoanapis), 63 neglecta (Nephropsis), 38
n. sp. 2 (Piesocorynus), 164 nanaria (Cyclophora), 335 neglecta (Phyllophaga), 122
n. sp. 20 (Ormiscus), 166 naniki (Margarettia), 369 neglecta (Saissetia), 276
n. sp. 21 (Ormiscus), 166 nanion (Hapithus), 370 Nehalennia, 362
n. sp. 22 (Ormiscus), 166 Nannohelea, 196 neiba (Calisto), 343
n. sp. 23 (Ormiscus), 166 Nannoparce, 344 neiba (Cyrtaphe), 88
n. sp. 24 (Ormiscus), 166 Nannosquilla, 28 neiba (Decorosa), 135
n. sp. 25 (Ormiscus), 166 NANNOSQUILLIDAE, 28 neiba (Eurycotis), 184
n. sp. 26 (Ormiscus), 166 Nanoctenus†, 69 neiba (Leucania), 356
n. sp. 27 (Ormiscus), 166 Nanolachesilla, 381 neiba (Podiscodesmus), 88
n. sp. 28 (Ormiscus), 166 Nanolycus, 129 neiba (Tilloclytus), 150
n. sp. 29 (Ormiscus), 166 Nanus, 176 neibae (Cazierius), 83
n. sp. 3 (Brachycorynus), 165 nanus (Anthonomus), 169 neibae (Kiskeya), 158
n. sp. 3 (Holostilna), 164 nanus (Antillotettix), 365 neibae (Tityus), 82
n. sp. 3 (Toxonotus), 165 nanus (Zeadolopus), 110 neibaense (Nesocyrtosoma), 147
n. sp. 30 (Ormiscus), 166 nanus† (Araneus?), 66 neibaensis (Lophocampa), 349
n. sp. 31 (Ormiscus), 166 naomi (Psorthaspis), 308 neibaensis (Notaepytus), 139
n. sp. 32 (Ormiscus), 166 Napata, 348 Neibamastax, 366
n. sp. 33 (Ormiscus), 166 naranjo (Ctenus), 69 neibana (Argyrotaenia), 332
n. sp. 34 (Ormiscus), 166 naranjo (Rowlandius), 81 neibanus (Acridurus), 367
n. sp. 35 (Ormiscus), 166 narrans (Ophiusa), 354 Neivamyrmex, 299
n. sp. 36 (Ormiscus), 166 nasalis (Gasterophilus), 219 Nelasa, 141
n. sp. 37 (Ormiscus), 166 nascens† (Trientoma), 144 Nelphe, 348
n. sp. 38 (Ormiscus), 166 nasidens (Pachodynerus), 309 neltumae (Tricorynus), 135
n. sp. 39 (Ormiscus), 166 nasoni (Lispe), 218 nemesis nemesis (Siderone), 341
n. sp. 4 (Brachycorynus), 165 Nasutitermes, 189 Nemobiopsis, 373
n. sp. 4 (Holostilna), 164 nasutus (Belopherus), 168 Nemocephalus, 168
n. sp. 4 (Toxonotus), 165 nasutus (Belorhynchus), 168 Nemomydas, 211
n. sp. 40 (Ormiscus), 166 Nathalis, 338 Nemorhinus, 168
n. sp. 41 (Ormiscus), 166 NAUCORIDAE, 240 Nemorimomyza, 224
n. sp. 51 (Ormiscus), 166 Naupactus, 175 Neoacanthoneura, 223
n. sp. 6 (Homocloeus), 165 Nauphoeta, 185 neoalagitans (Drosophila), 230
n. sp. 7 (Homocloeus), 165 Naushonia, 38 Neoathyreus, 119
n. sp. 8 (Homocloeus), 165 Nausibius, 138 Neobidessus, 100
n. sp. 9 (Homocloeus), 165 nautica (Neoscona), 67 Neobisnius, 117
n. sp. 9 (Ormiscus), 165 navarroi (Parhaplopus), 377 Neoblattella, 187
n. sp. nr. fossoria (Phyllophaga), 122 navis‡ (Cativella), 27 neobrevipes (Dysmicoccus), 279
n. sp.‡ (Caudites), 24 nealcesalis (Radara), 354 Neocallichirus, 43
n. sp.‡ (Cyprideis), 21 Neapion, 167 Neocaudites, 24
NABIDAE, 245 nearctica (Nomophila), 326 Neocenchrea, 260
Nabis, 245 nearnsi (Elaphidion), 151 neochma (Calisto), 343
nabokovi (Hesperia), 336 nearnsi (Nesocyrtosoma), 147 Neoclytus, 150
naboria† (Anoplotermes), 189 neascripta (Joruma), 269 Neococytius, 344
Nacaeus, 112 Nebridia, 74 Neoconis, 359
Nacarina, 359 nebulonemi (Canthochilum), 121 Neoconocephalus, 368
nacrealis (Cliniodes), 324 nebulosa (Cladochaeta), 229 Neocorynura, 315
nadleri (Cesonia), 72 nebulosa (Cymatothes), 144 neodarlingtoni (Tytthonyx), 132
naevifera (Polyortha), 331 nebulosa (Drosophila), 230 Neodiachipteryx, 102
Nahublattella, 187 nebulosus (Valenzuela), 381 Neodiploclampta, 211
naindaimei (Euseius), 53 necopinum (Caribophyllum), 368 Neoelmis, 126
nakos (Eriastichus), 294 Necrobia, 136 Neoerythromma, 362
Nallachius, 360 Nectopsyche, 390 Neogalea, 357
nana (Colpoptera), 261 necydalea (Methia), 152 Neogonodactylus, 28

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 630
Neographium, 332 neotropica (Brachydeutera), 233 Nessorhinus, 270
Neogriphoneura, 224 neotropica† (Mesochria), 201 NESTICIDAE, 61
Neohadria, 265 neotropicalis (Neopamera), 251 Nesticodes, 62
Neohannia, 69 neotropicus (Enicospilus), 377 Nesticus, 61
Neoheterocerus, 127 neotropicus (Trichobius), 217 nesydrion (Cycloptilum), 376
Neohybos, 213 neotropicus† (Leptomyrmex), 298 Nettarhinus, 176
Neohydatothrips, 384 Neoulosomus, 171 Neuraphes, 117
Neohypnus, 117 Neowadotes, 69 neuricella (Bema), 328
Neokrisna, 266 Neoxantholinus, 117 New genus F, nr. Goniocloeus, 165
Neolargulus, 252 Neoxenus, 164 Newportia, 85
Neolasioptera, 202 Nephaspis, 141 newtoni† (Dolichoxenus), 113
Neolema, 161 Nepheloleuca, 333 nexilis† (Pseudomyrmex), 306
Neoleucinodes, 326 Nephila, 67 Nezara, 257
Neolimonia, 194 nephilae (Argyrodes), 61 ni (Trichoplusia), 357
Neoliodes, 47 Nephotettix, 267 Nialaphodius, 119
NEOLIODIDAE, 47 NEPHROPIDAE, 38 niarchos† (Leptosalda), 238
Neomachilellus, 96 Nephrops, 38 nicaeense (Trischizostoma), 31
Neomamersa, 49 Nephropsis, 38 Nicentrus, 170
Neomastix, 169 Nephrotoma, 194 nicetaria (Thysanopyga), 334
neomedialis† (Lindneromyia), 215 Neplas, 215 nicippe (Abaeis), 338
Neomegalotomus, 253 nepos (Electropteron), 129 nicippiformis (Abaeis), 338
Neomida, 146 Neralsia, 289 nicoaria (Patalene), 334
Neoninus, 249 nerii (Aphis), 274 NICOLETIIDAE, 97
Neopamera, 251 nerii (Aspidiotus), 277 Nicrophorus, 111
Neopamphantus, 249 NERIIDAE, 221 Niesthrea, 254
Neopanope, 44 nerisaria (Pero), 334 nigellus (Ommatius), 210
Neopetissius, 251 Nerius, 221 niger (Araptus), 180
Neopisinus, 62 nervosa (Gynacantha), 363 niger (Chondrotropis), 88
Neoplea, 241 Nesagapostemon†, 316 niger (Ufens), 297
Neopleurophora, 214 Nesanoplium, 151 nigerrimus (Euconnus), 116
Neopoeciloderma, 152 Nesara, 358 nigli (Lamponius), 378
Neopomphale, 293 Nesepyris, 310 nigra (Mimeoma), 123
Neoproxius, 248 nesicos (Orphulella), 367 nigra (Nothomyia), 206
Neorhipidius, 142 Nesidiocoris, 243 nigra (Parasaissetia), 276
Neorileya, 295 Nesidiphilus, 85 nigra (Tainocampa), 347
Neortholomus, 250 Nesiosphaerion, 151 nigra† (Holocompsa), 187
Neoscapteriscus, 376 Nesiostrymon, 339 nigrescens (Anadenobolus), 92
neoschwarzi (Urosigalphus), 285 nesiota (Drosophila), 230 nigrescens (Marilia), 390
neoschwarzi† (Scolytodes), 179 nesiotes (Ctenophilus), 86 nigrescens (Strumigenys), 302
Neoscona, 67 nesiotes (Hypasclera), 148 nigriabdominalis (Tetraneura), 275
Neoseiulus, 53 nesiotes (Loxa), 256 nigricans (Grotiusomyia), 294
Neosibinia, 172 nesiotes (Polycricus), 85 nigricans† (Alumeda), 245
Neosilba, 222 nesiotica (Caribovia), 264 nigricephala (Encarsia), 290
Neosmerinthothrips, 385 nesiotus (Hypothenemus), 179 nigriceps (Cercyon), 109
Neosorius, 112 nesium (Trichapion), 167 nigriceps (Crassomicrodus), 285
Neospintharus, 62 neslihanae (Lychnacris), 130 nigriceps (Perigona), 104
Neostasina, 73 Nesobolus, 92 nigricollis (Heterophotinus), 131
Neotanygastrella, 230 nesocoris (Cymophyes), 250 nigridens (Fecelia), 256
Neotanypeza, 222 Nesocyrtosoma, 147 nigrifrons (Graminella), 266
Neotermes, 188 Nesogenes, 258 nigrifrons (Nacaeus), 112
neotheis (Neoscona), 67 Nesogerax†, 183 nigrifrons (Robopus), 131
Neotheronia, 287 Nesolinoceras, 287 nigrigaster (Yelicones), 287
NEOTHYRIDAE, 54 Nesomylacris, 186 nigriguttata (Polyortha), 331
Neotrachys, 124 Nesonana, 212 nigrilabris (Gyretes), 99
Neotrichia, 389 Nesosphecodes, 315 nigrimana (Leptopsilopa), 232
neotropica (Allograpta), 216 Nesothrips, 385 nigrina (Silis), 132

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nigrina (Sparatta), 183 nitidimacula (Gonodonta), 352 Notiobiella, 360
nigripalpus (Culex), 200 nitidula (Holocompsa), 187 Notioplusia, 358
nigripennis (Calocosmus), 155 NITIDULIDAE, 137 Notiphila, 234
nigripennis (Centrocorisa), 241 nitiduloides (Tisiphone), 137 notitipenne (Hoplocheiloma), 221
nigripennis (Laevicephalus), 266 nitidulus (Chydorus), 14 notius (Mimetus), 60
nigripes (Trichrous), 152 nitidulus (Coproporus), 118 Notocyphus, 306
nigriscripta (Laphygma), 352 nitidum (Pseudolathra), 114 NOTODONTIDAE, 346
nigrisensilla (Helicopsyche), 391 nitidum (Trichopelma), 57 Notomicrus, 100
nigrita (Haitiana), 258 nitidus (Centruroides), 81 Notonecta, 241
nigrita (Lebia), 104 nitidus (Hispanotettix), 367 NOTONECTIDAE, 241
nigrita (Nialaphodius), 119 nitidus taino (Centruroides), 81 nova† (Trichosia), 205
nigritarsa (Hamataliwa), 68 nivea (Panchlora), 186 Novantinoe, 153
nigritarsis (Calocosmus), 155 niveiciliata (Desmia), 325 novemdecimpunctatus (Tropirhinus), 174
nigritubus (Wyeomyia), 201 niveicollis (Ptinus), 135 Novohorus, 79
nigriventris (Erythrolychnia), 130 niveipennis (Minilimosina), 228 nr. eburneus‡ (Amphibalanus), 16
nigrivestis (Stenotarsus), 141 niveivenosa (Nostima), 234 nr. erythrocerus (Dyschirius), 106
nigrivittata (Grallipeza), 221 niveus (Lophoturus), 86 nr. Goniocloeus sp. 1 (New genus F), 165
nigroaenea (Spalangia), 296 nivosella (Ethmia), 321 nr. Goniocloeus sp. 2 (New genus F), 165
nigroclarus (Ardistomis), 106 noacki (Cales), 290 nr. Goniocloeus tuberculatus (New genus
nigrocoeruleus (Neobisnius), 117 noar (Pheidole), 303 F), 165
nigrocoxalis (Hadrosomus), 249 nobilis (Actenodes), 125 nubicola (Lebia), 104
nigrodorsata (Elytrogona), 160 nobilis (Alaus), 127 nubila (Eumecosomyia), 223
nigrofasciatus (Styloleptus), 153 nobilis (Aralichus), 51 nubila (Pseudatomoscelis), 244
nigrohalteralis (Paralauterborniella), nobilitella (Cydosia), 349 nubilipennis (Acerophagus), 291
198 noctilucus (Pyrophorus), 128 nubilus (Emesopsis), 246
nigrolateralis (Telephanus), 137 noctis (Mioptachys), 107 nubilus (Heterophotinus), 131
nigrolineata tolteca (Neotheronia), 287 noctuata (Leptostales), 335 nucicolora (Elaphria), 355
nigromaculata (Chaetoclusia), 225 NOCTUIDAE, 349 nuda (Oedenops), 233
nigromaculatum (Metachroma), 163 noctuiformis noctuiformis (Cautethia), nuda (Rhysida), 84
nigronervosa (Pentalonia), 274 346 nudatus (Haplocyclodesmus), 91
nigrorosea (Boenasa), 349 noda (Apogonalia), 264 nudinervosa (Leptosciarella), 205
nigrorubra (Vulsirea), 257 nodifrons (Menippe), 44 nudiseta (Synthesiomyia), 218
nigrosetosus (Klapperiches), 46 noeli (Erythmelus), 295 nudistyla† (Corethrella), 199
nigrosignatus (Eurypedus), 160 NOGODINIDAE, 262 nudus (Plagiodontochirus), 51
nigrostriatus (Leptolycus), 129 Nola, 357 nuezana (Argyrotaenia), 332
nigroviride (Metachroma), 163 Nomada, 318 numeria (Hemeroblemma), 353
nikitai (Fallceon), 235 Nomophila, 326 numerus (Neomastix), 169
Nilakantha, 76 nonseriatus (Selenophorus), 103 Numia, 333
Nilaparvata, 259 Nops, 59 nunezi (Curtara), 268
Nilio, 143 noralis (Odilla), 324 nunezi (Phyllophaga), 122
NINIDAE, 250 Normaltica, 158 nunezi (Purenleon), 361
Ninyas, 250 Nostima, 234 Nusalala, 360
nipae (Nipaecoccus), 279 notabilis (Caupolicana), 314 nutriaria (Patalene), 334
Nipaecoccus, 279 notabilis (Cymoninus), 249 nutrix (Gonodonta), 352
nipeensis‡ (Caudites), 24 Notaepytus, 139 nuttingi (Micropanope), 44
nishei (Pasiphaea), 37 Notapictinus, 248 nutua† (Elaver), 71
nisibonensis (Triepeolus), 317 notapuncta (Lobopoda), 145 nychios (Cophella), 374
nitaina (Tainacris), 366 notata (Bertamyia), 215 nyctalemonaria (Epimecis), 333
nitaino† (Anoplotermes), 189 notata (Cyclocephala), 123 Nyctelius, 336
Nitela, 312 notatalis (Nephopteryx), 328 nyctelius nyctelius (Nyctelius), 336
nitens (Dermacentor), 52 notatella (Ethmia), 321 Nyctibora, 186
nitens (Tetragnatha), 65 notatus (Tupiocoris), 243 nyctinomos (Anurogryllus), 372
nitida (Athyroglossa), 232 NOTERIDAE, 99 Nyctiplanctus, 159
nitidalis (Diaphania), 325 Nothodelphax, 259 Nyctosyles, 106
nitidicollis (Hilarotes), 125 Nothomyia, 206 Nylanderia, 300
nitidicutis (Pseudomus), 172 notidanus (Campylothorax), 95 NYMPHALIDAE, 340

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 632
nymphidiata (Lobocleta), 335 obscurus (Metacynortoides), 78 Ochthera, 232
Nymphocixia, 259 obscurus (Zeros), 234 Ochyrocera, 57
Nymphodromia, 187 obscurus† (Cephalotes), 301 OCHYROCERATIDAE, 57
Nyridela, 348 obsolescens (Thinodromus), 113 ocoa (Brasiella), 101
Nysius, 250 obsoleta (Helicoverpa), 356 ocola distipuncta (Panoquina), 336
Nysson, 312 obsoleta (Myospila), 218 Ocrepeira, 67
Nystalea, 347 obsoleta (Thyanta), 257 ocresia (Anastrepha), 223
oakleyi (Xantholinus), 118 obsoleta† (Chaetosciara), 204 Octhispa, 160
oarion‡ (Bairdia), 19 obsoletus (Exophthalmus), 173 octodecimguttatus (Cryptocephalus), 161
oaxacensis (Neoscona), 67 obsoletus (Rhytidoporus), 254 octolineata (Parahadria), 265
oberthuri (Actenodes), 125 obstinatus (Epidapus), 204 octomaculata (Polythrix), 337
oberthuri (Prosternodes), 156 obstipa (Amphiacusta), 374 octopustulata (Leucocera), 161
obesa (Ornidia), 215 obtectus (Acanthoscelides), 163 Octostruma, 301
obesula (Chaetocnema), 157 obtusa (Bradysia), 204 octotuberculatus (Brachyomus), 173
obesum (Metachroma), 163 obtusa (Melanagromyza), 224 ocularis (Ulomoides), 146
obesus (Haplocyclodesmus), 91 obtusiceps (Monobelus), 270 oculata (Apraea), 157
Obila, 335 obtusus (Amblyseius), 53 oculata (Cyclosa), 66
obliqua (Allograpta), 216 obvelatus (Acanthoscelides), 163 oculata (Pseudosquilla), 28
obliqua (Anastrepha), 223 obvertens (Massala), 353 oculatus† (Hymenorus), 145
obliqua (Empoasca), 269 Ocabaraja, 115 oculatus† (Modisimus), 58
obliqua (Lutosa), 376 ocampensis (Tergoceracris), 367 oculosa (Bradysia), 204
obliquella (Vindana), 328 ocampina (Leucochrysa), 359 ocululiter† (Corythalia), 75
obliquus (Ambrosiodmus), 180 Occemyia, 221 ocypete (Enyo), 345
obliquus (Sphyrocoris), 258 occidentalis (Frankliniella), 384 Ocypode, 42
obliterata (Antiblemma), 351 occidentalis (Lysathia), 158 OCYPODIDAE, 42
obliteratus (Urgleptes), 154 occidentalis (Neoconocephalus), 368 Ocyptamus, 216
oblitus (Rhinostomus), 173 occidentalis (Otostigmus), 84 odilae (Cosmosoma), 348
oblonga (Prytanes), 251 occidentalis (Phloeopora), 112 Odilla, 324
oblongata‡ (Bairdoppilata), 19 occidua (Tricharaea), 220 Odinia, 226
oblongostriata (Mnemosyne), 259 occulta (Melanagromyza), 224 ODINIIDAE, 226
oblongus (Loxostethus), 146 Occultocythereis, 27 odius odius (Historis), 342
oblongus (Strategus), 123 occultum (Acrolophus), 320 Odo, 70
oblongus‡ (Portunus), 43 occultum (Cycloptilum), 376 Odontacolus, 284
obovata (Ochrotrichia), 389 occultus (Achromoporus), 87 Odontamera†, 171
obovatus (Largus), 252 Oceanaspidiotus, 277 Odontobracon, 286
obrienae (Caribovia), 265 ocellatus (Calappa), 40 ODONTOCERIDAE, 390
obrieni (Dryinus), 311 ocellatus (Cerylcautomus), 139 Odontomachus, 305
obrieni† (Ogygius), 170 ocellea (Mothonica), 322 Odontota, 160
obscura (Calisto), 343 ochleros (Walkerana), 369 Odopoea, 263
obscura (Chrysometa), 65 Ochlerotatus, 199 odorata (Ascalapha), 351
obscura (Discocerina), 231 Ochlodes, 336 Oebalus, 257
obscura (Erinnyis), 345 ochraceum (Simulium), 201 Oecanthus, 373
obscura (Malacomorpha), 378 ochraceus (Enochrus), 108 Oecetis, 390
obscura (Myllaena), 111 ochreatus ochreatus (Dysdercus), 252 Oecleus, 259
obscura (Paralimna), 234 Ochrimnus, 249 Oeclidius, 261
obscura (Scatella), 233 ochrocerus (Entypus), 307 OECOBIIDAE, 60
obscura‡ (Platylambrus), 43 ochrochlaena (Claduncaria), 332 Oecobius, 60
obscurator (Althos), 253 ochrodes (Melipotis), 353 OECOPHORIDAE, 321
obscurella (Discocerina), 231 ochrodesma (Anabasis), 328 Oedancala, 250
obscurifrons (Hypothenemus), 179 ochropezus (Stenolophus), 103 OEDEMERIDAE, 148
obscurior (Brachymyrmex), 300 Ochrostomus, 250 Oedenops, 233
obscuriventris (Tetreuaresta), 222 ochrota (Eucereon), 347 Oedichirus, 114
obscurus (Agobardus), 74 Ochrotrichia, 389 oediloma† (Spiloconis), 360
obscurus (Elcaribe), 212 ochrura (Wyeomyia), 201 Oedionychus, 159
obscurus (Euschistus), 256 OCHTERIDAE, 241 oedipodalis (Epicorsia), 325
obscurus (Hypothenemus), 179 Ochthebius, 110 oedipus (Anthrax), 210

633 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Oedudes, 155 Omophoita, 158 opsia (Amphiacusta), 374
oemearia (Drepanodes), 334 Omophron, 106 optiva† (Styringomyia), 193
oenotrus (Erinnyis), 345 Omorgus, 119 optivus (Prosaspicera), 289
oerstedii (Neogonodactylus), 28 Omoteina, 160 oraapis (Anthonomus), 169
OESTRIDAE, 218 Omphale, 293 Oraesia, 354
oestrum (Cymothoa), 31 omphaleae (Dilophonota), 345 Orasema, 292
Oethecoctonus, 284 Oncerotrachelus, 247 Orasiopa, 231
Ogcodes, 206 Oncocephalus, 247 orbicularis (Spodoptera), 356
Ogulnius, 63 Oncolabis, 329 orbifera (Magusa), 358
Ogygius, 170 Oncopeltus, 250 orbis (Lychnacris), 131
Ogyrides, 36 oncora (Neokrisna), 267 orbis browni (Aphrissa), 338
OGYRIDIDAE, 36 oneitus (Psocus), 382 Orchestina, 59
Oidaematophorus, 330 ONISCIDAE, 32 orchidearum (Cerataphis), 275
Oiketicus, 320 Oniscus, 33 orchidii (Chaetanaphothrips), 384
oileus (Pyrgus), 337 Onthophagus, 120 orcus† (Micromimus), 170
Oithona, 17 onusta (Tramea), 364 ordinans (Isogona), 353
OITHONIDAE, 17 Onychotillus, 136 ordwayi (Portunus), 43
Okamninus, 134 oo (Pseudoplusia), 357 oreades (Hispanacris), 367
Olbiogaster, 201 Oodinus, 104 Orekthophora, 270
Olcella, 227 Oomorphus, 163 Oreodera, 154
oleae (Saissetia), 276 Oomyzus, 294 Orfelia, 202
oleivora (Phyllocoptruta), 49 OONOPIDAE, 59 ORIBATIDAE, 47
oleosa (Collaria), 244 Oosternum, 109 Oribotritia, 47
Olethreutes, 331 opaca (Euchroa), 106 ORIBOTRITIIDAE, 47
olfersi (Macrobrachium), 37 opaciceps (Hypoponera), 305 oriel (Ocyptamus), 216
Olfersia, 217 opacicollis (Ablechrus), 136 orientalis (Aonidiella), 277
Oliarus, 259 Opacifrons, 228 orientalis (Atherigona), 218
Oligembia, 235 opacior (Hypoponera), 305 orientalis (Atopsyche), 390
Oligochlora†, 315 opaculus (Catharthosilvanus), 138 orientalis (Blatta), 184
Oligocixia, 259 opaculus (Thinobius), 113 orientalis (Pteroptrix), 290
Oligomyrmex, 303 opacum (Tapinoma), 299 orientalis (Scarites), 107
Oligota, 112 opacus (Hypothenemus), 179 orientinum (Trypoxylon), 313
olima† (Platycorypha), 272 Opalimosina, 228 Orimarga, 194
olivacea (Alamis), 354 opalisans (Syllepte), 327 Orionina, 24
olivacea (Stenocrepis), 104 operculatus (Pylopagurus), 39 Oripoda, 47
olivaceella (Phycitodes), 330 operculella (Phthorimaea), 321 ORIPODIDAE, 47
oliveri (Culicoides), 196 opertanium (Cycloptilum), 376 Orius, 245
oliveri† (Antillopsyche), 387 opesta (Limnocythere), 25 orizaba (Tetragnatha), 65
olivieri (Erythrolychnia), 130 Opharus, 349 orizae (Caulophilus), 170
Ollanta, 263 Ophiomyia, 224 Orizaephilus, 138
Ollarianus, 266 Ophion, 288 orizicola (Tagosodes), 260
OLPIIDAE, 79 Ophisma, 354 Ormenana, 261
Olyca, 329 Ophiusa, 354 Ormenaria, 261
Omalium, 112 Ophrynopus, 282 ormenidis (Bloodiella), 297
Omalodes, 109 OPILIOACARIDAE, 52 Ormenina, 261
Omegalebra, 268 opimum (Homoschema), 158 Ormia, 220
omen† (Enicocephalus), 238 Opistacantha, 284 Ormiscus, 165
Omicron, 308 Opistacanthus, 83 ORMYRIDAE, 296
omicron (Ephyrodes), 352 opisthopus (Culex), 200 Ormyrus, 296
ominosa (Amphiacusta), 374 Opius, 286 ornata (Epinannolene), 93
Omiodes, 326 Opogona, 320 ornata (Heza), 246
omissa (Upogebia), 38 Opopaea, 59 ornata (Leptophysa), 158
Ommatius, 209 OPOSTEGIDAE, 319 ornatipennis (Drosophila), 230
Ommatospila, 326 Oppia, 47 ornatipennis (Espagnolopsis), 366
Omolicna, 260 OPPIIDAE, 47 ornatipennis (Thyreodon), 288
Omonadus, 149 Opsablechrus, 136 ornatrix ornatrix (Utetheisa), 347

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 634
ornatrix stretchii (Utetheisa), 347 OTOCEPHEIIDAE, 48 PACHYGRONTHIDAE, 250
ornatula (Chroantha), 256 OTOPHEIDOMENIDAE, 52 Pachylia, 345
ornatus (Aleurodicus), 273 Otostigmus, 84 Pachylioides, 345
ornatus (Atopocixius), 261 Otroacizzia†, 271 Pachyneuron, 296
ornatus (Callinectes), 43 ottenwalderi (Tityus), 82 Pachyolpium, 79
ornatus (Oebalus), 257 Ottenwalderia, 30 Pachyschelus, 124
ornatus (Reuteroscopus), 245 ottleyi (Ctenus), 69 Pachyteles, 107
ornatus (Thinobius), 113 otus (Nesocyrtosoma), 147 pacifica (Disorygma), 289
Ornebius, 376 ovale† (Theridion), 63 padovanii (Marshallella), 270
Ornidia, 215 ovalis (Apenes), 103 Paectes, 355
Ornithoctona, 217 ovata (Brachymeria), 291 Paederus, 114
Ornithodorus, 52 ovata (Caribovia), 265 paenevisus (Euconnus), 116
ornithogalli (Spodoptera), 356 ovata (Omoteina), 160 Pagiocerus, 177
ornithoglyphus (Toxomerus), 216 ovata‡ (Radimella), 24 Paguristes, 39
Orochirus, 371 ovatus (Achromoporus), 87 Pagurus, 39
orpha (Telomerina), 229 ovatus† (Cyptoxenus), 164 pahayokeensis (Manoa), 198
Orphulella, 367 oviedensis (Jaragua), 366 pala† (Phlegrata), 76
Orsa, 78 Oviedinus, 134 Palaehydropsyche†, 387
Orthellia, 203 oviedo (Agobardus), 74 PALAEMONIDAE, 37
Orthemis, 364 oviedo (Heteronebo), 83 palaenova† (Chimarra), 386
ORTHEZIIDAE, 280 oviedo (Selenops), 73 Palaeoanapis†, 63
Orthobelus, 270 oviedoensis (Robopus), 131 palaeobrevicristatus† (Thoracophorus),
Orthocladius, 198 oviedoensis (Zeadolopus), 110 113
Orthocomotis, 332 ovipes (Fennellia), 90 palaeobrevipteryx† (Spelobia), 229
Orthomegas, 155 ovis (Bovicola), 379 palaeodominicana† (Chimarra), 386
orthomera (Gonomyia), 192 ovisternus (Druciatus), 228 palaeodominicana† (Fallia), 141
orthonoides (Thermocyclops), 17 Oxacis, 148 palaeodominicanus† (Dryinus), 311
Orthopodomyia, 200 Oxidus, 89 Palaeodyctina†, 69
Orthoporus, 93 Oxybelus, 312 palaeoelegans† (Calosopsyche), 387
Orthotrichia, 389 OXYCARENIDAE, 250 Palaeoepeirotypus†, 63
ortizi (Phyllophaga), 122 Oxycarenus, 250 Palaeolathys†, 70
ortizorum (Oxyethira), 389 Oxycopis, 148 Palaeometopia†, 137
Orus, 114 Oxydia, 334 Palaeopatasson†, 295
ORUSSIDAE, 282 Oxydrepanus, 107 Palaeopus, 171
oryctes (Geogryllus), 372 Oxyethira, 389 palaeorchestes† (Tethorchestia), 30
Oryctometopia, 329 Oxyidris†, 303 palaeospinosa† (Xenolimosina), 229
oryctus† (Pseudomyrmex), 306 Oxyopes, 68 palaeothoracis† (Drosophila), 230
ORYIDAE, 85 OXYOPIDAE, 68 palaeus† (Zorotypus), 391
oryzae (Lasioseius), 52 Oxysarcodexia, 219 Palaminus, 114
oryzae (Latheticus), 145 Oxytelus, 113 palassaina (Kiskeyana), 247
oryzae (Sitophilus), 177 Oxyurella, 15 paleocaribis† (Neoconis), 360
oryzophilus (Lissorhoptrus), 175 Ozamia, 329 paleocircularis† (Dohrniphora), 213
Osachila, 40 Ozius, 44 paleodominica† (Anacaena), 108
Osbornellus, 267 Ozophora, 251 paleodominicanus† (Coptotermes), 188
osborni (Brachyufens), 297 pacatalis (Hypena), 357 Paleodoris†, 241
oscinidis (Chrysocharis), 293 Pachnaeus, 175 Paleoeuglossa†, 317
Osmia, 316 Pachodynerus, 309 Paleohemilophus†, 155
Osoriellus, 113 Pachybrachis, 162 paleolata (Macaria), 333
ossipellis (Oidaematophorus), 330 pachychaeta (Efferia), 208 paleoliberatus† (Glyptotermes), 188
ostreata (Pseudoparlatoria), 278 Pachycheles, 40 Paleomastacris†, 366
ostreum (Zaops), 43 Pachychernes, 80 paleomexicanus† (Dryinus), 311
othnius† (Pseudopilolabus), 167 pachyconcha‡ (Triangulocypris), 21 paleomystax† (Lindocerus), 312
otho druryi (Wallengrenia), 336 Pachycoris, 258 paleonesia† (Ululodes), 358
otiliae† (Pterolophosoma), 151 Pachygaster, 206 paleopestis† (Pintomyia), 195
otiosa (Perigea), 350 Pachygrapsus, 41 Paleophthorus†, 182
otiosus (Cryptorhynchus), 171 Pachygrontha, 250 Paleoploiariola†, 246

635 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Paleosinus†, 182 paludigenus (Ceraticelus), 64 Paracloeodes, 235
paleotownsendi† (Pintomyia), 195 palustris (Stenocrepis), 104 Paracoccus, 280
paleotrichia† (Pintomyia), 195 palvet (Modisimus), 58 paracubana (Adelphe), 310
Paleotrichius†, 124 Pammeces, 321 Paracyclops, 17
Palinurellus, 38 panamaensis (Soberocephala), 226 Paracypris, 20
PALINURIDAE, 38 panamense (Anteon), 310 Paracythere, 26
palisoti (Antillosciocoris), 257 panamensis (Microcyclops), 17 Paracytheridea, 25
palisoti (Hololepta), 109 panamensis (Scymnus), 141 PARACYTHERIDEIDAE, 25
palisoti (Proarna), 263 Panchlora, 186 Paracytheroma, 22
paliurus (Taraxippus), 378 pandalicola (Probopyrus), 32 Paradejeania, 220
Pallenopsis, 45 Pandeleteius, 175 Paradella, 33
pallens (Schistocerca), 367 pandens (Rhamphothrips), 385 paradoxa (Cariboptila), 388
pallescens (Ozophora), 252 paniceum (Stegobium), 134 PARADOXOSOMATIDAE, 89
pallescens (Rejanellus), 74 panicula (Phyllophaga), 122 Paradoxostoma, 26
pallescens (Tetragnatha), 65 panisea (Empoasca), 269 PARADOXOSTOMATIDAE, 26
pallida (Aedmon), 157 pannosa‡ (Jugosocythereis), 24 Paraglenanthe, 232
pallida (Cyptocephala), 256 pannosata (Obila), 335 Paragonatas, 251
pallida (Eidmanella), 61 pannychios (Hapithus), 370 Parahadria, 265
pallida (Loxa), 256 Panonychus, 50 parahageni (Helicopsyche), 391
pallida (Methia), 152 panopealis (Helvibotys), 325 Parahydriena, 262
pallida (Phoetalia), 185 Panopeus, 44 Parahymenorus, 145
pallida (Stantonia), 286 panops (Physemops), 233 paraiso (Modisimus), 58
pallidiceps (Parvitermes), 189 Panoquina, 336 Parakrithe, 24
pallidicornis (Myospila), 218 Panstrongylus, 247 Parakrithella, 24
pallidicornis (Neoclytus), 150 Pantala, 364 Paralabella, 183
pallidicosta (Rhyacionia), 332 panthalon panthalon (Horama), 347 Paralauterborniella, 198
pallidiventre† (Trypoxylon), 313 Panthea, 357 Paraleucolepidopa, 39
pallidoides (Pamphantus), 249 pantheratus pantheratus (Anetia), 341 Paraleyrodes, 273
pallidus (Achromoporus), 87 Pantographa, 326 Paralicus, 170
pallidus (Anthicus), 149 Panula, 354 Paralienates†, 238
pallidus (Antillocoris), 250 Panulirus, 38 Paralimna, 233
pallidus (Araptus), 180 panurgon (Cycloptilum), 376 parallela (Elabra), 268
pallidus (Delphastus), 141 papayae (Acerophagus), 291 parallela (Iomoides), 90
pallidus (Ectmesopus), 159 papayae (Empoasca), 269 parallela (Myrmarachne), 76
pallidus (Eretmocerus), 290 Papilio, 338 parallela (Parallelodera), 135
pallidus (Phytonemus), 50 PAPILIONIDAE, 337 parallelana (Coelostathma), 332
pallidus (Sapintus), 149 papinianus (Eantis), 337 parallelilobata (Silis), 132
pallidus (Syneches), 213 pappiaria (Drepanodes), 334 parallelocollis (Euconnus), 116
pallidus (Thymoites), 63 papulans (Corthylus), 181 Parallelodera, 135
pallipes (Buenoa), 241 paquetae (Merobruchus), 163 parallelos (Rhysotritia), 47
pallipes (Heterostylum), 211 Parabaeus, 283 parallelus (Platypus), 182
pallipes (Hydrochus), 108 Parabemisia, 273 Paramacrobiotus, 13
pallipes (Licodia), 376 Parabetaeus, 36 Parametriocnemus, 199
pallipes (Lytogaster), 234 Parabezzia, 196 Paramusonia, 190
pallipes (Saldula), 238 parabolocrata (Leocomia), 262 Paramycodrosophila, 230
Pallodes, 137 Paracamptopsis†, 171 Paranapiacaba, 159
palmae (Cerataphis), 275 Paracanace, 226 Parancistrocerus, 309
palmarum (Hemisphaerota), 160 Paracarniella, 243 parander (Streptopsyche), 388
palmarum (Tricorynus), 135 Paracarnus, 243 Parandraceps, 152
palmi (Thrips), 385 Paracerceis, 33 Paranesidea, 19
Palpada, 215 Parachelifer, 80 Paranocticola, 187
palpebratus† (Tropirhinus), 175 Parachernes, 80 Paraphidippus, 76
Palpita, 326 Parachironomus, 198 Paraphymata, 247
Palpomyia, 197 Parachmidia, 327 paraplacida (Grallipeza), 221
palton‡ (Cytheropteron), 22 Paraclius, 212 Paraplea, 241
paludicola (Rachispoda), 229 Paraclivina, 107 Paraponera, 399

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 636
Parapsyllopsis†, 272 parva† (Borinquena), 235 pavesii (Gyrostigma), 219
Paraputo, 280 parviceps (Tytthus), 244 pavida (Amphiacusta), 374
Pararcte, 354 parvicornis (Agromyza), 224 pawlowskii (Itaquascon), 13
pararemigio (Calliscelio), 283 parvidens (Acropyga), 302 Paxillus, 118
pararufescens (Bradysia), 204 parvior (Docodesmus), 90 payan (Huaca), 169
Parasaissetia, 276 parvipalpus (Zombiweckelia), 30 payasi (Nusalala), 360
parasetosipenis (Euconnus), 116 parvipatellaris† (Argyrodes), 61 paykulli (Plexippus), 76
Parasierola, 309 Parvitermes, 189 Peadus, 329
Parasphaerocera, 229 Parvocyrtusa, 110 peblique (Corythalia), 75
Parastenopa, 223 parvofoveata (Chrysobothris), 125 peckhami (Antillatus), 75
Paratachys, 107 Parvomygale†, 56 peckhami (Compsodecta), 75
Paratemnoides, 80 parvula (Abelona), 377 Peckhamia, 76
paratestaceus (Euconnus), 116 parvula (Brachystomella), 94 pecki (Hypselosomops), 239
Paratettix, 365 parvula (Chalcosicya), 162 pecki (Oosternum), 109
Paratheuma, 69 parvula (Corinna), 71 Peckia, 219
Parathiodina, 76 parvula (Frankliniella), 384 Peckidium, 288
Parathysanoes, 178 parvula (Lupettiana), 71 pectinata (Lepas), 15
Paratissa, 231 parvula (Rhabdomastix), 193 pectinata (Porphyrochroa), 213
Paratrechina, 300 parvulus (Folsomides), 95 Pectinifer, 231
Paratrikona, 160 parvulus (Hemipteroseius), 53 Pectinophora, 321
Paratyndaris, 126 parvulus (Idiocerus), 268 PECTOCYTHERIDAE, 26
Paraulosomus, 171 parvulus (Lathropus), 138 pectorina (Dalotia), 111
paravittata (Scaptomyza), 231 parvulus (Stenotabanus), 207 pedanos (Arphthicarus), 47
Parawithius, 80 parvum (Lasioglossum), 315 pedanus† (Drytotribus), 171
Parawixia, 68 parvus (Homodesmus), 89 pedernalensis (Cephalops), 214
paraxynticos (Hapithus), 371 parvus (Osbornellus), 267 pedernalensis (Erythrolychnia), 130
Parazaona, 80 Parydra, 233 pedernalensis (Lascelina), 329
Pardosa, 68 pascagoulaensis‡ (Cyprideis), 21 pedernales (Anadenobolus), 92
Parectopa, 320 Pasiphaea, 37 pedernales (Beatadesmus), 88
parens (Bolina), 353 PASIPHAEIDAE, 37 pedernales (Dibothrocricus), 92
paretii (Albunea), 39 PASSALIDAE, 118 pedernales (Euchroa), 106
Pareuchaetes, 349 Passalus, 118 pedernales (Phyllophaga), 122
Parhaplopus, 377 PASSANDRIDAE, 138 pedernales (Polycentropus), 387
Parhyale, 30 passiflorae (Corythaica), 242 pedernales (Zeadolopus), 110
Paridris, 284 passus (Lophocyclus), 91 pedernalesensis (Ischnoptera), 186
parilis (Pseudolycoriella), 205 pastinifer (Temnothorax), 302 pedernalesi (Euconnus), 116
parimpicta (Ozophora), 252 Patalene, 334 pedernalis (Lychnacris), 131
Parinesa, 141 patricius (Alaus), 127 pedersoni (Periclimenes), 37
parisiena (Drosophila), 230 Patu, 63 pedestris (Phurys), 352
parksi (Chrysocharis), 293 pauca (Paralimna), 233 pedetes (Walkerana), 369
Parlatoria, 277 paucella (Ethmia), 321 pedicularius (Phloeonomus), 112
Parmenonta, 154 paucidentatus (Callicentrus), 270 PEDICULIDAE, 379
Paroecanthus, 372 paucidentatus (Microtityus), 82 Pediculus, 379
Paromius, 251 paucipalpa (Keilbachia), 205 Pedicythere‡, 23
Paronella, 96 paucispinosa† (Ariadna), 59 pedinus† (Neoulosomus), 171
PARONELLIDAE, 96 Paucivena, 320 Pediobius, 293
Paropsylla†, 272 pauli (Calisto), 343 pediosomus† (Neoulosomus), 171
Paroxacis, 148 paulistus (Eretmocerus), 290 pedophila (Campsiophora), 388
Parribacus, 38 paullus (Hypanartia), 342 pedunculata† (Pulchellaranea), 68
parthenii (Phoenicoprocta), 348 paulmuelleri† (Eohomopterus), 106 pedunculatus (Metaniphargus), 30
Parthenope, 43 paulseni (Hispanioryctes), 123 Pegoscapus, 289
PARTHENOPIDAE, 43 Paululusus, 175 pelaus imerius (Heraclides), 338
parumpunctatus (Selenophorus), 103 paulus (Zeadolopus), 110 pelealis (Lamprosema), 325
parva (Neosconella), 66 paupera (Ormenina), 261 Pelegrina, 76
parva (Ptilomyia), 233 paupi (Tiphia), 308 pelejil (Modisimus), 58
parva (Temnoscheila), 135 pavens (Platynus), 105 pelempito (Barronopsis), 69

637 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Peleteria, 220 Perdita, 314 perpauca (Signiphora), 297
peleusalis (Botys), 326 perdita (Eustala), 66 perpendicularis (Melipotis), 353
Pelia, 42 perditaria (Lobocleta), 335 perpetua (Japanagromyza), 224
Pelidnota, 122 perditor (Thyanta), 257 perpilosus (Agobardus), 74
Pelitropis, 262 Peregrinus, 259 perplexa (Anasaitis), 74
pellax (Xysticus), 74 peregrinus (Robopus), 131 perplexa (Encarsia), 290
pellicia (Cylloceps), 244 peregrinus (Typhlodromalus), 53 perplexus (Araneus), 66
pelliciens (Hapithus), 370 Pereirapis, 315 perplexus (Chlaenius), 102
Pellobunus, 78 perexiguus (Pseudothysanoes), 178 perplexus (Diochus), 117
pellucidellus (Caristanius), 328 perezgelaberti (Compsodes), 187 perplexus (Plesiothrips), 384
Pellucistoma, 22 perezgelaberti (Mesepipsocoides), 380 perseae (Pseudacysta), 243
pelluscens (Fundella), 329 perezi (Euchroa), 106 persearum (Aspidiotus), 277
Pelmatosilpha, 184 perezi (Grallipeza), 221 Persephona, 41
Pelocoris, 240 perezi (Mediocampus), 256 persephone† (Anthophorula), 317
Pelonium, 136 perezi (Melanoplus), 367 persicae (Myzus), 274
Pelonomus, 126 perezi (Neosilba), 222 persicus (Argas), 51
pelops pelops (Antillea), 342 perezi (Sicoderus), 176 personalis (Psychonoctua), 330
peltifer (Ischnothyreus), 59 perezi† (Olbiogaster), 202 personata (Eurydice), 31
PENAEIDAE, 35 perfasciatus (Leptostylopsis), 153 personata (Mycetaspis), 277
Penaeopsis, 35 perforata (Dynamenella), 33 perspectalis (Hymenia), 325
pendator (Chalcis), 291 perforatum (Hoplocheiloma), 221 perspicua (Telphusa), 321
Penestragania, 268 pergandiella (Encarsia), 290 perstrialis (Carectocultus), 327
penetrans (Tunga), 382 pergandii (Parlatoria), 277 persubtilis (Remigia), 354
penicillata (Aristotelia), 321 Pericales, 136 pertinax (Ochlerotatus), 199
penicillata (Baris), 169 Perichares, 336 perturbata (Euphyia), 335
penicillatus (Piestus), 115 Periclimenes, 37 peruanus (Liohippelates), 227
peniculum (Hydrochasma), 231 Pericompsus, 107 peruensis (Rhysotritia), 47
peninsularis (Drosophila), 230 Peridinetus, 170 peruncta (Zale), 355
peninsularis (Haitimera), 77 perieri‡ (Campylocythere?), 27 Peruveleon, 361
peninsularis (Tiburonotus), 365 Perigona, 104 perviana (Botys), 324
penispina (Drosophila), 230 Perigonia, 345 petiolata (Ceratoneura), 294
Pensacolatus†, 76 Perileptus, 107 petioliventris (Conura), 291
pensilis (Selenops), 73 Perileucoptera, 323 petiverii (Acanthonyx), 42
Pentacora, 238 PERIPATIDAE, 13 Petrobia, 50
pentagona (Pseudaulacaspis), 278 Peripatus, 13 Petrochirus, 39
Pentagonica, 104 periphantos (Hapithus), 371 Petrolisthes, 40
Pentalonia, 274 Periplaneta, 184 Petrophila, 323
Pentaneura, 199 PERISCELIDIDAE, 226 petrophila (Eremoleon), 361
Pentanodes, 153 Periscelis, 226 Petrusa, 261
pentaphyllus (Paxillus), 118 perissochaeta (Bolivaridia), 96 Peucetia, 68
pentaspinosa‡ (Rathbunites), 44 Perissocytheridea, 22 pfeifferi† (Pseudosparianthis), 73
Pentatax, 49 Perithemis, 364 PHACOPTERONIDAE, 271
PENTATOMIDAE, 255 peritus† (Incisitermes), 188 Phaedropsis, 326
Penteporus, 90 peritus† (Ophrynopus), 282 Phaenobezzia, 197
Pentobesa, 347 peritus† (Pegoscapus), 289 Phaenonotum, 109
pepino (Agalliopsis), 264 periusalis (Pilemia), 326 Phaenotheriopsis, 165
Peplomicrus, 113 perkinsae idealus (Leptotes), 339 phaeopteralis (Herpetogramma), 325
Pepsis, 307 perlatus (Periclimenes), 37 phakellurata (Acratodes), 335
pequenita (Neotrichia), 389 PERLIDAE, 378 PHALACRIDAE, 138
peracuta peracuta (Gonomyia), 192 perlongisetus (Amblyseius), 53 phalacroides† (Liodema), 146
Perasia, 354 perlucida† (Atopsyche), 390 phalaenodectes (Dichrocheles), 52
peravia (Atopsyche), 390 permagna (Phyllophaga), 122 Phalaenomorpha, 261
peraviae (Saphenista), 332 permista (Steinovelia), 240 PHALANGODIDAE, 78
Percnon, 41 permixtella (Dialectica), 320 PHALANGOPSIDAE, 373
percussa (Peckia), 219 Pero, 334 phaleratus (Ectocyclops), 16
perdiligens (Tricolus), 181 Peronaemis, 125 Phaleria, 147

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 638
phallica (Hydrobiomorpha), 108 Phoebis, 338 Phymata, 247
Phalloniscus, 33 Phoebolampta, 368 phymatophorus (Alcaeorrhynchus), 255
Phamisus, 115 phoenicis (Brevipalpus), 50 Physcius, 149
phanasgalis (Bleptina), 356 Phoenicoprocta, 348 Physemops, 233
Phanerota, 112 Phoenicus, 153 Physocephala, 221
Phanerotoma, 285 Phoetalia, 185 Physoconops, 221
Phanocerus, 126 PHOLCIDAE, 57 Physocyclus, 58
Phaonia, 218 pholcoides† (Styposis), 63 Phytocoris, 244
Phaops, 173 Pholcomma, 62 Phytonemus, 50
pharaonis (Monomorium), 303 Pholcophora, 58 Phytoscutus, 53
pharaxalis (Bocchoropsis), 324 Pholeomyia, 227 PHYTOSEIIDAE, 53
phaselus (Selenops), 73 phorcaria (Leptostales), 335 Phytoseiulus, 53
phaseoli (Chaetostricha), 297 phorcys (Acanalonia), 258 Phytoseius, 53
phaseoli (Polynema), 295 PHORIDAE, 213 picciola (Dioxyna), 222
phasma (Eremoleon), 371 Phormictopus, 56 picea (Alisalia), 111
phasma (Hapithus), 371 phosphoreus (Ignelater), 128 picea (Coptodera), 103
PHASMATIDAE, 377 photinus (Longitarsus), 158 picea (Efferia), 208
phasmatis (Helius), 194 Photis, 30 picea (Perigona), 104
Phatnoma, 241 photon (Pseudoanthonomus), 169 piceoflava (Czernyola), 225
phedrus (Anchastomorphus), 129 phoxi (Anaxipha), 375 piceonotata (Lychnacris), 131
Pheidole, 303 PHOXICHILIDIIDAE, 45 pici (Philornis), 218
Phenacoccus, 280 phratrius (Hemiphileurus), 123 picicorne (Platydema), 146
Pheneps, 127 phrixonota (Chirotica), 287 picinus (Ataenius), 120
PHENGODIDAE, 129 Phrixus, 345 picipennis (Lobopoda), 145
Phereoeca, 320 Phronia, 203 picipes (Ataenius), 120
Pherhimius, 127 PHRUROLITHIDAE, 71 picipes (Dyscinetus), 123
Pheropsophus, 101 Phrurolithus, 71 picipes (Melanolestes), 246
phidilealis (Hellula), 324 phryganoides (Gonatista), 190 picipes (Phaleria), 147
Phidippus, 76 phryganoides (Olyca), 329 picipes (Toxorhynchus), 167
Philacarus, 78 Phrygionis, 334 picta (Phthiacnemia), 253
philea thalestris (Phoebis), 338 PHRYNIDAE, 54 picticornis (Pammeces), 322
Philephedra, 276 Phrynus, 54 picticornis (Stenotabanus), 207
philetes philetes (Perichares), 336 phthanatum† (Nesocyrtosoma), 147 pictipennis (Proceroplatus), 202
Phileurus, 124 Phthia, 253 pictipes (Metacyrba), 76
philipi (Apenes), 103 Phthiacnemia, 253 Picturaphis, 274
philipi (Brasiella), 101 PHTHIRIDAE, 379 picturata (Acinia), 222
philippi (Pericompsus), 107 phthisialis (Mursa), 356 pictus (Chondrotropis), 88
philipsi (Euconnus), 116 phthisicus (Myschocyttarus), 309 pictus (Eumenes), 308
philipsivieorum† (Canthochilum), 121 phthisicus (Stenogryllus), 371 pictus (Lathropus), 138
philocherda (Bactra), 331 Phthitia, 228 pictus (Oncopeltus), 250
PHILODROMIDAE, 73 Phthorimaea, 321 pictus (Physoconops), 221
Philonthus, 117 Phurys, 358 pictus (Toxomerus), 216
Philoponella, 60 Phycitodes, 330 piercei (Leucothrips), 384
PHILOPOTAMIDAE, 386 phylax (Venezillo), 33 PIERIDAE, 338
Philornis, 218 phyleus phyleus (Hylephila), 336 pierreti (Hypocala), 353
PHILOTARSIDAE, 381 phylladiphilus (Agobardus), 74 Piesocorynus, 164
Philothermus, 140 Phyllocnistis, 320 Piestus, 115
PHLAEOTHRIPIDAE, 385 Phyllocoptruta, 49 pieteri (Lycorea), 341
Phlegrata, 76 PHYLLOGNATHOPODIDAE, 18 Pieza, 211
Phloeonomus, 112 Phyllognathopus, 18 Piezobarra, 164
Phloeopora, 112 Phylloicus, 390 Piezodorus, 257
Phloeotribus, 177 Phyllomyza, 227 Piezosternum, 257
Phloeoxena, 104 Phyllophaga, 121 pigmalion bicolora (Phocides), 337
Phlugis, 368 Phyllovates, 190 pikna‡ (Abyssocypris), 26
Phlyctocythere, 25 Phylolestes, 362 pilatus (Apocephalus), 214
Phocides, 337 Phymacysta, 242 Pilemia, 326

639 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Pilica, 209 placida (Systellapha), 221 platypus (Scolopendra), 85
pilicrus (Mesoplia), 317 placida haitiensis (Mezira), 248 PLATYRHACIDAE, 89
pilifer (Nerius), 221 placidus (Antillatus), 75 Platyscytus, 244
pilifer (Paraclius), 212 Placonotus, 138 platysoma† (Geratozygops), 170
piliformis† (Oecobius), 60 Placopsidella, 232 platystegus† (Neoulosomus), 171
piliformis† (Scytodes), 57 Placosternus, 150 Platystethus, 113
piligera (Froeschneria), 251 Plaesius, 109 Platythyrea, 305
pilipes (Nomada), 318 Plagalebra, 268 Platyura, 203
pilosa (Caudellia), 328 plagata (Caribovia), 265 platyurus (Achromoporus), 87
pilosa† (Corythalia), 75 plagata (Ozamia), 329 Platyvelia, 240
pilosalis (Hypena), 357 Plagiodontochirus, 51 plaumanniana (Crovettia), 311
pilosula† (Strumigenys), 302 Plagusia, 41 Plaumannion, 284
pilosulus (Antillocoris), 250 plana (Adelina), 146 plauta‡ (Perissocytheridea), 22
pilosum (Acrepidopterum), 154 plana (Hispanodirphia), 344 plebeia (Empoasca), 269
pilosum (Elaphidion), 151 planaris (Corythaica), 242 plebeia (Pseudoligosita), 297
pilosus (Condylostylus), 212 Planctolpium, 79 plebeja (Erythemis), 363
pilosus (Culex), 200 Planes, 41 plebejana (Crocidosema), 331
pilosus (Metadorylas), 215 planeta† (Anoplius), 307 plebejus (Heraeus), 251
pilosus† (Nasutitermes), 189 planeticos (Nemobiopsis), 373 plebejus (Lachnopus), 174
Pilumnus, 44 Planicephalus, 266 plebius (Eulimnichus), 126
pimana (Cochylis), 332 planicollis (Nyctosyles), 106 Plecia, 202
pimentae (Hemigymnaspis), 277 planicollis (Styloleptus), 153 Plectoptera, 187
Pimpla, 287 planifrons (Lachnopus), 174 PLECTREURIDAE, 60
pinavorus (Pityophthorus), 181 Planinasus, 226 Plectreurys, 60
pinguis (Apogonalia), 264 planistylus† (Aphrastomyia), 202 Plectrocerum, 152
pinguis (Meromacrus), 215 planithorax† (Scytodes), 57 Plectromerus, 150
pini (Aspidiotus), 277 Planococcus, 280 PLEIDAE, 241
pini (Nylanderia), 300 Planodascalia, 261 Pleoticus, 35
pinicola (Neoconocephalus), 368 Planophileurus, 124 Plesiochrysa, 359
pinifoliae (Chionaspis), 278 planulum (Hesperophasma), 377 Plesioglymmius, 108
Pinnaspis, 278 planus (Wilcoxius), 208 Plesiomma, 210
PINNOTHERIDAE, 43 Platamus, 137 Plesiothrips, 384
pinnulifer (Chrysomphalus), 277 PLATASPIDAE, 257 pleuracanthus (Mithrax), 42
Pinophilus, 114 platicephala (Chaetocnema), 157 pleuralis (Craspedochaeta), 225
Pintalia, 259 platirostris (Munidopsis), 39 pleuralis (Gonomyia), 192
Pintaweckelia, 30 platnicki (Centruroides), 81 pleuralis (Hypasclera), 148
Pintomyia, 195 platops (Siphonophora), 91 Pleurambus†, 168
pionaria (Drepanodes), 334 PLATYARTHRIDAE, 33 pleuriticus (Sargus), 206
PIPUNCULIDAE, 214 platycera (Elcaribe), 212 Pleuroprucha, 335
Pirata, 68 platycheirus (Alpheus), 36 Pleuroptya, 326
PISAURIDAE, 68 platycnemis (Buenoa), 241 Plexippus, 76
pisina (Anasa), 253 platycnemis (Canthochilum), 121 plexippus megalippe (Danaus), 341
Pison, 313 Platycorypha, 272 plicata‡ (Gangamocytheridea?), 21
pisonii (Aratus), 41 Platycotis, 270 plicipennis (Chaetocnema), 157
pisorum (Bruchus), 163 platycotis (Ittys), 297 Plinthocoelium, 150
Pissodes, 176 Platycrepidius, 128 plinthopyga (Blaesoxipha), 219
Pissonotus, 259 Platydema, 146 Plocetes, 172
pistola (Alconeura), 269 platyfrons (Neotermes), 188 Plodia, 330
pistrix (Strandesia), 21 Platygaster, 283 plombyi (Camponotus), 300
Pitho, 42 PLATYGASTRIDAE, 283 ploralis (Desmia), 325
pithosalis (Bleptina), 356 Platylambrus, 43 plumatella (Soberocephala), 226
pittfieldae† (Leptofoenus), 296 Platylister, 110 plumbea (Rhagovelia), 240
pittfieldi† (Plectreurys), 60 Platynota, 332 plumbiceps (Paralimna), 233
Pityophthorus, 180 Platynus, 104 plumeriae (Hypothenemus), 178
placens (Arocera), 256 PLATYPEZIDAE, 215 plumicauda (Metaniphargus), 30
placida (Callichirus), 38 Platypus, 182 plumipes (Lachnopus), 174

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 640
plumipes (Peckia), 219 poinari† (Drosophila), 230 Ponsoonops, 59
plumosus (Neoseiulus), 53 poinari† (Dryinus), 311 PONTOCYPRIDIDAE, 26
pluridentatus (Callicentrus), 270 poinari† (Eohomopterus), 106 Pontocypris, 26
plurimella (Mesolia), 323 poinari† (Leptopharsa), 242 poolei (Pseudoeme), 150
plurispinosa (Atheta), 111 poinari† (Probethylus), 311 Popcornella, 76
plurivittatus (Nerius), 221 poinari† (Proceratium), 305 popovi† (Psallops), 245
Plusia, 357 poinari† (Prorhinopsenius), 118 Poratia, 90
Plusiodonta, 3544 poinari† (Scolytus), 177 porcata (Polycesta), 126
Plutella, 322 poinari† (Strumigenys), 302 Porcellana, 40
PLUTELLIDAE, 322 poinari† (Trachelas), 72 PORCELLANIDAE, 40
Pluto, 313 Polemius, 132 Porcellio, 33
pluto (Xylophanes), 346 polhemi† (Microvelia), 314 porcellus (Euscepes), 171
pluviosum (Cylindroxystus), 114 poliochaeta (Helicopsyche), 391 porphyrea (Monomacra), 158
Poaphila, 358 Polistes, 309 Porphyrochroa, 212
Pocadius, 137 polita (Ceratina), 318 porrectispinus (Arvelius), 256
pocsiana (Rotundabaloghia), 54 polita (Cytherella), 21 porrectus† (Veterator), 72
podagrica (Brachymeria), 291 polita‡ (Cytherella), 21 Porrerus, 361
podagricus (Ctenocolum), 163 polita‡ (Pterygocythereis), 27 portoplatensis (Tityus), 82
podagricus (Neoclytus), 150 Polites, 336 portoricensis (Alconeura), 269
podalirius (Microcrambus), 323 politifacies (Pholeomyia), 227 portoricensis (Colliuris), 104
Podiscodesmus, 88 politissimus (Placonotus), 138 portoricensis (Cosmocomoidea), 295
Podisus, 255 politus (Araptus), 180 portoricensis (Crenulaspidiotus), 277
Podium, 313 politus (Cazierius), 83 portoricensis (Emerita), 40
Podochela, 42 politus (Opsablechrus), 136 portoricensis (Encarsia), 290
poecila (Centris), 316 politus (Petrolisthes), 40 portoricensis (Eretmocerus), 290
Poecilanthrax, 211 politus (Toxomerus), 216 portoricensis (Galethalea), 347
Poecilantrops, 228 pollux (Solenopsis), 304 portoricensis (Metaphycus), 292
Poecilognathus, 211 Polyancistrus, 368 portoricensis (Moodnopsis), 329
Poecilopompilus, 308 POLYCENTROPODIDAE, 386 portoricensis (Naushonia), 38
Poecilosomella, 228 Polycentropus, 387 portoricensis (Pardosa), 68
poecilus (Oebalus), 257 Polycesta, 126 portoricensis (Stenochrus), 81
poeyi (Agapostemon), 315 Polycricus, 85 portoricensis (Toxorhynchites), 201
poeyi (Fallceon), 235 Polycyrtus, 287 portorium (Apion), 167
poeyi (Halictus), 315 polydamas polycrates (Battus), 338 portulacae (Neolasioptera), 202
poeyi (Leptostylopsis), 153 Polygonus, 337 PORTUNIDAE, 43
poeyi (Micratya), 37 Polygrammodes, 326 Portunus, 43
poeyi (Orthobelus), 270 polymechanus (Hapithus), 371 portusprospectuensis‡ (Cyprideis), 22
poeyi (Pelocoris), 240 Polymera, 193 positivum (Theridion), 63
poeyi (Platypus), 182 Polymorphomyia, 222 postdenticulata‡ (Cytherella), 21
poeyi (Rhopalosoma), 308 Polynema, 295 postfaciatus (Euscepes), 171
poeyi antillana (Isodontia), 314 Polyortha, 331 postica (Lychnacris), 131
poeyi haitiensis (Polistes), 309 Polypedilum, 198 posticatus (Copelatus), 100
poeyi poeyi (Nannoparce), 344 POLYPHAGIDAE, 187 posticus (Conoderus), 128
Pogonodaptus, 102 Polyphagotarsonemus, 50 posticus (Micromus), 360
Pogonodesmus, 88 polyporus‡ (Balanus), 15 postmajorella (Dioryctria), 329
Pogonomyrmex, 303 Polypsocus, 380 postoculidens (Euscelus), 166
Poinarembia†, 234 polystictus (Alpheus), 36 Potamocypris, 20
poinari† (Acanthognathus), 302 Polythrix, 337 potawatomi (Myllaena), 112
poinari† (Aglyptinus), 111 Polytrichophora, 232 Potimirin, 37
poinari† (Aphelopus), 310 polytrichum† (Nesogerax), 183 potrillo (Cabares), 337
poinari† (Arawakis), 135 Polyvena†, 331 Pozonia, 68
poinari† (Clinidium), 108 pomierense (Amphiacusta), 374 praeceps (Peckia), 219
poinari† (Coenosopsites), 216 pomorum (Bephratelloides), 295 praecipua (Streptopsyche), 388
poinari† (Cubanoptila), 388 POMPILIDAE, 306 praeclarus (Anelaphus), 151
poinari† (Didymia), 282 ponceana‡ (Cytheretta), 22 praeclarus (Macrolophus), 243
poinari† (Dohrniphora), 213 Ponometia, 349 praecognatus† (Episinus), 62

641 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Praecoris†, 246 priscus† (Ceratodacus), 222 progressata (Scotosia), 335
praecreolus† (Temnothorax), 302 priscus† (Coptotermes), 188 projectans (Mycodrosophila), 230
praecursor (Spelaeorhynchus), 54 priscus† (Dryinus), 311 projiciens (Selenisa), 358
praedohmi‡ (Costa), 27 priscus† (Feroseta), 361 prokopi† (Caccoleptus), 133
praedominicana† (Dipoenata), 62 Pristapenesia, 311 prolaminatus† (Dolichoderus), 298
praelonga (Praelongorthezia), 280 pristina† (Corynoptera), 204 prolatus (Cyphotylus), 89
Praelongorthezia, 280 pristina† (Gnamptogenys), 299 Proleucinodes, 326
praelongus (Ommatius), 210 pristina† (Laseola), 62 prolifera (Celaena), 350
praetextatus (Cercyon), 109 pristina† (Plecia), 202 prolixa‡ (Krithe), 24
praeusta (Charidotella), 160 pristina† (Tetragnatha), 65 prolixus (Apocephalus), 214
praevius (Glyphonyx), 129 pristinus (Merotettix), 365 prolixus (Arphthicarus), 47
Prairiana, 268 pristinus† (Dryinus), 311 Promecops, 173
prasinus (Dicyphus), 243 pristinus† (Lyssomanes), 76 prominens† (Stemmops), 63
prasinus (Tropocyclops), 17 pristinus† (Protopaussus), 106 prominuloides† (Brachypogon), 196
pratorum (Meromacrus), 215 prius† (Enicocephalus), 238 prona (Empoasca), 269
pratti (Rhipidia), 194 Proanaxipha†, 373 Proneoproxius†, 248
predaveterus† (Copelatus), 100 Proarna, 263 properans (Anomis), 351
Premnobius, 179 Proaspis, 89 properatos (Hapithus), 370
premordica (Kolourione), 32 proavus† (Pellobunus), 78 propinqua (Agroiconota), 159
prendinii (Microtityus), 82 probaporra (Phyllophaga), 122 propinqua (Heteropsylla), 271
prenolepidis (Myrmecophila), 376 Probaryconus, 284 propinqua (Metadelphax), 259
Prepops, 244 Probethylus, 311 Proplebeia†, 317
prepunctata‡ (Persephona), 41 problematica (Trigonodes), 355 Propontocypris, 20
Presbyolampis, 131 Problepharoneura†, 222 propria (Bulia), 351
pretiosa (Callida), 103 probohemica (Lispe), 218 Proprioseiopsis, 53
pretiosum (Trichogramma), 297 Procaeculus, 48 Proptomaphaginus, 111
pretiosus (Neplas), 215 Procambarus, 38 Prorhinopsenius, 118
preziosii† (Proceroplatus), 202 procax (Ectotrypa), 372 prorogata (Pantographa), 326
pria† (Metatrichia), 211 procellata (Pintalia), 259 prorsoperitrematus (Lasioseius), 52
priapus (Euhyloptera), 260 procera† (Platythyrea), 305 Prosapia, 262
priassalis (Bleptina), 356 Proceratium, 305 prosapia† (Euryconus), 272
priddyi (Eurema), 338 Proceroplatus, 202 Prosaspicera, 289
prima (Ephuta), 306 processa† (Dohrniphora), 213 proscorpio (Mastigoproctus), 83
prima (Pseudhomalopoda), 292 Procladius, 199 proserpina (Neoblattella), 187
primaeva† (Chymomyza), 229 Procoilodes†, 119 Prosierola, 309
primaevus† (Mycetomoellerius), 301 Procolobostema, 196 Prosopodesmus, 89
Primascena†, 271 Procryptotermes, 188 prosper (Stephanoderes), 179
primigenia† (Pheidole), 303 Proctacanthella, 208 prosplatos (Hapithus), 371
primitivus† (Dolichoderus), 298 Proctacanthus, 208 Prostemmiulus, 93
primitivus† (Termes), 190 Proctolaelaps, 52 Protalebra, 268
primus† (Tesserocerus), 182 Procula, 140 Protalebrella, 268
princeps (Heteroctenus), 82 Procythereis, 27 Protalges, 51
Priocnemis, 307 PRODIDOMIDAE, 72 Protambulyx, 344
Priocnessus, 307 prodigalis (Scopula), 327 Protandrena, 314
Prionapteryx, 323 proditata (Thysanopyga), 334 Proteides, 337
Prionocypris, 20 producta (Gonomyia), 192 Protelencholax, 383
Prionopelta, 298 producta (Phloeopora), 112 Proteoconcha, 27
prionura (Bogidiella), 29 producta‡ (Quadracythere), 27 protera† (Didymia), 282
Prionyx, 314 productum (Nesocyrtosoma), 147 proterpia (Pyrisitia), 338
prioris† (Pseudomyrmex), 306 productus (Caligus), 18 proterus† (Happlodontus), 83
prisca† (Corynoptera), 204 Proeces, 170 proteus (Clastoptera), 262
prisca† (Leucochrysa), 359 Proerginus, 78 proteus (Lachnopus), 174
Prisciba†, 262 profecta (Poaphila), 352 proteus (Parlatoria), 278
priscum† (Depressonotum), 229 progenies (Kakopoda), 353 proteus domingo (Urbanus), 337
priscus† (Amnestus), 254 prognatha (Drosophila), 230 Protheca, 134
priscus† (Cnemonyx), 177 Progomphus, 363 Protochymomyza†, 230

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 642
Protodarcia, 320 Pseudisobrachium, 310 pseudosolitarius (Dryocoetoides), 180
Protolepisma†, 97 Pseudleptomastix, 292 Pseudosparianthis, 73
Protoneura, 362 pseudoannulatus† (Antillostenochrus), 81 Pseudosphinx, 345
PROTONEURIDAE, 362 Pseudoanthonomus, 169 PSEUDOSPIROBOLELLIDAE, 93
Protopaussus, 106 Pseudoapseudomorpha, 29 Pseudospirobolellus, 93
Protophthiracarus, 47 PSEUDOARMADILLIDAE, 33 Pseudosquilla, 28
Protopulvinaria, 276 pseudoaztecus (Zacryptocerus), 301 pseudostigma (Eburia), 151
Protortalotrypeta†, 222 pseudobari† (Odontomachus), 305 pseudostriatolateralis (Sicoderus), 176
Protortonia, 279 pseudobrassicae (Rhopalosiphum), 274 Pseudosymploce, 187
Protosinus†, 182 pseudocalcaris (Phyllophaga), 122 pseudotestaceus (Euconnus), 116
Protosphaeroniscus†, 33 Pseudoceratina, 20 Pseudothamiaraea, 112
Prototama, 60 PSEUDOCHIRIDIIDAE, 80 PSEUDOTHELPHUSIDAE, 44
protransvena (Encarsia), 290 Pseudochiridium, 80 Pseudothysanoes, 178
provenzanoi (Pagurus), 39 Pseudochironomus, 198 Pseudoxenos, 383
proxima (Fecelia), 256 pseudochraceus (Enochrus), 108 Pseudozonitis, 148
proxima (Pelegrina), 76 Pseudochrysops, 339 psidii (Pulvinaria), 276
proxima (Siphonophora), 91 Pseudochthonius, 79 Psilanteris, 284
proximus (Cleogonus), 176 PSEUDOCOCCIDAE, 279 Psilochalcis, 291
proximus (Diaptomus), 16 pseudococcina (Hambletonia), 292 Psilonyx, 209
proximus (Laccophilus), 101 Pseudococcus, 280 Psilopa, 233
proximus (Lagynochthonius), 79 pseudocollaris (Ochrimnus), 250 psilus (Rhionaeshna), 363
proximus (Osbornellus), 267 Pseudocycnoides, 18 psittacinus (Artipus), 174
proximus (Pseudomus), 172 PSEUDOCYDNIDAE, 18 psittacus (Pachnaeus), 175
Proxys, 257 Pseudocythere, 20 Psochodesmus, 90
pruinosa (Soberocephala), 226 Pseudodisonycha, 158 PSOCIDAE, 382
pruinosus (Condylostylus), 212 pseudofulvovariegata (Conura), 291 Psocus, 382
pruinosus (Lachnopus), 174 Pseudogaurax, 227 Psomizopelma, 294
pruinosus (Neohybos), 213 Pseudognaptodon, 286 psophodes (Stenogryllus), 371
Prytanes, 251 Pseudolathra, 114 PSOQUILLIDAE, 380
Psalistops, 56 Pseudoligosita, 297 Psorolyma, 140
Psallodia, 263 Pseudolionothus, 110 Psorophora, 201
Psallops, 245 Pseudolycoriella, 205 PSOROPTIDAE, 51
Psara, 326 Pseudolynchia, 217 Psorthaspis, 308
psarocoris (Ozophora), 252 pseudomarginata (Youngszella), 269 PSYCHIDAE, 320
Psathyromyia, 195 Pseudomethoca, 306 Psychoda, 195
Psectrocladius, 199 Pseudomoides†, 171 psychoda (Microissus), 261
PSELLIODIDAE, 84 Pseudomorpha, 108 PSYCHODIDAE, 194
Pseneo, 313 pseudomundus (Selenophorus), 103 Psychonoctua, 330
PSEPHENIDAE, 127 Pseudomus, 172 Psygmophthalmus‡, 44
Psephenops, 127 Pseudomyrmex, 306 Psyllaephagus, 292
pseudacaciae (Tetraleurodes), 273 PSEUDONNANOLENIDAE, 93 PSYLLIDAE, 271
Pseudacteon, 214 Pseudopachybrachius, 251 Psyllipsocus, 380
Pseudacysta, 242 Pseudoparlatoria, 278 Psyllobora, 140
Pseudaonidia, 278 PSEUDOPHASMATIDAE, 378 Psyrassa, 152
Pseudapanteles, 286 Pseudophorellia, 223 Ptecticus, 206
Pseudaphycus, 292 pseudopiceus (Oodinus), 104 Ptenothrix, 96
Pseudaptinus, 102 Pseudopilolabus, 167 Pterocypha, 335
Pseudarachnocephalus†, 376 Pseudoplusia, 357 Pterogramma, 228
Pseudarmadillo, 33 Pseudoponera, 305 Pterogyne, 320
Pseudasopia, 330 Pseudopostega, 319 PTEROLICHIDAE, 51
Pseudatomoscelis, 244 Pseudopothea, 245 Pterolophosoma†, 151
Pseudaulacaspis, 278 Pseudopsammocythere, 22 PTEROMALIDAE, 296
Pseudechiniscus, 13 Pseudopsodexia, 217 Pteronemobius, 373
Pseudectroma, 292 Pseudosciara, 205 pterophoralis (Tomopteryx), 327
Pseudhomalopoda, 292 Pseudosinella, 95 PTEROPHORIDAE, 330
Pseudischnaspis, 278 Pseudosmittia, 199 Pteroptrix, 290

643 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

Pterosclerogibba†, 311 pulchellus (Nallachius), 360 punctatissimus (Pseudomus), 172
Pterygiodesmus, 89 pulchellus (Ochrostomus), 250 punctatus (Anoplischius), 128
Pterygocythereis, 27 pulchellus (Toxomerus), 216 punctatus (Apollophanes), 73
Pthirus, 379 pulcher (Lophoscutus), 247 punctatus (Calocosmus), 155
Ptichodis, 354 pulcher (Microdon), 216 punctatus (Corizus), 254
Ptilodexia, 220 pulcher† (Lyssomanes), 76 punctatus (Curius), 150
Ptilomyia, 233 pulcher† (Succinya), 70 punctatus (Flatoidinus), 261
Ptinus, 135 pulcherrima (Chrysso), 61 puncticeps (Barylaus), 105
Ptychamalia, 335 pulcherrimus (Epipompilus), 307 puncticollis (Amblygnathus), 102
Ptychoderes, 164 pulchra (Nostima), 234 puncticollis (Coenonica), 111
Ptycta, 382 pulchra (Rileya), 295 puncticollis (Curtomerus), 151
puberula (Agonoscelis), 256 pulchra† (Cernotina), 387 puncticollis (Diastolinus), 144
puberulum (Nesanoplium), 151 pulchra† (Phatnoma), 242 puncticosta (Litoprosopus), 353
puberulus (Philothermus), 140 pulchripes (Psilopa), 233 punctifer (Neoplea), 241
pubescens (Antillipeltis), 135 pulchritarsis (Pselliophora), 84 punctifera (Condica), 350
pubescens (Chydorus), 14 pulchrus (Phylloicus), 390 punctifera (Eurocalia), 260
pubescens (Dorynota), 160 Pulex, 382 punctifera (Glutophrissa), 339
pubescens (Euphorticus), 103 pulicaris (Euplatypus), 182 punctipalpus (Enochrus), 108
pubescens (Hypothenemus), 179 pulicarius (Anthonomus), 169 punctipennis (Atractomerus), 169
pubescens (Xyleborus), 180 PULICIDAE, 382 punctipennis (Bezzia), 197
pubiceps (Leptogenys), 305 Puliciphora, 214 punctipes (Geocoris), 249
pubis (Pthirus), 379 pulla (Pseudolycoriella), 205 punctirena (Dypterygia), 350
pudens (Hapithus), 371 Pullimosina, 228 punctulatus (Proxys), 257
pudens (Pityophthorus), 181 pullus (Alaetrinus), 145 pungens (Hypogeococcus), 279
pudibundella (Aristotelia), 321 pullus (Beatadesmus), 88 punicae (Andaspis), 278
pudibundus (Hepatus), 40 pullus (Spirobolellus), 92 puntacanaensis (Leptostylopsis), 153
puella (Paraplea), 241 pullus floridanus (Neobidessus), 100 puquiu (Huaca), 169
puer (Gonomyia), 192 pulverella (Parectopa), 320 pura (Phurys), 358
puera (Hyblaea), 323 pulvereus† (Stenommatus), 173 purchasi (Icerya), 279
puertoplatanus (Opius), 286 pulverius (Ommatius), 210 Purenleon, 361
puertoricense (Ideobisium), 79 Pulvinaria, 276 purgatus (Enicospilus), 288
puertoricensis (Carios), 51 pulvinariae (Diadiplosis), 202 Puriana, 24
puertoricensis (Comanopa), 268 pumila‡ (Perissocytheridea), 22 purisubrhomboidea‡ (Loxoconcha), 25
puertoricensis (Oxyethira), 389 pumilio† (Platythyrea), 305 purpurascens (Copestylum), 215
puertoricensis (Rhinacloa), 245 pumilis† (Stenotrupis), 170 purpurea (Colaspis), 162
puertoricensis (Theoborus), 180 Pumilocytheridea, 22 purpurella (Oedinychis), 157
puertoricensis (Urgleptes), 154 pumilus (Caribochernes), 80 purpureum (Nesocyrtosoma), 147
puertoricensis (Zelus), 246 pumilus (Heteronebo), 83 purpureus (Mastigodiaptomus), 16
puertoricoense (Eosentomon), 96 pumilus (Lophocyclus), 91 purpureus (Rhinotus), 87
pugilator (Uca), 42 pumilus† (Idiogaryops), 79 purpureus (Tylocerus), 132
pugione (Empyreuma), 347 puncta (Gyrophaena), 111 purpuriventris (Chrysis), 310
pugnans (Miogryllus), 372 punctata (Atya), 36 pusilla (Anyphaena), 71
pugnax (Exophthalmus), 173 punctata (Banasa), 256 pusilla (Mesoplophora), 47
pugnax (Oebalus), 257 punctata (Graminella), 266 pusilla (Musca), 220
pugnax (Uca), 42 punctata (Milichiella), 227 pusilla (Ozophora), 252
pulchella (Calisto), 343 punctata (Ormia), 220 pusillulum (Cycloptilum), 376
pulchella (Corinna), 71 punctata (Orphulella), 367 pusillus (Cryptolestes), 138
pulchella (Exomalopsis), 317 punctata (Persephona), 41 pusio (Amnestus), 254
pulchella (Microvelia), 240 punctata (Platythyrea), 305 pusio (Archisepsis), 224
pulchella (Spodoptera), 356 punctata (Rotundabaloghia), 54 pusio (Liohippelates), 227
pulchella darlingtoni (Calisto), 343 punctata (Syphraea), 158 pustulans (Russellaspis), 276
Pulchellaranea†, 68 punctata‡ (Xestoleberis), 28 pustulosa‡ (Puriana), 24
pulchellus (Cephaloplopus), 377 punctaticeps (Leptogenys), 305 puta† (Lasaeola), 62
pulchellus (Coccophagoides), 290 punctatissima (Banasa), 256 Puto, 280
pulchellus (Coproporus), 118 punctatissima (Hypoponera), 305 Pycnarthrum, 179
pulchellus (Lausus), 71 punctatissima (Leptophysa), 158 Pycnoderes, 243

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 644
Pycnomerus, 143 quadrimaculata (Tetraonyx), 148 Quisqueyaaphis†, 273
Pycnoscelus, 186 quadrimaculatus (Heterophotinus), 131 quisqueyae† (Tainosia), 262
Pycnosinga†, 68 quadrinotatus (Heterophotinus), 131 quisqueyana (Trogolaphysa), 96
PYGIDICRANIDAE, 183 quadripapillatus† (Rhipidandrus), 144 quisqueyanum (Strongylium), 147
Pygiopachymerus, 164 quadripilis (Iphiseiodes), 53 quisqueyanus (Apodrosus), 175
pygmaea (Plectoptera), 187 quadripunctata (Paradella), 33 quisqueyanus (Caribanosis), 144
pygmaea (Psorophora), 201 quadrisetosa† (Milichiella), 227 quisqueyanus (Tityus), 83
pygmaea (Solenopsis), 304 quadrisignata (Charidotella), 160 quisqueyensis (Anthonomus), 169
pygmaeus (Enochrus), 108 quadrispinosus (Hylocurus), 178 quisqueyus (Tachysphex), 313
pygmaeus (Piestus), 115 quadristigma (Brontaea), 218 Quisquicia, 88
pygmaeus† (Arachnolithulus), 57 quadristigmalis (Diaphania), 325 quitaespuela (Eurycotis), 184
pygmaeus† (Descangeles), 75 quadistrigmalis (Palpita), 326 quitaespuela (Quisquellacricus), 92
pygmaeus† (Serratochorus), 58 quadristriatus (Prostemmiulus), 93 Quizqueiplana†, 258
PYGMEPHORIDAE, 49 quadrituberculatus (Homophileurus), Rabela, 268
Pygmephorus, 49 124 raburni (Calisto), 343
Pylocheles, 40 quadriundulatus (Micronotus), 365 raccurti (Ataenius), 120
PYLOCHELIDAE, 40 quadrivittatus (Cosmocomoidea), 295 Rachispoda, 228
Pylopagurus, 39 quadrivittatus (Exophthalmus), 173 Radara, 354
Pyractonema, 130 quadrivittatus (Laccophilus), 101 Radaweckelia, 30
Pyragropsis, 183 quadruplex† (Palaeolathys), 70 radeki (Leptoteleia), 284
PYRALIDAE, 327 quantulella (Erelieva), 329 radialis (Amnestus), 254
pyraliformis (Epidromia), 352 quarta (Opacifrons), 228 radiata (Tamarixia), 294
Pyralis, 330 quasicornuta† (Dohrniphora), 213 radicicola (Cirolana), 31
pyraloides (Hemeroplanis), 353 quasiexquisitus† (Cenocephalus), 182 Radimella, 24
Pyrausta, 326 quasimaculata (Plecia), 202 Radionaspis, 278
pyrenealis (Trischistognatha), 323 Quasipirhidius, 143 ragonoti (Deuterolita), 327
Pyrgocorypha, 368 quassa (Azeta), 351 Raillietiella, 16
PYRGODESMIDAE, 89 Quatiella, 195 Rainieria, 221
PYRGOMATIDAE, 16 quatordecimsignatus (Cryptocephalus), rallum (Pelonium), 136
PYRGOMORPHIDAE, 366 161 ramburii (Ischnura), 362
Pyrgus, 337 quatuordecimmaculata (Elytrogona), 160 ramfis (Microscydmus), 116
pyriceps† (Masona), 286 Quedius, 118 ramfisensis (Euconnus), 116
pyriformis (Protopulvinaria), 276 quellita‡ (Kangarina), 23 ramicornis (Dicrepidius), 128
Pyrinia, 334 quesqueya (Canthonella), 121 ramireza (Cophella), 374
pyrirostris (Cossonus), 170 quietalis (Scoparia), 324 ramoni (Platynus), 105
Pyrisitia, 338 Quilessa, 262 Ramosella, 270
pyritosus (Selenophorus), 103 quinculus (Delphastus), 141 ramosi (Allecula), 145
pyro (Pyrisitia), 338 quinquefasciatus (Culex), 200 ramosi (Cicadettana), 263
Pyroderces, 322 quinquemaculata quinquemaculata (Ozo- ramosi (Sicoderus), 176
Pyrophorus, 128 phora), 252 ramphastonis (Stilbometopa), 217
Pyropus, 172 quinquemaculatus (Acanthinus), 149 Ramphocorixa, 241
Pyrrhocalles, 336 quinquenotata (Erythrolychnia), 130 ramulorum (Myrmelachista), 300
PYRRHOCORIDAE, 252 quinquenotatus (Elateropsis), 156 ramulorum vernulus (Camponotus), 300
Quadracythere, 26 quinquepunctata (Leucocera), 161 ramulus (Anadenobolus?), 93
Quadrastichus, 294 quinquepunctatus (Stiretrus), 255 rana (Hamataliwa), 68
quadrata (Ocypode), 42 quinquespinosa (Myropsis), 41 rangelana (Cyclocephala), 123
quadrata (Pepsis), 307 quinteca (Cedusa), 260 rantalis (Achyra), 324
quadratum (Tetragonoschema), 125 quinti† (Dohrniphora), 213 Ranzovius, 244
quadratus (Petrolisthes), 40 quisqueiana (Epilampra), 185 Raoiella, 50
quadricavum (Amblyomma), 52 quisqueiana (Tainacris), 366 rapax (Amblydromalus), 53
quadricollis (Catharthus), 138 Quisquellacricus, 92 rapax (Uca), 42
quadricolor (Condylostylus), 212 quisquellanus‡ (Neocallichirus?), 43 raphanus (Nysius), 250
quadridentata (Hololepta), 109 quisqueya (Cyclocephala), 123 raphiaster (Eurycyde), 45
quadrifoveatus (Strategus), 123 quisqueya (Schistocerca), 367 Raphirhynchus, 168
quadriguttata (Leucocera), 161 quisqueya† (Anoplotermes), 189 rapoporti (Metasinella), 95
quadriguttatus (Tetragonoderus), 102 quisqueya† (Notaepytus), 139 raptor (Megalostrata), 72

645 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

rarior (Anadenobolus?), 93 regificus (Hapithus), 370 resumens (Pachylioides), 345
rasa (Schacontia), 324 regillus (Hapithus), 370 reta (Rugosana), 268
Rasahus, 246 regina (Augochlora), 315 reticulata (Agallia), 263
rasenum (Tapinoma), 299 regina (Cyarda), 260 reticulata (Bradysia), 204
rastnitsyni† (Dryinus), 311 reginae (Polytrichophora), 232 reticulata (Gonatista), 190
rathbunae (Alpheus), 36 reginae† (Electromyrmococcus), 279 reticulata (Trichocorixa), 241
rathbunae (Lysmata), 36 regius (Phidippus), 76 reticulata (Xyphon), 265
Rathbunella‡, 44 regnyi (Leucauge), 65 reticulatus (Agenodesmus), 89
Rathbunites‡, 44 regularis (Polycesta), 126 reticulatus (Iridopagurus), 39
ravida (Aysha), 71 regulator (Culex), 200 reticulatus (Ozius), 44
ravilla (Eriophora), 66 regulus (Antillosorius), 112 retifera (Sphaeropthalma), 306
Ravilops, 83 regulus (Bryoporus), 118 Retiro, 70
Ravinia, 219 rehni (Colapteroblatta), 185 Retithrips, 384
rawlinsi (Apenes), 103 rehni (Eurycotis), 185 retroversa (Lucilia), 217
rawlinsi (Arilophia), 330 rehni (Miogryllus), 372 Reuteroscopus, 245
rawlinsi (Atteva), 323 reichardtiana (Trochosa), 68 reversa (Tannea), 113
rawlinsi (Brasiella), 101 reicheoides (Oxydrepanus), 107 reversa‡ (Krithe), 24
rawlinsi (Calocosmus), 155 reidae (Mesocyclops), 16 rex (Phyllophaga), 122
rawlinsi (Edessa), 255 reini (Microtityus), 82 rex (Phytoseius), 53
rawlinsi (Gypona), 268 Rejanellus, 74 rex† (Tyrannasorus), 119
rawlinsi (Horama), 348 Rekoa, 339 Rhabdodryas, 338
rawlinsi (Leucania), 356 relapsa (Baniana), 351 Rhabdomastix, 193
rawlinsi (Microborus), 179 relicta (Aphaenogaster), 397 rhabdophloeoides (Lathropus), 138
rawlinsi (Ozophora), 252 relicta epinotalis (Aphaenogaster), 303 rhadamanthus† (Philonthus), 118
rawlinsi (Phrygionis), 334 remanens (Remigia), 354 rhaebus (Dilophonota), 345
rawlinsi (Phyllophaga), 122 remigio (Calliscelio), 283 RHAGIONIDAE, 206
rawlinsi (Streptopsyche), 388 remigis (Aquarius), 239 Rhagoletotrypeta, 223
rawlinsi (Trichomyia), 195 Remosa, 262 Rhagovelia, 240
rawlinsi (Zamagiria), 330 remotus (Thaloe), 85 Rhammatocerus, 367
rawlinsoni (Cryptorhopalum), 133 remus (Telenomus), 284 Rhamphomyia, 213
raygilli (Anagrus), 295 rena (Lachesilla), 381 Rhamphothrips, 385
recorta (Phyllophaga), 122 renardii (Zelus), 246 Rhantus, 100
recta (Eublemma), 349 renelauerae† (Heteroctenus), 82 Rhaptinus, 169
recta‡ (Phlyctocythere), 25 rengus (Hemeroblemma), 353 Rhegmoclemina, 196
rectangularis‡ (Caudites), 24 Renia, 334 Rheumatobates, 239
rectangulatus (Aglaocoris), 248 Renocila, 32 Rhinacloa, 245
rectangulum (Trogoxylon), 134 Renodes, 354 Rhinanisus, 170
rectangulus (Loxandrus), 106 repanda (Bagisara), 351 rhinoceroide† (Cenocephalus), 182
recticollis (Glyphonyx), 129 repanda (Mocis), 354 rhinocerontis (Gyrostigma), 219
rectirostris (Stylodactylus), 37 repandana (Platynota), 332 RHINOCRICIDAE, 92
recurva (Tyrissa), 355 repentina† (Bradysiopsis), 204 Rhinoprosopa, 216
recurvalis (Spoladea), 327 repentinus (Ectotrypa), 372 Rhinostomus, 173
Recurvaria, 321 reperta (Ozophora), 252 Rhinotermes, 188
redeuns (Chironomus), 197 repleta (Drosophila), 230 RHINOTERMITIDAE, 188
redonda (Chimarra), 386 reposta† (Tainarys), 271 Rhinotus, 87
reducta (Heteropsylla), 271 reptans (Herpetocypris), 20 Rhinton, 336
reducta (Hogna), 68 republicanus (Tytthonyx), 133 Rhionaeshna, 363
reducta (Panthea), 357 repugnalis (Azeta), 351 Rhipicephalus, 52
reductum (Lobrathium), 114 resinae† (Cephalotes), 301 Rhipidandrus, 144
reductus (Jalysus), 248 resinae† (Chimarra), 386 Rhipidia, 194
REDUVIIDAE, 245 resinae† (Phrynus), 54 Rhizoecus, 280
reflexa† (Krocarites), 268 resinipes (Sphex), 314 Rhizoglyphus, 51
reflexus (Badister), 104 resinocapta† (Setodes), 390 Rhizoperta, 134
reflexus† (Wattius), 145 restituta (Geria), 351 rhizophora (Rachispoda), 229
refracta (Ptichodis), 354 restitutana (Platynota), 332 Rhodesiella, 227
regalis (Tetrabothynus), 174 restorata (Acidalia), 333 rhodocera (Xylophanes), 346

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 646
Rhombognathus, 49 RIPIPTERYGIDAE, 364 rostrata (Neolasioptera), 202
RHOPALIDAE, 254 rithrusaria (Alicia?), 333 rostratus (Pogonodaptus), 102
Rhopalogaster, 209 riveroi (Munidopsis), 39 rostratus† (Miophlugis), 368
Rhopalopterum, 227 rivularis (Amphonyx), 344 rota† (Poinarembia), 234
Rhopalosiphum, 274 robertae† (Acaricoris), 248 rothi (Aspiduchus), 185
Rhopalosoma, 308 roberti (Armases), 41 rothi (Eurycotis), 185
RHOPALOSOMATIDAE, 308 robini (Rhizoglyphus), 51 rothschildi (Ctenolepisma), 97
Rhopalum, 312 Robinsonia, 349 rotunda (Emersonella), 293
rhopheticos (Walkerana), 369 Robopus, 131 Rotundabaloghia, 54
Rhopobota, 331 robusta (Aedmon), 157 rotundalis (Physematia), 326
Rhyacionia, 331 robusta (Apogonalia), 264 rotundata (Antachara), 350
RHYNCHITIDAE, 182 robusta (Brachymeria), 291 rotundatus (Spissistilus), 271
Rhynchitobius, 182 robusta (Exoma), 260 rotundicephalus† (Nasutitermes), 189
RHYNCHOCINETIDAE, 44 robusta (Nerius), 221 rotundiceps (Cryptotermes), 188
Rhyncophoromyia, 214 Robustagramma, 229 rotundicollis (Hypothenemus), 179
Rhyopsocus, 380 robustum (Hydrochasma), 231 rotundifrons (Brepholoxa), 257
Rhyparobia, 185 robustus (Achromoporus), 88 rotundipenne (Elaphidion), 151
RHYPAROCHROMIDAE, 250 robustus (Acridurus), 367 rotunditelson (Arubolana), 31
Rhyparus, 120 robustus (Arganotus), 78 rotundocephala (Rhamphocorixa), 241
Rhypasma, 143 robustus (Calocosmus), 155 rotundostyla (Pseudolycoriella), 205
RHYSCOTIDAE, 33 robustus (Coccotrypes), 179 roumaini (Modisimus), 58
Rhyscotoides, 33 robustus (Pleoticus), 35 Rowlandius, 81
Rhyscotus, 33 robustus† (Toxorhynchus), 167 rozanovi† (Stephanitis), 243
Rhysida, 84 rochina (Ceraeochrysa), 359 rubedinella (Ufa), 330
Rhysocoris, 248 Rocinela, 31 rubellus (Chilaphrodesmus), 89
RHYSODIDAE, 108 Rodolia, 140 rubellus (Haplocyclodesmus), 91
Rhysotritia, 47 roederi (Neodiploclampta), 211 rubens (Beatadesmus), 88
Rhyssomatus, 176 rogatalis (Hellula), 324 rubens (Ceroplastes), 275
rhyticephalus (Ataenius), 120 rogeri (Strumigenys), 302 ruber (Mithrax), 42
Rhytidophthia, 253 Rogeria, 304 ruber† (Cladoctonus), 177
Rhytidoporus, 254 rogersi (Marava), 183 rubescens (Omoteina), 161
rica (Lucilia), 217 rohweri (Trypoxylon), 313 rubetra (Cleogonus), 176
rica (Pero), 334 rolstoni (Tricorythodes), 235 rubicunda (Callopisma), 130
rica (Phaenicia), 217 ROMALEIDAE, 367 rubida (Caribovia), 265
RICHARDIIDAE, 222 romana (Phyllophaga), 122 rubidum (Lathrobium), 114
richmondi (Lepidopa), 39 romanus (Metacynortoides), 78 rubidus (Reduvius), 246
richtersi (Paramacrobiotus), 14 romingeri (Ocabaraja), 115 rubiginosa (Gasteracantha), 66
ricini (Theoborus), 180 romingeri (Oxybelus), 312 rubipectus (Clystea), 347
rickstanleyi (Atimiola), 154 Romphaea, 62 rubra (Pepsis), 307
rickwesti (Antillena), 56 rontaleri† (Puliciphora), 214 rubrella (Carcelia), 220
ricordi (Armases), 41 rosacea (Gonionota), 321 rubricator rubricator (Zicca), 254
riegeli (Lachesilla), 381 rosae (Aulacaspis), 278 rubriclavus (Calliscelio), 283
Riegerochterus†, 241 rosae (Crypticerya), 279 rubricus (Cryptocephalus), 162
riehli (Baconia), 109 rosae (Macrosiphum), 274 rubripennis (Ambulyx), 344
rigens (Cinetorhynchus), 44 rosae† (Komisumena), 73 rubripes? (Metasesarma), 41
Rileya, 295 rosea (Balclutha), 266 rubrocinctus (Selenothrips), 384
rileyi (Convexocoleus), 160 rosea (Echinisca), 15 rubrofasciata (Triatoma), 247
rileyi (Pyroderces), 322 roseicollis (Diomus), 141 rubrolinea (Ozophora), 252
rimolii (Calliasmata), 36 roseimargo (Erythrolychnia), 130 rubropicta (Phthia), 253
rimosus (Cyphomyrmex), 301 roseipes (Exophthalmus), 173 rucilla (Cyrtognatha), 65
rimosus (Hydaticus), 100 rosenbergii (Macrobrachium), 37 rucillum (Bembidion), 107
rimosus flavescens (Cyphomyrmex), 301 roseni† (Procolobostema), 196 ruderata (Stelidota), 137
rimosus molitor (Isognathus), 345 roseomaculata (Dellia), 366 rudimentaria (Pleuroprucha), 335
RIODINIDAE, 340 rosescens (Eublemma), 349 rudis (Limnichites), 126
riparia (Labidura), 183 rostralatus (Belopherus), 168 rudisana (Agrophila), 350
RIPIPHORIDAE, 142 rostrana (Platynota), 332 ruetzleri (Glenanthe), 232

647 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

rufa (Callopisma), 130 runa‡ (Loxoconcha), 25 salarium (Trichapion), 167
rufa (Dasyllis), 208 runyoni (Chrysotus), 212 Salbia, 327
rufa rufa (Ischnoptera), 186 ruricola (Gecarcinus), 41 SALDIDAE, 238
rufescens (Pachycondyla), 305 rusci (Ceroplastes), 275 Saldula, 238
rufescentalis (Nephopteryx), 330 rusellae (Crenulaspidiotus), 277 Salebria, 330
rufiabdominale (Rhopalosiphum), 274 Russellaspis, 276 salebrosa (Cyprideis), 22
ruficaudum (Andrenosoma), 208 russelli (Ommatius), 210 salebrosa‡ (Cyprideis), 22
ruficollis (Pallodes), 137 russoi (Anthicus), 149 salei (Cupiennius), 69
ruficornis (Cerotoma), 159 russoi (Crematogaster), 302 Salia, 357
ruficornis (Pepsis), 307 russoi (Euceropsylla), 271 Salina, 96
rufidens (Conoderus), 128 russoi (Mabra), 326 sallei (Amblycerus), 163
rufidiscalis (Azochis), 324 rustica (Phyllophaga), 122 sallei (Aphodius), 119
rufifrons (Coproica), 228 rustica cubana (Manduca), 344 sallei (Beatadesmus), 88
rufilabris (Chrysoperla), 359 Rutela, 122 sallei (Demotispa), 160
rufinans (Antiblemma), 351 ruthae (Eurycotis), 185 sallei (Haroldiellus), 119
rufinervis (Brachycentrotus), 270 Rutherfordia, 278 sallei (Ischnodemus), 248
rufinervis (Dythemis), 363 rutilans (Tetracha), 101 sallei (Temnothorax), 302
rufipennis (Megachile), 316 rutilus (Coproporus), 118 sallei (Tricorynus), 135
rufipes (Mesoplia), 317 rutilus (Diplochaetus), 107 sallei (Trientoma), 144
rufipes (Necrobia), 136 ryani (Hemiphileurus), 123 sallei (Xerolinus), 144
rufipes (Neomegalotomus), 253 Rymosia, 203 sallei opacinoda (Macromischa), 302
rufipes (Nesticodes), 62 ryticephala (Trientoma), 144 Salmoneus, 36
rufipes (Sphex), 314 sabalella (Homaledra), 322 salmus (Ophiusa), 354
rufithorax (Macroteleia), 284 sabalis (Comstockiella), 278 SALPINGIDAE, 149
rufithorax (Silis), 132 sabana (Chrysometa), 65 salpiscinalis (Cyathura), 31
rufiventre (Silidiscodon), 132 Sabelo, 369 SALTICIDAE, 74
rufocaerulea (Lychnacris), 131 Sabethes, 201 saltuella (Balclutha), 266
rufoclarus (Ardistomis), 106 sabroskyi† (Stenomicra), 226 salutator (Antillosorius), 112
rufofascialis (Mimoschimia), 324 Sabulodes, 334 samana (Calisto), 343
rufoflavus (Opius), 286 sabulosa (Epilampra), 185 samana (Eurycotis), 185
rufomaculatus (Sennius), 164 sabulosa (Hypoconcha), 40 samana (Tainonia), 58
rufoviolacea (Callopisma), 130 saccharalis (Diatraea), 323 samanae (Uhleroides), 263
rufulella (Davara), 328 sacchari (Aspidiella), 277 samanense (Hapithus), 371
rufulus (Apalonychus), 119 sacchari (Melanaphis), 274 samanensis (Metacrangonyx), 30
rufus (Oxydrepanus), 107 sacchari (Saccharicoccus), 280 Sambomorpha, 124
rugata‡ (Perissocytheridea), 22 Saccharicoccus, 280 Samea, 327
rugiceps (Caccodes), 132 saccharina (Lepisma), 97 SAMOIDAE, 77
rugicollis (Brasa), 264 saccharinus (Nicentrus), 170 sanborni (Macrosiphoniella), 274
rugicollis (Cardiastethus), 245 saccharivora (Saccharosydne), 260 sanchezensis (Lasioseius), 52
rugicollis (Minthea), 134 Saccharosydne, 260 sanctaecrucis (Dialectica), 320
rugifer (Gyrinus), 99 Sacculobates, 46 Sanctanus, 266
rugifrons (Trientoma), 144 sacer‡ (Caudites), 24 sanctidomingi (Ululodes), 358
Rugilus, 114 saepis† (Dohrniphora), 213 sanctidomini (Pelidnota), 122
rugimanus (Pachycheles), 40 safranota (Omphale), 293 sandae (Tiphia), 308
ruginodis (Odontomachus), 305 sagarus (Osbornellus), 267 sandbergi‡ (Pumilocytheridea), 22
rugosa (Macroteleia), 284 sagitta (Podisus), 255 sandersi (Proprioseiopsis), 53
rugosa (Solenoptera), 156 sagittifera (Xyphon), 265 Sandersonella, 115
rugosa‡ (Loxoconcha), 25 sagra (Eupyrrhoglossum), 345 sandersoni (Lachesilla), 381
Rugosana, 268 sagrae (Scolopendra), 85 sandersoni (Lobopoda), 145
rugosus (Rhysocoris), 248 Saida, 26 sandersoni (Toxorhynchus), 167
rugulopygus (Hispanioryctes), 123 SAIDIDAE, 26 sandersoni (Urgleptes), 154
ruibali (Enthocythere), 23 saileri (Paraphymata), 247 Sandomingia‡, 42
ruidipunctus† (Caulophilus), 171 Saissetia, 276 sanguinarius sanguinarius (Dysdercus),
rulomus (Carpelimus), 113 sajoma (Hapithus), 371 252
rulomus (Stilosaurus), 115 salacia salacia (Melete), 339 sanguinea limbifer (Cycloneda), 140
rulomus (Xantholinus), 118 salaco† (Oeclidius), 261 sanguineus (Rhipicephalus), 52

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 648
sanguinicollis (Chalepus), 160 Scapsipedus, 372 schoenherrii (Belopherus), 168
sanguinipennis (Phoenicus), 153 Scaptomyza, 231 schrammi (Tityus), 83
sanguinipes (Melanoplus), 367 Scaptosciara, 205 schrankiae (Acanthoscelides), 163
sanguinipes (Protoneura), 362 scapularis (Ochlerotatus), 199 schultzei (Frankliniella), 384
santachloe (Phyllophaga), 122 SCARABAEIDAE, 119 schusteri (Metophthalmus), 142
santaclara (Cedusa), 260 Scarites, 107 schwartzi (Calisto), 343
sanum (Cycloptilum), 376 Scatella, 233 schwartzi (Choranthus), 336
Saphenista, 332 Scathophaga, 220 schwarzi (Amblycerus), 163
Saphirella, 18 SCATHOPHAGIDAE, 220 schwarzi (Phloeoxena), 104
sapidus (Callinectes), 43 scatophaga (Gymnodia), 218 schwarzi (Psyllobora), 140
Sapintus, 149 Scatopse, 196 schwarzi (Rhipidia), 194
sapotae (Conotrachelus), 176 SCATOPSIDAE, 195 SCIARIDAE, 203
sapphirinus (Pyropus), 172 scaurus (Prostemmiulus), 93 scinax (Paroecanthus), 372
sarasini (Ablechroides), 136 Scelianoma, 174 scindens (Isogona), 353
Sarcodexia, 219 Scelio, 284 scintilla (Lychnacris), 131
Sarcofahrtiopsis, 219 SCELIONIDAE, 283 Sciocharis, 114
Sarcophaga, 219 Sceliphron, 314 SCIOMYZIDAE, 224
SARCOPHAGIDAE, 219 Sceloenopla, 161 Sciophila, 203
Sarcoptes, 51 SCENOPINIDAE, 211 Scirtothrips, 385
SARCOPTIDAE, 51 schach (Aspidoglossa), 106 scissa† (Corythalia), 75
Sargus, 206 Schacontia, 324 scissalis (Hypena), 357
Sarifer, 155 schaefferi (Cicindela), 101 scita (Poaphila), 358
Sarinda, 76 schaffneri (Rhinacloa), 245 scitula (Melanophthalma), 142
sarmenticola (Habroxenus), 164 schawallfussi† (Borinquena), 235 scitulus (Acanthinus), 149
SARSIELLIDAE, 19 SCHENDYLIDAE, 86 scitulus (Beatadesmus), 88
sata (Curtara), 268 scheveni† (Boreofairchildia), 194 scitus (Dallasiellus), 254
Sathria, 327 scheveni† (Harpactosphecion), 311 Sclerochilus, 20
sativae (Liriomyza), 225 scheveni† (Mezira), 248 Sclerocoelus, 229
saturata (Thermesia), 354 scheveni† (Succinarisemus), 195 SCLEROGIBBIDAE, 311
SATURNIIDAE, 344 schildi (Nostima), 234 SCLEROPACTIDAE, 33
saucius (Pogonomyrmex), 303 Schildia, 209 SCLEROSOMATIDAE, 78
saukros (Hapithus), 370 Schistocerca, 367 scobifera scobifera (Apenes), 103
saundersi‡ (Munseyella), 26 Schizaphis, 274 scobina (Spelaeala), 369
sauroides (Lantanacoccus), 279 Schizogenius, 107 scobinellus (Anadenobolus?), 93
saussurei (Camponotus), 299 Schizopera, 18 SCOLEBYTHIDAE, 311
saussurei (Trypoxylon), 313 Schizophragma, 295 SCOLIIDAE, 308
sayana (Porcellana), 40 Schizoptera, 239 scolopaiae scolopaiae (Epimecis), 333
sayi (Calosoma), 101 SCHIZOPTERIDAE, 238 scolopaiae transitaria (Epimecis), 333
sayi (Dicromantispa), 361 Schizuroides, 346 Scolopendra, 84
sayi (Pilumnus), 44 schleei† (Protelencholax), 383 SCOLOPENDRIDAE, 84
sayi (Portunus), 43 schleei† (Psathyromyia), 195 SCOLOPOCRYPTOPIDAE, 85
Sayomyia, 199 schleeorum† (Strumigenys), 302 Scolopocryptops, 85
scabiei (Sarcoptes), 51 schmitti (Azygopleon), 32 Scolytocis, 142
scabra (Atya), 36 schmitti (Gnamptogenys), 299 Scolytodes, 179
scabrellus (Ataenius), 120 schmitti (Nannosquilla), 28 Scolytogenes, 179
scabricauda (Lysiosquilla), 28 schmitti (Pallenopsis), 45 Scolytus, 177
scabrosum (Nesocyrtosoma), 147 schmitti (Pogonomyrmex), 303 Scopaeus, 114
scalaris (Lestes), 362 schmitti aterrimus (Pogonomyrmex), 303 Scopelobates, 56
Scalaventer, 175 schmitti darlingtoni (Pogonomyrmex), Scopoides, 72
Scalenarthrus, 116 303 Scopula, 335
scaloida† (Helicopsyche), 391 schmitti sublaevigatus (Pogonomyrmex), scopulata (Alydda), 333
scalprum† (Platythyrea), 305 303 scopulepes (Hemeroplanis), 353
scambosomus† (Neoulosomus), 171 schmittii (Litopenaeus), 35 scopulina (Oxyethira), 389
scambus† (Neoulosomus), 171 schoenachii (Neotheronia), 287 scorbutica (Anasa), 253
Scapanea, 364 schoenherri (Sicoderus), 176 scorpio (Anotylus), 113
Scaphioides, 59 schoenherri (Tyloderma), 172 SCORPIONIDAE, 83

649 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

scottae‡ (Neocaudites), 24 seibo (Ciba), 69 sennae sennae (Phoebis), 338
Scraptia, 142 seicherti† (Neorhipidius), 143 Sennius, 164
SCRAPTIIDAE, 142 seini (Holopothrips), 385 senticosus (Pityophthorus), 181
scriptellus (Platynus), 105 Seira, 96 separata† (Meioneta), 64
scriptura (Speocropia), 350 SEIRIDAE, 96 separatus (Icius), 76
scriptus (Platynus), 105 Seirocastnia, 350 Sepedomerus, 224
scrobiculatus (Euscelus), 166 seldeni† (Stenoonops), 60 Sepedophilus, 118
scrupulosa (Teleonemia), 243 selector (Catorhintha), 254 SEPSIDAE, 224
scrupulosa haytiensis (Teleonemia), 243 Selenaspidus, 278 septempunctata (Shannonomyia), 193
scrupulosa‡ (Puriana), 24 Selenisa, 358 septentrionalis (Tityus), 82
scrutator (Rhinostomus), 173 Selenophorus, 102 septifera (Helicopsyche), 391
scudderi (Orphulella), 367 SELENOPIDAE, 72 septima (Triacanthagyna), 363
sculptilis (Hispanioryctes), 123 Selenops, 72 septiporus (Henicomus), 90
sculptus (Mithrax), 42 SELENORIBATIDAE, 48 serenalis (Megaphysa), 327
sculptus (Oxytelus), 113 Selenothrips, 384 serenus (Progomphus), 363
sculptus‡ (Lobonotus), 44 Selenyphantes, 64 Sergiolus, 72
scutellaris (Ataenius), 120 sella (Pero), 334 serialis serialis (Schistocerca), 367
scutellaris (Coccophagus), 290 sellata (Kapogea), 67 seriata (Pseudatomoscelis), 244
scutellata (Solenoptera), 156 selleanus (Bradycellus), 102 seriatus (Cryptocarenus), 178
scutellatus (Euscelus), 166 sellensis (Platynus), 105 seriatus (Hypothenemus), 179
scutellatus (Hemiphileurus), 124 sellnicki (Macrodinychus), 51 sericata (Gnaphosa), 72
scutellatus (Nysius), 250 Semaeomyia, 288 sericata (Lucilia), 217
scutellatus (Stenopus), 35 Semaeopus, 335 sericatus (Anaulacus), 102
SCUTELLERIDAE, 258 SEMATURIDAE, 336 sericatus (Conoderus), 128
Scutellista, 296 Semiardistomis, 107 sericea (Dendrobiella), 134
scuticollis (Silvanoprus), 138 semiaurata (Pereirapis), 315 sericea (Eburia), 151
scutiformis (Antillatus), 75 semicaeca† (Erucoblatta), 187 sericeiventris (Elateropsis), 156
Scutigerella, 94 semicaerulea (Plagiodera), 161 sericeum (Callipogon), 155
SCUTIGERELLIDAE, 94 Semiclivina, 107 sericeus (Paguristes), 39
SCUTIGERIDAE, 84 semicribratus (Pseudomus), 172 sericeus (Scopelobates), 56
SCUTOVERTICIDAE, 48 Semicytherura, 23 Sericoceros, 282
Scydmaenus, 117 Semidalis, 360 Sericothrips, 385
Scylla, 38 semidecora (Acidalia), 335 serina zetes (Dynamine), 340
SCYLLARIDAE, 38 semiferox (Gnamptogenys), 299 serius (Eretmocerus), 290
Scyllarides, 38 semifurca (Celaena), 350 serpentifera (Eudocima), 352
Scymnus, 141 semilimbatus (Oncopeltus), 250 serpentinus (Conotrachelus), 176
Scyphophorus, 177 semilutea (Paratissa), 231 serra (Ochrotrichia), 389
Scytodes, 57 semimarginatus (Calocosmus), 155 serrallesi (Ocrepeira), 67
SCYTODIDAE, 57 seminudus† (Nasutitermes), 189 serranus (Jaragua), 366
seabrai (Sarifer), 155 semiopaca (Geotrupes), 119 serrata (Apoweckelia), 29
sebae (Achelous), 43 Semiothisa, 334 serrata (Dunhevedia), 14
secernalis (Udea), 327 semiplumosa (Philoponella), 60 serrata (Eperigone), 64
secluda (Rhagovelia), 240 semirufus (Bombylius), 211 serrata (Kramerolidia), 265
secunda (Lithocharis), 114 semirugosa† (Oligochlora), 315 serrata (Parthenope), 43
secundus (Apocephalus), 214 semiseptus (Docodesmus), 90 serrata (Penaeopsis), 35
securigerus (Pseudothysanoes), 178 semisulcatus (Acilius), 100 serrata† (Prisciba), 262
secutor (Culex), 200 Semnorhynchus, 172 serraticrus† (Lachnopus), 174
sedentarius (Pirata), 68 semnos (Hapithus), 371 Serratochorus, 58
sedula (Megachile), 316 sempiternus dilloni (Anastrus), 337 serratum (Nesocyrtosoma), 147
seeversi† (Prorhinopsenius), 118 senectum† (Pycnarthrum), 179 serratus (Caryedon), 163
SEGESTRIIDAE, 59 senectus† (Cenocephalus), 182 serratus (Harmostes), 254
segmentata (Vanduzea), 271 senescens (Leucania), 356 serratus (Plectromerus), 150
segmentatus† (Nops), 59 senicula† (Neomida), 146 serratus† (Cephalotes), 301
segmentidentatus (Cacodemonius), 80 seniculus† (Enicocephalus), 238 serrei (Jonga), 37
seguin (Modisimus), 58 senilis (Chalcosicya), 162 serricorne (Lasioderma), 135
seiba (Ochrotrichia), 389 senilis (Geron), 211 serricornis (Daimon), 270

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 650
serricornis (Tricholabus), 287 SICARIIDAE, 57 simillima (Centris), 317
serripes (Tainonia), 58 sicca (Corynoptera), 204 simillima (Euhyloptera), 260
serriradiellus (Crambus), 327 Sicoderus, 176 simillimum (Tetramorium), 304
serrula (Efferia), 208 sidae (Calycomyza), 225 simoni (Drymusa), 57
serrulata‡ (Orionina), 24 sidae (Niesthrea), 254 simplex (Bythinogaster), 115
serrulatus (Eucyclops), 16 side (Conura), 291 simplex (Dolichowithius), 80
serrulatus oreiotes (Polyancistrus), 368 siderea (Atteva), 323 simplex (Ellipodesmus), 88
serrulatus serrulatus (Polyancistrus), 368 Siderone, 341 simplex (Miathyria), 364
servillei (Acanalonia), 258 sideroxyli (Ceropsylla), 272 simplex (Nesocyrtosoma), 147
servillei (Hyptia), 288 SIDIDAE, 14 simplex (Opogona), 320
servillei (Sphex), 314 Sierratettix, 365 simplex (Pseudomyrmex), 306
servus (Hister), 109 sigillata (Thermesia), 352 simplex (Singhiella), 273
setadebilia (Aptunga), 328 sigillatus (Gryllodes), 372 simplex (Spalangia), 296
setariae (Hysteroneura), 274 sigillatus (Spirobolellus), 92 simplex (Thrips), 385
Sethenira, 254 signata (Rocinela), 31 simplex (Tricharaea), 220
seticornis (Stenorhynchus), 42 signata (Stictia), 312 simplex (Xanthandrus), 216
setiger (Pteronemobius), 373 signaticolle (Trichophoroides), 152 simplicicollis (Erythemis), 363
setiger (Stenoxenus), 197 signatura (Calolampra), 185 simplicula (Microsepsis), 224
setiger (Trematodesmus), 89 signatus (Lasiomerus), 245 simulacrifer (Plocetes), 172
Setodes, 391 signatus (Peridinetus), 170 simulans (Drosophila), 230
Setomima, 195 signatus (Spirobolellus), 92 simulatrix (Oxyethira), 389
setosa (Calosota), 294 signifer (Canthon), 121 simulatus (Euwallacea), 180
setosa (Cophella), 374 signifera (Orthopodomyia), 200 SIMULIIDAE, 201
setosa (Neostasina), 73 signifera (Prodenia), 356 Simulium, 201
setosella (Chalcosicya), 162 signiferana (Agrophila), 349 sinclairi (Pieza), 211
setosipenis (Euconnus), 116 significans (Baniana), 351 Singhiella, 273
setosus (Haplocyclodesmus), 91 Signiphora, 296 singularis (Caccodes), 132
setosus (Hypothenemus), 179 SIGNIPHORIDAE, 296 singularis (Dissomphalus), 310
setosus (Neohybos), 213 signoreti (Pentacora), 238 singularis (Fennahiella), 243
setosus (Prostemmiulus), 94 sigsbeiana (Porcellana), 40 singularis insolata (Euphyes), 336
setosus† (Microcerotermes), 190 sikesi (Canthonella), 121 singularis† (Palaeodyctina?), 69
setulosa (Coproica), 228 silbermanni (Chalcolepidius), 127 sinuatus (Cryptophagus), 139
setulosa (Polytrichophora), 232 silendum (Trichapion), 167 sinuatus (Selenophorus), 103
sewelli (Elaphoidella), 17 silhavyi† (Hummelinckiolus), 77 sinuosa (Efferia), 208
sexguttatus (Camponotus), 299 silicalis (Pleuroptya), 326 Sipha, 273
sexmaculata (Hermetia), 206 Silidiscodon, 132 Sipho, 369
sexmaculatum (Copestylum), 215 Silis, 132 Siphonocybe, 91
sexoculata (Masteria), 56 SILPHIDAE, 111 Siphonophora, 91
sexpilis (Phytoscutus), 53 SILVANIDAE, 137 SIPHONOPHORIDAE, 91
sexsetosa (Meropliosepsis), 224 Silvanoprus, 138 SIPHONOTIDAE, 87
sexspinosa (Dohrniphora), 213 silvestrii (Strumigenys), 302 Siphunculina, 227
sexta paphus (Manduca), 344 simaethis (Chlorostrymon), 339 Siproeta, 342
shakespearei (Goetheana), 293 simialis (Omiodes), 326 Sirex, 282
Shannonomyia, 193 simile (Plesiomma), 210 SIRICIDAE, 282
sharpi (Lapethus), 140 similis (Cyphoderus), 95 Siriella, 29
sharpi (Notomicrus), 100 similis (Drosophila), 230 siro (Acarus), 51
sharpi (Oosternum), 109 similis (Exomalopsis), 317 Sirthenea, 246
sheldoni (Aceria), 48 similis (Halicyclops), 16 sisalanus (Phenacoccus), 280
sheylae (Dominicophus), 374 similis (Hapithus), 370 sisyphoides (Tidarren), 63
SIALIDAE, 358 similis (Hortensia), 265 sisyphus (Anthonomus), 169
Sialis, 358 similis (Leptopsilopa), 232 sitellaria (Racheospila), 334
sibelius (Euseius), 53 similis (Micrathena), 67 Sitobium, 274
sibillalis sibillalis (Glyphodes), 325 similis (Plagalebra), 268 Sitophilus, 177
Sibinia, 172 similis (Streptocephalus), 14 Sitotroga, 321
sibylleae (Euconnus), 116 similis† (Palaeolathys), 70 skelleyi (Nesocyrtosoma), 147
sibyllensis (Euconnus), 116 similis‡ (Cyprideis), 22 skelly (Spintharus), 62

651 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

skinneri (Cressonomyia), 232 sororcula (Wyeomyia), 201 sp. (Einfeldia), 198
slipinskii (Catogenus), 138 sororius (Eurhabdus), 209 sp. (Elasmus), 294
slosonella (Hypargyria), 329 sorpresus (Cyclargus), 339 sp. (Elpidium), 24
slossonae (Lipochaeta), 232 soteriodes (Cophella), 374 sp. (Eremochrysa), 359
slossonae (Nostima), 234 sotiusalis (Mursa), 356 sp. (Euaresta), 223
slossonae (Notodelphax), 259 soulouquii (Omalodes), 109 sp. (Euderus), 293
smaragdula (Colaspis), 162 soulouquina (Somomyia), 217 sp. (Eugeonemus), 174
SMARIDIDAE, 49 sp. (Acallepitrix), 156 sp. (Eukoenenia), 78
Smaris, 49 sp. (Ageniaspis), 291 sp. (Eusicyonia), 35
smetanai (Dryinus), 311 sp. (Allotropa), 283 sp. (Exomalopsis), 317
Smicridea, 387 sp. (Amorbia), 332 sp. (Fageicera), 57
SMICRIPIDAE, 137 sp. (Anaplecta), 186 sp. (Fidiobia), 283
Smicrips, 137 sp. (Anaulacomera), 368 sp. (Franklinothrips), 384
SMINTHURIDAE, 96 sp. (Anopsilana), 31 sp. (Glyptapanteles), 286
smithi (Metapenaeopsis), 35 sp. (Antillocladius), 198 sp. (Gonomyia), 192
smithi (Phlepsius), 267 sp. (Argon), 336 sp. (Habrobracon), 285
smithi (Thinodromus), 113 sp. (Arrenurus), 49 sp. (Helops), 144
smithi (Ululodes), 358 sp. (Artemia), 14 sp. (Heterotermes), 188
smithiana (Strepsicrates), 332 sp. (Asellus), 32 sp. (Holopothrips), 385
smithii (Mycocepurus), 301 sp. (Atoniomyia), 208 sp. (Hoplandria), 111
smithsoni (Pentobesa), 347 sp. (Atopsyche), 390 sp. (Hylobius), 176
Smodicum, 152 sp. (Augochlora), 315 sp. (Isotomurus), 95
Sobarocephala, 226 sp. (Bolostromus), 56 sp. (Kunzeana), 269
sobrina infuscata (Tetracha), 101 sp. (Bolothrips), 385 sp. (Lamponius), 377
socialis (Uranotaenia), 201 sp. (Brachycarpus), 37 sp. (Lepidocyrtus), 95
sofiae (Bonfilsus), 157 sp. (Brevipalpus), 50 sp. (Lepidopa), 39
Sogatella, 260 sp. (Bryocamptus), 17 sp. (Leptus), 49
sokolowskii (Oomyzus), 294 sp. (Bryocyclops), 16 sp. (Leucothoe), 30
sola (Cerotainia), 208 sp. (Caenis), 235 sp. (Leurolophus), 271
solani (Aulacorthum), 274 sp. (Camillina), 72 sp. (Lirceus), 32
solegladi (Microtityus), 82 sp. (Camptopteroides), 295 sp. (Liriomyza), 225
Solenocera, 35 sp. (Campylothorax), 95 sp. (Litoblatta), 186
SOLENOCERIDAE, 35 sp. (Cavariella), 274 sp. (Lophoblatta), 187
Solenopsis, 304 sp. (Cephalops), 214 sp. (Lutzomiops?), 201
solenopsis (Phenacoccus), 280 sp. (Characoma), 357 sp. (Lyroda), 312
Solenoptera, 156 sp. (Chironomus), 197 sp. (Macrobiotus), 13
solida (Amblystira), 242 sp. (Chthamalus), 16 sp. (Macronema), 387
solitaria (Acidalia), 335 sp. (Cirrospilus), 294 sp. (Macrothrix), 15
solitaria (Megachile), 316 sp. (Cobanus), 75 sp. (Masteria), 56
solitariceps (Theoborus), 180 sp. (Compsodes), 187 sp. (Mazax), 72
solitarius (Ceraticelus), 64 sp. (Copidosoma), 291 sp. (Meleoma), 359
solitarius (Lembodes), 171 sp. (Corethrella), 199 sp. (Melissodes), 317
sollicitans (Ochlerotatus), 200 sp. (Corinna), 71 sp. (Mesodixa), 201
solo (Boettcheria), 219 sp. (Cothonaspis), 289 sp. (Metopina), 214
solus (Homoeotarsus), 114 sp. (Creophilus), 117 sp. (Metrichia), 389
sondrae (Enicospilus), 288 sp. (Cryptobables), 328 sp. (Metriocnemus), 198
sonora (Graminella), 266 sp. (Cryptocephalus), 162 sp. (Microchironomus), 198
sonroja (Leucania), 356 sp. (Cryptopygus), 95 sp. (Microdipoena), 64
sophia (Encarsia), 290 sp. (Cubanochirus), 51 sp. (Micromus), 360
sordida (Hoplistocelis), 245 sp. (Cybocephalus), 137 sp. (Micropsephodes), 141
sordida (Lispe), 218 sp. (Darwinula), 23 sp. (Moinodaphnia), 15
sordida (Olfersia), 217 sp. (Dasiops), 222 sp. (Molosoma), 112
sordidus (Cosmopolites), 177 sp. (Deopalpus), 220 sp. (Monodella), 29
sorghicola (Stenodiplosis), 202 sp. (Dichomeris), 321 sp. (Monopelopia), 199
sorghiella (Nola), 357 sp. (Diglyphus), 293 sp. (Mysidium), 29
soriae† (Euryconus), 272 sp. (Dixella), 201 sp. (Nacarina), 359

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sp. (Neocenchrea), 260 sp. (Syngamia), 327 sp. 1‡ (Xestoleberis), 28
sp. (Neochrysocharis), 293 sp. (Talanus), 147 sp. 2 (Cariblatta), 186
sp. (Neohannia), 69 sp. (Tanytarsus), 198 sp. 2 (Colapteroblatta), 185
sp. (Neopanope?), 44 sp. (Temelucha), 287 sp. 2 (Cricotopus), 198
sp. (Neosmerinthothrips), 385 sp. (Thienemannimyia), 199 sp. 2 (Darwinula), 23
sp. (Neotrichia), 389 sp. (Tibellus), 73 sp. 2 (Elpidium), 24
sp. (Oithona), 17 sp. (Trybomia), 385 sp. 2 (Epilampra), 185
sp. (Oligota), 112 sp. (Uloma), 145 sp. 2 (Eurycotis), 184
sp. (Ollarianus), 266 sp. (Uropelma), 294 sp. 2 (Euthlastoblatta), 187
sp. (Omolicna), 260 sp. (Utetes), 286 sp. 2 (Goeldichironomus), 198
sp. (Ophion), 288 sp. (Xenylla), 94 sp. 2 (Heraeus), 251
sp. (Ormenana), 261 sp. (Xestochironomus), 198 sp. 2 (Labrundinia), 199
sp. (Ormenaria), 261 sp. (Xiphocentron), 387 sp. 2 (Megaris), 255
sp. (Ormyrus), 296 sp. (Zelotes), 72 sp. 2 (Melanagromyza), 224
sp. (Orthocladius), 198 sp. “A” (Ceranisus), 293 sp. 2 (Orthocladiinae), 198
sp. (Oxyethira), 389 sp. 1 (Cariblatta), 186 sp. 2 (Neoblattella), 187
sp. (Paracloeodes), 235 sp. 1 (Colapteroblatta), 185 sp. 2 (Nesomylacris), 186
sp. (Paracyclops), 17 sp. 1 (Compterosmittia), 198 sp. 2 (Ophiomyia), 225
sp. (Parametriocnemus), 199 sp. 1 (Cricotopus), 198 sp. 2 (Paranocticola), 187
sp. (Parastenopa), 223 sp. 1 (Epilampra), 185 sp. 2 (Pentaneura), 199
sp. (Paronella), 96 sp. 1 (Euthlastoblatta), 187 sp. 2 (Polypedilum), 198
sp. (Patu), 63 sp. 1 (Goeldichironomus), 198 sp. 2 (Stenochironomus), 198
sp. (Peleteria), 221 sp. 1 (Heraeus), 251 sp. 2 (Symploce), 186
sp. (Pelmatosilpha), 184 sp. 1 (Labrundinia), 199 sp. 2 (Thaumatomyrmex), 305
sp. (Phamisus), 115 sp. 1 (Megaris), 255 sp. 2 (Wyeomyia), 201
sp. (Philotrypesis), 296 sp. 1 (Melanagromyza), 224 sp. 2† (Cariblatta), 187
sp. (Phlugis), 368 sp. 1 (Orthocladiinae), 198 sp. 2† (Cyphoderus), 95
sp. (Pholcomma), 62 sp. 1 (Neoblattella), 187 sp. 2† (Isotoma), 95
sp. (Pilica), 209 sp. 1 (Nesomylacris), 186 sp. 2† (Neosorius), 113
sp. (Pissodes), 176 sp. 1 (Nyctibora), 186 sp. 2† (Opalimosina), 228
sp. (Planodascalia), 261 sp. 1 (Ophiomyia), 225 sp. 2† (Platydema), 146
sp. (Potamocypris), 20 sp. 1 (Paranocticola), 187 sp. 2† (Ululodes), 358
sp. (Prionocypris), 20 sp. 1 (Pentaneura), 199 sp. 2‡ (Argilloecia), 26
sp. (Protalebra), 268 sp. 1 (Polypedilum), 198 sp. 2‡ (Macrocypris), 25
sp. (Protalges), 51 sp. 1 (Procladius), 199 sp. 2‡ (Paradoxostoma), 26
sp. (Protortonia), 279 sp. 1 (Psectrocladius), 199 sp. 2‡ (Phlyctocythere), 25
sp. (Pseudleptomastix), 292 sp. 1 (Stenochironomus), 198 sp. 2‡ (Xestoleberis), 28
sp. (Pseudochironomus), 198 sp. 1 (Symploce), 186 sp. 3 (Epilampra), 185
sp. (Pseudopsodexia), 217 sp. 1 (Syscia), 299 sp. 3 (Eurycotis), 185
sp. (Pseudosmittia), 199 sp. 1 (Telomerina), 229 sp. 3 (Labrundinia), 199
sp. (Psilochalcis), 291 sp. 1 (Thaumatomyrmex), 305 sp. 3 (Melanagromyza), 224
sp. (Raillietiella), 16 sp. 1 (Wyeomyia), 201 sp. 3 (Neoblattella), 187
sp. (Rhizoecus), 280 sp. 1† (Cariblatta), 187 sp. 3 (Paranocticola), 187
sp. (Saphirella), 18 sp. 1† (Cyphoderus), 95 sp. 3 (Polypedilum), 198
sp. (Sayomyia), 199 sp. 1† (Dahlimosina), 228 sp. 3† (Platydema), 146
sp. (Scelio), 284 sp. 1† (Isotoma), 95 sp. 3‡ (Argilloecia), 26
sp. (Sciophila), 203 sp. 1† (Neosorius), 113 sp. 3‡ (Portunus), 43
sp. (Scymnus), 141 sp. 1† (Opalimosina), 228 sp. 3‡ (Xestoleberis), 28
sp. (Sericothrips), 385 sp. 1† (Platydema), 146 sp. 4 (Eurycotis), 184
sp. (Sphecodes), 315 sp. 1† (Ululodes), 358 sp. 4 (Melanagromyza), 224
sp. (Sphyrotheca), 96 sp. 1‡ (Argilloecia), 26 sp. 4 (Neoblattella), 187
sp. (Stenothoe), 30 sp. 1‡ (Macrocypris), 25 sp. 4 (Polypedilum), 198
sp. (Stigmacoccus), 280 sp. 1‡ (Paradoxostoma), 26 sp. 4‡ (Xestoleberis), 28
sp. (Stigmus), 313 sp. 1‡ (Phlyctocythere), 25 sp. 5 (Neoblattella), 187
sp. (Strandesia), 21 sp. 1‡ (Portunus), 43 sp. 5 (Polypedilum), 198
sp. (Strotarchus), 70 sp. 1‡ (Uroleberis), 28 sp. A (Anoplotermes), 189

653 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

sp. A (Ochrotrichia), 389 sp. D‡ (Eucytherura), 23 sp.† (Arthrovertex), 48
sp. A† (Drosophila), 230 sp. E‡ (Eucytherura), 23 sp.† (Asteia), 225
sp. A† (Eudorylas), 215 sp. E‡ (Xestoleberis), 28 sp.† (Atopsyche), 390
sp. A† (Nannohelea), 196 sp. G‡ (Cytherura), 23 sp.† (Atractocerus), 135
sp. A† (Stilobezzia), 196 sp. G‡ (Eucytherura), 23 sp.† (Atrichopogon), 197
sp. A‡ (Campylocythere?), 27 sp. H‡ (Cytherura), 23 sp.† (Aulacigaster), 225
sp. A‡ (Cytherella), 21 sp. H‡ (Xestoleberis), 28 sp.† (Australolimnophila), 193
sp. A‡ (Cytherura), 23 sp. I‡ (Cytherura), 23 sp.† (Baetis), 235
sp. A‡ (Eucytherura), 23 sp. indet.† (Ischnothele), 56 sp.† (Baeus), 283
sp. A‡ (Loxoconcha), 25 sp. indet.† (Meioneta), 64 sp.† (Barroeuplectoides), 115
sp. A‡ (Perissocytheridea), 22 sp. indet.† (Miagrammopes), 60 sp.† (Baryconus), 283
sp. A‡ (Semicytherura), 23 sp. indet.† (Orchestina), 59 sp.† (Basalys), 288
sp. aff. abyssicola‡ (Kangarina), 23 sp. indet.† (Sarinda), 76 sp.† (Beckeriella), 232
sp. aff. bichense‡ (Cytheropteron), 23 sp. indet.‡ (Callinectes), 43 sp.† (Beckerina), 214
sp. aff. caudata‡ (Cytheromorpha?), 25 sp. indet.‡ (Parthenope), 43 sp.† (Bembidion), 107
sp. aff. cicatricosa‡ (Aurila), 24 sp. indet.‡ (Portunus), 43 sp.† (Blaste), 382
sp. aff. cubensis‡ (Trachyleberidea), 27 sp. n. (Heterosarus), 314 sp.† (Blastopsocus), 382
sp. aff. decora‡ (Macrocyprina), 19 sp. n. (Nomada), 318 sp.† (Boletina), 203
sp. aff. E. crenatus‡ (Eurypanopeus), 44 sp. n. 1 (Lasioglossum), 315 sp.† (Borophaga), 214
sp. aff. forda‡ (Loxoconcha), 25 sp. nr. acutus (Oecleus), 259 sp.† (Brachinus), 101
sp. aff. formosa‡ (Bairdia), 19 sp. nr. crudus (Haplaxius), 259 sp.† (Brachyglutina), 115
sp. aff. fortificata‡ (Bairdia), 19 sp. poss. bella (Euaresta), 223 sp.† (Brachymeria), 291
sp. aff. fortificata‡ (Paranesidia), 19 sp.? (prope primigenia)† (Pheidole), 303 sp.† (Brachymyrmex), 300
sp. aff. hispida‡ (Paracytheridea), 26 sp.† (Aaroniella), 381 sp.† (Bracon), 285
sp. aff. howei‡ (Cushmanidea), 21 sp.† (Acerotella), 283 sp.† (Brontes), 137
sp. aff. howei‡ (Heinia), 5 sp.† (Achrysocharoides), 292 sp.† (Caenocholax), 383
sp. aff. laevicula‡ (Bairdia), 19 sp.† (Acrolophus), 320 sp.† (Calliscelio), 283
sp. aff. magniventra‡ (Pellucistoma), 22 sp.† (Aedes), 199 sp.† (Calotelea), 283
sp. aff. moriahensis‡ (Cativella), 27 sp.† (Agallia), 263 sp.† (Camponotus), 299
sp. aff. palton‡ (Cytheropteron), 23 sp.† (Agyneta), 64 sp.† (Camptoptera), 295
sp. aff. pandora‡ (Cytherella), 21 sp.† (Alethaxius), 162 sp.† (Canthidium), 120
sp. aff. pulchra‡ (Cytherella), 21 sp.† (Alienates), 238 sp.† (Carabodes?), 46
sp. aff. russelli‡ (Brachycythere), 27 sp.† (Allecula), 145 sp.† (Caribbomerus), 150
sp. aff. semitalis‡ (Cytherella), 21 sp.† (Allodia), 202 sp.† (Catolethrus), 170
sp. aff. swaini‡ (Cytherura), 23 sp.† (Allonothrus), 48 sp.† (Ceraphron), 297
sp. aff. tarentina‡ (Bosquetina), 27 sp.† (Allostemma), 283 sp.† (Ceratobaeus), 283
sp. aff. Uleberis sp. 1‡ (Uroleberis), 28 sp.† (Amblyomma), 52 sp.† (Cernotina), 387
sp. aff. victrix‡ (Bairdia), 19 sp.† (Americhernes), 80 sp.† (Chalcosicya), 162
sp. aff. vulgata‡ (Cytherella), 21 sp.† (Anacaena), 108 sp.† (Choeras), 286
sp. aff. woodwardsensis‡ (Loxoconcha), sp.† (Anaplecta), 186 sp.† (Chonocephalus), 214
25 sp.† (Anaulacus), 102 sp.† (Chrysopa), 359
sp. B (Anoplotermes), 189 sp.† (Anaxipha), 375 sp.† (Chrysopilus), 206
sp. B (Ochrotrichia), 389 sp.† (Anoncia?), 322 sp.† (Cladiopsocus), 380
sp. B† (Drosophila), 230 sp.† (Anthomyza), 225 sp.† (Cloeodes), 235
sp. B† (Eudorylas), 215 sp.† (Anthonomus), 169 sp.† (Clusiodes), 225
sp. B‡ (Cytherella), 21 sp.† (Apanteles), 286 sp.† (Clystopsenella), 311
sp. B‡ (Cytherelloidea), 21 sp.† (Apharus), 115 sp.† (Coccophagus), 290
sp. B‡ (Eucytherura), 23 sp.† (Aphelinus), 289 sp.† (Coelosia), 203
sp. B‡ (Loxoconcha), 25 sp.† (Apiomerus), 246 sp.† (Colaspoides?), 162
sp. B‡ (Paracypris), 20 sp.† (Apsectus), 134 sp.† (Colydium), 143
sp. B‡ (Perissocytheridea), 22 sp.† (Apsilophrys), 291 sp.† (Condylostylus), 212
sp. B‡ (Xestoleberis), 28 sp.† (Apterostigma), 300 sp.† (Conicera), 213
sp. C‡ (Cytherura), 23 sp.† (Aradophagus), 283 sp.† (Coniopteryx), 360
sp. C‡ (Eucytherura), 23 sp.† (Archipsocus), 381 sp.† (Conostigmus), 298
sp. cf. alexanderi‡ (Argilloecia), 26 sp.† (Arthmius), 115 sp.† (Contarinia), 202
sp. D‡ (Cytherura), 23 sp.† (Arthrostictus), 102 sp.† (Conura), 291

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 654
sp.† (Copestylum), 215 sp.† (Hemipsocus), 382 sp.† (Nitela), 312
sp.† (Copidosoma), 291 sp.† (Heteromyia), 197 sp.† (Nothomyia), 206
sp.† (Cordyla), 203 sp.† (Heteronebo), 83 sp.† (Nysson), 312
sp.† (Corethrella), 199 sp.† (Himatium), 170 sp.† (Octostruma), 301
sp.† (Crabro), 312 sp.† (Holoteleia), 284 sp.† (Odinia), 226
sp.† (Crematogaster (Acrocoelia)), 302 sp.† (Hypoponera), 305 sp.† (Odontacolus), 284
sp.† (Crematogaster (Orthocrema)), 302 sp.† (Idris), 284 sp.† (Oethecoctonus), 284
sp.† (Crossocerus), 312 sp.† (Indiopsocus), 382 sp.† (Oligomyrmex), 303
sp.† (Cryptops), 84 sp.† (Inodesmus?), 90 sp.† (Opistacantha), 284
sp.† (Cryptopygus), 95 sp.† (Inostemma), 283 sp.† (Oppia?), 47
sp.† (Culiseta), 201 sp.† (Iomus?), 90 sp.† (Orfelia), 202
sp.† (Cylindromyrmex), 298 sp.† (Isthmopsocus), 380 sp.† (Oribotritia), 47
sp.† (Dasyodontus), 89 sp.† (Jubus), 115 sp.† (Oripoda), 47
sp.† (Delocrania), 160 sp.† (Keroplatus), 202 sp.† (Ornithodorus), 52
sp.† (Dendrocerus), 298 sp.† (Kimula?), 77 sp.† (Orthopodomyia), 200
sp.† (Diachus), 162 sp.† (Latheticomyia), 221 sp.† (Osoriellus), 113
sp.† (Dicrotendipes), 197 sp.† (Leia), 203 sp.† (Pachychernes), 80
sp.† (Diolcogaster), 286 sp.† (Lepidocyrtus), 95 sp.† (Pachyteles), 107
sp.† (Diostracus), 212 sp.† (Leptanilloides), 300 sp.† (Parabaeus), 283
sp.† (Diplonevra), 213 sp.† (Leptogaster), 209 sp.† (Parahymenorus), 145
sp.† (Discothyrea), 305 sp.† (Leptonema), 387 sp.† (Parasierola), 309
sp.† (Docodesmus), 90 sp.† (Leptonesiotes), 159 sp.† (Paratrechina), 300
sp.† (Dohrniphora), 213 sp.† (Leptosalda), 238 sp.† (Parawithius?), 80
sp.† (Dolicheremaeus), 48 sp.† (Leptoteleia), 284 sp.† (Paronella), 96
sp.† (Duta), 283 sp.† (Leucochrysa), 359 sp.† (Pentanodes), 153
sp.† (Eburia), 151 sp.† (Liberiana), 112 sp.† (Phanerotoma), 285
sp.† (Echmepteryx), 379 sp.† (Limonia), 194 sp.† (Phloeoxena), 104
sp.† (Ectaetia), 196 sp.† (Liodes), 47 sp.† (Pholeomyia), 227
sp.† (Elaphidion), 151 sp.† (Liposcelis), 380 sp.† (Phronia), 203
sp.† (Elephantomyia), 193 sp.† (Litargus), 143 sp.† (Phyllomyza), 227
sp.† (Embidobia), 283 sp.† (Lithoseopsis), 380 sp.† (Platamus), 137
sp.† (Embioctonus), 283 sp.† (Lobopoda), 145 sp.† (Platygaster), 283
sp.† (Enderleinella), 381 sp.† (Loewimyia), 225 sp.† (Platyura), 203
sp.† (Epinannolene), 93 sp.† (Lophodesmus?), 90 sp.† (Plecia), 202
sp.† (Epipsocus), 380 sp.† (Lophoturus?), 86 sp.† (Poecilognathus), 211
sp.† (Eremaeozetes), 46 sp.† (Macrocentrus), 286 sp.† (Pogonomyrmex), 303
sp.† (Eretmocoris), 248 sp.† (Macrocera), 203 sp.† (Polymera), 193
sp.† (Erioptera), 192 sp.† (Macroteleia), 284 sp.† (Polypedilum), 198
sp.† (Eualebra), 268 sp.† (Mastigus), 116 sp.† (Prenolepis), 300
sp.† (Euceroplatus), 203 sp.† (Megophthalmidia), 203 sp.† (Prionopelta), 298
sp.† (Euconnus), 116 sp.† (Metachroma), 163 sp.† (Probably Bdella), 48
sp.† (Euderus), 293 sp.† (Methia), 152 sp.† (Probably Caeculus), 48
sp.† (Eugamandus?), 153 sp.† (Microscydmus), 116 sp.† (Probaryconus), 284
sp.† (Eurhopalothrix), 301 sp.† (Mochlozetes), 47 sp.† (Pseudoponera), 305
sp.† (Euthlastoblatta), 187 sp.† (Monelata), 288 sp.† (Pseudosinella), 95
sp.† (Exechia), 203 sp.† (Monopelopia), 199 sp.† (Pseudosymploce), 187
sp.† (Exechiopsis), 203 sp.† (Mycetobia), 201 sp.† (Psilanteris), 284
sp.† (Fidiobia), 283 sp.† (Mycetophila), 203 sp.† (Psilochalcis), 291
sp.† (Forcipomyia), 197 sp.† (Mycomyia), 203 sp.† (Psochodesmus), 90
sp.† (Genus nr. Fallopia), 49 sp.† (Myriophora), 214 sp.† (Psychoda), 195
sp.† (Geranomyia), 194 sp.† (Myrmelachista), 300 sp.† (Psyllipsocus), 380
sp.† (Glomeridesmus), 86 sp.† (Mythicomyia), 211 sp.† (Ptycta), 382
sp.† (Glyptoscelis), 162 sp.† (Neopleurophora), 214 sp.† (Pyroderces?), 322
sp.† (Gnophomyia), 192 sp.† (Neoxantholinus), 117 sp.† (Rainieria), 221
sp.† (Gryon), 284 sp.† (Nephila), 67 sp.† (Rasahus), 246
sp.† (Hemerobius), 360 sp.† (Newportia), 85 sp.† (Rhamphomyia), 213

655 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

sp.† (Rhegmoclemina), 196 sp.‡ (Ambocythere), 27 SPARASSIDAE, 73
sp.† (Rhinanisus), 170 sp.‡ (Argilloecia), 26 Sparatta, 183
sp.† (Rhinotus), 87 sp.‡ (Bythoceratina), 20 sparsum (Bembidion), 107
sp.† (Rhopalopsyllus), 382 sp.‡ (Bythocypris), 19 sparsuta (Pachybrachis), 162
sp.† (Rhyncophoromyia), 214 sp.‡ (Calappa), 40 Spartocera, 254
sp.† (Rhyopsocus), 380 sp.‡ (Calappella?), 40 spartocerana (Eubule), 254
sp.† (Rhyparus), 120 sp.‡ (Campylocythere?), 27 Spasalus, 118
sp.† (Rymosia), 203 sp.‡ (Cushmanidea), 21 spathifrons (Cryptotermes), 188
sp.† (Sacculobates), 46 sp.‡ (Cyclocypris), 20 spathor (Sarcofahrtiopsis), 219
sp.† (Salina), 96 sp.‡ (Cypretta), 20 spathula (Cossonus), 170
sp.† (Scalenarthrus), 116 sp.‡ (Cypria), 20 spatulata (Craspedisia), 61
sp.† (Scatopse), 196 sp.‡ (Cypridopsis), 20 spatulata (Micromyrmex), 172
sp.† (Scelio), 284 sp.‡ (Cytherella), 21 species A (Toxomerus), 216
sp.† (Scolopocryptops), 85 sp.‡ (Cytherelloidea), 21 speciosus (Evagoras), 246
sp.† (Scraptia), 142 sp.‡ (Cytheropteron), 23 Spectralia, 125
sp.† (Seira), 96 sp.‡ (Darwinula), 23 spectrum (Hapithus), 371
sp.† (Selenophorus), 103 sp.‡ (Dolerocypris), 20 specula† (Polymera), 193
sp.† (Selenops), 73 sp.‡ (Eucytherura), 23 Spelaeala, 369
sp.† (Siphonocybe), 91 sp.‡ (Glyptobairdia), 19 SPELAEORHYNCHIDAE, 54
sp.† (Siphonophora), 91 sp.‡ (Haplocytheridea), 22 Spelaeorhynchus, 54
sp.† (Smaris), 49 sp.‡ (Hemicytherura), 23 Spelobia, 229
sp.† (Solenopsis (Euophthalma)), 304 sp.‡ (Heterocypris), 20 Speocropia, 350
sp.† (Solenopsis (Solenopsis)), 304 sp.‡ (Iliacantha), 41 sperata (Diestota), 111
sp.† (Sphyrotheca), 96 sp.‡ (Limnocythere?), 25 sphacelata (Pentacora), 238
sp.† (Spiniteleia), 284 sp.‡ (Macrocyprina), 19 sphacelatus (Exophthalmus), 173
sp.† (Spurostigma), 380 sp.‡ (Macrocypris), 25 Sphacelodes, 334
sp.† (Stegana), 231 sp.‡ (Microxestoleberis), 28 SPHAERIODESMIDAE, 91
sp.† (Stenochironomus), 198 sp.‡ (Mithrax), 42 Sphaerocera, 229
sp.† (Stenognathus), 104 sp.‡ (Munseyella), 26 SPHAEROCERIDAE, 227
sp.† (Stylogaster), 221 sp.‡ (Panopeus), 44 Sphaerognathium, 146
sp.† (Sympherobius), 360 sp.‡ (Paracypris), 20 sphaeroidea‡ (Strandesia), 21
sp.† (Synapha), 203 sp.‡ (Paracythere), 26 Sphaeroma, 33
sp.† (Tachyporus), 118 sp.‡ (Paracytheridea), 26 SPHAEROMATIDAE, 33
sp.† (Tanypus), 199 sp.‡ (Paracytheroma), 22 Sphaeropthalma, 306
sp.† (Telenomus), 285 sp.‡ (Paradoxostoma), 26 SPHECIDAE, 313
sp.† (Teleoliodes), 47 sp.‡ (Parakrithella), 24 Sphecius, 312
sp.† (Telephanus), 137 sp.‡ (Pellucistoma), 22 Sphecodes, 315
sp.† (Telmatoscopus), 195 sp.‡ (Physocypria), 20 Sphendononema, 84
sp.† (Termitaradus), 275 sp.‡ (Pilumnus), 44 sphenobathra (Ardeutica), 331
sp.† (Termitodius), 120 sp.‡ (Pontocypris), 26 Sphenoceros, 181
sp.† (Theope), 340 sp.‡ (Potamocypris), 20 Sphenophorus, 177
sp.† (Thesium), 116 sp.‡ (Procythereis?), 27 Sphex, 314
sp.† (Toxorhina), 193 sp.‡ (Propontocypris), 20 SPHINDIDAE, 136
sp.† (Trichadenotecnum), 382 sp.‡ (Pseudocythere), 20 SPHINGIDAE, 344
sp.† (Trichonta), 203 sp.‡ (Schlerochilus), 20 sphingis (Telenomus), 285
sp.† (Trichopria), 288 sp.‡ (Uroleberis), 28 Sphingonotus, 367
sp.† (Trichorhina), 33 sp.‡ (Xiphichilus), 26 Sphinx, 344
sp.† (Tricimba), 227 sp.‡ (Zanthopsis), 44 Sphyrocoris, 258
sp.† (Triteleia), 284 sp.‡ cf. vespa (Argilloecia), 26 Sphyrotheca, 96
sp.† (Tyloderma), 172 spadix (Acrotaenia), 222 spiculum† (Palaeodyctina), 70
sp.† (Walterianella), 159 Spalacopsis, 154 spiculus (Catenes), 329
sp.† (Wasmannia), 301 Spalangia, 296 spiculus (Platystethus), 113
sp.† (Wilcoxia), 208 Spalangiopelta, 296 Spiloconis, 360
sp.† (Xanthocaecilius), 381 Spangbergiella, 266 spilogaster (Nostima), 234
sp.† (Xestocephalus), 269 spanglerorum (Dagus), 233 spilogastrata (Macaria), 333
sp.† (Zaischnopsis), 294 spanna (Epargyreus), 337 spiloleuca (Plaxia), 351

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 656
spilomela (Micratethis), 350 spinulosus (Neogonodactylus), 28 Steatonyssus, 52
spilota (Leucocera), 161 spinulosus (Xyleborus), 180 Stegana, 231
spilotrachela (Disonycha), 157 spio (Dismorphia), 339 Stegasta, 321
spina (Gypona), 268 spira (Cladochaeta), 229 stegmaieri (Calycomyza), 225
spinatus† (Cenocephalus), 182 spiraecola (Aphis), 274 stegmaieri (Mosillus), 232
spinea (Bradysia), 204 spirembolus† (Araneometa), 66 Stegobium, 134
spinicarpus (Portunus), 43 SPIROBOLELLIDAE, 91 Stegomyia, 200
spinicauda (Crangoweckelia), 29 Spirobolellus, 91 stegosoma† (Enischnomyia), 217
spinicollis (Malacomorpha), 378 SPIROSTREPTIDAE, 93 steinheili (Crematogaster), 302
spinicorne (Elaphidion), 151 Spissistilus, 271 steinheili (Nylanderia), 300
spinicorne (Plectrocerum), 152 splendida (Geotrupes), 119 Steinovelia, 240
spinicosta (Acanthocheila), 242 splendida (Rhytidophthia), 253 Steirastoma, 154
spinicrus (Citharacanthus), 56 splendidum (Calosoma), 101 stelenes stelenes (Siproeta), 342
spinicrus (Lasiomerus), 245 splendidum (Elaphidion), 151 Stelidota, 137
spinidens (Andrallus), 255 Spodoptera, 356 stellata (Acanthepeira), 65
spinifer (Mucrotettix), 365 Spoladea, 327 stellata (Disclisioprocta), 335
spinifer† (Odontomachus), 305 SPONGIPHORIDAE, 183 stellata (Gymnetis), 124
spiniferus (Aleurocanthus), 272 Spongovostox, 183 stellidaria (Polysemia), 335
spinifex (Euscelus), 166 sporadicos (Paroecanthus), 372 stellifera (Vinsonia), 276
spiniformis (Popcornella), 76 spp. (Cephalosphaera), 214 stellus (Elcaribe), 212
spiniger (Chlorion), 314 spp. (Eretmocerus), 290 STEMMIULIDAE, 93
Spinilimosina, 229 spp. (Rhodesiella), 227 Stemmops, 62
spinimanus (Portunus), 43 spp.† (Orthellia), 203 stena (Neokrisna), 267
spinimanus (Psilopus), 212 spp.† (Xestochironomus), 198 Stenacris, 367
spinipalpus† (Succinya), 70 Spragueia, 350 Stenapion, 167
spinipes (Haplopus), 377 spretum (Bembidion), 107 Stenaspidiotus†, 161
spinipes (Trujillina), 69 spretus (Lachnopus), 174 stenaspis (Triaeris), 60
spinipes† (Electrobates), 239 spumarius (Lestes), 362 stenelytrus† (Toxorhynchus), 167
spinipes† (Pensacolatus), 76 spurium‡ (Pellucistoma?), 22 Steneotarsonemus, 50
spinipes† (Wulfila), 71 Spurostigma, 380 stenis† (Paracamptopsis), 171
spiniscapula (Dasymutilla), 306 Squamanotoxus, 149 Stenocaecilius, 381
Spiniteleia, 284 squamatus‡ (Dardanus), 39 stenocalypus† (Neoulosomus), 171
spiniventris (Cercyon), 109 squamifera† (Micracites), 182 Stenochironomus, 198
spinki (Steneotarsonemus), 50 squamigera (Metrichia), 389 Stenochrus, 81
spinoalata‡ (Loxoconcha), 25 squamipennis (Aedeomyia), 200 Stenocionops, 42
spinolae (Melanaethus), 254 squamipennis (Exophthalmus), 173 Stenocoris, 253
spinosa (Bacteria), 377 squamosus† (Cephalotes), 301 Stenocrepis, 104
spinosa (Lupettiana), 71 squamosus† (Pseudochthonius), 79 stenoculus (Apodrosus), 175
spinosa (Ptilodexia), 220 squamula† (Diaprepes), 173 Stenodiplosis, 202
spinosa† (Palaeolathys), 70 Squilla, 28 Stenodontes, 155
spinosissima (Stenocionops), 42 SQUILLIDAE, 28 Stenognatha, 348
spinossisimus (Mithrax), 42 stabile (Apion), 167 Stenognathus, 104
spinosum† (Monoblemma?), 59 Stagmomantis, 190 Stenogryllus, 371
spinosus (Belopherus), 168 stainesorum† (Sceloenopla), 161 Stenolemus, 246
spinosus (Bicyrtes), 311 Stamnoderus, 114 Stenolophus, 103
spinosus (Chirothrips), 384 stangei (Compsocryptus), 287 Stenomicra, 226
spinosus (Hypasclera), 148 stangei (Osmia), 316 STENOMIDAE, 322
spinosus (Oceanaspidiotus), 277 stani (Leptoteleia), 284 Stenommatus, 172
spinosus (Phrurolithus), 71 Stantonia, 286 Stenomorphus, 103
spinosus (Steatonyssus), 52 STAPHYLINIDAE, 111 Stenoonops, 60
spinosus (Tytthonyx), 133 staplini‡ (Limnocythere), 25 Stenopoda, 247
Spintharus, 62 starki (Drosophila), 230 STENOPODIDAE, 35
spinula (Efferia), 208 Statira, 143 stenopterus (Corynothrips), 384
spinulifera baoruco (Chimarra), 386 statira hispaniolae (Aphrissa), 338 Stenopus, 35
spinulifera galalcha (Chimarra), 386 Stator, 164 Stenorhynchus, 42
spinulifera spinulifera (Chimarra), 386 Steatoda, 62 stenosoma† (Geratozygops), 170

657 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

stenostegus† (Semnorhynchus), 172 Strabala, 158 Strongylopsalis, 183
Stenotabanus, 207 strachani (Pinnaspis), 278 strongylus† (Pleurambus), 168
Stenotarsus, 141 stramineum (Menacanthus), 379 Strotarchus, 70
Stenothoe, 30 stramineus (Ommatius), 210 Strumigenys, 302
STENOTHOIDAE, 30 Strandesia, 21 strumosus (Pterygiodesmus), 89
Stenotrupis, 170 Strategus, 123 Strymon, 339
Stenoxenus, 197 STRATIOMYIDAE, 205 stuarti (Monobeloides), 270
Stenus, 118 Stratiorrhynchus, 168 studiosus (Anelosimus), 61
STEPHANIDAE, 282 strator (Cubachelifer), 80 sturmi† (Archeatelura), 97
Stephanitis, 243 Stratus, 116 sturtevanti (Drosophila), 230
stephanoderis (Cephalonomia), 309 straubae (Drosophila), 230 stygia (Nyctibora), 186
stephensoni‡ (Hemicyprideis), 22 Strebla, 217 STYGIOMYSIDAE, 29
stercoraria (Scathophaga), 220 STREBLIDAE, 217 Stygiomysis, 29
stercorea (Ancylostomia), 328 streblopa (Streblopotamia), 331 stylipennis (Drosophila), 230
Stereodermus, 168 Streblopotamia, 331 STYLODACTYLIDAE, 37
STERNOPHORIDAE, 79 strena (Amphiacusta), 374 Stylodactylus, 37
sterope (Antiblemma), 351 strenua (Eumorpha), 346 Stylogaster, 221
Stethorus, 141 strenua (Platythyrea), 305 Styloleptus, 153
Sthenaridea, 244 strenua (Ptilodexia), 220 stylolepus† (Neoulosomus), 171
Stichotrema, 383 strenuana (Epiblema), 331 Stylopalpia, 330
stictella (Australephestiodes), 328 Strepsicrates, 332 Styposis, 63
Stictia, 312 STREPTOCEPHALIDAE, 14 Styraxodesmus, 90
sticticollis (Agalliana), 263 Streptocephalus, 14 Styringomyia, 193
Sticticomus, 149 Streptopsyche, 387 stysi† (Phymacista), 242
sticticus (Conoderus), 128 striata (Cyclocephala), 123 subaeneus (Brachyplatys), 257
stictigramma (Phaedropsis), 326 striata (Cypriacis), 125 subaequale (Phoebolampta), 368
sticturus (Monocrepidius), 128 striata (Galerita), 102 subalipes (Trichopoda), 220
stigma (Alabagrus), 285 striata boxi (Myzine), 308 subanticus (Micromus), 360
stigma (Eburia), 151 striata haitiensis (Cyclocephala), 123 subapicalis (Cropia), 350
stigma (Pseudoponera), 305 striatifrons (Neogriphoneura), 224 subapicana (Oleuthreutes), 331
Stigmacoccus, 280 striatipes (Mangora), 67 subarcana† (Archicratyna), 203
stigmaterus (Microdus), 285 striatolateralis (Sicoderus), 176 subarmata (Pheidole), 303
stigmatias (Euxesta), 223 striatopennatus (Bolacothrips), 384 subater (Pseudomyrmex), 306
stigmatophora (Pyroderces), 322 striatopunctatus (Selenophorus), 103 subcaudalis (Eophrixus), 32
Stigmus, 313 striatula (Gnamptogenys), 299 subcelata (Geometra), 334
Stilbometopa, 217 striatulus (Scolytodes), 179 subcerdoso (Antillostenochrus), 81
Stilbus, 138 striatus (Heterophotinus), 131 subcinerascens (Antiblemma), 351
stillmani (Burca), 337 striatus (Hymenorus), 145 subclararia (Choerodes), 334
stillmani (Copaeodes), 336 striatus (Hypothenemus), 179 subcolumbata (Aspilates), 333
Stilobezzia, 196 stribalus (Berosus), 108 subconcentralis (Pityophthorus), 181
Stilomedon, 115 stricta‡ (Phlyctocythere), 25 subcordens (Platynus), 105
Stilosaurus, 115 stridulans† (Scytodes), 57 subcostatus (Platypus), 182
stimulans (Plusiodonta), 354 strigicauda (Ataenius), 120 subcyanea (Ctenuchidia), 348
stipata (Graphomya), 218 strigicollis (Cnesinus), 177 subdegeneratus (Matsucoccus), 280
stipes† (Dipoenata), 62 strigicollis (Cryptocephalus), 162 subdepressa (Stenocrepis), 104
Stirellus, 266 strigicollis (Stratiorrhynchus), 168 subducta (Plaxia), 351
Stiretrus, 255 strigifera (Magusa), 358 subelegans (Valenzuela), 381
Stizocera, 152 strigifera (Melipotis), 353 subequus‡ (Pilumnus), 44
Stobaera, 260 strigilis strigilis (Protambulyx), 344 suberosus (Omorgus), 119
stocki (Jehaia), 32 strigipennis (Odopoea), 263 subfasciatus (Zabrotes), 164
stocki (Strandesia), 21 strigosa (Calidota), 348 subferrugineus (Amnestus), 254
stolida† (Quilessa), 262 strigosella (Cryptolechia), 321 subferrugineus (Podisus), 255
Stomatomiris†, 244 strigulata (Lithocharodes), 117 subflexa (Heliothis), 356
Stomoxys, 218 striolata (Siphunculina), 227 subglobosa (Iliacantha), 41
stonei (Anastrepha), 223 Strongylaspis, 155 subidalis (Hypena), 357
stonei (Guttipsilopa), 231 Strongylium, 147 subimpressum (Trypoxylon), 313

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 658
subimpressus (Zelus), 246 succini† (Pintomyia), 195 Symmerista, 347
subita† (Diclidophlebia), 271 succinus† (Pseudomyrmex), 306 Sympetrum, 364
sublinea (Acanalonia), 258 Succinya†, 70 Sympherobius, 360
sublineatus (Stilbus), 138 sucinum† (Harpactosphecion), 311 Symphylus, 258
submutata (Eggyna), 349 sucinus† (Cephalotes), 301 SYMPHYTOGNATHIDAE, 63
suboblita (Atheta), 111 Sufetula, 327 Symploce, 186
subopalaria (Sabulodes), 334 sufficiens (Condica), 350 synalphei (Bopyrione), 32
subordinata (Poaphila), 354 suffusa (Odopoea), 263 synalphei (Hemiarthrus), 32
subosseella (Lipographis), 329 suffusa (Thermesia), 352 Synalpheus, 36
subquadraregularis‡ (Cyprideis), 22 suffuscus (Novohorus), 79 Synapha, 203
subquadratus (Selenophorus), 103 suillus (Anchonus), 176 Synapte, 336
subreticulata‡ (Ambocythere), 27 suillus (Pseudechiniscus), 13 Synchlora, 334
subreticulatum‡ (Cytheropteron), 23 suis (Haematopinus), 379 Synclera, 327
subrostratus (Felicola), 379 sulcata (Saldula), 238 Syndesus, 124
subrugosa‡ (Perissocytheridea), 22 sulcatus (Prostemmiulus), 94 Synecheporus, 87
subsignata (Charidotella), 160 sulcatus (Rhipidandrus), 144 Syneches, 213
subspinea (Bradysia), 204 sulci† (Fossilopaea), 59 Syngamia, 327
subspinipes (Scolopendra), 85 Sulciana†, 271 SYNLESTIDAE, 362
subspinosum subspinosum (Belostoma), sulcimargo (Lapethus), 140 Synoditella, 284
240 sulphureicornis (Entypus), 307 Synsynella, 32
substitutum (Pterogramma), 228 sulphureus (Tabanus), 207 Synthesiomyia, 218
substriatus (Conoderus), 128 summermanae (Mordella), 142 Syphraea, 158
substriatus (Lobopoda), 145 sunia (Neogalea), 357 syriacus (Retithrips), 384
subterranea (Agrotis), 357 Suniophacis, 115 Syringogaster, 221
subtessellatus (Xestocephalus), 269 Sunius, 115 SYRINGOGASTRIDAE, 221
subtilis (Aleuroglandulus), 272 Supella, 187 SYRPHIDAE, 215
subtilis (Anthicus), 149 superba (Phymata), 247 Syrphus, 216
subtilis (Antillitermes), 189 superba (Silis), 132 Syscia, 299
subtilis (Bolina), 353 superior (Cloeodes), 235 Syspira, 70
subtilis (Embolemus), 311 Suphis, 100 Systellapha, 221
subtilis (Farfantepenaeus), 35 Suphisellus, 100 Systena, 158
subtilis (Parvitermes), 189 suramerica (Acanthoscelides), 163 Szelenyia, 297
subtilis (Pityophthorus), 181 surieli (Achromoporus), 88 szuetsi (Rotundabaloghia), 54
subtilissimus (Hypselodesmus), 88 surieli (Cerceris), 313 T (Pepsis), 314
subtropica (Typhlodromina), 53 surinamensis (Aphelopus), 310 tabaci (Bemisia), 272
subtropicalis (Aphaereta), 285 surinamensis (Orizaephilus), 138 tabaci (Rhaptinus), 169
subtus† (Ommatius), 210 surinamensis (Pycnoscelus), 186 tabaci (Thrips), 385
subulatum (Piezosternum), 257 surinamum (Achryson), 150 tabaciphilus (Telephanus), 137
subulatus (Prostemmiulus), 94 surrepens (Coronta), 354 tabacivora (Encarsia), 290
Subulitermes, 189 susannae (Pheidole), 303 TABANIDAE, 206
subusta (Atethmia), 351 suspensa (Anastrepha), 223 Tabanus, 207
subvectaria (Geometra), 333 suspiciosa (Efferia), 208 tabebuia (Holopothrips), 385
subvertens (Ululodes), 358 sutor (Condica), 350 tabebuiae (Rabela), 269
subvitreus† (Enicocephalus), 238 suturalis (Achenomorphus), 113 tabulatum (Uvaroviella), 374
succedanea (Pseudoponera), 305 suturalis (Anacaena), 108 TACHINIDAE, 220
Succinarisemus, 195 suturalis (Euryscelis), 150 tachydromos (Amphiacusta), 374
succinctus succinctus (Trachyderes), 153 suturalis hebraea (Cylindera), 101 Tachypompilus, 308
succinea (Solenopsis), 304 suturalis suturalis (Cylindera), 101 Tachyporus, 118
succineus (Lithophorus), 139 swensoni† (Caulophilus), 171 Tachys, 107
succineus† (Apocephalus), 214 SYARINIIDAE, 79 Tachysphex, 313
succineus† (Dallasiellus), 254 sybaris (Composia), 348 Tachyta, 107
succini† (Anthrax), 210 syces insularis (Pachylia), 345 Tachytes, 313
succini† (Chimarra), 386 Syllectra, 355 taczanowskii (Tobias), 74
succini† (Dominicaphis), 275 syllepsicus (Mimetus), 60 Taeniaptera, 221
succini† (Drosophila), 230 Syllepte, 327 taeniatum (Doru), 183
succini† (Phaonia), 218 sylvicola woodruffi (Panoquina), 336 Taeniogonalos, 282

659 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

taeniola (Metacyrba), 76 taracte (Amphiacusta), 374 teneriffana (Pheidole), 303
taeniola (Zicca), 254 Taraxippus, 378 tenerrima (Emesa), 246
taeniorhynchus (Ochlerotatus), 200 tardigradum (Milnesium), 14 tennesseni (Progomphus), 363
Tafalisca, 372 tarsalis (Desmometopa), 227 tensicollis (Corticeus), 146
Tagosodes, 260 tarsalis (Notaepytus), 139 tentativa (Atheta), 111
taina (Atopsyche), 390 TARSONEMIDAE, 50 tenuicornis (Frankliniella), 384
taina (Proctacanthella), 208 tasajera (Calisto), 343 tenuicornis (Hapithus), 371
Tainacris, 366 tatila tatilista (Eunica), 340 Tenuicornus†, 296
Tainarys, 271 tau (Podium), 314 TENUIPALPIDAE, 50
tainensis (Dryinus), 311 tauralis (Radara), 354 Tenuipalpus, 50
tainicum† (Protolepisma), 97 tauricornis (Wagneriana), 68 tenuipes (Spirobolellus), 92
taino (Adelphe), 310 Tauriphila, 364 tenuis (Centruroides), 81
taino (Amphicranus), 181 taurus (Myopsocus), 382 tenuis (Hibana), 71
taino (Cercyon), 109 taylori (Trepobates), 240 tenuis (Hylastes), 177
taino (Ectemnius), 312 teapae (Caenodelphax), 259 tenuis (Nesidiocoris), 243
taino (Loxosceles), 57 teapensis (Sapintus), 149 tenuis† (Nephila), 67
taino (Neostasina), 73 Technomyrmex, 299 tenuis‡ (Portunus), 43
taino (Proceratium), 305 tectorum (Lachesilla), 381 tenuissima (Tetragnatha), 65
taino (Thoronidea), 284 tega (Oxyethira), 389 tenuivittatus (Sphenophorus), 177
taino† (Anoplotermes), 189 tegularia (Pyrinia), 333 TEPHRITIDAE, 222
taino† (Cephalotes), 301 telarius (Tetranychus), 50 Terastia, 327
taino† (Cyphomyrmex), 301 Telebasis, 362 TERATEMBIIDAE, 234
Tainocampa, 356 teleboas (Archimestra), 340 terebintharia (Numia), 333
Tainonia, 58 telemachusi (Encarsia), 290 terebrans (Sphaeroma), 33
Tainophasma, 378 Telenomus, 284 Teredon, 282
Tainopompilus†, 308 Teleoliodes, 47 terena (Libytheana), 341
tainosa (Asteriza), 160 Teleonemia, 243 teres (Chalcosicya), 162
Tainosia†, 262 Teleophrys, 42 Tergoceracris, 367
Talanus, 147 Telephanus, 137 Termes, 190
TALITRIDAE, 30 tellesaria (Drepanodes), 334 terminalis (Homalotylus), 292
talus (Astraptes), 337 Telmatoscopus, 195 terminalis (Homoptera), 358
Tamarixia, 294 Telocricus, 85 terminalis (Kapala), 292
tamatavensis (Amblyseius), 53 Telomerina, 229 terminifera (Bolina), 353
Tanaostigma, 297 Teloplatypus, 182 TERMITAPHIDIDAE, 275
TANAOSTIGMATIDAE, 297 Telphusa, 321 Termitaradus, 275
Tanaostigmodes, 297 TELYPHONIDAE, 83 TERMITIDAE, 188
Tangella, 262 Temelucha, 287 Termitodius, 120
Tangiopsis, 262 Teminius, 70 termitophila† (Palaeodyctina), 70
Tangyria, 262 Temnoscheila, 135 termophila (Trichorhina), 33
Tannea, 113 Temnothorax, 302 terraria (Almodes), 335
tantalus (Aeollopos), 345 tempestiva (Cyrtoxipha), 375 terresi (Pheidole), 303
tantilla (Hogna), 68 tempestiva† (Cratyna), 204 terresi (Pseudisobrachium), 310
tantilla† (Proplebeia), 317 temporarius† (Pityophthorus), 181 terresi (Sunius), 115
tanyacaena (Dominimyza), 224 tenax (Melanaspis), 277 terresi illota (Pheidole), 303
TANYPEZIDAE, 222 tenebricosa (Hemiblabera), 185 terribilis (Scaptosciara), 205
Tanypus, 199 tenebricosa† (Castianeira), 71 terricola (Lebia), 104
tanyrhinus† (Neoulosomus), 171 TENEBRIONIDAE, 143 terricola (Solenopsis), 304
tanyrhinus† (Rhynchitobius), 182 Tenebroides, 135 tersa tersa (Xylophanes), 346
tanyrhinus† (Semnorhynchus), 172 tenebrosus (Ataenius), 120 tertia† (Euryconus), 272
tanyrhinus† (Stenommatus), 172 tenella (Oxycopis), 148 TERTIAPATIDAE†, 13
Tanytarsus, 198 tenella (Tannea), 113 Tertiapatus†, 13
Taphrocerus, 124 tenellus (Circulifer), 267 tertiaria† (Archaeopolietes), 218
Tapinoma, 298 tenellus (Malacopterus), 152 tertiaria† (Lechytia), 79
Tapinopa, 64 tenellus (Metacanthus), 248 tertiaricola† (Forcipomyia), 197
taprobanicus (Mymar), 295 tenellus (Nysius), 250 tertiarius† (Corticeus), 147
Tarachidia, 350 tener (Hepomadus), 35 tertiarius† (Plectromerus), 150

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 660
tertiarius† (Protosphaeroniscus), 33 tetricus (Acanthoscelides), 163 thinophila (Phaleria), 147
terueli (Dominicophus), 374 tetricus (Bruchus), 163 Thionia, 261
TESSARATOMIDAE, 257 TETRIGIDAE, 365 Thiotricha, 321
Tessaropa, 152 tetrio (Pseudosphinx), 346 thoe (Doxocopa), 340
tesselaris (Halysidota), 348 tetrus (Osbornellus), 267 Tholymis, 364
tesselata (Achaearanea), 61 tettigonaris (Forcipomyia), 197 thomae (Adaina), 330
tessellatus (Eucalymnatus), 276 TETTIGONIIDAE, 368 thomae (Dioxyna), 223
tesseratus (Duplaspidiotus), 277 Tetyra, 258 thomae (Paralimna), 233
Tesserocerus, 182 texana (Ophiomyia), 225 thomae (Prionyx), 314
testacea (Centris), 317 texanella (Anadelosemia), 327 thomasi (Atopsyche), 390
testacea (Discothyrea), 305 texanus (Thicanus), 149 thomasi (Calisto), 343
testacea (Ozophora), 252 texense (Trypoxylon), 313 thomasi (Hoplicnema), 141
testacea (Phaleria), 147 texensis (Dysmicoccus), 279 thomasi (Leptostylopsis), 153
testacea (Thyanta), 257 THAEROCYTHERIDAE, 26 thomasi (Murgantia), 257
testaceipennis (Carpelimus), 113 Thalassozetes, 48 thomasi (Plectromerus), 151
testaceipennis (Neoxantholinus), 117 thalia (Alisotrichia), 388 thomasi (Sicoderus), 176
testaceipes (Homoeotarsus), 114 thalialis (Botys), 324 thomasi (Silidiscodon), 132
testaceipes (Lisyphlebus), 285 thalioculus† (Derelomus), 172 THOMISIDAE, 73
testaceomarginata (Chelymorpha), 160 thalli (Ramosella), 270 thompsoni (Telmatoscopus), 195
testaceomimus (Caccodes), 132 Thaloe, 71 thompsonii (Cryptorhopalum), 133
testaceovarius (Ablechrus), 136 Thalpius, 106 Thonalmus, 129
testaceum (Nesiosphaerion), 152 thamaluncus (Argyrotaenia), 332 thonalmus (Calocosmus), 155
testaceus (Ablechrus?), 136 thanatos† (Idarnes), 289 thoracica (Centris), 317
testaceus (Caccodes), 132 thaumasalis (Lophophora), 356 thoracica (Chrysopa), 359
testaceus (Loberus), 139 Thaumasia, 68 thoracica (Oxycopis), 148
testaceus (Lophoscutus), 247 thaumasta† (Notiobiella), 360 thoracica (Suniophacis), 115
testaceus (Pseudoanthonomus), 169 THAUMASTOCORIDAE, 241 thoracicus (Discoderus), 102
testudinea (Acrotaenia), 222 Thaumatodryinus, 311 thoracicus† (Tyrtaeus), 146
testulalis (Maruca), 326 Thaumatomyrmex, 305 Thoracophorus, 113
tethepa† (Pheidole), 303 Thaumatosoma, 316 thorates (Calliomma), 346
Tethina, 226 thayeri (Uca), 42 Thoronidea, 284
Tethorchestia, 30 thecolor† (Pseudomyrmex), 306 thraso (Achlyodes), 337
tethys (Ileopeltus), 266 theischingeri† (Molophilus), 192 THRIPIDAE, 384
Tethysbaena, 29 themis (Schacontia), 324 Thrips, 385
TETRABLEMMIDAE, 59 Theoborus, 237 Throscinus, 126
Tetrabothynus, 174 theobromae (Theoborus), 180 Thrypticus, 212
tetracantha (Gasteracantha), 66 theodori† (Milichiella), 227 thurberiella (Bucculathrix), 323
Tetraceratium, 261 Theope, 340 Thyanta, 257
Tetracha, 101 THERAPHOSIDAE, 56 thymicon (Cycloptilum), 376
Tetragnatha, 65 THEREVIDAE, 211 Thymoites, 63
TETRAGNATHIDAE, 64 THERIDIIDAE, 61 Thyreodon, 288
tetragona (Acontia), 349 Theridion, 63 THYRIDIDAE, 323
tetragona (Athyrma), 354 Theridiosoma, 63 Thyridopteryx, 320
Tetragoneura, 203 THERIDIOSOMATIDAE, 63 thyrsisalis (Macalla), 327
Tetragonoderus, 102 Theridula, 63 Thysania, 355
Tetragonoschema, 125 thermarum (Gabronthus), 117 Thysanopyga, 334
Tetraleurodes, 273 therminieri (Pitho), 42 thysanus (Dissomphalus), 310
Tetramorium, 304 thermocyclopoides (Thermocyclops), 17 Tibellus, 73
Tetraneura, 275 Thermocyclops, 17 tibialis (Eurycotis), 185
TETRANYCHIDAE, 50 Thermonectus, 100 tibialis (Geranomyia), 194
Tetranychus, 50 Thesium, 116 tibialis (Hydrellia), 234
Tetraonyx, 148 Thicanus, 149 tibialis (Mastigocera), 348
Tetrapriocera, 134 Thienemannimyia, 199 tibialis (Pachodynerus), 309
Tetrapus, 289 Thinobius, 113 tibialis (Stenocrepis), 104
tetrastichus (Tangiopsis), 262 Thinocharis, 115 tibialis† (Orchestina), 59
Tetreuaresta, 222 Thinodromus, 113 tibialis† (Pensacolatus?), 76

661 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

tibiaseta† (Episinus), 62 torrida (Megachile), 316 Trematodesmus, 89
tibicen (Calcinus), 39 torticornis (Ecdytolopha), 331 Trentepohila, 193
Tibraca, 257 tortilis (Ochlerotatus), 200 trepida (Hapithus), 371
tiburon (Modisimus), 58 tortilla (Cicadulina), 266 Trepobates, 240
tiburonica (Euchroa), 106 TORTRICIDAE, 331 treptona (Agalliopsis), 264
Tiburonotus, 365 tortuensis (Alethaxius), 162 Treseolobus, 90
Tidarren, 63 tortugae (Paguristes), 39 TRHYPOCHTHONIDAE, 48
Tildenia, 322 Tortyra, 322 triacantha (Schizopera), 18
Tilloclytus, 150 torva (Pseudolycoriella), 205 Triacanthagyna, 363
timais (Xanthopastis), 355 Torycus, 349 Triaeris, 60
timens (Walkerana), 369 TORYMIDAE, 297 Trialeurodes, 273
timida (Tetreuaresta), 222 toussainti (Apenes), 103 triangula‡ (Uroleberis), 28
tincta (Grazia), 256 toussainti (Camponotus), 299 triangularis (Dissomphalus), 310
tinctocorpus† (Apodrosus), 175 toussainti (Corinna), 71 triangularis (Metepeira), 67
tinctoides (Mouralia), 357 toussainti (Hemipepsis), 307 triangularis (Ozophora), 252
Tinea, 320 toussainti (Parvitermes), 189 triangularis (Setomima), 195
Tinearia, 195 toussainti (Strymon), 340 triangulifera (Silis), 132
TINEIDAE, 320 toussaintii (Isaloides), 73 Triangulocypris, 21
TINGIDAE, 241 townsendi (Signiphora), 297 Triatoma, 247
tinniens (Walkerana), 369 Toxomerus, 216 Tribolium, 145
Tinocallis, 275 Toxonotus, 165 tribullus† (Piesocorynus), 164
Tinus, 68 Toxonprucha, 355 tribuna (Seirocastnia), 350
Tiphia, 308 Toxoptera, 274 tricarinata (Booralana), 31
TIPHIIDAE, 308 Toxorhina, 193 Trichadenotecnum, 382
tipica‡ (Abyssocypris), 26 Toxorhynchites, 201 Trichapion, 167
tipoto (Platynus), 105 Toxorhynchus, 167 Tricharaea, 220
Tipula, 194 toxum (Gypona), 268 trichoarcana† (Archicratyna), 203
tipulalis (Ercta), 325 Trachelas, 72 Trichobius, 217
TIPULIDAE, 192 TRACHELIDAE, 72 trichocerus (Dagus), 233
tischeriella (Protodarcia), 320 trachoides (Aleurotrachelus), 272 Trichocorixa, 241
Titanogrypa, 219 trachycerus (Gymnandrosoma), 331 TRICHODECTIDAE, 379
Titanophilus, 85 Trachyderes, 153 Trichogramma, 297
titillans (Mansonia), 200 Trachyleberidea, 27 TRICHOGRAMMATIDAE, 297
Tityus, 82 TRACHYLEBERIDIDAE, 27 Tricholabus, 287
Tobias, 74 Trachyleberis, 27 Trichomyia, 195
tobias (Sticticomus), 149 Trachypus, 313 Trichomyrmex, 302
tocanum† (Elaphidion), 151 Trachythrips, 385 Trichonta, 203
tokionis (Lepidosaphes), 278 tragius (Calisto), 343 Trichopelma, 56
tolteca (Campsomeris), 308 trajectalis (Marimatha), 349 Trichophoroides, 152
tolteca (Drosophila), 230 Tramea, 364 Trichoplusia, 357
toma (Micropholcus), 58 tranquebarica (Chrysobothris), 125 Trichopoda, 220
toma (Modisimus), 58 transcibao (Platynus), 105 Trichopria, 288
tomaculus (Trachelas), 72 Transeius, 53 Trichoprionus, 155
Tomarus, 123 transferrana (Episinus), 331 Trichorhina, 33
tomentosa (Solenoptera), 156 transitella (Amyelois), 328 Trichosia, 205
tomentosum (Elaphidion), 151 translucendi† (Dohrniphora), 213 Trichrous, 152
Tominotus, 255 translucida (Acrobasis), 330 Tricimba, 227
Tomoplagia, 222 transmutata (Agrophila), 350 tricincta (Anastrepha), 223
Tomopteryx, 327 transversa (Sandersonella), 115 tricincta (Campsomeris), 308
Tomosvaryella, 215 transversalis (Metacynortoides), 78 tricolor (Acroricnus), 287
toni (Phyllophaga), 122 transversalis (Xylina), 350 tricolor (Alaus), 127
Toonglasa, 248 transversus (Pachygrapsus), 41 tricolor (Anthophora), 316
torquata (Lampetis), 125 trapezalis (Marasmia), 326 tricolor (Clibanarius), 39
torquata (Xylomeira), 134 trapezoidea (Caribecandona), 20 tricolor (Lintneria), 344
torquata‡ (Uroleberis), 28 trapherus (Photis), 30 tricolor (Megacerus), 163
torquatus (Rhaptinus), 169 tredecimnotata (Parawixia), 68 tricolor (Sphinx), 344

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 662
tricolorata (Silis), 132 trinitatis (Aphelopus), 310 TROPIDUCHIDAE, 262
tricoloripes (Neotanygastrella), 230 trinitatis (Hypolestes), 362 Tropirhinus, 174
Tricolus, 181 trinitatis (Lechytia), 79 Tropisternus, 108
tricornata‡ (Loxoconcha), 25 Trioacantha†, 272 Tropocyclops, 17
tricornis (Excorallana), 31 trionyx (Clinidium), 108 trossulella (Aristotelia), 321
Tricorynus, 135 Triops, 14 Trox, 119
Tricorythodes, 235 triops (Neoconocephalus), 368 Trujillina, 69
tricuspis (Anthonomus), 169 TRIOPSIDAE, 14 truncata (Chalcosicya), 162
TRIDACTYLIDAE, 364 TRIOZIDAE, 272 truncaticornis (Orthobelus), 270
tridens (Tetrapriocera), 134 tripartita (Tettigonia), 265 truncatipennis (Sicoderus), 176
tridens (Xylomeira), 134 tripartitus (Caristanius), 328 Truncattus, 76
tridentata (Cyathura), 31 Triphosa, 335 Truncotettix, 365
tridentiger (Xeropigo), 72 triplex (Micrathetis), 350 trunculus (Pseudothysanoes), 178
tridigitatus (Prostemmiulus), 94 triplistratus‡ (Neocaudites), 24 truncus (Crenulaspidiotus), 277
Triebelina, 19 Tripudia, 355 Trupanea, 223
Trientoma, 143 tripunctata (Lygropia), 326 Trybomia, 385
Triepeolus, 317 tripunctus tripunctus (Cymaenes), 336 Trypanaeus, 110
trifasciata (Argiope), 66 triquetrus (Orthoporus), 93 Trypanalebra, 269
trifasciata (Campsomeris), 308 Trischistognatha, 323 Trypanidius, 154
trifasciata (Cicindela), 101 Trischizostoma, 30 trypetina (Neoacanthoneura), 223
trifasciatus (Hamiltoniauletes), 167 TRISCHIZOSTOMATIDAE, 30 Trypoxylon, 313
trifasciatus (Liris), 313 triseta (Scatella), 233 tschirnhausi (Aphelopus), 310
trifida (Triacanthagyna), 363 trishae (Insulatitan), 212 tschoppi‡ (Hermanites), 24
trifilum (Tytthonyx), 133 trisicifer (Anthonomus), 169 tschoppi‡ (Paracytheridea), 26
trifolii (Liriomyza), 225 trispinosus (Collodes), 42 tubercularis (Aulacaspis), 278
trigemmis (Areva), 349 Trissolcus, 285 tuberculata (Platycotis), 270
TRIGONALIDAE, 282 tristani (Pyragropsis), 183 tuberculata (Tomosvaryella), 215
TRIGONIDIIDAE, 375 tristicta (Bagisara), 351 tuberculatus (Corthylus), 181
TRIGONIULIDAE, 93 tristis (Anasa), 253 tuberculatus (Haematopinus), 379
Trigoniulus, 93 tristis (Galerita), 102 tuberculatus (Haitianotettix), 365
Trigonodes, 355 tritax (Elaphropus), 107 tuberculatus (Phaenotheriopsis), 165
trigonodontos (Halictophagus), 383 trite watsoni (Rhabdodryas), 338 tuberculatus† (Pseudoarmadillo), 33
Trigonuropoda, 54 Triteleia, 284 tuberculosa (Philephedra), 276
TRIGONUROPODIDAE, 54 trivittatus (Hedriodiscus), 206 tuberis (Epitrix), 157
triguttatus (Heraeus), 251 TROCHANTERIIDAE, 72 tuberosa (Osachila), 40
trilinea (Laphygma), 351 troche† (Tapinoma), 299 tuberosus (Venezillo), 33
trilineata (Olcella), 227 trochilus (Cleogonus), 176 tuberosus† (Episinus), 62
trilineatus (Enicospilus), 288 Trochosa, 68 tuberosus† (Modisimus), 58
trilineatus (Metophthalmus), 142 TROCTOPSOCIDAE, 380 tubifer (Liriomyza), 225
trilineatus (Polycyrtus), 287 Troctopsocoides, 380 tubifex (Leucotrichia), 389
trilineatus (Uloborus), 60 TROGIDAE, 119 Tubomixdea, 49
trilobitiformis (Pseudaonidia), 278 troglodyta (Anea), 340 tubus (Osbornellus), 267
trilobus (Berosus), 108 troglodytes (Carcinops), 109 Tugana, 70
trimaculata (Caribovia), 265 Trogoderma, 133 tulipae (Aceria), 48
trimaculata (Fannia), 218 Trogolaphysa, 96 tumida (Phymacista), 242
trimaculata (Nelphe), 348 TROGOSSITIDAE, 135 tumidirostris† (Promecops), 173
trimaculatus (Amnestus), 254 Trogoxylon, 134 tumidulus (Cronius), 43
trimaculatus (Henicospilus), 287 Tromatobia, 287 tumidus (Callinectes), 43
trimaculatus (Pamphantus), 249 Trombicula, 50 tumidus (Metrobates), 240
trimarginatus (Elateropsis), 156 TROMBICULIDAE, 50 tumidus (Wilcoxius), 208
Trinemurodes, 97 tropicalis (Drosophila), 230 tumulus† (Chrosiothes), 61
trinidadensis (Brevipalpus), 50 tropicalis (Mesopenaeus), 35 Tunga, 382
trinidadensis (Culicoides), 196 tropicalis (Ophisma), 354 TUNGIDAE, 382
trinidadensis‡ (Abyssocythere), 27 tropicalis cubana (Drosophila), 230 tunicatus (Drapetes), 129
trinidadensis‡ (Cytheropteron?), 23 Tropicharis, 293 Tupiocoris, 243
trinidadensis‡ (Krithe), 24 Tropicus, 127 turbata (Corynoptera), 204

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turbatus (Brentus), 168 ULOBORIDAE, 60 UPOGEBIIDAE, 38
turbatus (Estenorhinus), 168 Uloborus, 60 URANIIDAE, 332
turbinata (Cyclosa), 66 Uloma, 145 Uranotaenia, 201
Turbopsebius, 206 Ulomoides, 146 Urbanus, 337
turbulenta (Hapithus), 371 ulricusalis (Bleptina), 356 urbicola (Pulvinaria), 276
turgida (Bradysia), 204 ultima (Alisotrichia), 388 Urgleptes, 154
turnbowi (Antillonirvana), 267 ultima (Amphiacusta), 374 urichii (Paraleyrodes), 273
turnbowi (Araptus), 180 ulula (Lembodes), 171 urnulum (Hydrochasma), 231
turnbowi (Parandraceps), 152 Ululodes, 358 URODIDAE, 321
turnbowi (Plectromerus), 151 Umbonia, 270 Urodus, 321
turnbowi (Sicoderus), 176 umbraculata (Acanalonia), 258 Uroleberis, 28
Turpilia, 368 umbrata (Erythrodiplax), 363 Uroleucon, 274
turrifera (Omoteina), 161 umbratipalpis (Acrolophus), 320 Uropelma, 294
turturaria (Macaria?), 333 umbronatus (Phaenotheriopsis), 165 UROPODIDAE, 54
tussia (Ptichodis), 354 umbrosa (Guttipsilopa), 231 Urosigalphus, 285
Tuta, 322 umbrosa (Ozophora), 252 ursula (Ageniella), 307
Tutaibo, 64 una (Toxomerus), 216 urticae (Tetranychus), 50
tutrix (Sabelo), 369 Unaspis, 278 urus (Orthobelus), 270
tweedyanus (Neoathyreus), 119 uncicornis (Cryptolestes), 138 urvillei (Platylister), 110
tychaeos (Hapithus), 370 uncimanus (Nemocephalus), 168 ustulatus (Teloplatypus), 182
Tylocerus, 132 uncinata (Clasiopella), 231 ustus (Camponotus), 300
Tyloderma, 172 uncinata (Pyrgocorypha), 368 ustus depolitus (Camponotus), 300
Tylospilus, 255 uncinatus (Camptocercus), 14 ustus furnissi (Camponotus), 300
Tylozygus, 265 uncula (Agalliopsis), 264 ustus sublautus (Camponotus), 300
typhaeus (Merocoris), 253 undata (Cymatothes), 144 ustus ulysses (Camponotus), 300
Typhlatya, 37 undatifascia (Pachylia), 345 uterella (Phereoeca), 320
Typhlocybella, 269 undecimtuberculata (Wagneriana), 68 Utetes, 286
Typhlocypris, 21 undifraga (Acrocercops), 320 Utetheisa, 347
Typhlodromalus, 53 undosa (Parectopa), 320 utilis (Ceroplastes), 275
Typhlodromina, 53 undulans (Caularis), 350 utilis (Ganaspidium), 289
Typhlodromips, 53 undulifera (Chabora), 351 utilis (Pseudaphycus), 292
Typhlodromus, 53 Unerus, 266 utinanus (Opius), 286
Typhloseiopsis, 53 unguiculata (Cariblatta), 187 utowani (Beatadesmus), 88
Typopsilopa, 234 unica (Gonodonta), 352 uvae (Coccobius), 290
tyralis (Pyrausta), 327 unicarinata (Gnathopleura), 285 Uvaroviella, 374
Tyrannasorus†, 119 unicinctus (Amblypsilopus), 212 uviferae (Euscelus), 166
tyrannicos (Hapithus), 370 unicolor (Alcidion), 153 uxoria (Gonodonta), 353
Tyrannochelifer, 80 unicolor (Brachypremna), 194 uzeli (Gynaikothrips), 385
Tyrannochthonius, 79 unicolor (Erythrolychnia), 130 uzinuri (Ferrisia), 279
Tyrannocoris, 255 unicolor (Galerita), 102 vacaensis (Laccophilus), 101
Tyrissa, 355 unicolor (Hyalochloria), 244 vacillans (Leptobasis), 362
Tyrtaeus, 146 unicolor (Loxostethus), 146 vacillans (Odopoea), 263
Tytthonyx, 132 unicolor (Smicridea), 387 Vacusus, 149
Tytthus, 244 Unicorniella, 368 vaderi (Diastolinus), 144
ubica (Chinavia), 256 unicornis† (Chrosiothes), 61 vagabundus (Loganius), 177
Uca, 42 unicus (Atomothenemus), 178 vagabundus (Nemobiopsis), 373
Ucides, 42 unidens (Rhopobota), 331 v-album (Cynortoides), 78
Udea, 329 unidentata (Marava), 183 valdiviaensis (Austrophthicarus), 47
Ufa, 330 unifasciata (Signiphora), 297 valencius (Arrenurus), 49
Ufens, 297 uniformis (Nanus), 176 valens (Dryinus), 311
ufeus (Desmia), 325 unimaculatus (Cephalotes), 301 valens (Paroecanthus), 372
uhleri (Bubaces), 251 unipuncta (Mythimna), 356 valentinei (Pycnomerus), 143
uhleri (Geogryllus), 372 unispina† (Palaeodyctina), 70 valentinei (Toxorhynchus), 167
Uhleroides, 263 univitella (Anypsipyla), 328 valentinei (Tytthonyx), 133
ul (Synalpheus), 36 unnamed species 5 (Drosophila), 230 valentinorum (Pachyschelus), 124
ULIDIIDAE, 223 Upogebia, 38 Valenzuela, 381

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 664
Valeseguya, 196 vega (Heteroonops), 59 versatiliaria (Hyperythra), 333
VALESEGUYIDAE, 196 vegaensis (Presbyolampis), 131 versicolor (Ataenius), 120
valga (Marilia), 390 vegana (Rhagovelia), 240 versicolor (Corthylus), 181
valgus (Phileurus), 124 vegas (Wagneriana), 68 verticalis verticalis (Trichocorixa), 241
valida (Ceraeochrysa), 470 vegensis (Mnemosyne), 259 verticordia (Memphis), 341
valida (Thermesia), 352 vegeta (Eustala), 66 vesciculosa (Erythemis), 363
validus (Aleuroplatus), 272 Velapia, 373 vespa‡ (Argilloecia), 26
validus (Heteroonops), 59 Velatis†, 176 vespera (Amphiacusta), 374
vallenuevo (Achromoporus), 88 velezi† (Siphonophora), 91 vespertilionis (Hesperochernes), 80
vallenuevo (Wagneriana), 68 VELIIDAE, 240 vespertina (Cyrtoxipha), 375
vallinovae (Robopus), 131 Vella, 361 VESPIDAE, 308
Valtissius, 251 Velocitermes, 189 vespiformis (Franklinothrips), 384
vancei (Priocnessus), 307 velox (Hibana), 71 Vestitecola, 77
vanderpooli (Polycentropus), 387 velox (Thaumasia), 68 vestitipennis (Anopheles), 200
vanderwulpi (Belvosia), 220 velozae (Robopus), 131 vestitum (Neapion), 167
Vanduzea, 271 velozi (Robopus), 131 vesulia transponens (Oxydia), 334
vanduzei (Gargaphia), 242 velteni† (Anelaphus), 151 vetehae† (Glyphembia), 234
Vanessa, 342 velteni† (Coryssocnemis), 57 Veterator, 72
vanillae insularis (Agraulis), 341 velteni† (Ischnura), 362 vetula† (Chrysopa?), 359
vannamei (Litopenaeus), 35 velteni† (Phylloicus), 390 vetula† (Nylanderia), 300
vapida (Cladochaeta), 229 velteni† (Rowlandius), 81 vetusalis (Pyralis), 330
vaporariorum (Trialeurodes), 273 velutinus (Corinthiscus), 136 vetusta† (Oligembia), 235
vaquitarum (Aprostocetus), 294 venatoria (Heteropoda), 73 vetusta† (Proplebeia), 317
varaderoensis‡ (Propontocypris), 21 venatus vestitus (Sphenophorus), 177 vetustalis (Hypena), 357
varenna (Dexosarcophaga), 219 Venezillo, 33 vetustaria (Azelina), 334
varia (Aeneolamia), 262 venosa† (Paleoploiariola), 246 vetustus† (Bocchus), 311
variabilis (Cerataphis), 275 venosus (Dardanus), 39 vetustus† (Dryinus), 311
variabilis (Neoxantholinus), 117 venticola (Pseudopostega), 319 veve (Toxomerus), 216
variabilis (Trialeurodes), 273 ventosa (Scaptosciara), 205 vexans (Elcaribe), 212
variabilis‡ (Pedicythere), 23 ventralis (Holopyga), 310 vexans (Walkerana), 369
varicans (Microcyclops), 17 ventralis (Oncocephalus), 247 vexus (Tenuipalpus), 50
varicolor (Murgantia), 257 ventralis (Philonthus), 118 vibrans (Anurogryllus), 372
varicolor (Oncopeltus), 250 venusta (Acanthepeira), 65 vibrissae† (Dominocephala), 296
varicornis (Carpelimus), 113 venusta (Leucauge), 65 vicarius (Neopamera), 251
variegata (Encarsia), 290 venusta (Melanophthalma), 142 vicina (Leptopharsa), 242
variegata (Omoteina), 161 venusta (Ozophora), 252 vicina† (Laseola), 62
variegatus (Chrysops), 207 venusta (Physocephala), 221 Vicinilura†, 271
variegatus (Diolcus), 258 venustalis (Leucinodes), 326 vicinoides† (Laseola), 62
variegatus (Ischnodemus), 248 venustella (Leucauge), 65 vicinus (Vacusus), 149
variegatus (Neopetissius), 251 venustula (Cardiocondyla), 302 vicinus† (Pseudomyrmex), 306
varilobatus (Augesmus), 89 venustula (Tarachidia), 350 vicksburgensis‡ (Jugosocythereis), 24
variolosa (Ophiusa), 352 venustulus (Anisopilothrips), 384 vicksburgensis‡ (Pseudopsammo-
variolosa (Spodoptera), 356 venustus (Anisepyris), 309 cythere), 22
variosoma† (Theridion), 63 venustus (Chondrotropis), 88 victor (Proxys), 257
varipes (Alabagrus), 285 venustus (Osbornellus), 267 victori (Calisto), 343
varipes (Anthonomus), 169 venustus (Prostemmiulus), 94 victori (Ebanodesmus), 88
varipes (Parasphaerocera), 229 venustus (Rejanellus), 74 victori (Triepeolus), 317
varius stygius (Cephalops), 214 verda (Ochrotrichia), 389 vidua (Cyclocephala), 123
Varneria, 330 verruca (Argyrogramma), 357 vidua (Cypridopsis), 20
Varroa, 54 Verrucosa, 68 vidua‡ (Cypridopsis), 20
VARROIDAE, 54 verrucosa (Aedmon), 157 viduata (Exoma), 261
Vatiga, 243 verrucosa† (Enacrosoma), 66 viduus (Cenophorus), 144
vaughani‡ (Orionina), 24 verrucosus (Mithrax), 42 viduus† (Veterator), 72
vecordalis (Botys), 325 verrucosus (Orthobelus), 270 vieja† (Meoneura), 226
vega (Aptunga), 328 verrucosus (Phaenotheriopsis), 165 viettei (Palpita), 326
vega (Cubanops), 59 verruculosus (Aeolus), 128 vigil (Hapithus), 370

665 · Zootaxa 4749 (1) © 2020 Magnolia Press PEREZ-GELABERT

vigilans (Coelioxys), 316 viridissima (Dellia), 366 volvulus (Brentus), 168
vigilax (Hapithus), 370 viriditas (Coryphaeschna), 363 volvulus (Xyleborus), 180
vigilax (Metepeira), 67 viridium (Apodrosus), 175 vulgaris (Sthenaridea), 244
viguerasi (Carios), 52 viridius (Chlorotettix), 266 vulnerata (Gyretes), 99
vigueri (Phyllognathopus), 18 viridula (Nezara), 257 vulnerata (Spangbergiella), 266
Villa, 211 viridulus sp. group (Agapostemon), 315 vulnerata (Telebasis), 362
villosa (Chalcosicya), 162 virkkii† (Leptonesiotes), 159 vulnerata† (Grimalantha), 216
villosoantennata† (Cratyna), 204 Virbia, 347 vulpes (Lychnacris), 131
villosus (Ululodes), 358 visite (Aculepeira), 65 vulpes (Nessorhinus), 270
vincenti (Antillisa), 346 visite (Tainonia), 58 Vulsirea, 257
vincentiae (Ataenius), 120 visitor (Platynus), 105 vultus (Apachekolos), 209
vinctus (Pseudopachybrachius), 251 viticola (Diaspidiotus), 277 Wagneriana, 68
vinetorum (Palpada), 215 vitis vitis (Eumorpha), 346 walckenaeri (Cyclosa), 66
vineus (Syneches), 213 vitraci (Copelatus), 100 walcotti (Leptopharsa), 242
vinosa (Hyalurga), 348 vitrata (Maruca), 326 waldneri (Renocila), 32
vinosus (Cephalotes), 302 vitrea† (Hyalistata), 231 Walkerana, 369
Vinsonia, 276 vitreus (Ommatius), 210 Walkerella, 369
violaceipennis (Trichrous), 152 vitrina (Catabena), 350 Wallengrenia, 336
violaceipes (Ageniella), 307 vittata (Caribovia), 265 Wanderbiltiana, 159
violaceotincta (Silis), 132 vittata (Ercta), 325 wapleri (Platycrepidius), 128
violaceus (Canthon), 121 vittata (Hentzia), 76 wappesi (Elaphidion), 151
violaceus (Thrypticus), 212 vittata (Oxycopis), 148 wasmanni (Hyperlasion), 205
violaceus (Toxomerus), 216 vittata (Solenoptera), 156 Wasmannia, 301
virens (Plinthocoelium), 150 vittatifrons (Zygothrica), 231 waterhousei (Apalonychus), 119
virescens (Heliothis), 356 vittatipenne (Acmopolynema), 295 waterhousei (Echiaster), 114
virescens (Idioderma), 271 vittatum (Mionochroma), 150 waterhousei (Epierus), 110
virescens (Kleidocerys), 249 vittatus (Clibanarius), 39 watsoni (Cryptohelea), 320
virga (Calliscelio), 283 vittatus (Eobates), 239 watsoni (Pyractonema), 130
virgata (Ferrisia), 279 vittatus (Heterophotinus), 131 watsoni (Toxomerus), 216
virgata (Pseudolycoriella), 205 vittatus (Modisimus), 58 watsoni (Volucella), 215
virginiae (Microtityus), 82 vittatus (Onychotillus), 136 watsoni watsoni (Allothrips), 385
virginiae (Rowlandius), 81 vittatus (Proctacanthus), 208 Wattius, 145
virginicus (Echiniscus), 13 vitticollis (Pyractonema), 130 wawasita† (Wanderbiltiana), 159
virginiensis (Chalcophora), 125 vittiger (Tachys), 107 websteri (Asphondylia), 202
virginiensis (Vanessa), 342 vittiger guatemalanus (Tabanus), 207 weithi (Hyptia), 288
virgo (Peckia), 219 vittipalpis (Hippia), 347 weitschati† (Chimarra), 386
virgo† (Polymera), 193 vittula (Pentagonica), 104 weitschati† (Cyrtognatha), 65
viridans (Peucetia), 69 vivafosile (Brachypsectra), 127 Wendilgarda, 63
viridari (Alpheus), 36 vivax (Anaxipha), 375 Werneckiella, 379
viride (Cnodalon), 147 vivax (Knyella), 369 Westindia, 349
viridescens (Xanthia), 356 vivianae (Chinaria), 263 westwoodi antillarum (Elephantomyia),
viridicolor (Heterophotinus), 131 vivida (Carylla), 369 193
viridicomus (Leptostylopsis), 153 vividula antillana (Paratrechina), 300 wetherbeei (Calisto), 343
viridiflavescens (Cicindela), 101 vivus (Hapithus), 370 wetherbeei (Epilobocera), 44
viridiguttatus (Ischionoplus), 174 vivus (Ommatius), 210 wetherbeei (Ravilops), 83
viridipennis (Alagoasa), 157 vocator (Uca), 42 whartoni (Pseudognatodon), 286
viridis (Acanalonia), 258 vockerothi (Johnrichardia), 222 wheeleri† (Neotanygastrella), 230
viridis (Coccus), 275 vockerothi (Syrphus), 216 whiteheadi (Parinesa), 141
viridis (Lebia), 104 vodou (Cupiennius), 69 wierzejskii (Moina), 15
viridis (Lemphus), 136 voigti† (Helicopsyche), 391 Wilcoxia, 208
viridis (Loxa), 256 voldroguea (Tubomixdea), 49 Wilcoxius, 208
viridis (Lyssomanes), 76 Voldroguella, 49 williamsi (Frankliniella), 384
viridis (Panchlora), 186 volkswageni† (Dohrniphora), 213 williamsi (Microcephalops), 215
viridis (Protoneura), 362 Voltinia, 340 willisensis‡ (Bairdia), 19
viridis (Xerophloea), 268 volubilis (Apachekolos), 209 willistoni (Beskia), 220
viridissima (Antianthe), 270 volucripes (Epeira), 67 willistoni (Drosophila), 230

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willistoni (Tethina), 226 Xantholinus, 118 yawi (Anagrus), 295
wirthi (Dagus), 233 Xanthopastis, 355 yayitas (Acridurus), 367
wirthi (Lemnaphila), 234 xanthopoda (Tethina), 226 Yelicones, 287
wirthi (Parabezzia), 196 Xanthorhoe, 335 yenli† (Diphleps), 244
wirthi† (Phaenobezzia), 197 xanthosoma (Telenomus), 285 Yingaresca, 159
wirthorum† (Palpomyia), 197 xantotricha (Chondromorpha), 89 ylem (Anthonomus), 169
WITHIIDAE, 80 xehohypopia (Nostima), 234 yolandae† (Dipoenata), 62
Witica, 68 Xenasphondylia, 202 yoni‡ (Caribbella), 26
wittmeri (Hispanioryctes), 123 xenisma (Paradejeania), 220 Yoshimotoana, 295
wittmeri (Lobopoda), 145 Xenochalepus, 161 yothersi (Brevipalpus), 50
woglumi (Aleurocanthus), 272 Xenolimosina, 229 youngi (Apenes), 103
wolcotti (Anysepyris), 309 xenopodium (Clinidium), 108 youngi (Balclutha), 266
wolcotti (Chrysopa), 359 xenops (Prostemmiulus), 94 youngi (Boreofairchildia), 194
wolcotti (Inopeplus), 149 Xenorhipis, 125 youngi (Brasiella), 101
wolcotti (Jassus), 265 xenoverpa (Nesonana), 212 youngi (Caribojosia), 346
wolcotti (Metachroma), 163 Xenylla, 94 youngi (Cryptorhopalum), 133
wolcotti (Myzine), 308 xericum (Canthochilum), 121 youngi (Curtara), 268
wolcotti (Orthobelus), 270 Xerolinus, 144 youngi (Haliplus), 99
wolcotti (Tetraceratium), 261 Xeromelecta, 317 youngi (Insulatitan), 212
wolcotti (Tiphia), 308 Xerophloea, 268 youngi (Metadorylas), 215
wolcotti† (Lorelus), 143 Xeropigo, 72 youngi (Nesonana), 212
wolla (Platynus), 105 xerozoyphius (Endroedianibe), 123 youngi (Pityophthorus), 181
wolverinei (Bromeloecia), 228 Xestocephalus, 269 youngi† (Dominicicada), 263
woodburyi (Phytoseius), 53 Xestochironomus, 198 Youngszella, 269
woodleyi (Argopistes), 157 XESTOLEBERIDIDAE, 28 Yoyuteris, 374
woodleyi (Efferia), 208 Xestoleberis, 28 yponomeutaria (Eumacrodes), 335
woodleyi (Elateropsis), 156 Xiphichilus, 26 YPONOMEUTIDAE, 322
woodleyi (Psyrassa), 152 XIPHOCARIDIDAE, 37 ypsilon (Lophoscutus), 247
woodruffi (Ablechrus), 136 Xiphocaris, 37 ypsilongriseus (Oebalus), 257
woodruffi (Alisotrichia), 388 Xiphocentron, 387 yucatanicus (Periclimenes), 37
woodruffi (Atrichopogon), 197 XIPHOCENTRONIDAE, 387 yuma (Beatadesmus), 88
woodruffi (Brachypogon), 196 Xiphopeneus, 35 yumai† (Paridris), 284
woodruffi (Chaetophloeus), 178 Xixuthrus, 155 yunax (Elaphropus), 107
woodruffi (Gypona), 268 xuthocolus† (Rhynchitobius), 182 yurabia (Atomosia), 208
woodruffi (Mediocampus), 256 Xyalaspis, 289 Zabrotes, 164
woodruffi (Ninyas), 250 Xyleborinus, 180 zagluli (Anthonomus), 169
woodruffi (Ozophora), 252 Xyleborus, 180 Zagrammosoma, 294
woodruffi (Pelonium), 136 Xyleus, 368 Zaischnopsis, 294
woodruffi (Purenleon), 361 Xylocopa, 318 Zale, 355
woodruffi (Strongylium), 147 Xylomeira, 134 Zamagiria, 330
woodruffi† (Forcipomyia), 197 Xylophanes, 346 Zanthopsis, 44
woodruffi† (Stenotabanus), 207 Xylosandrus, 180 Zanysson, 312
woodruffi† (Zophodetus), 297 xylostella (Plutella), 322 Zaops, 43
woodsi (Calisto), 344 Xyonysius, 250 zapoides (Empoasca), 269
worki† (Diceratobasis), 362 Xyphon, 265 zappii† (Heterachthes), 152
Wulfila, 71 Xysticus, 74 zaptlana (Megachile), 316
wunderlichi† (Theridion), 63 Yabisi, 60 Zaraza, 29
Wyeomyia, 201 yamini† (Cryptotermes), 188 zarucco (Erynnis), 337
wygodzinskyi (Chinavia), 256 yanomami (Parachironomus), 198 Zatania, 300
wygodzinskyi† (Malacopus), 246 yapchoontecki† (Craspedisia), 61 zatheos (Hapithus), 370
Xanthaciura, 223 yaque† (Grossoxipha), 373 zayasi (Conotelus), 137
Xanthandrus, 216 yaquiensis (Ogyrides), 36 zayasi† (Plesioglymmius), 108
XANTHIDAE, 44 yaquiensis‡ (Sandomingia), 42 zayensis (Pullimosina), 228
Xantho, 44 yarari (Margarettia), 369 zea (Helicoverpa), 356
Xanthocaecilius, 3811 yaseeni (Psyllaephagus), 292 Zeadolopus, 110
Xanthocomus, 138 yasumatsui (Aulacaspis), 278 zeamais (Sitophilus), 177

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zebratus (Dicranocentrus), 95
Zelotes, 72
Zelus, 246
zemi (Strumigenys), 302
zenobia (Thysania), 355
zephodes (Ephyriades), 337
zephyrus (Progomphus), 363
zephyrus (Psilonyx), 209
Zerene, 338
Zeros, 234
Zeta, 309
zeta (Trichogramma), 297
zeteki (Banasa), 256
zetes (Papilio), 338
Zethus, 309
zetides (Battus), 338
Zeugomantispa, 361
Zicca, 254
Zilus, 141
zimbabweensis† (Archicratyna), 203
Zimiris, 72
zinckenella (Etiella), 329
ziziphi (Parlatoria), 278
zomata (Ghinallelia), 246
zombia (Hentzia), 76
Zombiweckelia, 30
zonaria (Neographium), 337
zonata (Dichrogaster), 287
ZOPHERIDAE, 143
Zophobas, 145
Zophodetus†, 297
ZOROTYPIDAE, 391
Zorotypus, 391
Zosis, 60
zuluagai (Iphiseiodes), 53
Zuphium, 106
Zygoribatula, 47
Zygothrica, 231

CHECKLIST OF HISPANIOLAN ARTHROPODS Zootaxa 4749 (1) © 2020 Magnolia Press · 668