Ecology has a complex origin dating back to ancient Greek philosophers like Aristotle who made early observations of natural history. Modern ecology emerged in the late 19th century as a more rigorous science. Key figures included Ernst Haeckel who coined the term "ecology" and Charles Darwin whose theory of evolution was a cornerstone of ecological thought. Ecology is defined as the scientific study of the interactions between organisms and their environment, and includes variables like species distribution, abundance, and changing states within ecosystems. It is a multidisciplinary field with applications in conservation, natural resource management, and human social systems.